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<mods:title id="12ACAA8A60658F51B2D1B011640D4B9D">Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations</mods:title>
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Key to species of
<taxonomicName id="4C584D357C04CD31264D9D2AFD66FA36" authority="(Females)" authorityName="Mickel" authorityYear="1936" baseAuthorityName="Females" box="[365,644,1398,1424]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAA47C04CD31264D9D2AFDE0FA36" bold="true" box="[365,514,1398,1424]" italics="true" pageId="2" pageNumber="103">Lomachaeta</emphasis>
(Females)
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<paragraph id="8BE736B67C04CD3127B79DE1FA79FA49" blockId="2.[151,1436,1469,2018]" pageId="2" pageNumber="103">
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1. Mesosoma and T2 disc with matching thickened posteriorly-directed subparallel appressed silvery to golden setae (
<figureCitation id="13632A337C04CD3122609DE1FA75FA74" box="[1344,1431,1469,1490]" captionStart="FIGURES 14" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[166,1419,189,1961]" captionTargetId="figure@7.[166,1419,189,1961]" captionTargetPageId="7" captionText="FIGURES 14. Lomachaeta females; dorsal habitus. 1) L. argenta; 2) L. vacamuerta, LaSalle County, Texas; 3) L. vacamuerta, Kennedy County, Texas; 4) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588772/files/figure.png" pageId="2" pageNumber="103">Figs 13</figureCitation>
; North America)...................................................................................... 2
</keyLead>
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<paragraph id="8BE736B67C04CD3127B79DABFA79F9E1" blockId="2.[151,1436,1469,2018]" pageId="2" pageNumber="103">
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- Mesosoma and T2 not having matching appressed posteriorly-directed silvery golden setae; dorsum usually without any appressed thickened setae, but if thick setae present on mesonotum, then they are not predominantly subparallel and the T2 disc has erect sparse setae only (e.g.
<figureCitation id="13632A337C04CD3125719E6EFD50F9E1" box="[593,690,1586,1607]" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="2" pageNumber="103">Figs 9, 12</figureCitation>
,
<figureCitation id="13632A337C04CD31259C9E6EFD34F9E1" box="[700,726,1586,1607]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="2" pageNumber="103">21</figureCitation>
,
<figureCitation id="13632A337C04CD3125C09E6EFD19F9E1" box="[736,763,1586,1607]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="2" pageNumber="103">33</figureCitation>
; North or South America).................................. 3
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<paragraph id="8BE736B67C04CD3127B79E13FA74F924" blockId="2.[151,1436,1469,2018]" pageId="2" pageNumber="103">
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2(1). Mesosoma elongate, thoracic dorsum nearly as long as wide; head wider (~1.2×) than mesosoma (
<figureCitation id="13632A337C04CD3123BC9E13FB10F9C2" box="[1180,1266,1615,1636]" captionStart="FIGURES 14" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[166,1419,189,1961]" captionTargetId="figure@7.[166,1419,189,1961]" captionTargetPageId="7" captionText="FIGURES 14. Lomachaeta females; dorsal habitus. 1) L. argenta; 2) L. vacamuerta, LaSalle County, Texas; 3) L. vacamuerta, Kennedy County, Texas; 4) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588772/files/figure.png" pageId="2" pageNumber="103">Figs 2, 3</figureCitation>
,
<figureCitation id="13632A337C04CD3123DD9E13FAC1F9C2" box="[1277,1315,1615,1636]" captionStart="FIGURES 58" captionStartId="8.[151,264,1065,1088]" captionTargetBox="[166,1419,157,1043]" captionTargetId="figure@8.[166,1419,157,1043]" captionTargetPageId="8" captionText="FIGURES 58. Lomachaeta females; lateral habitus. 5) L. argenta; 6) L. vacamuerta, Kennedy County, Texas; 7) L. vacamuerta, LaSalle County, Texas; 8) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588774/files/figure.png" pageId="2" pageNumber="103">6, 7</figureCitation>
;
<collectingRegion id="499CF8547C04CD31220D9E13FA8BF9C2" box="[1325,1385,1615,1636]" country="United States of America" name="Texas" pageId="2" pageNumber="103">Texas</collectingRegion>
)........................................................................
<taxonomicName id="4C584D357C04CD3123099E31FA74F924" authority="Williams &amp; Pitts, 2009" authorityName="Williams &amp; Pitts" authorityYear="2009" box="[1065,1430,1645,1666]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="vacamuerta">
<emphasis id="B92CEAA47C04CD3123099E31FB51F924" box="[1065,1203,1645,1666]" italics="true" pageId="2" pageNumber="103">L. vacamuerta</emphasis>
<bibRefCitation id="EFC94B477C04CD31239A9E31FA74F924" author="Williams, K. A. &amp; Pitts, J. P." box="[1210,1430,1645,1666]" pageId="2" pageNumber="103" pagination="227 - 243" refId="ref25741" refString="Williams, K. A. &amp; Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227 - 243." type="journal article" year="2009">Williams &amp; Pitts, 2009</bibRefCitation>
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- Mesosoma more compact, thoracic dorsum shorter (0.85×) than wide; head as wide as mesosoma (
<figureCitation id="13632A337C04CD3123FB9ED6FAF9F939" box="[1243,1307,1674,1695]" captionStart="FIGURES 14" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[166,1419,189,1961]" captionTargetId="figure@7.[166,1419,189,1961]" captionTargetPageId="7" captionText="FIGURES 14. Lomachaeta females; dorsal habitus. 1) L. argenta; 2) L. vacamuerta, LaSalle County, Texas; 3) L. vacamuerta, Kennedy County, Texas; 4) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588772/files/figure.png" pageId="2" pageNumber="103">Figs 1</figureCitation>
,
<figureCitation id="13632A337C04CD31220B9ED6FADBF939" box="[1323,1337,1674,1695]" captionStart="FIGURES 58" captionStartId="8.[151,264,1065,1088]" captionTargetBox="[166,1419,157,1043]" captionTargetId="figure@8.[166,1419,157,1043]" captionTargetPageId="8" captionText="FIGURES 58. Lomachaeta females; lateral habitus. 5) L. argenta; 6) L. vacamuerta, Kennedy County, Texas; 7) L. vacamuerta, LaSalle County, Texas; 8) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588774/files/figure.png" pageId="2" pageNumber="103">5</figureCitation>
;
<collectingRegion id="499CF8547C04CD3122689ED6FA7AF939" box="[1352,1432,1674,1695]" country="United States of America" name="Arizona" pageId="2" pageNumber="103">Arizona</collectingRegion>
,
<collectingRegion id="499CF8547C04CD3127D19EFBFEDFF91A" box="[241,317,1703,1724]" country="United States of America" name="Nevada" pageId="2" pageNumber="103">Nevada</collectingRegion>
,
<collectingRegion id="499CF8547C04CD3126669EFBFE94F91A" box="[326,374,1703,1724]" country="United States of America" name="Utah" pageId="2" pageNumber="103">Utah</collectingRegion>
)..............................................................
<taxonomicName id="4C584D357C04CD3123459EFBFA7EF91A" authority="Pitts &amp; Manley, 2004" authorityName="Pitts &amp; Manley" authorityYear="2004" box="[1125,1436,1703,1724]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="argenta">
<emphasis id="B92CEAA47C04CD3123459EFBFB2BF91A" box="[1125,1225,1703,1724]" italics="true" pageId="2" pageNumber="103">L. argenta</emphasis>
<bibRefCitation id="EFC94B477C04CD3123EF9EFBFA7EF91A" author="Pitts, J. P. &amp; Manley, D. G." box="[1231,1436,1703,1724]" pageId="2" pageNumber="103" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. &amp; Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts &amp; Manley, 2004</bibRefCitation>
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3(1). Areolations on frons and vertex broad, most transverse intervals obliterated, leaving variously defined apparent longitudinal carinae; lateral pronotal and propodeal faces with areolations having intervals more clearly defined, at least one complete areolation on mesopleuron and lateral propodeal face; body variegated dark and light brown or primarily dark-brown (
<figureCitation id="13632A337C04CD3122799EA3FEE9F894" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="2" pageNumber="103">Figs 9, 10</figureCitation>
,
<figureCitation id="13632A337C04CD3126359F41FEB1F894" box="[277,339,1821,1842]" captionStart="FIGURES 1520" captionStartId="14.[151,264,1557,1580]" captionTargetBox="[167,1420,193,1536]" captionTargetId="figure@14.[167,1420,193,1536]" captionTargetPageId="14" captionText="FIGURES 1520. Lomachaeta females; lateral habitus. 15) L. hyphantria, Brazil; 16) L. hyphantria, Venezuela; 17) L. ibarrai, Holotype; 18) L. meloi, sp. nov.; 19) L. vianai, Argentina, Holotype; 20) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588778/files/figure.png" pageId="2" pageNumber="103">15, 16</figureCitation>
; Central and South America only)...................................
<taxonomicName id="4C584D357C04CD3123669F41FA79F894" authority="Pitts &amp; Manley, 2004" authorityName="Pitts &amp; Manley" authorityYear="2004" box="[1094,1435,1821,1842]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="hyphantria">
<emphasis id="B92CEAA47C04CD3123669F41FB2AF894" box="[1094,1224,1821,1842]" italics="true" pageId="2" pageNumber="103">L. hyphantria</emphasis>
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<keyLead id="30A995837C04CD3127B79F66FA79F82C" pageId="2" pageNumber="103">- Areolations on frons and vertex variable, either complete or with obliterated edges not oriented to leave longitudinal carinae; lateral pronotal and propodeal faces often with areolations largely obliterated; body often predominantly orange-brown (North, Central, or South America)...................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C04CD3127B79FCEFA79F844" pageId="2" pageNumber="103">
<paragraph id="8BE736B67C04CD3127B79FCEFA7EF801" blockId="2.[151,1436,1469,2018]" box="[151,1436,1938,1959]" pageId="2" pageNumber="103">
<keyLead id="30A995837C04CD3127B79FCEFA7EF801" box="[151,1436,1938,1959]" pageId="2" pageNumber="103">
4(3). South American species; mesosomal areolations dense, intervals narrow and even, not scabrous (
<figureCitation id="13632A337C04CD3123B09FCEFB1CF801" box="[1168,1278,1938,1959]" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="2" pageNumber="103">Figs 1214</figureCitation>
)........... 5
</keyLead>
</paragraph>
<paragraph id="8BE736B67C04CD3127B79FF3FA79F844" blockId="2.[151,1436,1469,2018]" pageId="2" pageNumber="103">
<keyLead id="30A995837C04CD3127B79FF3FA79F844" pageId="2" pageNumber="103">
- North and Central American species; mesosomal sculpture variable, areolations often sparser with intervals partially obliterated, sometimes appearing scabrous (e.g.
<figureCitation id="13632A337C04CD3125A09F91FD0FF844" box="[640,749,1997,2018]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="2" pageNumber="103">Figs 22, 23</figureCitation>
,
<figureCitation id="13632A337C04CD3125D79F91FCD6F844" box="[759,820,1997,2018]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="2" pageNumber="103">33, 34</figureCitation>
)................................................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B798CBFA79FEA2" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B798CBFA7EFF6C" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B798CBFA7EFF6C" pageId="3" pageNumber="104">
5(4). Mesosoma, metasoma, and legs black, head orange-brown; entire body except T2 disc with appressed setae thickened, white, contrasting with integument (
<figureCitation id="13632A337C05CD30252A98E9FDB7FF6C" box="[522,597,181,202]" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="3" pageNumber="104">Figs 12</figureCitation>
,
<figureCitation id="13632A337C05CD30257F98E9FD9BFF6C" box="[607,633,181,202]" captionStart="FIGURES 1520" captionStartId="14.[151,264,1557,1580]" captionTargetBox="[167,1420,193,1536]" captionTargetId="figure@14.[167,1420,193,1536]" captionTargetPageId="14" captionText="FIGURES 1520. Lomachaeta females; lateral habitus. 15) L. hyphantria, Brazil; 16) L. hyphantria, Venezuela; 17) L. ibarrai, Holotype; 18) L. meloi, sp. nov.; 19) L. vianai, Argentina, Holotype; 20) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588778/files/figure.png" pageId="3" pageNumber="104">18</figureCitation>
;
<collectingRegion id="499CF8547C05CD3025A398E9FD04FF6C" box="[643,742,181,202]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="104">São Paulo</collectingRegion>
,
<collectingCountry id="F34F76267C05CD3025D098E9FCCEFF6C" box="[752,812,181,202]" name="Brazil" pageId="3" pageNumber="104">Brazil</collectingCountry>
).............................
<taxonomicName id="4C584D357C05CD3023B798E9FAA1FF6C" authority="Williams" authorityName="Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts" authorityYear="2019" box="[1175,1347,181,202]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="meloi" status="sp. nov.">
<emphasis id="B92CEAA47C05CD3023B798E9FB07FF6C" box="[1175,1253,181,202]" italics="true" pageId="3" pageNumber="104">L. meloi</emphasis>
Williams
</taxonomicName>
,
<emphasis id="B92CEAA47C05CD30226F98E8FA7EFF6C" bold="true" box="[1359,1436,180,202]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A21F57DF7C05CD30226F98E8FA7EFF6C" box="[1359,1436,180,202]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B7988EFA79FEA2" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B7988EFA79FEA2" pageId="3" pageNumber="104">- Head and mesosoma reddish brown, often darkened laterally, metasoma blackish, always darker than head; body setae predominantly brown.................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79951FA7EFEDC" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79951FA7EFE99" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79951FA7EFE99" pageId="3" pageNumber="104">
6(5). Propodeal spiracle prominently dentiform; scape as long as pedicel and F13 combined (
<figureCitation id="13632A337C05CD30236E9951FB7AFE84" box="[1102,1176,269,290]" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="3" pageNumber="104">Figs 11</figureCitation>
,
<figureCitation id="13632A337C05CD3023839951FB5FFE84" box="[1187,1213,269,290]" captionStart="FIGURES 1520" captionStartId="14.[151,264,1557,1580]" captionTargetBox="[167,1420,193,1536]" captionTargetId="figure@14.[167,1420,193,1536]" captionTargetPageId="14" captionText="FIGURES 1520. Lomachaeta females; lateral habitus. 15) L. hyphantria, Brazil; 16) L. hyphantria, Venezuela; 17) L. ibarrai, Holotype; 18) L. meloi, sp. nov.; 19) L. vianai, Argentina, Holotype; 20) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588778/files/figure.png" pageId="3" pageNumber="104">17</figureCitation>
;
<collectingRegion id="499CF8547C05CD3023E99951FAAFFE84" box="[1225,1357,269,290]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="104">Buenos Aires</collectingRegion>
,
<collectingCountry id="F34F76267C05CD3022789951FEF5FE99" name="Argentina" pageId="3" pageNumber="104">Argentina</collectingCountry>
)..............................................................................
<taxonomicName id="4C584D357C05CD3023E79976FA7EFE99" authority="Casal, 1969" authorityName="Casal" authorityYear="1969" box="[1223,1436,298,319]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="ibarrai">
<emphasis id="B92CEAA47C05CD3023E79976FAC6FE99" box="[1223,1316,298,319]" italics="true" pageId="3" pageNumber="104">L. ibarrai</emphasis>
Casal, 1969
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B7991BFA7EFEDC" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B7991BFA7EFEDC" pageId="3" pageNumber="104">
- Propodeal spiracle lowly rounded, not dentiform; scape shorter than pedicel and F13 combined (
<figureCitation id="13632A337C05CD3023B7991BFAE7FEFA" box="[1175,1285,327,348]" captionStart="FIGURES 914" captionStartId="13.[151,264,1956,1979]" captionTargetBox="[167,1420,373,1935]" captionTargetId="figure@13.[167,1420,373,1935]" captionTargetPageId="13" captionText="FIGURES 914. Lomachaeta females; dorsal habitus. 9) L. hyphantria, Brazil; 10) L. hyphantria, Venezuela; 11) L. ibarrai, Holotype; 12) L. meloi, sp. nov.; 13) L. vianai, Argentina, Holotype; 14) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588776/files/figure.png" pageId="3" pageNumber="104">Figs 13, 14</figureCitation>
,
<figureCitation id="13632A337C05CD302230991BFAACFEFA" box="[1296,1358,327,348]" captionStart="FIGURES 1520" captionStartId="14.[151,264,1557,1580]" captionTargetBox="[167,1420,193,1536]" captionTargetId="figure@14.[167,1420,193,1536]" captionTargetPageId="14" captionText="FIGURES 1520. Lomachaeta females; lateral habitus. 15) L. hyphantria, Brazil; 16) L. hyphantria, Venezuela; 17) L. ibarrai, Holotype; 18) L. meloi, sp. nov.; 19) L. vianai, Argentina, Holotype; 20) L. vianai, Paraguay. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588778/files/figure.png" pageId="3" pageNumber="104">19, 20</figureCitation>
;
<collectingCountry id="F34F76267C05CD302278991BFEFAFEDC" name="Argentina" pageId="3" pageNumber="104">Argentina</collectingCountry>
,
<collectingCountry id="F34F76267C05CD3026019939FEBCFEDC" box="[289,350,357,378]" name="Brazil" pageId="3" pageNumber="104">Brazil</collectingCountry>
, and
<collectingCountry id="F34F76267C05CD3026B09939FE09FEDC" box="[400,491,357,378]" name="Paraguay" pageId="3" pageNumber="104">Paraguay</collectingCountry>
).............................................................
<taxonomicName id="4C584D357C05CD3023EE9939FA7EFEDC" authority="Casal, 1969" authorityName="Casal" authorityYear="1969" box="[1230,1436,357,378]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="vianai">
<emphasis id="B92CEAA47C05CD3023EE9939FAC6FEDC" box="[1230,1316,357,378]" italics="true" pageId="3" pageNumber="104">L. vianai</emphasis>
Casal, 1969
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B799DEFA79FDC2" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B799DEFA7EFE49" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B799DEFA7EFE49" pageId="3" pageNumber="104">
7(4). Mandible with baso-ventral subhyaline lamella, mandible narrowed beyond lamella (
<figureCitation id="13632A337C05CD30230499DEFB8EFE31" box="[1060,1132,386,407]" captionStart="FIGURES 6572" captionStartId="32.[151,264,1932,1955]" captionTargetBox="[191,1396,193,1898]" captionTargetId="figure@32.[191,1396,193,1898]" captionTargetPageId="32" captionText="FIGURES 6572. Lomachaeta line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65) L. hicksi; 66) L. beadugrimi; 6768) L. calamondin, sp. nov.; 6970) L. eironeia, sp. nov.; 7172) L. juanita, sp. nov." httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="3" pageNumber="104">Fig. 65</figureCitation>
); pronotal and propodeal spiracles swollen and apparently tuberculate (
<figureCitation id="13632A337C05CD30255B99C3FD09FE12" box="[635,747,415,436]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 22, 23</figureCitation>
); T1 shape sub-disciform, often with dorsal and anterior faces set apart by angle or incomplete carina; T2 fringe usually composed of thick silvery setae (
<figureCitation id="13632A337C05CD30232499E1FB7BFE74" box="[1028,1177,445,466]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 21, 2326</figureCitation>
); mesosomal dorsum often with dense areolations, at least anteriorly (
<figureCitation id="13632A337C05CD3025A69986FD14FE49" box="[646,758,474,495]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 21, 24</figureCitation>
); frequently encountered in trap nests......................... 8
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B799ABFA79FDC2" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B799ABFA79FDC2" pageId="3" pageNumber="104">
- Mandible without basal lamella, if weak lamella apparently present, it does not interrupt ventral mandible contour (
<figureCitation id="13632A337C05CD30226899ABFA72FDAA" box="[1352,1424,503,524]" captionStart="FIGURES 6572" captionStartId="32.[151,264,1932,1955]" captionTargetBox="[191,1396,193,1898]" captionTargetId="figure@32.[191,1396,193,1898]" captionTargetPageId="32" captionText="FIGURES 6572. Lomachaeta line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65) L. hicksi; 66) L. beadugrimi; 6768) L. calamondin, sp. nov.; 6970) L. eironeia, sp. nov.; 7172) L. juanita, sp. nov." httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="3" pageNumber="104">Fig. 66</figureCitation>
); pronotal and propodeal spiracles usually not so swollen (e.g.
<figureCitation id="13632A337C05CD30246E9A49FC5DFD8C" box="[846,959,533,554]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 3336</figureCitation>
); T1 shape sub-sessile, dorsal and anterior faces evenly rounded together; T2 fringe usually composed of sparse slender setae (e.g.
<figureCitation id="13632A337C05CD30233E9A6EFB6DFDE1" box="[1054,1167,562,583]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 3335</figureCitation>
); mesosomal dorsum never with areolations dense; not known from trap nests......................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79A31FA79FCB2" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79A31FA7EFD1A" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79A31FA7EFD1A" pageId="3" pageNumber="104">
8(7). Head and mesosoma reddish brown, metasomal segments 26 uniform blackish; mesosomal dorsum with tight subuniform areolations; T6 with dense thickened long setae that are black basally and whitish apically (
<figureCitation id="13632A337C05CD3023469AD6FB53FD39" box="[1126,1201,650,671]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 21</figureCitation>
,
<figureCitation id="13632A337C05CD30239C9AD6FB35FD39" box="[1212,1239,650,671]" captionStart="FIGURES 2732" captionStartId="16.[151,264,1939,1962]" captionTargetBox="[167,1420,503,1898]" captionTargetId="figure@16.[167,1420,503,1898]" captionTargetPageId="16" captionText="FIGURES 2732. Lomachaeta females; lateral habitus. 27) L. chionothrix; 28) L. cirrhomeris, California; 29) L. cirrhomeris, Arizona; 30) L. hicksi, Coconino Co., Arizona; 31) L. hicksi, Alachua Co., Florida; 32) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588782/files/figure.png" pageId="3" pageNumber="104">27</figureCitation>
; subtropical
<collectingCountry id="F34F76267C05CD3022729AD6FA79FD39" box="[1362,1435,650,671]" name="Mexico" pageId="3" pageNumber="104">Mexico</collectingCountry>
to
<collectingCountry id="F34F76267C05CD3026299AFBFE91FD1A" box="[265,371,679,700]" name="Costa Rica" pageId="3" pageNumber="104">Costa Rica</collectingCountry>
)...........................................................
<taxonomicName id="4C584D357C05CD3023609AFBFA7EFD1A" authority="Pitts &amp; Manley, 2004" authorityName="Pitts &amp; Manley" authorityYear="2004" box="[1088,1436,679,700]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="chionothrix">
<emphasis id="B92CEAA47C05CD3023609AFBFB2BFD1A" box="[1088,1225,679,700]" italics="true" pageId="3" pageNumber="104">L. chionothrix</emphasis>
<bibRefCitation id="EFC94B477C05CD3023EF9AFBFA7EFD1A" author="Pitts, J. P. &amp; Manley, D. G." box="[1231,1436,679,700]" pageId="3" pageNumber="104" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. &amp; Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts &amp; Manley, 2004</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79A99FA79FCB2" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79A99FA79FCB2" pageId="3" pageNumber="104">- Variably colored, usually variegapted light and dark brown, but if metasoma predominantly blackish, then head and mesosoma also dark; mesosomal dorsum usually with fewer areolations that are often broader posteriorly; T6 lacking long thick bi-colored setae (widespread in North America)............................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79B41FA77FC01" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79B41FA7EFCCA" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79B41FA7EFCCA" pageId="3" pageNumber="104">
9(8). Body unicolorous pale orange-brown, at most with T6 largely blackened and/or legs and antennae at most darkened apically; T36 setae mostly blackish (
<figureCitation id="13632A337C05CD3025239B66FD90FCE9" box="[515,626,826,847]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 22, 23</figureCitation>
,
<figureCitation id="13632A337C05CD30255B9B66FD58FCE9" box="[635,698,826,847]" captionStart="FIGURES 2732" captionStartId="16.[151,264,1939,1962]" captionTargetBox="[167,1420,503,1898]" captionTargetId="figure@16.[167,1420,503,1898]" captionTargetPageId="16" captionText="FIGURES 2732. Lomachaeta females; lateral habitus. 27) L. chionothrix; 28) L. cirrhomeris, California; 29) L. cirrhomeris, Arizona; 30) L. hicksi, Coconino Co., Arizona; 31) L. hicksi, Alachua Co., Florida; 32) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588782/files/figure.png" pageId="3" pageNumber="104">28, 29</figureCitation>
; southwestern
<collectingCountry id="F34F76267C05CD3024699B66FC9BFCE9" box="[841,889,826,847]" name="United States of America" pageId="3" pageNumber="104">USA</collectingCountry>
and northwestern
<collectingCountry id="F34F76267C05CD30230A9B66FB97FCE9" box="[1066,1141,826,847]" name="Mexico" pageId="3" pageNumber="104">Mexico</collectingCountry>
)..............................................................................................
<taxonomicName id="4C584D357C05CD30231F9B0BFA7EFCCA" authority="Pitts &amp; Manley, 2004" authorityName="Pitts &amp; Manley" authorityYear="2004" box="[1087,1436,855,876]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="cirrhomeris">
<emphasis id="B92CEAA47C05CD30231F9B0BFB2BFCCA" box="[1087,1225,855,876]" italics="true" pageId="3" pageNumber="104">L. cirrhomeris</emphasis>
<bibRefCitation id="EFC94B477C05CD3023EF9B0BFA7EFCCA" author="Pitts, J. P. &amp; Manley, D. G." box="[1231,1436,855,876]" pageId="3" pageNumber="104" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. &amp; Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts &amp; Manley, 2004</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79B29FA77FC01" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79B29FA77FC01" pageId="3" pageNumber="104">
- Body generally darker, variegated with orange, brown, and blackish, legs and antennae variably blackened; T36 with interspersed silvery and blackish setae (
<figureCitation id="13632A337C05CD3025619BCEFD52FC01" box="[577,688,914,935]" captionStart="FIGURES 2126" captionStartId="15.[151,264,1841,1864]" captionTargetBox="[209,1399,186,1801]" captionTargetId="figure@15.[209,1399,186,1801]" captionTargetPageId="15" captionText="FIGURES 2126. Lomachaeta females; dorsal habitus. 21) L. chionothrix; 22) L. cirrhomeris, California; 23) L. cirrhomeris, Arizona; 24) L. hicksi, Alachua Co., Florida; 25) L. hicksi, Coconino Co., Arizona; 26) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588780/files/figure.png" pageId="3" pageNumber="104">Figs 2426</figureCitation>
,
<figureCitation id="13632A337C05CD3025999BCEFD1BFC01" box="[697,761,914,935]" captionStart="FIGURES 2732" captionStartId="16.[151,264,1939,1962]" captionTargetBox="[167,1420,503,1898]" captionTargetId="figure@16.[167,1420,503,1898]" captionTargetPageId="16" captionText="FIGURES 2732. Lomachaeta females; lateral habitus. 27) L. chionothrix; 28) L. cirrhomeris, California; 29) L. cirrhomeris, Arizona; 30) L. hicksi, Coconino Co., Arizona; 31) L. hicksi, Alachua Co., Florida; 32) L. hicksi, Cache Co., Utah. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588782/files/figure.png" pageId="3" pageNumber="104">3032</figureCitation>
; widespread in North America)............
<taxonomicName id="4C584D357C05CD30239E9BCEFA77FC01" authority="Mickel, 1936" authorityName="Mickel" authorityYear="1936" box="[1214,1429,914,935]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="hicksi">
<emphasis id="B92CEAA47C05CD30239E9BCEFAF2FC01" box="[1214,1296,914,935]" italics="true" pageId="3" pageNumber="104">L. hicksi</emphasis>
<bibRefCitation id="EFC94B477C05CD3022359BCEFA77FC01" author="Mickel, C. E." box="[1301,1429,914,935]" pageId="3" pageNumber="104" pagination="289 - 297" refId="ref25326" refString="Mickel, C. E. (1936) Two new genera and five new species of Mutillidae. Annals of the Entomological Society of America, 29, 289 - 297. https: // doi. org / 10.1093 / aesa / 29.2.289" type="journal article" year="1936">Mickel, 1936</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79BF3FA79FBBA" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79BF3FA7EFC44" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79BF3FA7EFC44" pageId="3" pageNumber="104">
10(7). Mesosoma elongate, thoracic length subequal to width; mesosomal areolations deep and distinct, with all intervals distinct; frons mostly with complete areolations (
<figureCitation id="13632A337C05CD3025539B91FD50FC44" box="[627,690,973,994]" captionStart="FIGURES 14" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[166,1419,189,1961]" captionTargetId="figure@7.[166,1419,189,1961]" captionTargetPageId="7" captionText="FIGURES 14. Lomachaeta females; dorsal habitus. 1) L. argenta; 2) L. vacamuerta, LaSalle County, Texas; 3) L. vacamuerta, Kennedy County, Texas; 4) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588772/files/figure.png" pageId="3" pageNumber="104">Figs 4</figureCitation>
,
<figureCitation id="13632A337C05CD30259C9B91FD28FC44" box="[700,714,973,994]" captionStart="FIGURES 58" captionStartId="8.[151,264,1065,1088]" captionTargetBox="[166,1419,157,1043]" captionTargetId="figure@8.[166,1419,157,1043]" captionTargetPageId="8" captionText="FIGURES 58. Lomachaeta females; lateral habitus. 5) L. argenta; 6) L. vacamuerta, Kennedy County, Texas; 7) L. vacamuerta, LaSalle County, Texas; 8) L. osita, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588774/files/figure.png" pageId="3" pageNumber="104">8</figureCitation>
; Madrean Archipelago in
<collectingRegion id="499CF8547C05CD3024E59B91FBF6FC44" box="[965,1044,973,994]" country="United States of America" name="Arizona" pageId="3" pageNumber="104">Arizona</collectingRegion>
)...........
<taxonomicName id="4C584D357C05CD3023BA9B91FAA7FC44" authority="Williams" authorityName="Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts" authorityYear="2019" box="[1178,1349,973,994]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="osita" status="sp. nov.">
<emphasis id="B92CEAA47C05CD3023BA9B91FB07FC44" box="[1178,1253,973,994]" italics="true" pageId="3" pageNumber="104">L. osita</emphasis>
Williams
</taxonomicName>
,
<emphasis id="B92CEAA47C05CD30226F9B90FA7EFC44" bold="true" box="[1359,1436,972,994]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A21F57DF7C05CD30226F9B90FA7EFC44" box="[1359,1436,972,994]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79BB6FA79FBBA" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79BB6FA79FBBA" pageId="3" pageNumber="104">
- Mesosoma not so elongate, thoracic length less than 0.9 × width; mesosomal areolations with margins less distinct and often partly obliterated; frons generally with obscure punctures and shallow rounded intervals (e.g.
<figureCitation id="13632A337C05CD3023539C5BFB02FBBA" box="[1139,1248,1031,1052]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 3336</figureCitation>
)............. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79C79FA79FB6A" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79C79FA74FBD2" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79C79FA74FBD2" pageId="3" pageNumber="104">
11(10). T2 sculpture mostly smooth between punctures (microreticulations not visible at 30 × magnification, obscure apparent reticulations sometimes visible at&gt; 80 × magnification); dorsal bristles of mesonotum and T2 blackish, many bristles nearly as long as scape (
<figureCitation id="13632A337C05CD30265D9C03FE2AFBD2" box="[381,456,1119,1140]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 35</figureCitation>
,
<figureCitation id="13632A337C05CD3026F29C03FE0EFBD2" box="[466,492,1119,1140]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="3" pageNumber="104">39</figureCitation>
; Central Valley and Coast Ranges,
<collectingRegion id="499CF8547C05CD30241E9C03FC43FBD2" box="[830,929,1119,1140]" country="United States of America" name="California" pageId="3" pageNumber="104">California</collectingRegion>
)....................
<taxonomicName id="4C584D357C05CD3023809C03FA74FBD2" authority="(Mickel), 1964" authorityName="Mickel" authorityYear="1964" baseAuthorityName="Mickel" box="[1184,1430,1119,1140]" class="Insecta" family="Mutillidae" genus="Smicromutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B92CEAA47C05CD3023809C03FAE2FBD2" box="[1184,1280,1119,1140]" italics="true" pageId="3" pageNumber="104">L. powelli</emphasis>
(
<bibRefCitation id="EFC94B477C05CD30222C9C03FA74FBD2" author="Mickel, C. E." box="[1292,1430,1119,1140]" pageId="3" pageNumber="104" pagination="163 - 171" refId="ref25412" refString="Mickel, C. E. (1964) Synonymical notes on Neotropical Mutillidae (Hymenoptera). Proceedings of the Royal Entomological Society of London (B), 33 (9 - 10), 163 - 171. https: // doi. org / 10.1111 / j. 1365 - 3113.1964. tb 01635. x" type="journal article" year="1964">Mickel), 1964</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79C21FA79FB6A" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79C21FA79FB6A" pageId="3" pageNumber="104">
- T2 sculpture generally microreticulate between punctures (visible at 30 × magnification); if microreticulations obscure, then dorsal bristles of mesonotum and T2 mostly brown or yellowish, generally much shorter than scape (arid and semi-arid habitats South or East of Sierra
<collectingRegion id="499CF8547C05CD3026D89CEBFDA3FB6A" box="[504,577,1207,1228]" country="United States of America" name="Nevada" pageId="3" pageNumber="104">Nevada</collectingRegion>
and related ranges)...................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79C89FA79FA82" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79C89FA79FAA1" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79C89FA79FAA1" pageId="3" pageNumber="104">
12(11). Antennae and forelegs entirely blackish, middle and hind legs mostly black (
<figureCitation id="13632A337C05CD3024EE9C89FBFBFB4C" box="[974,1049,1237,1258]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 33</figureCitation>
,
<figureCitation id="13632A337C05CD3023039C89FBDCFB4C" box="[1059,1086,1237,1258]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="3" pageNumber="104">37</figureCitation>
;
<collectingRegion id="499CF8547C05CD3023679C89FAE1FB4C" box="[1095,1283,1237,1258]" country="Mexico" name="Baja California Sur" pageId="3" pageNumber="104">
Baja
<collectingRegion id="499CF8547C05CD3023599C89FB38FB4C" box="[1145,1242,1237,1258]" country="United States of America" name="California" pageId="3" pageNumber="104">California</collectingRegion>
Sur
</collectingRegion>
)....................................................................................
<taxonomicName id="4C584D357C05CD30237E9CAEFA79FAA1" authority="Williams &amp; Pitts, 2009" authorityName="Williams &amp; Pitts" authorityYear="2009" box="[1118,1435,1266,1287]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="hedera">
<emphasis id="B92CEAA47C05CD30237E9CAEFB58FAA1" box="[1118,1210,1266,1287]" italics="true" pageId="3" pageNumber="104">L. hedera</emphasis>
<bibRefCitation id="EFC94B477C05CD30239F9CAEFA79FAA1" author="Williams, K. A. &amp; Pitts, J. P." box="[1215,1435,1266,1287]" pageId="3" pageNumber="104" pagination="227 - 243" refId="ref25741" refString="Williams, K. A. &amp; Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227 - 243." type="journal article" year="2009">Williams &amp; Pitts, 2009</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79D53FA79FA82" blockId="3.[151,1436,151,1434]" box="[151,1435,1295,1316]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79D53FA79FA82" box="[151,1435,1295,1316]" pageId="3" pageNumber="104">
- Antennae and legs largely orange-brown, at least scape and fore coxa lighter brown (
<figureCitation id="13632A337C05CD30233C9D53FB69FA82" box="[1052,1163,1295,1316]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 34, 36</figureCitation>
,
<figureCitation id="13632A337C05CD3023B49D53FB30FA82" box="[1172,1234,1295,1316]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="3" pageNumber="104">38, 40</figureCitation>
).............. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC2E137C05CD3027B79D71FA7EFA3C" pageId="3" pageNumber="104">
<paragraph id="8BE736B67C05CD3027B79D71FA79FAF9" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79D71FA79FAF9" pageId="3" pageNumber="104">
13(12). Body orange-brown, at most with T6 blackened (
<figureCitation id="13632A337C05CD3025E79D71FCF3FAE4" box="[711,785,1325,1346]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 34</figureCitation>
,
<figureCitation id="13632A337C05CD30243B9D71FCD4FAE4" box="[795,822,1325,1346]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="3" pageNumber="104">38</figureCitation>
; Mojave and Sonoran Deserts in
<collectingRegion id="499CF8547C05CD3023549D71FB37FAE4" box="[1140,1237,1325,1346]" country="United States of America" name="California" pageId="3" pageNumber="104">California</collectingRegion>
and
<collectingRegion id="499CF8547C05CD3022249D71FAB1FAE4" box="[1284,1363,1325,1346]" country="United States of America" name="Arizona" pageId="3" pageNumber="104">Arizona</collectingRegion>
)..........................................................................
<taxonomicName id="4C584D357C05CD30230E9D16FA79FAF9" authority="(Pitts &amp; Manley, 2004)" baseAuthorityName="Pitts &amp; Manley" baseAuthorityYear="2004" box="[1070,1435,1354,1375]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="beadugrimi">
<emphasis id="B92CEAA47C05CD30230E9D16FB5BFAF9" box="[1070,1209,1354,1375]" italics="true" pageId="3" pageNumber="104">L. beadugrimi</emphasis>
(
<bibRefCitation id="EFC94B477C05CD3023E59D16FA77FAF9" author="Pitts, J. P. &amp; Manley, D. G." box="[1221,1429,1354,1375]" pageId="3" pageNumber="104" pagination="1 - 27" refId="ref25528" refString="Pitts, J. P. &amp; Manley, D. G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1 - 27. https: // doi. org / 10.11646 / zootaxa. 474.1.1" type="journal article" year="2004">Pitts &amp; Manley, 2004</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE736B67C05CD3027B79D3BFA7EFA3C" blockId="3.[151,1436,151,1434]" pageId="3" pageNumber="104">
<keyLead id="30A995837C05CD3027B79D3BFA7EFA3C" pageId="3" pageNumber="104">
- Femora and T25 variably darkened brown, in part clearly darker than orange-brown mesosoma (
<figureCitation id="13632A337C05CD3023BB9D3BFB04FADA" box="[1179,1254,1383,1404]" captionStart="FIGURES 3336" captionStartId="19.[151,264,1960,1983]" captionTargetBox="[180,1414,193,1938]" captionTargetId="figure@19.[180,1414,193,1938]" captionTargetPageId="19" captionText="FIGURES 3336. Lomachaeta females; dorsal habitus. 33) L. hedera; 34) L. beadugrimi; 35) L. powelli; 36) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588784/files/figure.png" pageId="3" pageNumber="104">Figs 36</figureCitation>
,
<figureCitation id="13632A337C05CD3023D09D3BFAE9FADA" box="[1264,1291,1383,1404]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="3" pageNumber="104">40</figureCitation>
;
<collectingRegion id="499CF8547C05CD3022359D3BFA8CFADA" box="[1301,1390,1383,1404]" country="United States of America" name="Colorado" pageId="3" pageNumber="104">Colorado</collectingRegion>
Plateau and Sonoran Desert in
<collectingRegion id="499CF8547C05CD3026DA9DD9FDABFA3C" box="[506,585,1413,1434]" country="United States of America" name="Arizona" pageId="3" pageNumber="104">Arizona</collectingRegion>
)..............................................
<taxonomicName id="4C584D357C05CD3023A09DD9FAA7FA3C" authority="Williams" authorityName="Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts" authorityYear="2019" box="[1152,1349,1413,1434]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="warneri" status="sp. nov.">
<emphasis id="B92CEAA47C05CD3023A09DD9FB07FA3C" box="[1152,1253,1413,1434]" italics="true" pageId="3" pageNumber="104">L. warneri</emphasis>
Williams
</taxonomicName>
,
<emphasis id="B92CEAA47C05CD30226F9DD8FA7EFA3C" bold="true" box="[1359,1436,1412,1434]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A21F57DF7C05CD30226F9DD8FA7EFA3C" box="[1359,1436,1412,1434]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>