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<document id="6D1A110E4134C5D95495ECA542D9B4B7" ID-DOI="10.1649/0010-065X-72.4.665" ID-ISSN="1938-4394" ID-Zenodo-Dep="5384479" ID-ZooBank="urn:lsid:zoobank.org:pub:754D1387-0B53-4C3F-AAD7-01591A9ED3AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299362533" checkinUser="diego" docAuthor="Paucar-Cabrera, Aura &amp; Ratcliffe, Brett C." docDate="2018" docId="03F187B81C6719568CA9C6F20866F604" docLanguage="en" docName="0010-065X-72.4.665.pdf" docOrigin="The Coleopterists Bulletin 72 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-72.4.665" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Ancognatha uncinata Paucar-Cabrera and Ratcliffe 2018, new species" docType="treatment" docVersion="3" lastPageNumber="683" masterDocId="FFC8FFC01C7519458C18C206090DF53E" masterDocTitle="The Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) of Ecuador, with Description of a New Species" masterLastPageNumber="687" masterPageNumber="665" pageNumber="682" updateTime="1699219988748" updateUser="plazi">
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<mods:title id="44B6F710445B0A6CF81A61BB701DB81E">The Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) of Ecuador, with Description of a New Species</mods:title>
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<mods:namePart id="B65F81FE813F91C26C943F25F55F2C14">Paucar-Cabrera, Aura</mods:namePart>
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<mods:namePart id="2C81F860F71D4FEFBD513945E48E8D4F">Ratcliffe, Brett C.</mods:namePart>
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<mods:title id="2EB662398C37D1E100D7A733572264A0">The Coleopterists Bulletin</mods:title>
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<mods:date id="91A4D820629831BFF04F96FF21CF976F">2018</mods:date>
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<paragraph id="8BE736AE1C6719578CA9C6F208FAF01D" blockId="18.[160,629,1268,1395]" pageId="18" pageNumber="682">
<heading id="D0AF81C21C6719578CA9C6F208FAF01D" centered="true" fontSize="9" level="2" pageId="18" pageNumber="682" reason="2">
<emphasis id="B92CEABC1C6719578CA9C6F208FAF01D" bold="true" pageId="18" pageNumber="682">
<taxonomicName id="4C584D2D1C6719578CA9C6F2087BF01D" authority="Paucar-Cabrera and Ratcliffe" authorityName="Paucar-Cabrera and Ratcliffe" authorityYear="2018" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="682" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="B92CEABC1C6719578CA9C6F20887F037" bold="true" box="[177,394,1268,1289]" italics="true" pageId="18" pageNumber="682">Ancognatha uncinata</emphasis>
Paucar-Cabrera and Ratcliffe
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C71C6719578D99C70808FAF01D" box="[385,503,1294,1315]" pageId="18" pageNumber="682" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C34265251C6719578D52C72F0B78F04D" pageId="18" pageNumber="682" type="description">
<paragraph id="8BE736AE1C6719578D52C72F0B78F04D" blockId="18.[160,629,1268,1395]" pageId="18" pageNumber="682">
(
<figureCitation id="13632A2B1C6719578D49C72F08C8F000" box="[337,453,1321,1343]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="18" pageNumber="682">Figs. 3234</figureCitation>
) Zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C34265251C6719578CB7C78F0A01F3FC" pageId="18" pageNumber="682" type="materials_examined">
<paragraph id="8BE736AE1C6719578CB7C78F085EF336" blockId="18.[151,638,1417,1730]" pageId="18" pageNumber="682">
<emphasis id="B92CEABC1C6719578CB7C78F0853F0A0" bold="true" box="[175,350,1417,1438]" pageId="18" pageNumber="682">Type Material.</emphasis>
<materialsCitation id="3B303CF31C6719578D60C78F0B12F0D3" collectingDate="1990-02-15" collectorName="S. Sandoval" country="Ecuador" location="Fall San Rafael" pageId="18" pageNumber="682" specimenCount="2" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="54E3880C1C6719578D60C78F08D9F0A0" box="[376,468,1417,1438]" pageId="18" pageNumber="682" type="holotype">Holotype</typeStatus>
male labeled “
<collectingCountry id="F34F763E1C6719578CBAC7A50815F086" box="[162,280,1443,1464]" name="Ecuador" pageId="18" pageNumber="682">ECUADOR</collectingCountry>
/
<collectingRegion id="499CF84C1C6719578D34C7A50863F086" box="[300,366,1443,1464]" country="Ecuador" name="Napo" pageId="18" pageNumber="682">NAPO</collectingRegion>
/
<location id="8E8760751C6719578D9BC7A50B11F086" LSID="urn:lsid:plazi:treatment:03F187B81C6719568CA9C6F20866F604:8E8760751C6719578D9BC7A50B11F086" box="[387,540,1443,1464]" country="Ecuador" name="Fall San Rafael" pageId="18" pageNumber="682" stateProvince="Napo">Fall San Rafael</location>
/
<date id="FFE6106E1C6719578E28C7A509C5F0ED" pageId="18" pageNumber="682" value="1990-02-15">
<collectingDate id="EFA2E9861C6719578E28C7A509C5F0ED" pageId="18" pageNumber="682" value="1990-02-15">15-Feb- 1990</collectingDate>
</date>
/ Legit:
<collectorName id="26AD53781C6719578D31C7B808AEF0ED" box="[297,419,1470,1491]" pageId="18" pageNumber="682">S. Sandoval</collectorName>
// 3545 //
<collectingRegion id="499CF84C1C6719578E01C7B80B60F0ED" box="[537,621,1470,1491]" country="Ecuador" name="QCAZ I" pageId="18" pageNumber="682">QCAZ I</collectingRegion>
/ 224571” and with our red
<typeStatus id="54E3880C1C6719578D80C7DE08E5F0D3" box="[408,488,1496,1517]" pageId="18" pageNumber="682" type="holotype">holotype</typeStatus>
label
</materialsCitation>
.
<materialsCitation id="3B303CF31C6719578E31C7DE0842F336" collectionCode="QCAZ" country="Ecuador" pageId="18" pageNumber="682" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="54E3880C1C6719578E31C7DE0B73F0D3" box="[553,638,1496,1517]" pageId="18" pageNumber="682" type="holotype">Holotype</typeStatus>
deposited at
<collectionCode id="ED49AE6B1C6719578D15C7F50842F336" box="[269,335,1523,1544]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="18" pageNumber="682" type="Museum">QCAZ</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE736AE1C6719578CB7C40B0A01F3FC" blockId="18.[151,638,1417,1730]" lastBlockId="18.[685,1173,1098,1730]" pageId="18" pageNumber="682">
<emphasis id="B92CEABC1C6719578CB7C40B0828F31C" bold="true" box="[175,293,1549,1570]" pageId="18" pageNumber="682">Decription.</emphasis>
<typeStatus id="54E3880C1C6719578D37C40B0886F31C" box="[303,395,1549,1570]" pageId="18" pageNumber="682" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="13632A2B1C6719578DCDC40B0B2EF31C" box="[469,547,1549,1570]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="18" pageNumber="682">Fig. 32</figureCitation>
). Length
<quantity id="4CA09B4B1C6719578C8FC42E09FBF303" box="[151,246,1576,1597]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="18" pageNumber="682" unit="mm" value="24.3">24.3 mm</quantity>
; width across humeri
<quantity id="4CA09B4B1C6719578DC3C42E0B34F303" box="[475,569,1576,1597]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="18" pageNumber="682" unit="mm" value="10.3">10.3 mm</quantity>
. Color dark testaceous with black, elongate spot on each humerus and behind scutellum either side of sutural line; tarsi, apices of femora, and tibiae black, pygidium and abdominal sternites piceous.
<emphasis id="B92CEABC1C6719578E26C4940B73F399" bold="true" box="[574,638,1682,1703]" pageId="18" pageNumber="682">Head:</emphasis>
Frons and clypeus with small, moderately dense punctures. Clypeal shape parabolic, apex narrowly rounded and slightly reflexed. Interocular width equals 3.0 transverse eye diameters. Antennae with 10 antennomeres, club slightly longer than antennomeres 27. Mentum with apex deeply emarginated (about half length of mentum).
<emphasis id="B92CEABC1C6719578831C6C90D99F1DA" bold="true" box="[1065,1172,1231,1252]" pageId="18" pageNumber="682">Pronotum:</emphasis>
Surface with small, moderately dense punctures similar to those on head. Base lacking marginal bead.
<emphasis id="B92CEABC1C6719578EE9C7180A37F00D" bold="true" box="[753,826,1310,1331]" pageId="18" pageNumber="682">Elytra:</emphasis>
Surface weakly punctate-striate, striae shallow, indistinct; punctures moderate in size, weakly ocellate.
<emphasis id="B92CEABC1C6719578F62C7520AD2F057" bold="true" box="[890,991,1364,1385]" pageId="18" pageNumber="682">Pygidium:</emphasis>
Surface with moderately large, moderately dense, setigerous punctures; setae long, dense, pale. In lateral view, surface convex, especially on apical half.
<emphasis id="B92CEABC1C6719578843C7A50D9EF086" bold="true" box="[1115,1171,1443,1464]" pageId="18" pageNumber="682">Legs:</emphasis>
Protibia tridentate, teeth subequally spaced from each other. Protarsus enlarged, median large claw narrowly split at apex.
<emphasis id="B92CEABC1C6719578F97C7F50AD6F336" bold="true" box="[911,987,1523,1544]" pageId="18" pageNumber="682">Venter:</emphasis>
Prosternal process long, apex flattened into transverse oval surrounded by fringe of long, pale setae and with raised, transversely oval “button” on anterior half; posterior margin of shaft near base with weak swelling.
<emphasis id="B92CEABC1C6719578EB5C47E0A2BF3B3" bold="true" box="[685,806,1656,1677]" pageId="18" pageNumber="682">Parameres:</emphasis>
With long, flange-like tooth at base [best seen in anterolateral and lateral views] (
<figureCitation id="13632A2B1C6719578EADC4AB0A0CF3FC" box="[693,769,1709,1730]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="18" pageNumber="682">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34265251C6619568CDDC2C40803F418" pageId="19" pageNumber="683" type="distribution">
<paragraph id="8BE736AE1C6619568CDDC2C40803F418" blockId="19.[173,660,193,826]" pageId="19" pageNumber="683">
<emphasis id="B92CEABC1C6619568CDDC2C4084BF5E9" bold="true" box="[197,326,194,215]" pageId="19" pageNumber="683">Distribution.</emphasis>
<taxonomicName id="4C584D2D1C6619568D57C2C70B26F5E9" authorityName="Paucar-Cabrera and Ratcliffe" authorityYear="2018" box="[335,555,193,215]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="B92CEABC1C6619568D57C2C70B26F5E9" box="[335,555,193,215]" italics="true" pageId="19" pageNumber="683">Ancognatha uncinata</emphasis>
</taxonomicName>
is known from the area of the Cascada San Rafael (the highest waterfall in
<collectingCountry id="F34F763E1C6619568D64C2F108DCF432" box="[380,465,247,268]" name="Ecuador" pageId="19" pageNumber="683">Ecuador</collectingCountry>
) in
<collectingRegion id="499CF84C1C6619568DE4C2F10B9EF432" box="[508,659,247,268]" country="Ecuador" name="Napo" pageId="19" pageNumber="683">Napo province</collectingRegion>
(
<figureCitation id="13632A2B1C6619568CACC317080FF418" box="[180,258,273,294]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="19" pageNumber="683">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34265251C6619568CDDC32A0B0AF4AF" pageId="19" pageNumber="683" type="etymology">
<paragraph id="8BE736AE1C6619568CDDC32A0B0AF4AF" blockId="19.[173,660,193,826]" pageId="19" pageNumber="683">
<emphasis id="B92CEABC1C6619568CDDC32A083AF47F" bold="true" box="[197,311,300,321]" pageId="19" pageNumber="683">Etymology.</emphasis>
The adjectival epithet
<taxonomicName id="4C584D2D1C6619568E3DC32A0B75F47F" authorityName="Paucar-Cabrera and Ratcliffe" authorityYear="2018" box="[549,632,300,321]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="B92CEABC1C6619568E3DC32A0B75F47F" box="[549,632,300,321]" italics="true" pageId="19" pageNumber="683">uncinata</emphasis>
</taxonomicName>
is from the Latin
<emphasis id="B92CEABC1C6619568D52C341088FF462" box="[330,386,327,348]" italics="true" pageId="19" pageNumber="683">uncus</emphasis>
, meaning a hook, barb, or angle, in reference to the unique hook-like flange at the base of the parameres (
<figureCitation id="13632A2B1C6619568DAAC37A08F6F4AF" box="[434,507,380,401]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="19" pageNumber="683">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34265251C6619568CDDC3900866F604" pageId="19" pageNumber="683" type="diagnosis">
<paragraph id="8BE736AE1C6619568CDDC3900886F7D4" blockId="19.[173,660,193,826]" pageId="19" pageNumber="683">
<emphasis id="B92CEABC1C6619568CDDC3900821F495" bold="true" box="[197,300,406,427]" pageId="19" pageNumber="683">Diagnosis.</emphasis>
<taxonomicName id="4C584D2D1C6619568D2DC3930B09F495" authorityName="Paucar-Cabrera and Ratcliffe" authorityYear="2018" box="[309,516,405,427]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="B92CEABC1C6619568D2DC3930B09F495" box="[309,516,405,427]" italics="true" pageId="19" pageNumber="683">Ancognatha uncinata</emphasis>
</taxonomicName>
is identical to
<taxonomicName id="4C584D2D1C6619568CB5C3B60811F4F8" authorityName="Arrow" authorityYear="1911" box="[173,284,432,454]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B92CEABC1C6619568CB5C3B60811F4F8" box="[173,284,432,454]" italics="true" pageId="19" pageNumber="683">A. vulgaris</emphasis>
</taxonomicName>
in form, color, and sculpturing (including the densely setose pygidium) except for the different parameres (compare
<figureCitation id="13632A2B1C6619568DD1C3E00B18F4C5" box="[457,533,486,507]" captionStart="Figs" captionStartId="18.[175,214,1015,1034]" captionTargetBox="[163,1167,196,997]" captionTargetId="figure-312@18.[154,1169,194,999]" captionTargetPageId="18" captionText="Figs. 3234. Ancognatha uncinata. 32) Habitus; 33) Parameres; 34) Distribution." figureDoi="http://doi.org/10.5281/zenodo.5384509" httpUri="https://zenodo.org/record/5384509/files/figure.png" pageId="19" pageNumber="683">Figs. 33</figureCitation>
and
<figureCitation id="13632A2B1C6619568E5BC3E00B53F4C5" box="[579,606,486,507]" captionStart="Figs" captionStartId="21.[197,236,1004,1023]" captionTargetBox="[185,1188,196,986]" captionTargetId="figure-433@21.[175,1190,194,988]" captionTargetPageId="21" captionText="Figs. 3840. Ancognatha vulgaris. 38) Habitus; 39) Parameres; 40) Distribution in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.5384513" httpUri="https://zenodo.org/record/5384513/files/figure.png" pageId="19" pageNumber="683">39</figureCitation>
). The large, flange-like hook at the base of the parameres of
<taxonomicName id="4C584D2D1C6619568CDCC01C083DF70E" authorityName="Paucar-Cabrera and Ratcliffe" authorityYear="2018" box="[196,304,538,560]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="B92CEABC1C6619568CDCC01C083DF70E" box="[196,304,538,560]" italics="true" pageId="19" pageNumber="683">A. uncinata</emphasis>
</taxonomicName>
is unique among
<taxonomicName id="4C584D2D1C6619568DC9C01C0B99F70E" box="[465,660,538,560]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92CEABC1C6619568DC9C01C0B49F70E" box="[465,580,538,560]" italics="true" pageId="19" pageNumber="683">Ancognatha</emphasis>
species.
</taxonomicName>
Like
<taxonomicName id="4C584D2D1C6619568CF8C0330846F775" authorityName="Arrow" authorityYear="1911" box="[224,331,565,587]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B92CEABC1C6619568CF8C0330846F775" box="[224,331,565,587]" italics="true" pageId="19" pageNumber="683">A. vulgaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C584D2D1C6619568D66C03308F7F775" box="[382,506,565,587]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B92CEABC1C6619568D66C03308F7F775" box="[382,506,565,587]" italics="true" pageId="19" pageNumber="683">A. humeralis</emphasis>
</taxonomicName>
, the mentum is deeply cleft at its apex for at least half of its length. In Endrödi s (1985) key, this species will go as far as couplet 19 based upon the character of the deeply cleft mentum. The form of the parameres will then distinguish it from
<taxonomicName id="4C584D2D1C6619568D7DC0BC08C3F7EE" authorityName="Arrow" authorityYear="1911" box="[357,462,698,720]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B92CEABC1C6619568D7DC0BC08C3F7EE" box="[357,462,698,720]" italics="true" pageId="19" pageNumber="683">A. vulgaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C584D2D1C6619568DE7C0BC0B74F7EE" box="[511,633,698,720]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="683" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B92CEABC1C6619568DE7C0BC0B74F7EE" box="[511,633,698,720]" italics="true" pageId="19" pageNumber="683">A. humeralis</emphasis>
</taxonomicName>
in the following couplets.
</paragraph>
<paragraph id="8BE736AE1C6619568CDDC0F60866F604" blockId="19.[173,660,193,826]" pageId="19" pageNumber="683">
<emphasis id="B92CEABC1C6619568CDDC0F6087EF63B" bold="true" box="[197,371,752,773]" pageId="19" pageNumber="683">Natural History.</emphasis>
Based upon only a single specimen, nothing is known about the natural history of this species.
</paragraph>
</subSubSection>
</treatment>
</document>