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<document id="A94826699CAB3013C0EED3979A55B581" ID-CLB-Dataset="156940" ID-DOI="10.11646/zootaxa.5190.4.7" ID-GBIF-Dataset="39f670af-d883-4a0b-9956-d6e29af453c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138572" ID-ZooBank="763C975C-7D97-497B-BC7C-8A94CD92FBD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664793912077" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="03F14B1A5B42BA37FF24EDF41D5A4FB8" docLanguage="en" docName="zootaxa.5190.4.7.pdf" docOrigin="Zootaxa 5190 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum grandidentatum Shavrin 2022, sp.n." docType="treatment" docVersion="7" lastPageNumber="580" masterDocId="FFC833625B41BA32FFB3E9751C754B19" masterDocTitle="New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="583" masterPageNumber="575" pageNumber="578" updateTime="1699362515216" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="1C6E383F89FB51E8E0EB33AA1DA82E17" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8353929A99704E221702C86EC0025756">
<mods:title id="75C2ACF1A55D7D6FD29568F3390A0FE8">New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
</mods:titleInfo>
<mods:name id="C69899FA01207BFCCBAA9FA2A8DD15CD" type="personal">
<mods:role id="A9F46471006945FA05CE8560EBFAF092">
<mods:roleTerm id="C96BD0959F4FE1450B3DD1B5BB4A86E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A54EC23256F248996D79436E4F4FB0B5">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C042E86712D5AFF95AA3BA7069906DF6">text</mods:typeOfResource>
<mods:relatedItem id="B8AB9961EB2BC539E145F8B01BD469DB" type="host">
<mods:titleInfo id="A1A8F9CBC0B642D04E63059B60BD1906">
<mods:title id="A3A7A4A4170D0660EC029C5B9DCF10E5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2AD509BF924CDF994880E2B673092D7D">
<mods:date id="AFE0A2234D8D77661959E37DD082BA1F">2022</mods:date>
<mods:detail id="75E3EAFED7B90288BE4AECD526C218EA" type="pubDate">
<mods:number id="6E63D0F30485EE1663E8FFB7AABE540A">2022-09-30</mods:number>
</mods:detail>
<mods:detail id="D2B4D6ECE79F5775C70CF38612B99B10" type="volume">
<mods:number id="E83ECCC34074ECCDA4C5CD91E56CEDE3">5190</mods:number>
</mods:detail>
<mods:detail id="D36A5B575511C610F49E11D371321E1C" type="issue">
<mods:number id="533204F0BAF2E20825667F64C525DED0">4</mods:number>
</mods:detail>
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<mods:start id="9B8D4BDEDCF1AA6AB4E111D3AFC962CE">575</mods:start>
<mods:end id="D101173D2AECF04DCB04CF3E174CC081">583</mods:end>
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<mods:classification id="54243BAD05BFDA2DE1A316DF9717824B">journal article</mods:classification>
<mods:identifier id="151113C6B93FC1CD77EF75CF1E704D34" type="CLB-Dataset">156940</mods:identifier>
<mods:identifier id="DD756D3ED7CF9316387AA48481515973" type="DOI">10.11646/zootaxa.5190.4.7</mods:identifier>
<mods:identifier id="9DE5B03488C345D3983AE36ACA9FD963" type="GBIF-Dataset">39f670af-d883-4a0b-9956-d6e29af453c3</mods:identifier>
<mods:identifier id="D4EABE93843FEBCCAA1073FAB8DD143D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7A0815B0E5B3FBCFB34C0012CD977DDD" type="Zenodo-Dep">7138572</mods:identifier>
<mods:identifier id="1D4125754E2B2DB077BE7E796BEF3419" type="ZooBank">763C975C-7D97-497B-BC7C-8A94CD92FBD1</mods:identifier>
</mods:mods>
<treatment id="03F14B1A5B42BA37FF24EDF41D5A4FB8" ID-DOI="http://doi.org/10.5281/zenodo.7138576" ID-GBIF-Taxon="201582201" ID-Zenodo-Dep="7138576" LSID="urn:lsid:plazi:treatment:03F14B1A5B42BA37FF24EDF41D5A4FB8" httpUri="http://treatment.plazi.org/id/03F14B1A5B42BA37FF24EDF41D5A4FB8" lastPageId="5" lastPageNumber="580" pageId="3" pageNumber="578">
<subSubSection id="C342A9875B42BA31FF24EDF41E264F82" box="[151,595,1153,1180]" pageId="3" pageNumber="578" type="nomenclature">
<paragraph id="8BE7FA0C5B42BA31FF24EDF41E264F82" blockId="3.[151,595,1153,1215]" box="[151,595,1153,1180]" pageId="3" pageNumber="578">
<heading id="D0AF4D605B42BA31FF24EDF41E264F82" bold="true" box="[151,595,1153,1180]" fontSize="11" level="1" pageId="3" pageNumber="578" reason="1">
<emphasis id="B92C261E5B42BA31FF24EDF41E264F82" bold="true" box="[151,595,1153,1180]" pageId="3" pageNumber="578">
<taxonomicName id="4C58818F5B42BA31FF24EDF41E644F85" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,529,1153,1180]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="grandidentatum" status="sp. nov.">
<emphasis id="B92C261E5B42BA31FF24EDF41E644F85" bold="true" box="[151,529,1153,1180]" italics="true" pageId="3" pageNumber="578">Amphichroum grandidentatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F9B655B42BA31FDABEDF41E264F82" box="[536,595,1153,1179]" pageId="3" pageNumber="578" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B42BA31FF24EDD01D4E4FA6" box="[151,315,1189,1215]" pageId="3" pageNumber="578" type="description">
<paragraph id="8BE7FA0C5B42BA31FF24EDD01D4E4FA6" blockId="3.[151,595,1153,1215]" box="[151,315,1189,1215]" pageId="3" pageNumber="578">
(
<figureCitation id="1363E6895B42BA31FF2CEDD01D7E4FA6" box="[159,267,1189,1215]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1983,2007]" captionTargetBox="[161,1424,355,1952]" captionTargetId="figure-103@4.[151,1436,345,1959]" captionTargetPageId="4" captionText="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="3" pageNumber="578">Figs. 7–8</figureCitation>
,
<figureCitation id="1363E6895B42BA31FEA4EDD01D474FA6" box="[279,306,1189,1215]" captionStart="FIGURES 11–12" captionStartId="6.[151,264,1738,1762]" captionTargetBox="[159,1427,563,1704]" captionTargetId="figure-109@6.[151,1435,556,1714]" captionTargetPageId="6" captionText="FIGURES 11–12. Habitus of Amphichroum: 11–A. grandidentatum, 12–A. reticulatum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138580" httpUri="https://zenodo.org/record/7138580/files/figure.png" pageId="3" pageNumber="578">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B42BA31FF24ED9D1E524D3B" pageId="3" pageNumber="578" type="materials_examined">
<paragraph id="8BE7FA0C5B42BA31FF24ED9D1D124E8B" blockId="3.[151,1437,1256,2039]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF24ED9D1DB74E1B" bold="true" box="[151,450,1256,1282]" pageId="3" pageNumber="578">Type material examined.</emphasis>
<materialsCitation id="3B30F0515B42BA31FE78ED9D1D124E8B" ID-GBIF-Occurrence="3920575304" collectingDate="2019-07-05" collectionCode="NMPC" collectorName="D. Kral &amp; J. Ruzicka" country="China" county="Gongshan Co." elevation="3900" latitude="28.075" location="Biluo Mts. Pass" longLatPrecision="123" longitude="98.76" municipality="Biluo Mts. Pass" pageId="3" pageNumber="578" specimenCount="1" specimenCount-male="1" stateProvince="YUNNAN PROv." typeStatus="holotype">
<typeStatus id="54E344AE5B42BA31FE78ED9D1E464E1B" box="[459,563,1256,1282]" pageId="3" pageNumber="578" type="holotype">Holotype</typeStatus>
<specimenCount id="9D5E31855B42BA31FD8FED9C1E244E1A" box="[572,593,1257,1283]" pageId="3" pageNumber="578" type="male">♂</specimenCount>
(
<figureCitation id="1363E6895B42BA31FDD1ED9D1EC34E1B" box="[610,694,1256,1282]" captionStart="FIGURES 11–12" captionStartId="6.[151,264,1738,1762]" captionTargetBox="[159,1427,563,1704]" captionTargetId="figure-109@6.[151,1435,556,1714]" captionTargetPageId="6" captionText="FIGURES 11–12. Habitus of Amphichroum: 11–A. grandidentatum, 12–A. reticulatum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138580" httpUri="https://zenodo.org/record/7138580/files/figure.png" pageId="3" pageNumber="578">Fig. 11</figureCitation>
; left antennomeres 10–11 missing): ‘
<collectingCountry id="F34FBA9C5B42BA31FBEAED9D18CE4E1B" box="[1113,1211,1256,1282]" name="China" pageId="3" pageNumber="578">CHINA</collectingCountry>
:
<collectingRegion id="499C34EE5B42BA31FB75ED9D19FE4E1B" box="[1222,1419,1256,1282]" country="China" name="Yunnan" pageId="3" pageNumber="578">YUNNAN PROv.</collectingRegion>
, |
<collectingCounty id="628682805B42BA31FF24EC791D4E4E3F" box="[151,315,1292,1318]" pageId="3" pageNumber="578">Gongshan Co.</collectingCounty>
, |
<collectingMunicipality id="6B8360765B42BA31FEE4EC791E784E3E" box="[343,525,1292,1319]" pageId="3" pageNumber="578">Biluo Mts. Pass</collectingMunicipality>
,
<quantity id="4CA057E95B42BA31FDA9EC791ECC4E3E" box="[538,697,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="3.91" metricValueMin="3.8899999999999997" pageId="3" pageNumber="578" unit="m" value="3900.0" valueMax="3910.0" valueMin="3890.0">
<elevation id="00751D3F5B42BA31FDA9EC791ECC4E3E" box="[538,697,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="3.91" metricValueMin="3.8899999999999997" pageId="3" pageNumber="578" unit="m" value="3900.0" valueMax="3910.0" valueMin="3890.0">3890–3910 m</elevation>
</quantity>
, |
<geoCoordinate id="EE6C9CCB5B42BA31FD67EC781F3F4E31" box="[724,842,1293,1320]" degrees="28" direction="north" minutes="04.5" orientation="latitude" pageId="3" pageNumber="578" precision="92" value="28.075">28°04.5′N</geoCoordinate>
,
<geoCoordinate id="EE6C9CCB5B42BA31FCE4EC781FA24E31" box="[855,983,1293,1320]" degrees="098" direction="east" minutes="45.6" orientation="longitude" pageId="3" pageNumber="578" precision="92" value="98.76">098°45.6′E</geoCoordinate>
, |
<collectorName id="26AD9FDA5B42BA31FC41EC7818904E3E" box="[1010,1253,1292,1319]" pageId="3" pageNumber="578">D. Král &amp; J. Růžička</collectorName>
leg.’ &lt;printed&gt;, ‘
<date id="FFE6DCCC5B42BA31FF2FEC451D614E52" box="[156,276,1328,1355]" pageId="3" pageNumber="578" value="2019-07-05">
<collectingDate id="EFA225245B42BA31FF2FEC451D614E52" box="[156,276,1328,1355]" pageId="3" pageNumber="578" value="2019-07-05">5.vii.2019</collectingDate>
</date>
, sift #06 [Y06], | mixed forest with dominant |
<taxonomicName id="4C58818F5B42BA31FCACEC451F284E53" box="[799,861,1328,1354]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="578" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B42BA31FCACEC451F284E53" box="[799,861,1328,1354]" italics="true" pageId="3" pageNumber="578">Abies</emphasis>
</taxonomicName>
and
<taxonomicName id="4C58818F5B42BA31FC20EC45184D4E53" box="[915,1080,1328,1354]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="578" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B42BA31FC20EC45184D4E53" box="[915,1080,1328,1354]" italics="true" pageId="3" pageNumber="578">Rhododendron</emphasis>
</taxonomicName>
shrubs, | litter under shrubs and along | fallen trunks’ &lt;printed&gt;, ‘HOLOTYPE |
<taxonomicName id="4C58818F5B42BA31FD1AEC211F324E77" authorityName="Kraatz" authorityYear="1857" box="[681,839,1364,1390]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B92C261E5B42BA31FD1AEC211F324E77" box="[681,839,1364,1390]" italics="true" pageId="3" pageNumber="578">Amphichroum</emphasis>
</taxonomicName>
|
<taxonomicName id="4C58818F5B42BA31FCEBEC21187E4E77" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[856,1035,1364,1390]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="grandidentatum" status="sp. nov.">
<emphasis id="B92C261E5B42BA31FCEBEC21187E4E77" box="[856,1035,1364,1390]" italics="true" pageId="3" pageNumber="578">grandidentatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F9B655B42BA31FBA2EC2018304E76" box="[1041,1093,1365,1391]" pageId="3" pageNumber="578" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. des. 2022’ &lt;red, printed&gt; (
<collectionCode id="ED4962C95B42BA31FEB6EC0D1D2E4E8B" box="[261,347,1400,1426]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="578">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74ECE81E524D3B" blockId="3.[151,1437,1256,2039]" pageId="3" pageNumber="578">
<typeStatus id="54E344AE5B42BA31FF74ECE81D4D4EAE" box="[199,312,1437,1463]" pageId="3" pageNumber="578" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5E31855B42BA31FEFBECE91DF94EAE" box="[328,396,1436,1463]" pageId="3" pageNumber="578" type="male">3 ♂♂</specimenCount>
(
<specimenCount id="9D5E31855B42BA31FE13ECE91E394EAF" box="[416,588,1436,1462]" pageId="3" pageNumber="578" type="generic">two specimens</specimenCount>
dissected;
<specimenCount id="9D5E31855B42BA31FD60ECE81F044EAF" box="[723,881,1436,1463]" pageId="3" pageNumber="578" type="generic">one specimen</specimenCount>
without left antennomeres 7–11),
<specimenCount id="9D5E31855B42BA31FAB8ECE9193A4EAE" box="[1291,1359,1436,1463]" pageId="3" pageNumber="578" type="female">5 ♀♀</specimenCount>
(
<specimenCount id="9D5E31855B42BA31FAD1ECE91D794EC3" pageId="3" pageNumber="578" type="generic">three specimens</specimenCount>
without left antennomeres 10–11, 7–11 or 11 respectively): same data as the holotype, with additional red printed label: ‘
<typeStatus id="54E344AE5B42BA31FED6EC911D804EE7" box="[357,501,1508,1534]" pageId="3" pageNumber="578" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C58818F5B42BA31FDB8EC911EDC4EE7" authorityName="Kraatz" authorityYear="1857" box="[523,681,1508,1534]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B92C261E5B42BA31FDB8EC911EDC4EE7" box="[523,681,1508,1534]" italics="true" pageId="3" pageNumber="578">Amphichroum</emphasis>
</taxonomicName>
|
<taxonomicName id="4C58818F5B42BA31FD0DEC911F044EE7" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[702,881,1508,1534]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="grandidentatum" status="sp. nov.">
<emphasis id="B92C261E5B42BA31FD0DEC911F044EE7" box="[702,881,1508,1534]" italics="true" pageId="3" pageNumber="578">grandidentatum</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B42BA31FCCAEC911FC44EE7" bold="true" box="[889,945,1508,1534]" pageId="3" pageNumber="578">
<taxonomicNameLabel id="A21F9B655B42BA31FCCAEC911FC44EE7" box="[889,945,1508,1534]" pageId="3" pageNumber="578" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="ED4962C95B42BA31FB8DEC9118214EE7" box="[1086,1108,1508,1534]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="578" type="Museum">V</collectionCode>
. des. 2022’ &lt;red, printed&gt; (
<specimenCount id="9D5E31855B42BA31FA23EC911CDB4D3A" pageId="3" pageNumber="578" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5E31855B42BA31FF09EF7D1C934D3A" box="[186,230,1544,1571]" pageId="3" pageNumber="578" type="female">1 ♀</specimenCount>
: cSh;
<specimenCount id="9D5E31855B42BA31FE9FEF7D1D184D3A" box="[300,365,1544,1571]" pageId="3" pageNumber="578" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D5E31855B42BA31FECAEF7D1DCF4D3A" box="[377,442,1544,1571]" pageId="3" pageNumber="578" type="female">4 ♀♀</specimenCount>
:
<collectionCode id="ED4962C95B42BA31FE74EF7D1E694D3B" box="[455,540,1544,1570]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="578">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B42BA37FF74EF591D4E48F4" lastPageId="5" lastPageNumber="580" pageId="3" pageNumber="578" type="description">
<paragraph id="8BE7FA0C5B42BA31FF74EF59180C4D97" blockId="3.[151,1437,1256,2039]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF74EF591D2C4D5F" bold="true" box="[199,345,1580,1606]" pageId="3" pageNumber="578">Description.</emphasis>
Measurements (n=9): HW: 0.64–0.68; HL: 0.41–0.44; AL(
<typeStatus id="54E344AE5B42BA31FBB3EF5918124D5F" box="[1024,1127,1580,1606]" pageId="3" pageNumber="578" type="holotype">holotype</typeStatus>
): 1.68; OL: 0.16–0.20; PL: 0.56–0.67;PW:0.98–1.17; ESL:1.26–1.30;EW:1.31–1.37;AW:1.19–1.29;MTbL(
<typeStatus id="54E344AE5B42BA31FB9EEF2518E64D73" box="[1069,1171,1616,1642]" pageId="3" pageNumber="578" type="holotype">holotype</typeStatus>
):0.72;MTrL(
<typeStatus id="54E344AE5B42BA31FA9FEF2519E44D73" box="[1324,1425,1616,1642]" pageId="3" pageNumber="578" type="holotype">holotype</typeStatus>
): 0.61 (MTrL 1–4: 0.32; MTrL 5: 0.29); AedL: 0.55–0.58; TL: 2.95–3.70 (
<typeStatus id="54E344AE5B42BA31FC76EF0118584D97" box="[965,1069,1652,1678]" pageId="3" pageNumber="578" type="holotype">holotype</typeStatus>
: 3.32).
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EFED1E9E4CEE" blockId="3.[151,1437,1256,2039]" pageId="3" pageNumber="578">
Habitus as in
<figureCitation id="1363E6895B42BA31FED8EFED1DB54DAB" box="[363,448,1688,1714]" captionStart="FIGURES 11–12" captionStartId="6.[151,264,1738,1762]" captionTargetBox="[159,1427,563,1704]" captionTargetId="figure-109@6.[151,1435,556,1714]" captionTargetPageId="6" captionText="FIGURES 11–12. Habitus of Amphichroum: 11–A. grandidentatum, 12–A. reticulatum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138580" httpUri="https://zenodo.org/record/7138580/files/figure.png" pageId="3" pageNumber="578">Fig. 11</figureCitation>
. Head, antennomeres 4–11, pronotum and abdomen brown or reddish-brown (frontal portion and infraorbital ridges of head and sometimes paratergites distinctly paler); lateral and basal portions of pronotum and elytra yellow-brown; mouthparts, antennomeres 1–3 and legs yellow (two preapical segments of maxillary palpi darker, brown or reddish-brown). Head with very dense transverse microsculpture, sometimes finer in middle and coarser on infraorbital ridges; neck with dense isodiametric microreticulation; pronotum with dense isodiametric microsculpture, missing in mediobasal third in some
<typeStatus id="54E344AE5B42BA31FC3AEE381F8D4C7E" box="[905,1016,1869,1895]" pageId="3" pageNumber="578" type="paratype">paratypes</typeStatus>
; visible part of scutellum with dense isodiametric meshes. Head with sparse and moderately large punctation, denser and deeper in middle, finer, sparser and sometimes indistinct on infraorbital ridges; pronotum with moderately dense punctation, slightly finer than that in middle portion of head, sometimes sparser in middle and mediobasal portions; punctation of elytra denser, larger and deeper than that on pronotum, finer along suture.
</paragraph>
<paragraph id="8BE7FA0C5B45BA36FF74E9E2184A4A58" blockId="4.[151,1437,150,321]" pageId="4" pageNumber="579">Head 1.5 times as broad as long; anteocellar foveae narrow and very deep, diagonally stretching toward supraantennal elevation. Ocelli very large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palpi slightly longer than precending segment, from basal portion strongly narrowed toward acute apex.Antennomeres 5–7 about as long as and slightly broader than 4, 8–10 distinctly shorter and slightly broader than 7, apical antennomere 1.5–1.7 times as long as 10.</paragraph>
<caption id="DF27AA845B45BA36FF24EECA19504CE3" ID-DOI="http://doi.org/10.5281/zenodo.7138578" ID-Zenodo-Dep="7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="4" pageNumber="579" startId="4.[151,264,1983,2007]" targetBox="[161,1424,355,1952]" targetPageId="4">
<paragraph id="8BE7FA0C5B45BA36FF24EECA19504CE3" blockId="4.[151,1436,1983,2043]" pageId="4" pageNumber="579">
<emphasis id="B92C261E5B45BA36FF24EECA1D3F4CCE" bold="true" box="[151,330,1983,2007]" pageId="4" pageNumber="579">FIGURES 3–10.</emphasis>
Aedeagus of
<taxonomicName id="4C58818F5B45BA36FE6EEECA1E194CCE" authorityName="Kraatz" authorityYear="1857" box="[477,620,1983,2007]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B92C261E5B45BA36FE6EEECA1E194CCE" box="[477,620,1983,2007]" italics="true" pageId="4" pageNumber="579">Amphichroum</emphasis>
</taxonomicName>
(Figs. 3–5:
<taxonomicName id="4C58818F5B45BA36FD5DEECA1F154CCE" authorityName="Shavrin" authorityYear="2022" box="[750,864,1983,2007]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B92C261E5B45BA36FD5DEECA1E884CCF" box="[750,765,1983,2006]" italics="true" pageId="4" pageNumber="579">A</emphasis>
.
<emphasis id="B92C261E5B45BA36FCBEEECA1F154CCE" box="[781,864,1983,2007]" italics="true" pageId="4" pageNumber="579">discolor</emphasis>
</taxonomicName>
; Fig. 6 (apical portion):
<taxonomicName id="4C58818F5B45BA36FBDBEECA188E4CCE" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1128,1275,1983,2007]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B92C261E5B45BA36FBDBEECA18024CCF" box="[1128,1143,1983,2006]" italics="true" pageId="4" pageNumber="579">A</emphasis>
.
<emphasis id="B92C261E5B45BA36FB35EECA188E4CCE" box="[1158,1275,1983,2007]" italics="true" pageId="4" pageNumber="579">rotundatum</emphasis>
</taxonomicName>
; Figs. 7–8:
<taxonomicName id="4C58818F5B45BA36FA35EECA1D4C4CE2" authorityName="Shavrin" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="species" species="grandidentatum">
<emphasis id="B92C261E5B45BA36FA35EECA19E04CCF" box="[1414,1429,1983,2006]" italics="true" pageId="4" pageNumber="579">A</emphasis>
.
<emphasis id="B92C261E5B45BA36FF24EE961D4C4CE2" box="[151,313,2019,2043]" italics="true" pageId="4" pageNumber="579">grandidentatum</emphasis>
</taxonomicName>
; Figs. 9–10:
<taxonomicName id="4C58818F5B45BA36FE0CEE961E3A4CE2" authorityName="Coiffait" authorityYear="1978" box="[447,591,2019,2043]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis id="B92C261E5B45BA36FE0CEE961DBB4CE3" box="[447,462,2019,2042]" italics="true" pageId="4" pageNumber="579">A</emphasis>
.
<emphasis id="B92C261E5B45BA36FE68EE961E3A4CE2" box="[475,591,2019,2043]" italics="true" pageId="4" pageNumber="579">reticulatum</emphasis>
</taxonomicName>
): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE7FA0C5B44BA37FF74E9E21DD64BE0" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">Pronotum transverse, 1.7 times as broad as long, 1.5–1.7 times as broad as head, disitnctly more narrowed anteriad than posteriad; widely rounded anterior angles distinctly protruded anteriad; lateral portions very wide, flattened and explanate.</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74E8761DB04A58" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">Elytra wide, distinctly broader than long, reaching basal portion of abdominal tergite IV, about or more than twice as long as pronotum.</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74E83E1F7B4A7C" blockId="5.[151,1437,150,1185]" box="[199,782,331,357]" pageId="5" pageNumber="580">Metatibia 1.1 times as long as very long metatarsus.</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74E81A1D0F4AB4" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">Abdomen distinctly narrower than elytra, with a pair of indistinct and very wide transverse tomentose spots in middle of tergite IV.</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74E8C21E6849B0" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">
Male. Medial margin of apical half of protibia with two parallel rows of several moderately long peg setae; mesotibia strongly curved mediad, with dense row of 18–20 short thorns. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII widely rounded. Aedeagus with very wide basal portion, strongly narrowed towards rounded apex; parameres moderately wide, long and slightly curved, significantly exceeding somewhat rounded apex of median lobe, with one short apical and one preapical setae; internal sac long, with a row of large sclerotized thorns in middle and four strongly sclerotized, very large teeth in basal portions (
<figureCitation id="1363E6895B44BA37FB46EB1E194D499C" box="[1269,1336,619,645]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1983,2007]" captionTargetBox="[161,1424,355,1952]" captionTargetId="figure-103@4.[151,1436,345,1959]" captionTargetPageId="4" captionText="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="5" pageNumber="580">Fig. 7</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="1363E6895B44BA37FE67EBFA1E6D49B0" box="[468,536,655,681]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1983,2007]" captionTargetBox="[161,1424,355,1952]" captionTargetId="figure-103@4.[151,1436,345,1959]" captionTargetPageId="4" captionText="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="5" pageNumber="580">Fig. 8</figureCitation>
.
</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74EBC61F9349E8" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">Female. Medial margin of protibia and mesotibia without modifications. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74EB8F1D4E48F4" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">
<emphasis id="B92C261E5B44BA37FF74EB8F1DC7480C" bold="true" box="[199,434,762,789]" pageId="5" pageNumber="580">Comparative notes.</emphasis>
<taxonomicName id="4C58818F5B44BA37FE08EB8E1F61480C" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[443,788,763,789]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="grandidentatum" status="sp. nov.">
<emphasis id="B92C261E5B44BA37FE08EB8E1F61480C" box="[443,788,763,789]" italics="true" pageId="5" pageNumber="580">Amphichroum grandidentatum</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B44BA37FCAFEB8E1F21480C" bold="true" box="[796,852,763,789]" pageId="5" pageNumber="580">
<taxonomicNameLabel id="A21F9B655B44BA37FCAFEB8E1F21480C" box="[796,852,763,789]" pageId="5" pageNumber="580" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from remaining species of the genus by the presence of large sclerotized teeth in basal portion of the internal sac of the aedeagus. Regarding the shape of the median lobe with rounded apex and the length of the parameres considerably exceeding apex of median lobe, it is somewhat similar to
<taxonomicName id="4C58818F5B44BA37FE33EA121F334898" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[384,838,871,897]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B92C261E5B44BA37FE33EA121DE44898" box="[384,401,871,897]" italics="true" pageId="5" pageNumber="580">A</emphasis>
.
<emphasis id="B92C261E5B44BA37FE2DEA121E554898" box="[414,544,871,897]" italics="true" pageId="5" pageNumber="580">rotundatum</emphasis>
<bibRefCitation id="EFC987FD5B44BA37FD95EA121F334898" author="Shavrin, A. V. &amp; Smetana, A." box="[550,838,871,897]" pageId="5" pageNumber="580" pagination="377 - 402" refId="ref5394" refString="Shavrin, A. V. &amp; Smetana, A. (2018) New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China. Zootaxa, 4508 (3), 377 - 402. https: // doi. org / 10.11646 / zootaxa. 4508.3.3" type="journal article" year="2018">Shavrin &amp; Smetana, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C58818F5B44BA37FCC8EA1218D84898" authority="Shavrin, 2019" authorityName="Shavrin" authorityYear="2019" box="[891,1197,871,897]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="feldmanni">
<emphasis id="B92C261E5B44BA37FCC8EA121FF94898" box="[891,908,871,897]" italics="true" pageId="5" pageNumber="580">A</emphasis>
.
<emphasis id="B92C261E5B44BA37FC29EA12187F4898" box="[922,1034,871,897]" italics="true" pageId="5" pageNumber="580">feldmanni</emphasis>
<bibRefCitation id="EFC987FD5B44BA37FBA3EA1218D84898" author="Shavrin, A. V." box="[1040,1197,871,897]" pageId="5" pageNumber="580" pagination="197 - 200" refId="ref5218" refString="Shavrin, A. V. (2019) A new species and records of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from Sichuan, China. Zootaxa, 4701 (2), 197 - 200. https: // doi. org / 10.11646 / zootaxa. 4701.2.7" type="journal article" year="2019">Shavrin, 2019</bibRefCitation>
</taxonomicName>
(both from
<collectingRegion id="499C34EE5B44BA37FA81EA1219E44898" box="[1330,1425,871,897]" country="China" name="Sichuan" pageId="5" pageNumber="580">Sichuan</collectingRegion>
), but can be distinguished from them by the different coloration of the body and the morphology of the aedeagus. Additionally, it differs from
<taxonomicName id="4C58818F5B44BA37FE62EADA1F7748D0" authority="Shavrin, 2019" authorityName="Shavrin" authorityYear="2019" box="[465,770,943,969]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="feldmanni">
<emphasis id="B92C261E5B44BA37FE62EADA1D9748D0" box="[465,482,943,969]" italics="true" pageId="5" pageNumber="580">A</emphasis>
.
<emphasis id="B92C261E5B44BA37FE43EADA1E1548D0" box="[496,608,943,969]" italics="true" pageId="5" pageNumber="580">feldmanni</emphasis>
<bibRefCitation id="EFC987FD5B44BA37FDD5EADA1F7748D0" author="Shavrin, A. V." box="[614,770,943,969]" pageId="5" pageNumber="580" pagination="197 - 200" refId="ref5218" refString="Shavrin, A. V. (2019) A new species and records of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from Sichuan, China. Zootaxa, 4701 (2), 197 - 200. https: // doi. org / 10.11646 / zootaxa. 4701.2.7" type="journal article" year="2019">Shavrin, 2019</bibRefCitation>
</taxonomicName>
by the smaller body, more transverse pronotum and slightly broader elytra.
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B44BA37FF74EA8319344F2C" pageId="5" pageNumber="580" type="distribution">
<paragraph id="8BE7FA0C5B44BA37FF74EA8319264F08" blockId="5.[151,1437,150,1185]" box="[199,1363,1014,1041]" pageId="5" pageNumber="580">
<emphasis id="B92C261E5B44BA37FF74EA831D144F09" bold="true" box="[199,353,1014,1040]" pageId="5" pageNumber="580">Distribution.</emphasis>
The species is at present known only from the
<typeStatus id="54E344AE5B44BA37FCC2EA821FD44F08" box="[881,929,1015,1041]" pageId="5" pageNumber="580">type</typeStatus>
locality in Biluo Mts.,
<collectingRegion id="499C34EE5B44BA37FB17EA8218884F08" box="[1188,1277,1015,1041]" country="China" name="Yunnan" pageId="5" pageNumber="580">Yunnan</collectingRegion>
,
<collectingCountry id="F34FBA9C5B44BA37FABBEA82193A4F08" box="[1288,1359,1015,1041]" name="China" pageId="5" pageNumber="580">China</collectingCountry>
.
</paragraph>
<paragraph id="8BE7FA0C5B44BA37FF74ED6F19344F2C" blockId="5.[151,1437,150,1185]" box="[199,1345,1050,1077]" pageId="5" pageNumber="580">
<emphasis id="B92C261E5B44BA37FF74ED6F1D3E4F2D" bold="true" box="[199,331,1050,1076]" pageId="5" pageNumber="580">Bionomics.</emphasis>
Specimens were collected in the same locality together with
<taxonomicName id="4C58818F5B44BA37FC4BED6E18014F2C" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1016,1140,1051,1077]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">
<emphasis id="B92C261E5B44BA37FC4BED6E187C4F2C" box="[1016,1033,1051,1077]" italics="true" pageId="5" pageNumber="580">A</emphasis>
.
<emphasis id="B92C261E5B44BA37FBABED6E18014F2C" box="[1048,1140,1051,1077]" italics="true" pageId="5" pageNumber="580">discolor</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B44BA37FBC9ED6E18C74F2C" bold="true" box="[1146,1202,1051,1077]" pageId="5" pageNumber="580">
<taxonomicNameLabel id="A21F9B655B44BA37FBC9ED6E18C74F2C" box="[1146,1202,1051,1077]" pageId="5" pageNumber="580" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B44BA37FF74ED4A1D5A4FB8" pageId="5" pageNumber="580" type="etymology">
<paragraph id="8BE7FA0C5B44BA37FF74ED4A1D5A4FB8" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="580">
<emphasis id="B92C261E5B44BA37FF74ED4A1D3B4F40" bold="true" box="[199,334,1087,1113]" pageId="5" pageNumber="580">Etymology.</emphasis>
The specific epithet is an adjective composed of the Latin adjectives
<emphasis id="B92C261E5B44BA37FB27ED4A189F4F40" box="[1172,1258,1087,1113]" italics="true" pageId="5" pageNumber="580">grandis</emphasis>
, -
<emphasis id="B92C261E5B44BA37FAB6ED4A19644F41" box="[1285,1297,1087,1112]" italics="true" pageId="5" pageNumber="580">e</emphasis>
(large) and
<emphasis id="B92C261E5B44BA37FF24ED161C834F64" box="[151,246,1123,1149]" italics="true" pageId="5" pageNumber="580">dentatus</emphasis>
, -
<emphasis id="B92C261E5B44BA37FEBEED161D6E4F65" box="[269,283,1123,1148]" italics="true" pageId="5" pageNumber="580">a</emphasis>
, -
<emphasis id="B92C261E5B44BA37FE81ED161D214F65" box="[306,340,1123,1148]" italics="true" pageId="5" pageNumber="580">um</emphasis>
(toothed). It alludes to the presence of large sclerotized teeth in basal portion of the internal sac of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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