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<document id="AD22EAFFBF81F2357B0DBEE4C1E41508" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D6310622FDB6FA42C1DAFE69" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor Frederiksen 2013" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="21" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EAD7FAD440CB2DF48FE3B5DA349EB2F0">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<subSubSection id="C342BEA4D6310623FDB6FA42C6C3FA63" box="[581,1007,1440,1467]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BE7ED2FD6310623FDB6FA42C6C3FA63" blockId="20.[581,1007,1440,1467]" box="[581,1007,1440,1467]" pageId="20" pageNumber="21">
<heading id="D0AF5A43D6310623FDB6FA42C6C3FA63" box="[581,1007,1440,1467]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
Genus
<taxonomicName id="4C5896ACD6310623FD65FA43C6C3FA63" ID-CoL="7NQQQ" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[662,1007,1440,1467]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6310623FD65FA43C63FFA62" bold="true" box="[662,787,1440,1466]" italics="true" pageId="20" pageNumber="21">Aquattuor</emphasis>
Frederiksen, 2013
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6310623FF4EFA01C163F9F2" pageId="20" pageNumber="21" type="type_taxon">
<paragraph id="8BE7ED2FD6310623FF4EFA01C474FA24" blockId="20.[189,344,1506,1532]" box="[189,344,1506,1532]" pageId="20" pageNumber="21">
<heading id="D0AF5A43D6310623FF4EFA01C474FA24" bold="true" box="[189,344,1506,1532]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B92C313DD6310623FF4EFA01C474FA24" bold="true" box="[189,344,1506,1532]" pageId="20" pageNumber="21">
<typeStatus id="54E3538DD6310623FF4EFA01C5D6FA24" box="[189,250,1506,1532]" pageId="20" pageNumber="21">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6310623FF4EF9F2C163F9F2" blockId="20.[189,1103,1552,1579]" box="[189,1103,1552,1579]" pageId="20" pageNumber="21">
<taxonomicName id="4C5896ACD6310623FF4EF9F2C601F9F3" ID-CoL="FZKY" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[189,813,1552,1579]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B92C313DD6310623FF4EF9F2C4E6F9F2" box="[189,458,1552,1579]" italics="true" pageId="20" pageNumber="21">Aquattuor denticulatus</emphasis>
Frederiksen, 2013 (
<collectingCountry id="F34FADBFD6310623FD46F9F3C609F9F3" box="[693,805,1552,1579]" name="Tanzania" pageId="20" pageNumber="21">Tanzania</collectingCountry>
)
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6310623FF4EF9B1C496F942" pageId="20" pageNumber="21" type="description">
<paragraph id="8BE7ED2FD6310623FF4EF9B1C4F4F9B4" blockId="20.[189,472,1618,1644]" box="[189,472,1618,1644]" pageId="20" pageNumber="21">
<heading id="D0AF5A43D6310623FF4EF9B1C4F4F9B4" bold="true" box="[189,472,1618,1644]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B92C313DD6310623FF4EF9B1C4F4F9B4" bold="true" box="[189,472,1618,1644]" pageId="20" pageNumber="21">Other included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6310623FF4EF963C496F942" blockId="20.[189,442,1664,1690]" box="[189,442,1664,1690]" pageId="20" pageNumber="21">
Nine (
<bibRefCitation id="EFC990DED6310623FEF6F963C482F942" author="Enghoff H." box="[261,430,1664,1690]" pageId="20" pageNumber="21" pagination="1 - 32" refId="ref47656" refString="Enghoff H. 2020. A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1 - 32. https: // doi. org / 10.5852 / ejt. 2020.626" type="journal article" year="2020">Enghoff 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6310623FF4EF922C6E9F8F6" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BE7ED2FD6310623FF4EF922C41AF903" blockId="20.[189,310,1729,1755]" box="[189,310,1729,1755]" pageId="20" pageNumber="21">
<heading id="D0AF5A43D6310623FF4EF922C41AF903" bold="true" box="[189,310,1729,1755]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B92C313DD6310623FF4EF922C41AF903" bold="true" box="[189,310,1729,1755]" pageId="20" pageNumber="21">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6310623FF4EF913C6E9F8F6" blockId="20.[189,1398,1776,1838]" pageId="20" pageNumber="21">
Differs from all other genera of
<taxonomicName id="4C5896ACD6310623FDA5F913C627F8D2" authorityName="Attems" authorityYear="1909" box="[598,779,1776,1802]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
by the large, rectangular, detachable limbus flaps (
<bibRefCitation id="EFC990DED6310623FF36F8F0C489F8F5" author="Frederiksen S. B." box="[197,421,1811,1837]" pageId="20" pageNumber="21" pagination="59 - 66" refId="ref48141" refString="Frederiksen S. B. 2013 b. East African odontopygid millipedes 3: Two new genera; Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3694 (1): 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen 2013b</bibRefCitation>
;
<bibRefCitation id="EFC990DED6310623FE42F8F0C62AF8F6" author="Frederiksen S. B. &amp; Enghoff H." box="[433,774,1811,1838]" pageId="20" pageNumber="21" pagination="541 - 560" refId="ref48260" refString="Frederiksen S. B. &amp; Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541 - 560. https: // doi. org / 10.11646 / zootaxa. 3926.4.5" type="journal article" year="2015">Enghoff &amp; Frederiksen 2015</bibRefCitation>
;
<bibRefCitation id="EFC990DED6310623FCE1F8F0C695F8F6" author="Enghoff H." box="[786,953,1811,1838]" pageId="20" pageNumber="21" pagination="1 - 32" refId="ref47656" refString="Enghoff H. 2020. A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1 - 32. https: // doi. org / 10.5852 / ejt. 2020.626" type="journal article" year="2020">Enghoff 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6310622FF4EF8B6C1DAFE69" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BE7ED2FD6310623FF4EF8B6C403F8B7" blockId="20.[189,303,1877,1903]" box="[189,303,1877,1903]" pageId="20" pageNumber="21">
<heading id="D0AF5A43D6310623FF4EF8B6C403F8B7" bold="true" box="[189,303,1877,1903]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B92C313DD6310623FF4EF8B6C403F8B7" bold="true" box="[189,303,1877,1903]" pageId="20" pageNumber="21">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6310623FF4EF860C644F83C" blockId="20.[188,1399,1923,2020]" pageId="20" pageNumber="21">
Eight of the ten species in the genus are Eastern Arc endemics.
<taxonomicName id="4C5896ACD6310623FC56F867C068F846" authority="(Attems, 1896)" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[933,1348,1923,1950]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B92C313DD6310623FC56F867C1A7F846" box="[933,1163,1924,1950]" italics="true" pageId="20" pageNumber="21">Aquattuor fasciatus</emphasis>
(
<bibRefCitation id="EFC990DED6310623FB68F867C010F846" author="Attems C. G." box="[1179,1340,1923,1950]" pageId="20" pageNumber="21" pagination="23 - 42" refId="ref46019" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]." type="journal article" year="1896">Attems, 1896</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C5896ACD6310623FF4EF844C7C1F819" authority="Enghoff &amp; Frederiksen, 2015" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[189,749,1958,1985]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="B92C313DD6310623FF4EF844C4BCF819" box="[189,400,1959,1985]" italics="true" pageId="20" pageNumber="21">A. claudiahempae</emphasis>
<bibRefCitation id="EFC990DED6310623FE64F844C7C1F819" author="Frederiksen S. B. &amp; Enghoff H." box="[407,749,1958,1985]" pageId="20" pageNumber="21" pagination="541 - 560" refId="ref48260" refString="Frederiksen S. B. &amp; Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541 - 560. https: // doi. org / 10.11646 / zootaxa. 3926.4.5" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
</taxonomicName>
have been found at various Tanzanian localities outside the Eastern Arc, and may by synonymous (
<bibRefCitation id="EFC990DED6310623FD40F829C670F83C" author="Enghoff H." box="[691,860,1994,2020]" pageId="20" pageNumber="21" pagination="1 - 32" refId="ref47656" refString="Enghoff H. 2020. A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1 - 32. https: // doi. org / 10.5852 / ejt. 2020.626" type="journal article" year="2020">Enghoff 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8BE7ED2FD6300622FF4EFEE9C1DAFE69" blockId="21.[189,1399,265,433]" pageId="21" pageNumber="22">
In the diagnosis of
<taxonomicName id="4C5896ACD6300622FE5AFEEAC7DBFEFB" authority="Enghoff (2020)" authorityName="Enghoff" authorityYear="2020" box="[425,759,265,291]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B92C313DD6300622FE5AFEEAC719FEFB" box="[425,565,265,291]" italics="true" pageId="21" pageNumber="22">A. fasciatus</emphasis>
<bibRefCitation id="EFC990DED6300622FDCCFEEAC7DBFEFB" author="Enghoff H." box="[575,759,265,291]" pageId="21" pageNumber="22" pagination="1 - 32" refId="ref47656" refString="Enghoff H. 2020. A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1 - 32. https: // doi. org / 10.5852 / ejt. 2020.626" type="journal article" year="2020">Enghoff (2020)</bibRefCitation>
</taxonomicName>
stated: “Shares a short (as broad as long, or slightly longer) gonopodal palette with
<taxonomicName id="4C5896ACD6300622FDBFFECEC60BFE9E" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[588,807,300,327]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="B92C313DD6300622FDBFFECEC60BFE9E" box="[588,807,300,327]" italics="true" pageId="21" pageNumber="22">A. claudiahempae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FCCEFECEC6E1FE9F" authorityName="Enghoff" authorityYear="2020" box="[829,973,301,327]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B92C313DD6300622FCCEFECEC6E1FE9F" box="[829,973,301,327]" italics="true" pageId="21" pageNumber="22">A. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FC10FECEC1BDFE9E" authorityName="Enghoff" authorityYear="2015" box="[995,1169,300,327]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="B92C313DD6300622FC10FECEC1BDFE9E" box="[995,1169,300,327]" italics="true" pageId="21" pageNumber="22">A. stereosathe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FB55FECEC011FE9E" authorityName="Enghoff" authorityYear="2015" box="[1190,1341,300,327]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="B92C313DD6300622FB55FECEC011FE9E" box="[1190,1341,300,327]" italics="true" pageId="21" pageNumber="22">A. submajor</emphasis>
</taxonomicName>
and
<emphasis id="B92C313DD6300622FF4EFEB3C4A0FEB2" box="[189,396,336,362]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C5896ACD6300622FF4EFEB3C4A4FEB2" authorityName="Enghoff" authorityYear="2015" box="[189,392,336,362]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="udzungwensis">A. udzungwensis</taxonomicName>
.
</emphasis>
” This, of course makes no sense it should have been “Shares a short (as broad as long, or slightly longer) gonopodal palette with
<taxonomicName id="4C5896ACD6300622FCD1FE97C6D9FE55" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[802,1013,371,398]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="B92C313DD6300622FCD1FE97C6D9FE55" box="[802,1013,371,398]" italics="true" pageId="21" pageNumber="22">A. claudiahempae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FBF5FE97C19BFE55" authorityName="Frederiksen" authorityYear="2013" box="[1030,1207,371,398]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B92C313DD6300622FBF5FE97C19BFE55" box="[1030,1207,371,398]" italics="true" pageId="21" pageNumber="22">A. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FB3BFE97C05CFE55" authorityName="Enghoff" authorityYear="2015" box="[1224,1392,371,398]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="B92C313DD6300622FB3BFE97C05CFE55" box="[1224,1392,371,398]" italics="true" pageId="21" pageNumber="22">A. stereosathe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5896ACD6300622FF4EFE74C461FE69" authorityName="Enghoff" authorityYear="2015" box="[189,333,407,433]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="B92C313DD6300622FF4EFE74C461FE69" box="[189,333,407,433]" italics="true" pageId="21" pageNumber="22">A. submajor</emphasis>
</taxonomicName>
and
<emphasis id="B92C313DD6300622FE76FE74C77DFE69" box="[389,593,407,433]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C5896ACD6300622FE76FE74C761FE69" authorityName="Enghoff" authorityYear="2015" box="[389,589,407,433]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="udzungwensis">A. udzungwensis</taxonomicName>
.
</emphasis>
” I am grateful to Carlos Martínez for having spotted this.
</paragraph>
</subSubSection>
</treatment>
</document>