214 lines
22 KiB
XML
214 lines
22 KiB
XML
<document id="9CDA185B05D34C9FB67D7A37CAEB9FBF" ID-CLB-Dataset="28429" ID-DOI="10.11646/zootaxa.4564.1.4" ID-GBIF-Dataset="6a6ca226-8343-4208-8cdb-f0bd627c3817" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588770" ID-ZooBank="D0A9801B-8049-4211-A4A7-D7792B9D6936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552176191024" checkinUser="plazi" docAuthor="Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes & Pitts, James P." docDate="2019" docId="03F187A07C1DCD2827B79AD3FE6FFACB" docLanguage="en" docName="zootaxa.4564.1.4.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lomachaeta ptilohyalus Pitts & Manley 2004" docType="treatment" docVersion="15" lastPageNumber="128" masterDocId="FFC8FFD87C06CD332720985CFFE2FFA6" masterDocTitle="Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations" masterLastPageNumber="136" masterPageNumber="101" pageNumber="128" updateTime="1698700719565" updateUser="ExternalLinkService">
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<mods:title id="161B3A516389BC80D1C73AEBBBB56DFA">Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations</mods:title>
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<mods:namePart id="3ADEC239F7DC22736DE4A10DFD9AC8C5">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="FDD81F399CB899D43F75F56DF72F8EA4">Cambra, Roberto A.</mods:namePart>
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<mods:namePart id="5456808118375260DE30703E2E20FF85">Bartholomay, Pedro R.</mods:namePart>
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<mods:namePart id="2C731E31D57912273BDD8E14288FA05E">Luz, David R.</mods:namePart>
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<mods:namePart id="28C4DCBC815B0CB6309A4763DB6E1381">Quintero, Diomedes</mods:namePart>
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<mods:namePart id="790049498C353B51F42831014DD881E9">Pitts, James P.</mods:namePart>
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<mods:title id="4FAD6F18108EB620043405ABF16B1593">Zootaxa</mods:title>
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<mods:date id="ED2B037D3D80E197A8219DB3D068A643">2019</mods:date>
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<mods:number id="64CD3B041C3D9C2E9A3049FBDB08971A">4564</mods:number>
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<treatment id="03F187A07C1DCD2827B79AD3FE6FFACB" ID-DOI="http://doi.org/10.5281/zenodo.3511557" ID-GBIF-Taxon="154366387" ID-Zenodo-Dep="3511557" LSID="urn:lsid:plazi:treatment:03F187A07C1DCD2827B79AD3FE6FFACB" httpUri="http://treatment.plazi.org/id/03F187A07C1DCD2827B79AD3FE6FFACB" lastPageNumber="128" pageId="27" pageNumber="128">
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<taxonomicName id="4C584D357C1DCD2827B79AD3FD25FD0C" authority="Pitts & Manley, 2004" authorityName="Pitts & Manley" authorityYear="2004" box="[151,711,655,682]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="ptilohyalus">
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<emphasis id="B92CEAA47C1DCD2827B79AD3FD25FD0C" bold="true" box="[151,711,655,682]" pageId="27" pageNumber="128">
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<emphasis id="B92CEAA47C1DCD2827B79AD3FE5BFD0F" bold="true" box="[151,441,655,681]" italics="true" pageId="27" pageNumber="128">Lomachaeta ptilohyalus</emphasis>
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<bibRefCitation id="EFC94B477C1DCD2826E09ACCFD25FD0C" author="Pitts, J. P. & Manley, D. G." box="[448,711,656,682]" pageId="27" pageNumber="128" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. & Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts & Manley, 2004</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C342653D7C1DCD2827B79AEEFF1BFD6D" box="[151,249,690,715]" pageId="27" pageNumber="128" type="description">
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<paragraph id="8BE736B67C1DCD2827B79AEEFF1BFD6D" blockId="27.[151,711,655,715]" box="[151,249,690,715]" pageId="27" pageNumber="128">
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(
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<figureCitation id="13632A337C1DCD2827BF9AEEFF13FD6D" box="[159,241,690,715]" captionStart="FIGURES 47–52" captionStartId="25.[151,264,1966,1989]" captionTargetBox="[167,1420,215,1918]" captionTargetId="figure@25.[167,1420,215,1918]" captionTargetPageId="25" captionText="FIGURES 47–52. Lomachaeta males; lateral habitus (47–49, 51, 52) and face view (50). 47) L. beadugrimi; 48) L. powelli; 49–50) L. calamondin, sp. nov.; 51) L. ptilohyalus; 52) L. crocopinna. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588790/files/figure.png" pageId="27" pageNumber="128">Fig. 51</figureCitation>
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)
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</paragraph>
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<subSubSection id="C342653D7C1DCD2827B79AA6FE12FC89" pageId="27" pageNumber="128" type="reference_group">
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<treatmentCitationGroup id="AB4811987C1DCD2827B79AA6FE12FC89" pageId="27" pageNumber="128">
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<taxonomicName id="4C584D357C1DCD2827B79AA6FD74FCB6" authority="Pitts & Manley, 2004: 12" authorityName="Pitts & Manley" authorityPageNumber="12" authorityYear="2004" box="[151,662,762,784]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="ptilohyalus">
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<emphasis id="B92CEAA47C1DCD2827B79AA6FE6EFCB6" box="[151,396,762,784]" italics="true" pageId="27" pageNumber="128">Lomachaeta ptilohyalus</emphasis>
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<bibRefCitation id="EFC94B477C1DCD2826B29AA6FD93FCB6" author="Pitts, J. P. & Manley, D. G." box="[402,625,762,784]" pageId="27" pageNumber="128" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. & Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts & Manley, 2004</bibRefCitation>
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: 12
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.
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<typeStatus id="54E388147C1DCD2825819AA6FCE0FCB6" box="[673,770,762,784]" pageId="27" pageNumber="128" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="9D5EFD3F7C1DCD28242C9AA7FCC3FCB6" box="[780,801,763,784]" pageId="27" pageNumber="128" type="male">♂</specimenCount>
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,
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<collectingCountry id="F34F76267C1DCD28240D9AA6FC9FFCB6" box="[813,893,762,784]" name="Mexico" pageId="27" pageNumber="128">Mexico</collectingCountry>
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,
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<collectingRegion id="499CF8547C1DCD2824A79AA6FC37FCB6" box="[903,981,762,784]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="128">Oaxaca</collectingRegion>
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,
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North of Huajuapan de Leon (CNCI).
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<bibRefCitation id="EFC94B477C1DCD2827E79B45FE4DFC89" author="Pitts, J. P. & Manley, D. G." box="[199,431,793,815]" pageId="27" pageNumber="128" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. & Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts & Manley (2004)</bibRefCitation>
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: Host.
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</treatmentCitationGroup>
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<subSubSection id="C342653D7C1DCD2827B79B00FA84FBA2" pageId="27" pageNumber="128" type="diagnosis">
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<paragraph id="8BE736B67C1DCD2827B79B00FA84FBA2" blockId="27.[151,1437,860,1389]" pageId="27" pageNumber="128">
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<emphasis id="B92CEAA47C1DCD2827B79B00FEF3FCD3" bold="true" box="[151,273,860,885]" pageId="27" pageNumber="128">Diagnosis.</emphasis>
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MALE. The following combination of characters is diagnostic for
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<taxonomicName id="4C584D357C1DCD2823339B01FB4DFCD3" authorityName="Pitts & Manley" authorityYear="2004" box="[1043,1199,860,885]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="ptilohyalus">
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<emphasis id="B92CEAA47C1DCD2823339B01FB4DFCD3" box="[1043,1199,860,885]" italics="true" pageId="27" pageNumber="128">L. ptilohyalus</emphasis>
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</taxonomicName>
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: the body is entirely black, except T2–3 are largely orange; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the mesoscutum has sparse punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the T2 fringe is composed of simple setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length
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<quantity id="4CA09B537C1DCD2822219BB0FA80FBA2" box="[1281,1378,1004,1029]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="27" pageNumber="128" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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.
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</paragraph>
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<subSubSection id="C342653D7C1DCD2827E79C68FBBAFBD6" pageId="27" pageNumber="128" type="materials_examined">
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<paragraph id="8BE736B67C1DCD2827E79C68FBBAFBD6" blockId="27.[151,1437,860,1389]" pageId="27" pageNumber="128">
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<emphasis id="B92CEAA47C1DCD2827E79C68FE52FBEB" bold="true" box="[199,432,1076,1101]" pageId="27" pageNumber="128">Material examined.</emphasis>
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<materialsCitation id="3B303CEB7C1DCD2826969C68FBB6FBD6" ID-GBIF-Occurrence="2012757477" collectingDate="2010-05-25" collectingDateMax="2010-06-03" collectingDateMin="2010-05-25" collectionCode="CSCA" collectorName="S. Arlington & M. E. Irwin" country="United States of America" county="Maricopa County" elevation="200" latitude="33.22167" location="Gila River" longLatPrecision="92" longitude="-112.758835" pageId="27" pageNumber="128" specimenCount="1" specimenCount-male="1" stateProvince="Arizona">
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<collectingCountry id="F34F76267C1DCD2826969C68FE10FBEA" box="[438,498,1076,1100]" name="United States of America" pageId="27" pageNumber="128">USA</collectingCountry>
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:
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<emphasis id="B92CEAA47C1DCD2826DD9C68FDB4FBED" box="[509,598,1076,1099]" italics="true" pageId="27" pageNumber="128">
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<collectingRegion id="499CF8547C1DCD2826DD9C68FDB4FBED" box="[509,598,1076,1099]" country="United States of America" name="Arizona" pageId="27" pageNumber="128">Arizona</collectingRegion>
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</emphasis>
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:
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<collectingCounty id="62864E3A7C1DCD2825449C68FCC8FBEA" box="[612,810,1076,1101]" pageId="27" pageNumber="128">Maricopa County</collectingCounty>
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,
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<location id="8E87606D7C1DCD2824149C68FC49FBEB" LSID="urn:lsid:plazi:treatment:03F187A07C1DCD2827B79AD3FE6FFACB:8E87606D7C1DCD2824149C68FC49FBEB" box="[820,939,1076,1101]" country="United States of America" county="Maricopa County" latitude="33.22167" longLatPrecision="92" longitude="-112.758835" name="Gila River" pageId="27" pageNumber="128" stateProvince="Arizona">Gila River</location>
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,
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<quantity id="4CA09B537C1DCD2824969C68FC1FFBEB" box="[950,1021,1076,1101]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="128" unit="km" value="10.0">10 km</quantity>
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<collectorName id="26AD53607C1DCD2823239C68FB72FBEB" box="[1027,1168,1076,1101]" pageId="27" pageNumber="128">S. Arlington</collectorName>
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, malaise on sand beach,
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<elevation id="0075D1857C1DCD2827B79C04FF02FBC9" box="[151,224,1112,1136]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="128" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
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,
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<date id="FFE610767C1DCD28252B9C0BFD30FBD6" box="[523,722,1111,1136]" pageId="27" pageNumber="128" value="2010-05-25" valueMax="2010-06-03" valueMin="2010-05-25">
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<collectingDate id="EFA2E99E7C1DCD28252B9C0BFD30FBD6" box="[523,722,1111,1136]" pageId="27" pageNumber="128" value="2010-05-25" valueMax="2010-06-03" valueMin="2010-05-25">25.V–03.VI.2010</collectingDate>
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</date>
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||
,
|
||
<collectorName id="26AD53607C1DCD2825FE9C04FC82FBD6" box="[734,864,1111,1136]" pageId="27" pageNumber="128">M. E. Irwin</collectorName>
|
||
(
|
||
<specimenCount id="9D5EFD3F7C1DCD2824519C04FC77FBD6" box="[881,917,1112,1136]" pageId="27" pageNumber="128" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE737C1DCD2824819C04FC0FFBD6" box="[929,1005,1112,1136]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="27" pageNumber="128" type="Museum">CSCA</collectionCode>
|
||
,
|
||
<figureCitation id="13632A337C1DCD2824D89C0BFBA9FBD6" box="[1016,1099,1111,1136]" captionStart="FIGURES 47–52" captionStartId="25.[151,264,1966,1989]" captionTargetBox="[167,1420,215,1918]" captionTargetId="figure@25.[167,1420,215,1918]" captionTargetPageId="25" captionText="FIGURES 47–52. Lomachaeta males; lateral habitus (47–49, 51, 52) and face view (50). 47) L. beadugrimi; 48) L. powelli; 49–50) L. calamondin, sp. nov.; 51) L. ptilohyalus; 52) L. crocopinna. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588790/files/figure.png" pageId="27" pageNumber="128">Fig. 51</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C1DCD2827E79C20FC91FB1E" pageId="27" pageNumber="128" type="distribution">
|
||
<paragraph id="8BE736B67C1DCD2827E79C20FC91FB1E" blockId="27.[151,1437,860,1389]" pageId="27" pageNumber="128">
|
||
<emphasis id="B92CEAA47C1DCD2827E79C20FE82FB33" bold="true" box="[199,352,1148,1173]" pageId="27" pageNumber="128">Distribution.</emphasis>
|
||
This species has an apparently disjunct distribution in the Mojave and Sonoran Deserts (
|
||
<collectingRegion id="499CF8547C1DCD2822609C20FA7EFB33" box="[1344,1436,1148,1173]" country="United States of America" name="Arizona" pageId="27" pageNumber="128">Arizona</collectingRegion>
|
||
and
|
||
<collectingRegion id="499CF8547C1DCD2827E69CC3FED9FB1E" box="[198,315,1183,1208]" country="United States of America" name="California" pageId="27" pageNumber="128">California</collectingRegion>
|
||
, (
|
||
<collectingCountry id="F34F76267C1DCD28266E9CFCFE6BFB1E" box="[334,393,1184,1208]" name="United States of America" pageId="27" pageNumber="128">USA</collectingCountry>
|
||
) and Balsas Dry Forests (
|
||
<collectingRegion id="499CF8547C1DCD2825899CFCFCE3FB1E" box="[681,769,1184,1208]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="128">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34F76267C1DCD28242D9CC3FC8AFB1E" box="[781,872,1183,1208]" name="Mexico" pageId="27" pageNumber="128">Mexico</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C1DCD2827E79C98FE6FFACB" pageId="27" pageNumber="128" type="discussion">
|
||
<paragraph id="8BE736B67C1DCD2827E79C98FE6FFACB" blockId="27.[151,1437,860,1389]" pageId="27" pageNumber="128">
|
||
<emphasis id="B92CEAA47C1DCD2827E79C98FED9FB7B" bold="true" box="[199,315,1220,1245]" pageId="27" pageNumber="128">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54E388147C1DCD2826559C98FE35FB7B" box="[373,471,1220,1245]" pageId="27" pageNumber="128" type="holotype">holotype</typeStatus>
|
||
from
|
||
<collectingRegion id="499CF8547C1DCD28253D9C98FD96FB7A" box="[541,628,1220,1244]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="128">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34F76267C1DCD2825A09C98FD37FB7B" box="[640,725,1220,1245]" name="Mexico" pageId="27" pageNumber="128">Mexico</collectingCountry>
|
||
is separated from the other known specimens of
|
||
<taxonomicName id="4C584D357C1DCD2822219C99FA7EFB7B" authorityName="Pitts & Manley" authorityYear="2004" box="[1281,1436,1220,1245]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="ptilohyalus">
|
||
<emphasis id="B92CEAA47C1DCD2822219C99FA7EFB7B" box="[1281,1436,1220,1245]" italics="true" pageId="27" pageNumber="128">L. ptilohyalus</emphasis>
|
||
</taxonomicName>
|
||
in hot deserts of the
|
||
<collectingCountry id="F34F76267C1DCD2826B39CB4FE29FAA6" box="[403,459,1256,1280]" name="United States of America" pageId="27" pageNumber="128">USA</collectingCountry>
|
||
by over
|
||
<quantity id="4CA09B537C1DCD28251C9CB4FD45FAA6" box="[572,679,1255,1280]" metricMagnitude="6" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="128" unit="km" value="2300.0">2300 km</quantity>
|
||
. Further collections in
|
||
<collectingCountry id="F34F76267C1DCD28249C9CBBFBF3FAA6" box="[956,1041,1255,1280]" name="Mexico" pageId="27" pageNumber="128">Mexico</collectingCountry>
|
||
may reveal other populations, or molecular data comparisons between Oaxacan and hot desert specimens may reveal that they are not conspecific. Either way, this is a compelling link between disjunct arid habitats of southern Pacific
|
||
<collectingCountry id="F34F76267C1DCD2823B99D73FB0CFAEE" box="[1177,1262,1327,1352]" name="Mexico" pageId="27" pageNumber="128">Mexico</collectingCountry>
|
||
and the North American hot deserts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |