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<mods:title id="DFC81176680C9E82DCA1F60497E588DD">The genus Leucophenga (Diptera, Drosophilidae), part VIII: twenty-one species from the Oriental region, with morphological and molecular evidence</mods:title>
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<mods:namePart id="3AD0D00D2CF720500F9DEA58313B4ED4">O’Grady, Patrick Michael</mods:namePart>
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<mods:namePart id="2771F9DC0E04877AD6314E62F8314164">Su, Yirui</mods:namePart>
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<mods:namePart id="17EC2F26DB8946D9FF5C05BBE7B13BD2">Chen, Hongwei</mods:namePart>
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<heading id="D0B08A1CFFF4FF89DE812326C862F9D4" bold="true" box="[151,587,1553,1580]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<taxonomicName id="4C4746F3FFF4FF89DE812326C862F9D4" ID-CoL="8CJFZ" authority="Duda, 1926" authorityName="Duda" authorityYear="1926" box="[151,587,1553,1580]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B933E162FFF4FF89DE812326C862F9D4" bold="true" box="[151,587,1553,1580]" pageId="25" pageNumber="26">
<emphasis id="B933E162FFF4FF89DE812326CB9EF9D3" bold="true" box="[151,439,1553,1579]" italics="true" pageId="25" pageNumber="26">Leucophenga jacobsoni</emphasis>
<bibRefCitation id="EFD64081FFF4FF89DFAB2325C862F9D4" author="Duda, O." box="[445,587,1554,1580]" pageId="25" pageNumber="26" pagination="42 - 116" refId="ref45409" refString="Duda, O. (1926) Fauna sumatrensis. (Beitrag Nr. 26). Drosophilidae (Dipt.). Supplementa Entomologica, 14, 42 - 116." type="journal article" year="1926">Duda, 1926</bibRefCitation>
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<paragraph id="8BF83D70FFF4FF89DE812303C819F9B5" blockId="25.[151,587,1553,1613]" box="[151,560,1588,1613]" pageId="25" pageNumber="26">
(
<figureCitation id="137C21F5FFF4FF89DE892303CB2AF9B5" box="[159,259,1588,1613]" captionStart="FIGURE 2" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[151,1436,637,1954]" captionTargetId="figure-187@15.[151,1436,637,1954]" captionTargetPageId="15" captionText="FIGURE 2. Wing of male: A. Leucophenga jacobsoni; B. L. kurahashii; C. L. setipalpis; D. L. sorii; E. L. spinifera; F. L. varinervis; G, H. L. acantha Huang &amp; Chen, sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297898" httpUri="https://zenodo.org/record/5297898/files/figure.png" pageId="25" pageNumber="26">Figs. 2A</figureCitation>
, 
<figureCitation id="137C21F5FFF4FF89DF182303CB7DF9B5" box="[270,340,1588,1613]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">5A–C</figureCitation>
, 
<figureCitation id="137C21F5FFF4FF89DF492303CBACF9B5" box="[351,389,1588,1613]" captionStart="FIGURE 9" captionStartId="22.[151,250,1964,1986]" captionTargetBox="[297,1287,213,1941]" captionTargetId="figure-81@22.[278,1307,193,1951]" captionTargetPageId="22" captionText="FIGURE 9. Colour patterns of abdominal tergites: A. Leucophenga jacobsoni; B, C. L. kurahashii; D, E. L. setipalpis; F. L. sorii; G, H. L. spinifera; I, J. L. varinervis; K. L. acantha Huang &amp; Chen, sp. nov.; L. L. alafumosa Huang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5297912" httpUri="https://zenodo.org/record/5297912/files/figure.png" pageId="25" pageNumber="26">9A</figureCitation>
, 
<figureCitation id="137C21F5FFF4FF89DF862303CBD6F9B5" box="[400,511,1588,1613]" captionStart="FIGURE 13" captionStartId="26.[151,250,1980,2002]" captionTargetBox="[347,1247,201,1969]" captionTargetId="figure-66@26.[347,1247,193,1969]" captionTargetPageId="26" captionText="FIGURE 13. Male terminalia (provided as a supplement for the line drawing of each species): A, B. Leucophenga jacobsoni; C, D. L. kurahashii; E, F. L. setipalpis; G, H. L. sorii; I, J. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297914" httpUri="https://zenodo.org/record/5297914/files/figure.png" pageId="25" pageNumber="26">13A, 13B</figureCitation>
, 
<figureCitation id="137C21F5FFF4FF89DC1D2303C80EF9B5" box="[523,551,1588,1613]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="25" pageNumber="26">17</figureCitation>
)
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<treatmentCitation id="0AE61B61FFF4FF89DE81234BC807F96A" author="Duda, O." box="[151,558,1660,1682]" page="50" pageId="25" pageNumber="26" year="1926">
<taxonomicName id="4C4746F3FFF4FF89DE81234BC807F96A" ID-CoL="8CJFZ" authority="Duda, 1926: 50" authorityName="Duda" authorityPageNumber="50" authorityYear="1926" box="[151,558,1660,1682]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B933E162FFF4FF89DE81234BCBA1F96A" box="[151,392,1660,1682]" italics="true" pageId="25" pageNumber="26">Leucophenga jacobsoni</emphasis>
<bibRefCitation id="EFD64081FFF4FF89DF98234BC807F96A" author="Duda, O." box="[398,558,1660,1682]" pageId="25" pageNumber="26" pagination="42 - 116" refId="ref45409" refString="Duda, O. (1926) Fauna sumatrensis. (Beitrag Nr. 26). Drosophilidae (Dipt.). Supplementa Entomologica, 14, 42 - 116." type="journal article" year="1926">Duda, 1926: 50</bibRefCitation>
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; 
<treatmentCitation id="0AE61B61FFF4FF89DC2E234BC8FFF96A" author="Okada, T." box="[568,726,1660,1682]" page="2" pageId="25" pageNumber="26" year="1976">
<bibRefCitation id="EFD64081FFF4FF89DC2E234BC8FFF96A" author="Okada, T." box="[568,726,1660,1682]" pageId="25" pageNumber="26" pagination="1 - 8" refId="ref46822" refString="Okada, T. (1976) New distribution records of the drosophilids in the Oriental Region. Acta Dipterologica, 8, 1 - 8." type="journal article" year="1976">Okada, 1976: 2</bibRefCitation>
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; 
<treatmentCitation id="0AE61B61FFF4FF89DCF7234BC914F96A" author="Okada, T." box="[737,829,1660,1682]" page="93" pageId="25" pageNumber="26" year="1987">
<bibRefCitation id="EFD64081FFF4FF89DCF7234BC914F96A" author="Okada, T." box="[737,829,1660,1682]" pageId="25" pageNumber="26" pagination="90 - 99" refId="ref46849" refString="Okada, T. (1987) The Leucophenga proxima Species Group of Southeast Asia and New Guinea (Diptera, Drosophilidae). Kontyu, 55, 90 - 99." type="journal article" year="1987">1987: 93</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BF83D70FFF4FF89DE812388CBEFF8D8" blockId="25.[151,1436,1727,2006]" pageId="25" pageNumber="26">
<emphasis id="B933E162FFF4FF89DE812388CB38F920" bold="true" box="[151,273,1727,1752]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
This species differs from the other species of the genus 
<taxonomicName id="4C4746F3FFF4FF89DD832388CE05F92F" box="[917,1068,1727,1751]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B933E162FFF4FF89DD832388CE05F92F" box="[917,1068,1727,1751]" italics="true" pageId="25" pageNumber="26">Leucophenga</emphasis>
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in having the surstylus caudally curved and narrow (
<figureCitation id="137C21F5FFF4FF89DF6923D4CBC3F904" box="[383,490,1763,1788]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="25" pageNumber="26">Fig. 17A</figureCitation>
); aedeagus apically acute, laterally with two long, acute processes, each bearing a sensillum at tip (
<figureCitation id="137C21F5FFF4FF89DF472230CB90F8D8" box="[337,441,1799,1824]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="25" pageNumber="26">Fig. 17D</figureCitation>
).
</paragraph>
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<paragraph id="8BF83D70FFF4FF8EDED1221CC86AFE76" blockId="25.[151,1436,1727,2006]" lastBlockId="30.[151,1436,151,650]" lastPageId="30" lastPageNumber="31" pageId="25" pageNumber="26">
<emphasis id="B933E162FFF4FF89DED1221CCB70F8BC" bold="true" box="[199,345,1835,1860]" pageId="25" pageNumber="26">Description.</emphasis>
Body small. Ocellar triangle dark brown, lacking seta above ocellar setae. Postocellar setae minute (
<figureCitation id="137C21F5FFF4FF89DEEC2278CB73F890" box="[250,346,1871,1896]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
). Frons yellow (
<figureCitation id="137C21F5FFF4FF89DC342278C8ABF890" box="[546,642,1871,1896]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
). Orb 
<subScript id="17C33F35FFF4FF89DCD12269C8E6F894" attach="left" box="[711,719,1886,1900]" fontSize="6" pageId="25" pageNumber="26">3</subScript>
and Orb 
<subScript id="17C33F35FFF4FF89DD2A2269C96DF894" attach="left" box="[828,836,1886,1900]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
parallel (
<figureCitation id="137C21F5FFF4FF89DDAC2278CE33F890" box="[954,1050,1871,1896]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
). Pedicel yellow (
<figureCitation id="137C21F5FFF4FF89DAE32278CF7CF890" box="[1269,1365,1871,1896]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
); first flagellomere yellowish (
<figureCitation id="137C21F5FFF4FF89DFA42242C838F876" box="[434,529,1909,1934]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
). Clypeus entirely yellow. Palpus yellow (
<figureCitation id="137C21F5FFF4FF89DA1E2242CE4EF876" box="[1032,1127,1909,1934]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5A</figureCitation>
). Mesonotum mostly dark yellow (
<figureCitation id="137C21F5FFF4FF89DEE322AECB79F84A" box="[245,336,1945,1970]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5B</figureCitation>
). Postpronotal lobe yellow (
<figureCitation id="137C21F5FFF4FF89DC8022AEC8D8F84A" box="[662,753,1945,1970]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5C</figureCitation>
). Acrostichal setulae in ca. 10–12 irregular rows (
<figureCitation id="137C21F5FFF4FF89DB2222AECFA6F84A" box="[1332,1423,1945,1970]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5B</figureCitation>
). Scutellum dark yellow, brownish laterally, yellow at tip (
<figureCitation id="137C21F5FFF4FF89DD05228AC942F82E" box="[787,875,1981,2006]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="25" pageNumber="26">Fig. 5B</figureCitation>
). Pleura, katepisternum, and mesopleuron mostly yellow (
<figureCitation id="137C21F5FFF3FF8EDEE225A0CB66FF48" box="[244,335,151,176]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="30" pageNumber="31">Fig. 5C</figureCitation>
). Wing (
<figureCitation id="137C21F5FFF3FF8EDFA525A0C838FF48" box="[435,529,151,176]" captionStart="FIGURE 2" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[151,1436,637,1954]" captionTargetId="figure-187@15.[151,1436,637,1954]" captionTargetPageId="15" captionText="FIGURE 2. Wing of male: A. Leucophenga jacobsoni; B. L. kurahashii; C. L. setipalpis; D. L. sorii; E. L. spinifera; F. L. varinervis; G, H. L. acantha Huang &amp; Chen, sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297898" httpUri="https://zenodo.org/record/5297898/files/figure.png" pageId="30" pageNumber="31">Fig. 2A</figureCitation>
): entirely hyaline; costal vein between R 
<subScript id="17C33F35FFF3FF8EDDF22591C9D5FF4C" attach="left" box="[996,1020,166,180]" fontSize="6" pageId="30" pageNumber="31">2+3</subScript>
and R 
<subScript id="17C33F35FFF3FF8EDA5F2591CE48FF4C" attach="left" box="[1097,1121,166,180]" fontSize="6" pageId="30" pageNumber="31">4+5</subScript>
distally with ca. 7 peg-like spinules on ventral surface; R 
<subScript id="17C33F35FFF3FF8EDFE625FAC821FF23" attach="left" box="[496,520,205,219]" fontSize="6" pageId="30" pageNumber="31">4+5</subScript>
and M 
<subScript id="17C33F35FFF3FF8EDC4825FAC84FFF23" attach="left" box="[606,614,205,219]" fontSize="6" pageId="30" pageNumber="31">1</subScript>
nearly parallel distally; halter entirely yellowish (
<figureCitation id="137C21F5FFF3FF8EDABD2589CF2EFF2F" box="[1195,1287,190,215]" captionStart="FIGURE 5" captionStartId="18.[151,250,1962,1984]" captionTargetBox="[173,1416,193,1944]" captionTargetId="figure-70@18.[173,1416,193,1944]" captionTargetPageId="18" captionText="FIGURE 5. Lateral view of head, dorsal and lateral views of thorax in males: A, B, C. Leucophenga jacobsoni; D, E, F. L. kurahashii; G, H, I. L. setipalpis; J, K, L. L. sorii; M, N, O. L. spinifera." figureDoi="http://doi.org/10.5281/zenodo.5297904" httpUri="https://zenodo.org/record/5297904/files/figure.png" pageId="30" pageNumber="31">Fig. 5B</figureCitation>
). Abdominal tergites brown, 2nd tergite with yellow patches medially, laterally and posteriorly, 3rd tergite entirely yellowish, setigerous only on posterior margin, 4th tergite yellow anteriorly in males (
<figureCitation id="137C21F5FFF3FF8EDDFA243ECE6EFEDA" box="[1004,1095,265,290]" captionStart="FIGURE 9" captionStartId="22.[151,250,1964,1986]" captionTargetBox="[297,1287,213,1941]" captionTargetId="figure-81@22.[278,1307,193,1951]" captionTargetPageId="22" captionText="FIGURE 9. Colour patterns of abdominal tergites: A. Leucophenga jacobsoni; B, C. L. kurahashii; D, E. L. setipalpis; F. L. sorii; G, H. L. spinifera; I, J. L. varinervis; K. L. acantha Huang &amp; Chen, sp. nov.; L. L. alafumosa Huang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5297912" httpUri="https://zenodo.org/record/5297912/files/figure.png" pageId="30" pageNumber="31">Fig. 9A</figureCitation>
). Male terminalia: Epandrium pubescent, with ca. 4 setae near posterior margin per side (
<figureCitation id="137C21F5FFF3FF8EDD45241AC9E9FEBE" box="[851,960,301,326]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="30" pageNumber="31">Fig. 17A</figureCitation>
). Cercus lacking pubescence (
<figureCitation id="137C21F5FFF3FF8EDB35241ACFB9FEBE" box="[1315,1424,301,326]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="30" pageNumber="31">Fig. 17A</figureCitation>
). Surstylus pubescent, with only a few setae along distal margin (
<figureCitation id="137C21F5FFF3FF8EDD602466C9C8FE92" box="[886,993,337,362]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="30" pageNumber="31">Fig. 17A</figureCitation>
). Paramere small, rounded basally and apically, lacking sensillum (
<figureCitation id="137C21F5FFF3FF8EDFC62442C81FFE76" box="[464,566,373,398]" captionStart="FIGURE 17" captionStartId="30.[151,250,1907,1929]" captionTargetBox="[152,1435,709,1896]" captionTargetId="figure-292@30.[151,1436,705,1897]" captionTargetPageId="30" captionText="FIGURE 17. Leucophenga jacobsoni Duda, 1926, male terminalia in lateral view. A. Epandrium (epand), surstylus (sur), and cercus (cerc); B. Hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed), and aedeagal apodeme (aed a); C. Paramere; D. Aedeagus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5297922" httpUri="https://zenodo.org/record/5297922/files/figure.png" pageId="30" pageNumber="31">Fig. 17C</figureCitation>
).
</paragraph>
<caption id="DF386DF8FFF7FF8ADE81228BC974F808" ID-DOI="http://doi.org/10.5281/zenodo.5297914" ID-Zenodo-Dep="5297914" httpUri="https://zenodo.org/record/5297914/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1980,2002]" targetBox="[347,1247,201,1969]" targetPageId="26">
<paragraph id="8BF83D70FFF7FF8ADE81228BC974F808" blockId="26.[151,1436,1980,2032]" pageId="26" pageNumber="27">
<emphasis id="B933E162FFF7FF8ADE81228BCB09F829" bold="true" box="[151,288,1980,2002]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
Male terminalia (provided as a supplement for the line drawing of each species): A, B. 
<taxonomicName id="4C4746F3FFF7FF8ADAB5228BCFBDF82A" authorityName="Duda" authorityYear="1926" box="[1187,1428,1980,2002]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B933E162FFF7FF8ADAB5228BCFBDF82A" box="[1187,1428,1980,2002]" italics="true" pageId="26" pageNumber="27">Leucophenga jacobsoni</emphasis>
</taxonomicName>
; C, D. 
<taxonomicName id="4C4746F3FFF7FF8ADEC522ECCB73F808" authorityName="Okada" authorityYear="1987" box="[211,346,2010,2032]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kurahashii">
<emphasis id="B933E162FFF7FF8ADEC522ECCAC8F808" box="[211,225,2011,2032]" italics="true" pageId="26" pageNumber="27">L</emphasis>
. 
<emphasis id="B933E162FFF7FF8ADEF822EDCB73F808" box="[238,346,2010,2032]" italics="true" pageId="26" pageNumber="27">kurahashii</emphasis>
</taxonomicName>
; E, F. 
<taxonomicName id="4C4746F3FFF7FF8ADF8A22ECC831F808" authorityName="Duda" authorityYear="1923" box="[412,536,2010,2032]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="setipalpis">
<emphasis id="B933E162FFF7FF8ADF8A22ECCB83F808" box="[412,426,2011,2032]" italics="true" pageId="26" pageNumber="27">L</emphasis>
. 
<emphasis id="B933E162FFF7FF8ADFA122EDC831F808" box="[439,536,2010,2032]" italics="true" pageId="26" pageNumber="27">setipalpis</emphasis>
</taxonomicName>
; G, H. 
<taxonomicName id="4C4746F3FFF7FF8ADC7622ECC880F808" authorityName="Kang, Lee &amp; Bhang" authorityYear="1965" box="[608,681,2011,2032]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sorii">
<emphasis id="B933E162FFF7FF8ADC7622ECC847F808" box="[608,622,2011,2032]" italics="true" pageId="26" pageNumber="27">L</emphasis>
. 
<emphasis id="B933E162FFF7FF8ADC6D22ECC880F808" box="[635,681,2011,2032]" italics="true" pageId="26" pageNumber="27">sorii</emphasis>
</taxonomicName>
; I, J. 
<taxonomicName id="4C4746F3FFF7FF8ADCF522ECC97EF808" authorityName="Okada" authorityYear="1987" box="[739,855,2010,2032]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B933E162FFF7FF8ADCF522ECC8D8F808" box="[739,753,2011,2032]" italics="true" pageId="26" pageNumber="27">L</emphasis>
. 
<emphasis id="B933E162FFF7FF8ADCEB22EDC97EF808" box="[765,855,2010,2032]" italics="true" pageId="26" pageNumber="27">spinifera</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF386DF8FFF6FF8BDE812280CF07F814" ID-DOI="http://doi.org/10.5281/zenodo.5297916" ID-Zenodo-Dep="5297916" httpUri="https://zenodo.org/record/5297916/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1975,1997]" targetBox="[346,1244,193,1965]" targetPageId="27">
<paragraph id="8BF83D70FFF6FF8BDE812280CF07F814" blockId="27.[151,1436,1975,2028]" pageId="27" pageNumber="28">
<emphasis id="B933E162FFF6FF8BDE812280CB08F834" bold="true" box="[151,289,1975,1997]" pageId="27" pageNumber="28">FIGURE 14.</emphasis>
Male terminalia: A, B. 
<taxonomicName id="4C4746F3FFF6FF8BDC0F2280C939F835" authorityName="Duda" authorityYear="1923" box="[537,784,1975,1997]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="varinervis">
<emphasis id="B933E162FFF6FF8BDC0F2280C939F835" box="[537,784,1975,1997]" italics="true" pageId="27" pageNumber="28">Leucophenga varinervis</emphasis>
</taxonomicName>
; C, D. 
<taxonomicName id="4C4746F3FFF6FF8BDD48228FCE5BF835" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[862,1138,1975,1997]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="acantha" status="sp. nov.">
<emphasis id="B933E162FFF6FF8BDD48228FC945F835" box="[862,876,1976,1997]" italics="true" pageId="27" pageNumber="28">L</emphasis>
. 
<emphasis id="B933E162FFF6FF8BDD6C2280C9E5F835" box="[890,972,1975,1997]" italics="true" pageId="27" pageNumber="28">acantha</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF6FF8BDA6B228FCEE6F835" bold="true" box="[1149,1231,1976,1997]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A2005C19FFF6FF8BDA6B228FCEE6F835" box="[1149,1231,1976,1997]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E, F. 
<taxonomicName id="4C4746F3FFF6FF8BDB00228FCB1AF814" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="alafumosa" status="sp. nov.">
<emphasis id="B933E162FFF6FF8BDB00228FCF0DF835" box="[1302,1316,1976,1997]" italics="true" pageId="27" pageNumber="28">L</emphasis>
. 
<emphasis id="B933E162FFF6FF8BDB242280CFB5F835" box="[1330,1436,1975,1997]" italics="true" pageId="27" pageNumber="28">alafumosa</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF6FF8BDF2B22E0CBA4F814" bold="true" box="[317,397,2007,2028]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A2005C19FFF6FF8BDF2B22E0CBA4F814" box="[317,397,2007,2028]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; G, H. 
<taxonomicName id="4C4746F3FFF6FF8BDFC022E0C8D2F814" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[470,763,2006,2028]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevipenis" status="sp. nov.">
<emphasis id="B933E162FFF6FF8BDFC022E0CBCDF814" box="[470,484,2007,2028]" italics="true" pageId="27" pageNumber="28">L</emphasis>
. 
<emphasis id="B933E162FFF6FF8BDFE622E1C871F814" box="[496,600,2006,2028]" italics="true" pageId="27" pageNumber="28">brevipenis</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF6FF8BDD1322E0C97FF814" bold="true" box="[773,854,2007,2028]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A2005C19FFF6FF8BDD1322E0C97FF814" box="[773,854,2007,2028]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I, J. 
<taxonomicName id="4C4746F3FFF6FF8BDD9922E0CEFAF814" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[911,1235,2006,2028]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevitabulata" status="sp. nov.">
<emphasis id="B933E162FFF6FF8BDD9922E0C9B4F814" box="[911,925,2007,2028]" italics="true" pageId="27" pageNumber="28">L</emphasis>
. 
<emphasis id="B933E162FFF6FF8BDDBC22E1CE18F814" box="[938,1073,2006,2028]" italics="true" pageId="27" pageNumber="28">brevitabulata</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<taxonomicNameLabel id="A2005C19FFF6FF8BDACB22E0CF07F814" box="[1245,1326,2007,2028]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF386DF8FFF1FF8CDE8122ABCACEF816" ID-DOI="http://doi.org/10.5281/zenodo.5297918" ID-Zenodo-Dep="5297918" httpUri="https://zenodo.org/record/5297918/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1948,1970]" targetBox="[354,1232,202,1926]" targetPageId="28">
<paragraph id="8BF83D70FFF1FF8CDE8122ABCACEF816" blockId="28.[151,1436,1948,2030]" pageId="28" pageNumber="29">
<emphasis id="B933E162FFF1FF8CDE8122ABCB09F849" bold="true" box="[151,288,1948,1970]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
Male terminalia: A, B. 
<taxonomicName id="4C4746F3FFF1FF8CDC0522ABC951F84A" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[531,888,1948,1970]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="delta" status="sp. nov.">
<emphasis id="B933E162FFF1FF8CDC0522ABC8FCF84A" box="[531,725,1948,1970]" italics="true" pageId="28" pageNumber="29">Leucophenga delta</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF1FF8CDD9422ABC9FAF849" bold="true" box="[898,979,1948,1969]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2005C19FFF1FF8CDD9422ABC9FAF849" box="[898,979,1948,1969]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C, D. 
<taxonomicName id="4C4746F3FFF1FF8CDA0922ABCF10F84A" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[1055,1337,1948,1970]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="forcipula" status="sp. nov.">
<emphasis id="B933E162FFF1FF8CDA0922ABCE04F849" box="[1055,1069,1948,1969]" italics="true" pageId="28" pageNumber="29">L</emphasis>
. 
<emphasis id="B933E162FFF1FF8CDA2C22ABCEBFF84A" box="[1082,1174,1948,1970]" italics="true" pageId="28" pageNumber="29">forcipula</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF1FF8CDB5222ABCFBDF849" bold="true" box="[1348,1428,1948,1969]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2005C19FFF1FF8CDB5222ABCFBDF849" box="[1348,1428,1948,1969]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E, F. 
<taxonomicName id="4C4746F3FFF1FF8CDEDD228CC82DF828" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[203,516,1978,2000]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fuscipalpula" status="sp. nov.">
<emphasis id="B933E162FFF1FF8CDEDD228CCAF0F828" box="[203,217,1979,2000]" italics="true" pageId="28" pageNumber="29">L</emphasis>
. 
<emphasis id="B933E162FFF1FF8CDEF3228DCB4BF828" box="[229,354,1978,2000]" italics="true" pageId="28" pageNumber="29">fuscipalpula</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF1FF8CDC1B228CC877F828" bold="true" box="[525,606,1979,2000]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2005C19FFF1FF8CDC1B228CC877F828" box="[525,606,1979,2000]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; G, H. 
<taxonomicName id="4C4746F3FFF1FF8CDCB3228CC9CBF828" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[677,994,1978,2000]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="glabtabulata" status="sp. nov.">
<emphasis id="B933E162FFF1FF8CDCB3228CC89AF828" box="[677,691,1979,2000]" italics="true" pageId="28" pageNumber="29">L</emphasis>
. 
<emphasis id="B933E162FFF1FF8CDCD6228DC96BF828" box="[704,834,1978,2000]" italics="true" pageId="28" pageNumber="29">glabtabulata</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF1FF8CDDFA228CCE15F828" bold="true" box="[1004,1084,1979,2000]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2005C19FFF1FF8CDDFA228CCE15F828" box="[1004,1084,1979,2000]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I, J. 
<taxonomicName id="4C4746F3FFF1FF8CDA62228CCFB1F828" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[1140,1432,1978,2000]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="helvipecta" status="sp. nov.">
<emphasis id="B933E162FFF1FF8CDA62228CCEABF828" box="[1140,1154,1979,2000]" italics="true" pageId="28" pageNumber="29">L</emphasis>
. 
<emphasis id="B933E162FFF1FF8CDA99228DCEDEF828" box="[1167,1271,1978,2000]" italics="true" pageId="28" pageNumber="29">helvipecta</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<taxonomicNameLabel id="A2005C19FFF1FF8CDE8122EECACEF816" box="[151,231,2009,2030]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF386DF8FFF0FF8DDE8122B8C8BAF81A" ID-DOI="http://doi.org/10.5281/zenodo.5297920" ID-Zenodo-Dep="5297920" httpUri="https://zenodo.org/record/5297920/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1935,1957]" targetBox="[428,1154,193,1916]" targetPageId="29">
<paragraph id="8BF83D70FFF0FF8DDE8122B8C8BAF81A" blockId="29.[151,1436,1935,2018]" pageId="29" pageNumber="30">
<emphasis id="B933E162FFF0FF8DDE8122B8CB09F85C" bold="true" box="[151,288,1935,1957]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
Male terminalia: A, B. 
<taxonomicName id="4C4746F3FFF0FF8DDC0222B8C99DF85D" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[532,948,1935,1957]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hyaloptera" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDC0222B8C938F85D" box="[532,785,1935,1957]" italics="true" pageId="29" pageNumber="30">Leucophenga hyaloptera</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF0FF8DDDA922B8CE39F85C" bold="true" box="[959,1040,1935,1956]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2005C19FFF0FF8DDDA922B8CE39F85C" box="[959,1040,1935,1956]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C, D. 
<taxonomicName id="4C4746F3FFF0FF8DDA4A22A7CF5AF85D" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[1116,1395,1935,1957]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oxyptera" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDA4A22A7CE43F85D" box="[1116,1130,1936,1957]" italics="true" pageId="29" pageNumber="30">L</emphasis>
. 
<emphasis id="B933E162FFF0FF8DDA6122A7CEF9F85D" box="[1143,1232,1936,1957]" italics="true" pageId="29" pageNumber="30">oxyptera</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF0FF8DDB6B22B8CAEAF83B" bold="true" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2005C19FFF0FF8DDB6B22B8CAEAF83B" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E, F. 
<taxonomicName id="4C4746F3FFF0FF8DDF132298C80DF83C" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[261,548,1966,1988]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="platypyga" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDF132298CB3AF83C" box="[261,275,1967,1988]" italics="true" pageId="29" pageNumber="30">L</emphasis>
. 
<emphasis id="B933E162FFF0FF8DDF092299CBAAF83C" box="[287,387,1966,1988]" italics="true" pageId="29" pageNumber="30">platypyga</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF0FF8DDC382299C856F83B" bold="true" box="[558,639,1966,1987]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2005C19FFF0FF8DDC382299C856F83B" box="[558,639,1966,1987]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; G, H. 
<taxonomicName id="4C4746F3FFF0FF8DDCD12298C9D6F83C" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[711,1023,1966,1988]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="serrateiceps" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDCD12298C8FCF83C" box="[711,725,1967,1988]" italics="true" pageId="29" pageNumber="30">L</emphasis>
. 
<emphasis id="B933E162FFF0FF8DDCF72298C974F83C" box="[737,861,1967,1988]" italics="true" pageId="29" pageNumber="30">serrateiceps</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF0FF8DDA1E2299CE70F83B" bold="true" box="[1032,1113,1966,1987]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2005C19FFF0FF8DDA1E2299CE70F83B" box="[1032,1113,1966,1987]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I, J. 
<taxonomicName id="4C4746F3FFF0FF8DDA842298CFB1F83C" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[1170,1432,1966,1988]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="valvata" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDA842298CE89F83C" box="[1170,1184,1967,1988]" italics="true" pageId="29" pageNumber="30">L</emphasis>
. 
<emphasis id="B933E162FFF0FF8DDABA2299CEDEF83C" box="[1196,1271,1966,1988]" italics="true" pageId="29" pageNumber="30">valvata</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<emphasis id="B933E162FFF0FF8DDE8122FACACEF81A" bold="true" box="[151,231,1997,2018]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2005C19FFF0FF8DDE8122FACACEF81A" box="[151,231,1997,2018]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; K, L. 
<taxonomicName id="4C4746F3FFF0FF8DDF2622FAC810F81A" authority="Huang &amp; Chen" authorityName="Huang &amp; Wang &amp; O’Grady &amp; Su &amp; Chen" authorityYear="2018" box="[304,569,1996,2018]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zebrina" status="sp. nov.">
<emphasis id="B933E162FFF0FF8DDF2622FACB17F81A" box="[304,318,1997,2018]" italics="true" pageId="29" pageNumber="30">L</emphasis>
. 
<emphasis id="B933E162FFF0FF8DDF5D22FBCBBEF81A" box="[331,407,1996,2018]" italics="true" pageId="29" pageNumber="30">zebrina</emphasis>
Huang &amp; Chen
</taxonomicName>
, 
<taxonomicNameLabel id="A2005C19FFF0FF8DDC5522FAC8BAF81A" box="[579,659,1997,2018]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF83D70FFF3FF8EDED124AEC876FDE6" blockId="30.[151,1436,151,650]" pageId="30" pageNumber="31">
<emphasis id="B933E162FFF3FF8EDED124AEC823FE4A" bold="true" box="[199,522,409,434]" pageId="30" pageNumber="31">Measurements and indices.</emphasis>
BL = 
<quantity id="4CBF9095FFF3FF8EDC4524AEC891FE49" box="[595,696,409,434]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="31" unit="mm" value="2.4">2.40 mm</quantity>
, ThL = 
<quantity id="4CBF9095FFF3FF8EDD0724AEC95EFE49" box="[785,887,409,434]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="30" pageNumber="31" unit="mm" value="1.17">1.17 mm</quantity>
, WL = 
<quantity id="4CBF9095FFF3FF8EDDDA24AECE1BFE49" box="[972,1074,409,434]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="30" pageNumber="31" unit="mm" value="2.18">2.18 mm</quantity>
, WW = 1.00 mm, arb = 7/4, avd = 0.90, adf = 2.21, flw = 1.71, FW/HW = 0.32, ch/o = 0.02, prorb = 0.067, rcorb = 0.74, vb = 0.44, dcl = 0.50, presctl = 0.55, sctl = 1.20, sterno = 0.64, orbito = 1.64, dcp = 0.20, sctlp = 1.00, C = 2.14, 4c = 1.24, 4v = 1.90, 5x = 1.36, ac = 2.25, M = 0.66, C3F = 0.75.
</paragraph>
<paragraph id="8BF83D70FFF3FF8EDED1271EC8A3FD9E" blockId="30.[151,1436,151,650]" pageId="30" pageNumber="31">
<emphasis id="B933E162FFF3FF8EDED1271ECB97FDBA" bold="true" box="[199,446,553,578]" pageId="30" pageNumber="31">Specimen examined.</emphasis>
<materialsCitation id="3B2F372DFFF3FF8EDFD1271EC8AFFD9E" collectingDate="2004-12-01" collectionCode="SCAU" collectorName="HW Chen" country="China" elevation="220" location="Ledong" municipality="Jianfengling" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<collectingCountry id="F3507DE0FFF3FF8EDFD1271EC80DFDBA" box="[455,548,553,578]" name="China" pageId="30" pageNumber="31">CHINA</collectingCountry>
: 
<specimenCount id="9D41F6F9FFF3FF8EDC27271EC87DFDBA" box="[561,596,553,578]" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
(
<collectionCode id="ED56A5B5FFF3FF8EDC73271EC89DFDBA" box="[613,692,553,578]" pageId="30" pageNumber="31">SCAU</collectionCode>
, No. 128504), 
<collectingMunicipality id="6B9CA70AFFF3FF8EDD74271EC9C6FDBA" box="[866,1007,553,578]" pageId="30" pageNumber="31">Jianfengling</collectingMunicipality>
, 
<location id="8E986BABFFF3FF8EDDEB271ECE7CFDBA" LSID="urn:lsid:plazi:treatment:03EE8C66FFF4FF8EDE812326CFB5FD72:8E986BABFFF3FF8EDDEB271ECE7CFDBA" box="[1021,1109,553,578]" country="China" municipality="Jianfengling" name="Ledong" pageId="30" pageNumber="31" stateProvince="Hainan">Ledong</location>
, 
<collectingRegion id="4983F392FFF3FF8EDA75271ECE9EFDBA" box="[1123,1207,553,578]" country="China" name="Hainan" pageId="30" pageNumber="31">Hainan</collectingRegion>
, 18˚41'N, 108˚52'E, alt. 
<quantity id="4CBF9095FFF3FF8EDED7277ACB2AFD9E" box="[193,259,589,614]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="30" pageNumber="31" unit="m" value="220.0">
<elevation id="006ADA43FFF3FF8EDED7277ACB2AFD9E" box="[193,259,589,614]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="30" pageNumber="31" unit="m" value="220.0">220m</elevation>
</quantity>
, 
<date id="FFF91BB0FFF3FF8EDF18277ACBABFD9E" box="[270,386,589,614]" pageId="30" pageNumber="31" value="2004-12-01">
<collectingDate id="EFBDE258FFF3FF8EDF18277ACBABFD9E" box="[270,386,589,614]" pageId="30" pageNumber="31" value="2004-12-01">1.xii.2004</collectingDate>
</date>
, ex tussock, 
<collectorName id="26B258A6FFF3FF8EDC042779C8AFFD9E" box="[530,646,589,614]" pageId="30" pageNumber="31">HW Chen</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D6EFBFFF3FF8EDED12746CFB5FD72" box="[199,1436,625,650]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BF83D70FFF3FF8EDED12746CFB5FD72" blockId="30.[151,1436,151,650]" box="[199,1436,625,650]" pageId="30" pageNumber="31">
<emphasis id="B933E162FFF3FF8EDED12746CB49FD72" bold="true" box="[199,352,625,650]" pageId="30" pageNumber="31">Distribution.</emphasis>
<collectingCountry id="F3507DE0FFF3FF8EDF702746CB80FD72" box="[358,425,625,650]" name="China" pageId="30" pageNumber="31">China</collectingCountry>
(
<collectingRegion id="4983F392FFF3FF8EDFAE2746C827FD72" box="[440,526,625,650]" country="China" name="Hainan" pageId="30" pageNumber="31">Hainan</collectingRegion>
), 
<collectingCountry id="F3507DE0FFF3FF8EDC362746C8ADFD72" box="[544,644,625,650]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
(Khao Yai), 
<collectingCountry id="F3507DE0FFF3FF8EDD052746C953FD72" box="[787,890,625,650]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
(Malaya), 
<collectingCountry id="F3507DE0FFF3FF8EDDE22746CE41FD72" box="[1012,1128,625,650]" name="Singapore" pageId="30" pageNumber="31">Singapore</collectingCountry>
, 
<collectingCountry id="F3507DE0FFF3FF8EDA652746CEC9FD72" box="[1139,1248,625,650]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
(
<collectingRegion id="4983F392FFF3FF8EDAF82746CF79FD72" box="[1262,1360,625,650]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">Sumatra</collectingRegion>
, 
<collectingRegion id="4983F392FFF3FF8EDB4C2746CFB9FD72" box="[1370,1424,625,650]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="31">Java</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>