378 lines
40 KiB
XML
378 lines
40 KiB
XML
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<mods:title id="C2F9B4D5EA54ABD71693582BB0F0CA85">Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations</mods:title>
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<mods:namePart id="F62A6231B7A851D6C2C9CC28498D5EFF">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="CB0A162A01A1534FCDFC581A76EC9C3D">Cambra, Roberto A.</mods:namePart>
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<mods:namePart id="8891CA045202BCDD4EB799FDB12C6BCD">Bartholomay, Pedro R.</mods:namePart>
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<mods:namePart id="D85B2DB6ABF904CAB565CC8377FBB7FC">Luz, David R.</mods:namePart>
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<mods:namePart id="C417CA0DA0945EFDF658A95CF4B4CCE4">Quintero, Diomedes</mods:namePart>
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<mods:namePart id="AFECDBCA283DF1F080D49473B174AA82">Pitts, James P.</mods:namePart>
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<mods:title id="D1E867C8C1336CA083189032E53AC7C6">Zootaxa</mods:title>
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<mods:date id="9FE4177CDA594ABB1300552CC3DBFA47">2019</mods:date>
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<treatment id="03F187A07C18CD1227B79F10FC40FC5E" ID-DOI="http://doi.org/10.5281/zenodo.5943949" ID-GBIF-Taxon="154366378" ID-Zenodo-Dep="5943949" LSID="urn:lsid:plazi:treatment:03F187A07C18CD1227B79F10FC40FC5E" httpUri="http://treatment.plazi.org/id/03F187A07C18CD1227B79F10FC40FC5E" lastPageId="33" lastPageNumber="134" pageId="30" pageNumber="131">
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<subSubSection id="C342653D7C18CD2D27B79F10FD97F8C0" box="[151,629,1868,1894]" pageId="30" pageNumber="131" type="nomenclature">
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<paragraph id="8BE736B67C18CD2D27B79F10FD97F8C0" blockId="30.[151,629,1868,1927]" box="[151,629,1868,1894]" pageId="30" pageNumber="131">
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<heading id="D0AF81DA7C18CD2D27B79F10FD97F8C0" bold="true" box="[151,629,1868,1894]" fontSize="11" level="1" pageId="30" pageNumber="131" reason="1">
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<emphasis id="B92CEAA47C18CD2D27B79F10FD97F8C0" bold="true" box="[151,629,1868,1894]" pageId="30" pageNumber="131">
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<taxonomicName id="4C584D357C18CD2D27B79F10FDE9F8C0" authority="Williams" authorityName="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts" authorityYear="2019" box="[151,523,1868,1894]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="warneri" status="sp. nov.">
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<emphasis id="B92CEAA47C18CD2D27B79F10FE70F8C0" bold="true" box="[151,402,1868,1894]" italics="true" pageId="30" pageNumber="131">Lomachaeta warneri</emphasis>
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Williams
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21F57DF7C18CD2D25389F10FD97F8C0" box="[536,629,1868,1894]" pageId="30" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342653D7C18CD2D27B79F32FEC5F821" box="[151,295,1902,1927]" pageId="30" pageNumber="131" type="description">
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<paragraph id="8BE736B67C18CD2D27B79F32FEC5F821" blockId="30.[151,629,1868,1927]" box="[151,295,1902,1927]" pageId="30" pageNumber="131">
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(
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<figureCitation id="13632A337C18CD2D27BF9F32FF17F821" box="[159,245,1902,1927]" captionStart="FIGURES 33–36" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 33–36. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="30" pageNumber="131">Figs 36</figureCitation>
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,
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<figureCitation id="13632A337C18CD2D26219F33FEFDF821" box="[257,287,1903,1927]" captionStart="FIGURES 37–40" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 37–40. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="30" pageNumber="131">40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C342653D7C18CD2C27B79FEAFB75FEE6" lastPageId="31" lastPageNumber="132" pageId="30" pageNumber="131" type="diagnosis">
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<paragraph id="8BE736B67C18CD2C27B79FEAFB75FEE6" blockId="30.[151,1436,1974,2036]" lastBlockId="31.[151,1437,151,320]" lastPageId="31" lastPageNumber="132" pageId="30" pageNumber="131">
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<emphasis id="B92CEAA47C18CD2D27B79FEAFEF3F869" bold="true" box="[151,273,1974,1999]" pageId="30" pageNumber="131">Diagnosis.</emphasis>
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FEMALE. This species can be recognized by the following combination of characters: the body is predominantly orange-brown, except the antennae, legs, and T3–6 are somewhat darker brown; the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the erect dorsal brachyplumose setae of the whole body are mostly yellow-brown; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the T2 disc is sparsely punctate with the intervals partly smooth and partly microreticulate; the T2 fringe is somewhat dense with slightly thickened silvery setae; and the S6 lateral carina is shallow, but distinct.
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</paragraph>
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</subSubSection>
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<caption id="DF27663E7C19CD2C27B79E40FF0FF9F7" httpUri="https://zenodo.org/record/2588796/files/figure.png" pageId="31" pageNumber="132" startId="31.[151,264,1564,1587]" targetBox="[167,1420,373,1543]" targetPageId="31">
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<paragraph id="8BE736B67C19CD2C27B79E40FF0FF9F7" blockId="31.[151,1435,1564,1617]" pageId="31" pageNumber="132">
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<emphasis id="B92CEAA47C19CD2C27B79E40FEB6F994" bold="true" box="[151,340,1564,1587]" pageId="31" pageNumber="132">FIGURES 61–64.</emphasis>
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<taxonomicName id="4C584D357C19CD2C26799E40FE34F994" authorityName="Mickel" authorityYear="1936" box="[345,470,1564,1586]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAA47C19CD2C26799E40FE34F994" box="[345,470,1564,1586]" italics="true" pageId="31" pageNumber="132">Lomachaeta</emphasis>
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</taxonomicName>
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males; lateral habitus. 61)
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<taxonomicName id="4C584D357C19CD2C25C89E42FCC5F994" authorityName="Williams & Pitts" authorityYear="2009" box="[744,807,1564,1587]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="ilex">
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<emphasis id="B92CEAA47C19CD2C25C89E42FCC5F994" box="[744,807,1564,1587]" italics="true" pageId="31" pageNumber="132">L. ilex</emphasis>
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</taxonomicName>
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; 62)
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<taxonomicName id="4C584D357C19CD2C247C9E42FC5EF994" authorityName="Williams & Pitts" authorityYear="2009" box="[860,956,1564,1587]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="hedera">
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<emphasis id="B92CEAA47C19CD2C247C9E42FC5EF994" box="[860,956,1564,1587]" italics="true" pageId="31" pageNumber="132">L. hedera</emphasis>
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</taxonomicName>
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; 63)
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<taxonomicName id="4C584D357C19CD2C24D29E42FB88F994" authorityName="Williams & Pitts" authorityYear="2009" box="[1010,1130,1564,1587]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="litosisyra">
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<emphasis id="B92CEAA47C19CD2C24D29E42FB88F994" box="[1010,1130,1564,1587]" italics="true" pageId="31" pageNumber="132">L. litosisyra</emphasis>
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</taxonomicName>
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; 64)
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<taxonomicName id="4C584D357C19CD2C23809E42FAD0F995" authorityName="Williams & Pitts" authorityYear="2009" box="[1184,1330,1566,1587]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="vacamuerta">
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<emphasis id="B92CEAA47C19CD2C23809E42FAD0F995" box="[1184,1330,1566,1587]" italics="true" pageId="31" pageNumber="132">L. vacamuerta</emphasis>
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</taxonomicName>
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. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="C342653D7C19CD1227E79E22FE78FE0B" lastPageId="33" lastPageNumber="134" pageId="31" pageNumber="132" type="description">
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<paragraph id="8BE736B67C19CD1227E79E22FC2FFE2E" blockId="31.[151,1436,1662,1975]" lastBlockId="33.[151,1437,151,1016]" lastPageId="33" lastPageNumber="134" pageId="31" pageNumber="132">
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<emphasis id="B92CEAA47C19CD2C27E79E22FEBBF931" bold="true" box="[199,345,1662,1687]" pageId="31" pageNumber="132">Description.</emphasis>
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FEMALE. Body length
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<quantity id="4CA09B537C19CD2C25529E22FD0AF930" box="[626,744,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="31" pageNumber="132" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
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.
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<emphasis id="B92CEAA47C19CD2C25D49E22FC8DF931" box="[756,879,1662,1687]" italics="true" pageId="31" pageNumber="132">Coloration</emphasis>
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. Body reddish-orange except F2–10 dark brown; femora, tibiae, and tarsi largely brown; T2 apex and T3–5 brown; and T6 dark brown. Tibial spurs white. Erect dorsal brachyplumose setae entirely white to pale brown, many setae lighter at base; except T6 with setae predominantly brown. Mesonotum with few scattered subappressed simple white setae; fringes of T2–5 composed of subappressed white setae, those of T2 silvery, thicker than surrounding erect setae. Ventral and pleural setae white, except S6 setae largely brown.
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<emphasis id="B92CEAA47C19CD2C25719F6EFD6FF8ED" box="[593,653,1842,1867]" italics="true" pageId="31" pageNumber="132">Head</emphasis>
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. Head width 1.1 × pronotal width. Frons, vertex and gena areolate, vertex with many intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with barely perceptible ventral lamella that does not interrupt ventral mandibular contour. Antennal scrobe without dorsal carina. Lengths of F1 and F2 each subequal to pedicel length.
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<emphasis id="B92CEAA47C27CD12253498C5FD6EFF16" box="[532,652,153,176]" italics="true" pageId="33" pageNumber="134">Mesosoma</emphasis>
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. Mesosoma compact; dorsal thoracic length 0.85 × width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; areolations clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate.
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<emphasis id="B92CEAA47C27CD1225B39975FCEAFEE6" box="[659,776,297,320]" italics="true" pageId="33" pageNumber="134">Metasoma</emphasis>
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. T1 shape sub-sessile, mostly smooth. T2 slightly longer than wide. Disc of T2 with sparse punctures, intervals smooth. T3–5 and S2–5 with separated punctures, intervals shagreened. T6 convex. Incomplete lateral S6 carina shallow but distinct.
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</paragraph>
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<caption id="DF27663E7C26CD1327B79FD0FEB4F846" httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="32" pageNumber="133" startId="32.[151,264,1932,1955]" targetBox="[191,1396,193,1898]" targetPageId="32">
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<paragraph id="8BE736B67C26CD1327B79FD0FEB4F846" blockId="32.[151,1436,1932,2016]" pageId="32" pageNumber="133">
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<emphasis id="B92CEAA47C26CD1327B79FD0FEB6F804" bold="true" box="[151,340,1932,1955]" pageId="32" pageNumber="133">FIGURES 65–72.</emphasis>
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<taxonomicName id="4C584D357C26CD13267B9FD1FE3AF805" authorityName="Mickel" authorityYear="1936" box="[347,472,1933,1955]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAA47C26CD13267B9FD1FE3AF805" box="[347,472,1933,1955]" italics="true" pageId="32" pageNumber="133">Lomachaeta</emphasis>
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</taxonomicName>
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line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65)
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<emphasis id="B92CEAA47C26CD13269D9FF1FDF9F864" box="[445,539,1964,1986]" italics="true" pageId="32" pageNumber="133">
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<taxonomicName id="4C584D357C26CD13269D9FF1FDF6F864" box="[445,532,1964,1986]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="hicksi">L. hicksi</taxonomicName>
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;
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</emphasis>
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66)
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<taxonomicName id="4C584D357C26CD1325699FF1FD3AF864" authorityName="Pitts & Manley" authorityYear="2004" box="[585,728,1964,1986]" class="Insecta" family="Mutillidae" genus="Smicromutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="beadugrimi">
|
||
<emphasis id="B92CEAA47C26CD1325699FF1FD3AF864" box="[585,728,1964,1986]" italics="true" pageId="32" pageNumber="133">L. beadugrimi</emphasis>
|
||
</taxonomicName>
|
||
; 67–68)
|
||
<emphasis id="B92CEAA47C26CD1324139FF1FC29F864" box="[819,971,1964,1986]" italics="true" pageId="32" pageNumber="133">
|
||
<taxonomicName id="4C584D357C26CD1324139FF1FC25F864" authority="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts, 2019" authorityName="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts" authorityYear="2019" box="[819,967,1964,1986]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="calamondin" status="sp. nov.">L. calamondin</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B92CEAA47C26CD1324F19FF0FBC3F867" bold="true" box="[977,1057,1964,1985]" pageId="32" pageNumber="133">
|
||
<taxonomicNameLabel id="A21F57DF7C26CD1324F19FF0FBC3F867" box="[977,1057,1964,1985]" pageId="32" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 69–70)
|
||
<emphasis id="B92CEAA47C26CD13235C9FF1FB12F864" box="[1148,1264,1965,1986]" italics="true" pageId="32" pageNumber="133">
|
||
<taxonomicName id="4C584D357C26CD13235C9FF1FB0FF864" authority="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts, 2019" authorityName="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts" authorityYear="2019" box="[1148,1261,1965,1986]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="eironeia" status="sp. nov.">L. eironeia</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B92CEAA47C26CD1323D69FF0FAA5F867" bold="true" box="[1270,1351,1964,1985]" pageId="32" pageNumber="133">
|
||
<taxonomicNameLabel id="A21F57DF7C26CD1323D69FF0FAA5F867" box="[1270,1351,1964,1985]" pageId="32" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 71–72)
|
||
<emphasis id="B92CEAA47C26CD1327B79F97FEE2F846" box="[151,256,1995,2016]" italics="true" pageId="32" pageNumber="133">
|
||
<taxonomicName id="4C584D357C26CD1327B79F97FF1EF846" authority="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts, 2019" authorityName="Williams & Cambra & Bartholomay & Luz & Quintero & Pitts" authorityYear="2019" box="[151,252,1995,2016]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="juanita" status="sp. nov.">L. juanita</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A21F57DF7C26CD1326269F97FEB4F846" box="[262,342,1995,2016]" pageId="32" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE736B67C27CD1227E799C8FE78FE0B" blockId="33.[151,1437,151,1016]" box="[199,410,404,429]" pageId="33" pageNumber="134">MALE. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C27CD1227E799EBFDCCFD15" pageId="33" pageNumber="134" type="materials_examined">
|
||
<paragraph id="8BE736B67C27CD1227E799EBFDCCFD15" blockId="33.[151,1437,151,1016]" pageId="33" pageNumber="134">
|
||
<emphasis id="B92CEAA47C27CD1227E799EBFE50FE76" bold="true" box="[199,434,439,464]" pageId="33" pageNumber="134">Material examined.</emphasis>
|
||
<materialsCitation id="3B303CEB7C27CD12269A99EBFF22FDB8" ID-GBIF-Occurrence="2012757484" collectingDate="2014-04-24" collectingDateMax="2014-05-25" collectingDateMin="2014-04-24" collectionCode="CSCA" collectorName="W. B. Warner" country="United States of America" county="Coconino County" latitude="36.922222" location="Coconino County" longLatPrecision="19" longitude="-112.25278" pageId="33" pageNumber="134" specimenCount="1" specimenCount-female="1" stateProvince="Arizona" typeStatus="holotype">
|
||
<emphasis id="B92CEAA47C27CD12269A99EBFDC2FE76" box="[442,544,439,464]" italics="true" pageId="33" pageNumber="134">
|
||
<typeStatus id="54E388147C27CD12269A99EBFDC2FE76" box="[442,544,439,464]" pageId="33" pageNumber="134" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<specimenCount id="9D5EFD3F7C27CD12251099E4FDA7FE69" box="[560,581,440,463]" pageId="33" pageNumber="134" type="female">♀</specimenCount>
|
||
,
|
||
<collectingCountry id="F34F76267C27CD12257499E4FD6DFE76" box="[596,655,440,464]" name="United States of America" pageId="33" pageNumber="134">USA</collectingCountry>
|
||
,
|
||
<emphasis id="B92CEAA47C27CD1225BC99E4FD17FE69" box="[668,757,440,463]" italics="true" pageId="33" pageNumber="134">
|
||
<collectingRegion id="499CF8547C27CD1225BC99E4FD17FE69" box="[668,757,440,463]" country="United States of America" name="Arizona" pageId="33" pageNumber="134">Arizona</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingCounty id="62864E3A7C27CD12242499EBFC28FE76" box="[772,970,439,464]" pageId="33" pageNumber="134">Coconino County</collectingCounty>
|
||
, Hwy. 89A,
|
||
<quantity id="4CA09B537C27CD12237D99E4FB4FFE76" box="[1117,1197,439,464]" metricMagnitude="3" metricUnit="m" metricValue="7.8857856" pageId="33" pageNumber="134" unit="mi" value="4.9">4.9 mi.</quantity>
|
||
[
|
||
<quantity id="4CA09B537C27CD12239C99E4FAF3FE76" box="[1212,1297,439,464]" metricMagnitude="3" metricUnit="m" metricValue="7.8" pageId="33" pageNumber="134" unit="km" value="7.8">7.8 km</quantity>
|
||
] ESE of jct. 389,
|
||
<geoCoordinate id="EE6C50717C27CD1227F19982FEB5FE51" box="[209,343,475,503]" direction="north" orientation="latitude" pageId="33" pageNumber="134" precision="15" value="36.922222">
|
||
36
|
||
<superScript id="7C2D9BFE7C27CD1227CD9987FF16FE4F" attach="none" box="[237,244,475,489]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
55’20”N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE6C50717C27CD1226419983FE1BFE51" box="[353,505,475,503]" direction="west" orientation="longitude" pageId="33" pageNumber="134" precision="15" value="-112.25278">
|
||
112
|
||
<superScript id="7C2D9BFE7C27CD1226AA9987FE73FE4F" attach="none" box="[394,401,475,489]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
15’10”W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFE610767C27CD1225289982FD29FE51" box="[520,715,478,503]" pageId="33" pageNumber="134" value="2014-04-24" valueMax="2014-05-25" valueMin="2014-04-24">
|
||
<collectingDate id="EFA2E99E7C27CD1225289982FD29FE51" box="[520,715,478,503]" pageId="33" pageNumber="134" value="2014-04-24" valueMax="2014-05-25" valueMin="2014-04-24">24.IV–25.V.2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26AD53607C27CD1225F999BCFC8DFE51" box="[729,879,480,503]" pageId="33" pageNumber="134">W.B. Warner</collectorName>
|
||
, barrier pitfall with black cups (
|
||
<collectionCode id="ED49AE737C27CD1223C49983FAD0FE51" box="[1252,1330,479,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="33" pageNumber="134" type="Museum">CSCA</collectionCode>
|
||
,
|
||
<figureCitation id="13632A337C27CD12221F9982FA75FE51" box="[1343,1431,478,503]" captionStart="FIGURES 33–36" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 33–36. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="33" pageNumber="134">Figs 36</figureCitation>
|
||
,
|
||
<figureCitation id="13632A337C27CD1227B79A5AFF55FDB8" box="[151,183,518,542]" captionStart="FIGURES 37–40" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 37–40. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="33" pageNumber="134">40</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B303CEB7C27CD1227EF9A5AFAA8FDE3" ID-GBIF-Occurrence="2012757476" collectingDate="2014-06-29" collectingDateMax="2014-07-26" collectingDateMin="2014-06-29" collectionCode="ASUT, FSCA" collectorName="W. B. Warner" country="United States of America" county="Coconino County" latitude="36.922222" location="Hwy." longLatPrecision="19" longitude="-112.25278" pageId="33" pageNumber="134" specimenCount="2" specimenCount-female="2" stateProvince="Arizona" typeStatus="paratype">
|
||
<emphasis id="B92CEAA47C27CD1227EF9A5AFEA2FDBB" box="[207,320,518,541]" italics="true" pageId="33" pageNumber="134">
|
||
<typeStatus id="54E388147C27CD1227EF9A5AFEA2FDBB" box="[207,320,518,541]" pageId="33" pageNumber="134" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="F34F76267C27CD1226769A5AFE70FDB8" box="[342,402,518,542]" name="United States of America" pageId="33" pageNumber="134">USA</collectingCountry>
|
||
:
|
||
<emphasis id="B92CEAA47C27CD1226829A5AFE19FDBB" box="[418,507,518,541]" italics="true" pageId="33" pageNumber="134">
|
||
<collectingRegion id="499CF8547C27CD1226829A5AFE19FDBB" box="[418,507,518,541]" country="United States of America" name="Arizona" pageId="33" pageNumber="134">Arizona</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingCounty id="62864E3A7C27CD12252E9A59FD3AFDB8" box="[526,728,517,542]" pageId="33" pageNumber="134">Coconino County</collectingCounty>
|
||
,
|
||
<location id="8E87606D7C27CD1225CB9A5AFCC5FDB8" LSID="urn:lsid:plazi:treatment:03F187A07C18CD1227B79F10FC40FC5E:8E87606D7C27CD1225CB9A5AFCC5FDB8" box="[747,807,518,542]" country="United States of America" county="Coconino County" latitude="36.922222" longLatPrecision="19" longitude="-112.25278" name="Hwy." pageId="33" pageNumber="134" stateProvince="Arizona">Hwy.</location>
|
||
89A,
|
||
<quantity id="4CA09B537C27CD1224569A5AFC28FDB8" box="[886,970,517,542]" metricMagnitude="3" metricUnit="m" metricValue="7.8857856" pageId="33" pageNumber="134" unit="mi" value="4.9">4.9 mi.</quantity>
|
||
[
|
||
<quantity id="4CA09B537C27CD1224FD9A5AFBD4FDB8" box="[989,1078,517,542]" metricMagnitude="3" metricUnit="m" metricValue="7.8" pageId="33" pageNumber="134" unit="km" value="7.8">7.8 km</quantity>
|
||
] ESE of jct. 389,
|
||
<geoCoordinate id="EE6C50717C27CD1222369A59FA79FDB8" box="[1302,1435,514,542]" direction="north" orientation="latitude" pageId="33" pageNumber="134" precision="15" value="36.922222">
|
||
36
|
||
<superScript id="7C2D9BFE7C27CD1222129A5EFADBFDB6" attach="none" box="[1330,1337,514,528]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
55’20”N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE6C50717C27CD1227B79A71FECDFDE3" box="[151,303,553,581]" direction="west" orientation="longitude" pageId="33" pageNumber="134" precision="15" value="-112.25278">
|
||
112
|
||
<superScript id="7C2D9BFE7C27CD1227E09A75FF25FD91" attach="right" box="[192,199,553,567]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
15’10”W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFE610767C27CD1226639A70FDFFFDE3" box="[323,541,556,581]" pageId="33" pageNumber="134" value="2014-06-29" valueMax="2014-07-26" valueMin="2014-06-29">
|
||
<collectingDate id="EFA2E99E7C27CD1226639A70FDFFFDE3" box="[323,541,556,581]" pageId="33" pageNumber="134" value="2014-06-29" valueMax="2014-07-26" valueMin="2014-06-29">29.VI–26.VII.2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26AD53607C27CD1225109A71FD29FDE2" box="[560,715,557,580]" pageId="33" pageNumber="134">W.B. Warner</collectorName>
|
||
, barrier pitfall with black cups (
|
||
<specimenCount id="9D5EFD3F7C27CD1223429A71FB65FDE3" box="[1122,1159,557,581]" pageId="33" pageNumber="134" type="female">2♀</specimenCount>
|
||
:
|
||
<collectionCode id="ED49AE737C27CD1223BA9A71FB06FDE3" box="[1178,1252,557,581]" country="USA" name="Frank M. Hasbrouck Insect Collection" pageId="33" pageNumber="134">ASUT</collectionCode>
|
||
,
|
||
<collectionCode id="ED49AE737C27CD1223D79A71FAA0FDE3" box="[1271,1346,557,581]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="33" pageNumber="134" type="Museum">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B303CEB7C27CD12227D9A70FDD5FD29" ID-GBIF-Occurrence="2012757481" collectingDate="2015-01" collectingDateMax="2015-07-03" collectingDateMin="2015-01" collectionCode="CSCA" country="United States of America" latitude="37.2649" location="Mills Rd." longLatPrecision="6" longitude="-113.3259" municipality="County" pageId="33" pageNumber="134" specimenCount="1" specimenCount-female="1" stateProvince="Utah" typeStatus="paratype">
|
||
<emphasis id="B92CEAA47C27CD12227D9A70FA77FDE3" box="[1373,1429,556,581]" italics="true" pageId="33" pageNumber="134">
|
||
<collectingRegion id="499CF8547C27CD12227D9A70FA77FDE3" box="[1373,1429,556,581]" country="United States of America" name="Utah" pageId="33" pageNumber="134">Utah</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="499CF8547C27CD1227B79A0FFEFEFDCA" box="[151,284,595,620]" country="United States of America" name="Washington" pageId="33" pageNumber="134">Washington</collectingRegion>
|
||
<collectingMunicipality id="6B83ACCC7C27CD1226049A0FFE95FDCD" box="[292,375,595,619]" pageId="33" pageNumber="134">County</collectingMunicipality>
|
||
,
|
||
<location id="8E87606D7C27CD1226A39A0FFE12FDCA" LSID="urn:lsid:plazi:treatment:03F187A07C18CD1227B79F10FC40FC5E:8E87606D7C27CD1226A39A0FFE12FDCA" box="[387,496,595,620]" country="United States of America" latitude="37.2649" longLatPrecision="6" longitude="-113.3259" municipality="County" name="Mills Rd." pageId="33" pageNumber="134" stateProvince="Utah">Mills Rd.</location>
|
||
,
|
||
<quantity id="4CA09B537C27CD1226DC9A0FFDA9FDCA" box="[508,587,595,620]" metricMagnitude="3" metricUnit="m" metricValue="2.5749504" pageId="33" pageNumber="134" unit="mi" value="1.6">1.6 mi.</quantity>
|
||
[
|
||
<quantity id="4CA09B537C27CD1225799A0FFD4FFDCA" box="[601,685,595,620]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="33" pageNumber="134" unit="km" value="2.6">2.6 km</quantity>
|
||
] S
|
||
<date id="FFE610767C27CD1225F29A0FFCE2FDCA" box="[722,768,595,620]" pageId="33" pageNumber="134" value="2015-01">
|
||
<collectingDate id="EFA2E99E7C27CD1225F29A0FFCE2FDCA" box="[722,768,595,620]" pageId="33" pageNumber="134" value="2015-01">I-15</collectingDate>
|
||
</date>
|
||
exit 27;
|
||
<geoCoordinate id="EE6C50717C27CD1224459A0FFC39FDCD" box="[869,987,591,620]" direction="north" orientation="latitude" pageId="33" pageNumber="134" precision="5" value="37.2649">
|
||
37.2649
|
||
<superScript id="7C2D9BFE7C27CD1224E09A13FC25FDFB" attach="right" box="[960,967,591,605]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE6C50717C27CD1224C39A0FFB8CFDCD" box="[995,1134,591,620]" direction="west" orientation="longitude" pageId="33" pageNumber="134" precision="5" value="-113.3259">
|
||
113.3259
|
||
<superScript id="7C2D9BFE7C27CD12236B9A13FBB0FDFB" attach="right" box="[1099,1106,591,605]" fontSize="6" pageId="33" pageNumber="134">o</superScript>
|
||
W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFE610767C27CD1223599A0FFAA7FDCA" box="[1145,1349,595,620]" pageId="33" pageNumber="134" value="2015-04-15" valueMax="2015-07-03" valueMin="2015-04-15">
|
||
<collectingDate id="EFA2E99E7C27CD1223599A0FFAA7FDCA" box="[1145,1349,595,620]" pageId="33" pageNumber="134" value="2015-04-15" valueMax="2015-07-03" valueMin="2015-04-15">15.IV–3.VII.2015</collectingDate>
|
||
</date>
|
||
, barrier pitfall with black cups (
|
||
<specimenCount id="9D5EFD3F7C27CD1226909A2BFE33FD29" box="[432,465,631,655]" pageId="33" pageNumber="134" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE737C27CD1226C19A2BFDCDFD29" box="[481,559,631,655]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="33" pageNumber="134" type="Museum">CSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
. Other Material.
|
||
<materialsCitation id="3B303CEB7C27CD1225DE9A2BFDC8FD15" ID-GBIF-Occurrence="2012757465" collectingDate="1963-04-12" collectionCode="EMUS" collectorName="G. I. Stage" country="United States of America" county="Pinal County" location="Stanfield" pageId="33" pageNumber="134" specimenCount="1" specimenCount-female="1" stateProvince="Arizona" typeStatus="paratype">
|
||
<collectingCountry id="F34F76267C27CD1225DE9A2BFCD8FD29" box="[766,826,631,655]" name="United States of America" pageId="33" pageNumber="134">USA</collectingCountry>
|
||
:
|
||
<emphasis id="B92CEAA47C27CD1224689A24FC43FD29" box="[840,929,632,655]" italics="true" pageId="33" pageNumber="134">
|
||
<collectingRegion id="499CF8547C27CD1224689A24FC43FD29" box="[840,929,632,655]" country="United States of America" name="Arizona" pageId="33" pageNumber="134">Arizona</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingCounty id="62864E3A7C27CD1224929A2BFBA4FD29" box="[946,1094,631,656]" pageId="33" pageNumber="134">Pinal County</collectingCounty>
|
||
,
|
||
<quantity id="4CA09B537C27CD1223779A2BFB76FD36" box="[1111,1172,631,656]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="33" pageNumber="134" unit="mi" value="4.0">4 mi.</quantity>
|
||
[
|
||
<quantity id="4CA09B537C27CD1223849A2BFB19FD36" box="[1188,1275,631,656]" metricMagnitude="3" metricUnit="m" metricValue="6.4" pageId="33" pageNumber="134" unit="km" value="6.4">6.4 km</quantity>
|
||
] W
|
||
<location id="8E87606D7C27CD1222109A2BFA7AFD36" LSID="urn:lsid:plazi:treatment:03F187A07C18CD1227B79F10FC40FC5E:8E87606D7C27CD1222109A2BFA7AFD36" box="[1328,1432,631,656]" country="United States of America" county="Pinal County" name="Stanfield" pageId="33" pageNumber="134" stateProvince="Arizona">Stanfield</location>
|
||
,
|
||
<date id="FFE610767C27CD1227B79AC7FEF7FD12" box="[151,277,667,692]" pageId="33" pageNumber="134" value="1963-04-12">
|
||
<collectingDate id="EFA2E99E7C27CD1227B79AC7FEF7FD12" box="[151,277,667,692]" pageId="33" pageNumber="134" value="1963-04-12">12.IV.1963</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26AD53607C27CD1226019AC7FE6FFD15" box="[289,397,667,691]" pageId="33" pageNumber="134">G.I. Stage</collectorName>
|
||
(
|
||
<specimenCount id="9D5EFD3F7C27CD1226809AC7FE20FD15" box="[416,450,667,691]" pageId="33" pageNumber="134" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE737C27CD1226EF9AC7FDC0FD15" box="[463,546,667,691]" pageId="33" pageNumber="134">EMUS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C27CD1227E79AE3FCF9FD7E" box="[199,795,703,728]" pageId="33" pageNumber="134" type="distribution">
|
||
<paragraph id="8BE736B67C27CD1227E79AE3FCF9FD7E" blockId="33.[151,1437,151,1016]" box="[199,795,703,728]" pageId="33" pageNumber="134">
|
||
<emphasis id="B92CEAA47C27CD1227E79AE3FE82FD7E" bold="true" box="[199,352,703,728]" pageId="33" pageNumber="134">Distribution.</emphasis>
|
||
<collectingRegion id="499CF8547C27CD1226479AE3FE2DFD7E" box="[359,463,703,728]" country="United States of America" name="Colorado" pageId="33" pageNumber="134">Colorado</collectingRegion>
|
||
Plateau in
|
||
<collectingRegion id="499CF8547C27CD12256D9AE3FD45FD7E" box="[589,679,703,728]" country="United States of America" name="Arizona" pageId="33" pageNumber="134">Arizona</collectingRegion>
|
||
and
|
||
<collectingRegion id="499CF8547C27CD1225FD9AE3FCF5FD7E" box="[733,791,703,728]" country="United States of America" name="Utah" pageId="33" pageNumber="134">Utah</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C27CD1227E79ABFFE91FC86" pageId="33" pageNumber="134" type="etymology">
|
||
<paragraph id="8BE736B67C27CD1227E79ABFFE91FC86" blockId="33.[151,1437,151,1016]" pageId="33" pageNumber="134">
|
||
<emphasis id="B92CEAA47C27CD1227E79ABFFEACFD5A" bold="true" box="[199,334,739,764]" pageId="33" pageNumber="134">Etymology.</emphasis>
|
||
Named in honor of William B. Warner, the collector of the
|
||
<typeStatus id="54E388147C27CD1223279ABFFB8BFD5A" box="[1031,1129,739,764]" pageId="33" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
and many other important mutillid specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342653D7C27CD1227E79B77FC40FC5E" pageId="33" pageNumber="134" type="discussion">
|
||
<paragraph id="8BE736B67C27CD1227E79B77FC40FC5E" blockId="33.[151,1437,151,1016]" pageId="33" pageNumber="134">
|
||
<emphasis id="B92CEAA47C27CD1227E79B77FED6FCE2" bold="true" box="[199,308,811,836]" pageId="33" pageNumber="134">Remarks</emphasis>
|
||
. This female apparently belongs to the
|
||
<taxonomicName id="4C584D357C27CD12242E9B70FC4DFCE5" authorityName="Pitts & Manley" authorityYear="2004" box="[782,943,812,835]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="134" phylum="Arthropoda" rank="species" species="crocopinna">
|
||
<emphasis id="B92CEAA47C27CD12242E9B70FC4DFCE5" box="[782,943,812,835]" italics="true" pageId="33" pageNumber="134">L. crocopinna</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<bibRefCitation id="EFC94B477C27CD1223489B77FA91FCE5" author="Williams, K. A. & Pitts, J. P." box="[1128,1395,811,836]" pageId="33" pageNumber="134" pagination="227 - 243" refId="ref25741" refString="Williams, K. A. & Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227 - 243." type="journal article" year="2009">Williams & Pitts 2009</bibRefCitation>
|
||
) in having the ventral mandibular lamella obliterated and the pronotal and propodeal spiracles lowly produced. Within this apparent group, the thickened T2 fringe setae are unique. The specimen from Pinal County
|
||
<collectingRegion id="499CF8547C27CD1223C89B2FFAA0FC2A" box="[1256,1346,883,908]" country="United States of America" name="Arizona" pageId="33" pageNumber="134">Arizona</collectingRegion>
|
||
has the cuticle somewhat darker, the dorsal setae longer and darker, and the intervals of T2 with the microreticulations more distinct. Although these differences are not great enough to justify describing it as a distinct species at this point, they are enough to justify removing it from the
|
||
<typeStatus id="54E388147C27CD1225D39BBCFCB6FC5E" box="[755,852,992,1016]" pageId="33" pageNumber="134" type="paratype">paratype</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |