treatments-xml/data/03/E7/87/03E787FA6708F17CFE88FC8E7028213A.xml

<document id="5D0DC226404D9C01CB0EEF0E3FA08D2C" ID-CLB-Dataset="6032" ID-DOI="10.11606/1807-0205/2020.60.56" ID-GBIF-Dataset="03bba471-dfa8-471a-933c-aff4a3e8779b" ID-ISSN="1807-0205" ID-Zenodo-Dep="4614061" ID-ZooBank="FCF6CE24-EFB5-4BD8-856F-032740367771" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615997364254" checkinUser="carolina" docAuthor="Araujo-Vieira, Katyuscia, Caramaschi, Ulisses, Novaes-e-Fagundes, Gabriel, Orrico, Victor G. D. &amp; Faivovich, Julián" docDate="2020" docId="03E787FA6708F17CFE88FC8E7028213A" docLanguage="en" docName="PapAvZoo.60.e20206056.pdf" docOrigin="Papéis Avulsos de Zoologia 60" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Scinax x-signatus" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFDEFF82670EF16EFFBDFFF472572709" masterDocTitle="A neotype for Hyla x-signata Spix, 1824 (Amphibia, Anura, Hylidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="7" updateTime="1698933916446" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
<mods:mods id="2CB023D581940A3C415871B586D03CBF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6E3BC1221AD8AEFECEE7DC8B71E0AF8">
<mods:title id="5EF3B4219B2305028E71F3F376574585">A neotype for Hyla x-signata Spix, 1824 (Amphibia, Anura, Hylidae)</mods:title>
</mods:titleInfo>
<mods:name id="32F1350C6FF04506D60D98B359534DF3" type="personal">
<mods:role id="275DC8263BBB83D508B2DDE70958F539">
<mods:roleTerm id="654959ED0D57CB6E2DC6F63E137D87F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3AF466A5A2BC5DD3E04B016B8DF3F2B2">Araujo-Vieira, Katyuscia</mods:namePart>
</mods:name>
<mods:name id="266B32539B8B70603F9A6A75CF0D3DF0" type="personal">
<mods:role id="2F67F0E0E316E4F7E6DE166A2A41E49C">
<mods:roleTerm id="CB2DA71C2A1A36DEF3D69B7274273275">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0669B35C0ED4B9F9B2E78100DB4E629">Caramaschi, Ulisses</mods:namePart>
</mods:name>
<mods:name id="9519A99C56C5DAFAE9B3E08C236DE81B" type="personal">
<mods:role id="3330B52B1B19003C90A62E834E614056">
<mods:roleTerm id="2E1EBFFC508BB3B6DEDE4D2678CC4317">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37BAA4E3937421F82982C5FA8BABCDA7">Novaes-e-Fagundes, Gabriel</mods:namePart>
</mods:name>
<mods:name id="8F2113FF0DE41F52A5056D02DA2E1651" type="personal">
<mods:role id="1630EF5D3271355DA23D3B9985A93DB6">
<mods:roleTerm id="787CC88AAC3014A40B0DD06EED941FDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D4C7C1C638D81C4C486F420C44A4E1A">Orrico, Victor G. D.</mods:namePart>
</mods:name>
<mods:name id="7E0E2FDDB9781A269AA13D7882C08997" type="personal">
<mods:role id="1453EF335051CA6B1E4B9212CC353386">
<mods:roleTerm id="6F28AD9B7B45E433CB49B8D54ABCBF9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="89893C9B73B4B5612E75A0F352628E98">Faivovich, Julián</mods:namePart>
</mods:name>
<mods:typeOfResource id="BB7B3A76B98AB899421F9D76FC76A223">text</mods:typeOfResource>
<mods:relatedItem id="B4941AD568717FA75F72DC4EC4663EED" type="host">
<mods:titleInfo id="FA8962FD4B678987D5692B672047EE5A">
<mods:title id="E2146469E6CA45D30EA25D98F81EA75E">Papéis Avulsos de Zoologia</mods:title>
</mods:titleInfo>
<mods:part id="E097F84293FD3E799A42B09E6E351940">
<mods:date id="28FCB5E8646CF418C104F068AD0BF9FB">2020</mods:date>
<mods:detail id="291E1AEDA7184F4B471DB3C4294EAA6B" type="pubDate">
<mods:number id="EF6C5E22E594E1646F12DA8B22CD1B87">2020-11-11</mods:number>
</mods:detail>
<mods:detail id="E810A2AB1716C63688945E4AB7A732A7" type="volume">
<mods:number id="C883E8219EB1EDB046CA95EBE8324B7C">60</mods:number>
</mods:detail>
<mods:extent id="DA4A5738D05D2B9378B6771113FC8F06" unit="page">
<mods:start id="8D40498D8BCC2FB356A599E533F898FD">1</mods:start>
<mods:end id="FC013696604BAE31A3184DFD79E9E4FE">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F92B0459C596E3178BEC7F142749898F">journal article</mods:classification>
<mods:identifier id="A2190FB70981AAB645C732989A7AD9FB" type="CLB-Dataset">6032</mods:identifier>
<mods:identifier id="CD430DC615C14991B44FC994EDAC842C" type="DOI">10.11606/1807-0205/2020.60.56</mods:identifier>
<mods:identifier id="95C84A6EFD82AAAE6370A03A89B4D9B7" type="GBIF-Dataset">03bba471-dfa8-471a-933c-aff4a3e8779b</mods:identifier>
<mods:identifier id="2D1F186E7068128E3668DD3B1CBF1EA4" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="3F5EC8FFB5DB1E02F62E194937711874" type="Zenodo-Dep">4614061</mods:identifier>
<mods:identifier id="39671B9327DB4EB7C8CDEB3E8AB221A2" type="ZooBank">FCF6CE24-EFB5-4BD8-856F-032740367771</mods:identifier>
</mods:mods>
<treatment id="03E787FA6708F17CFE88FC8E7028213A" ID-DOI="http://doi.org/10.5281/zenodo.4975570" ID-GBIF-Taxon="182952894" ID-Zenodo-Dep="4975570" LSID="urn:lsid:plazi:treatment:03E787FA6708F17CFE88FC8E7028213A" httpUri="http://treatment.plazi.org/id/03E787FA6708F17CFE88FC8E7028213A" lastPageId="18" lastPageNumber="18" pageId="6" pageNumber="7">
<subSubSection id="C35465676708F168FE88FC8E70D9249A" box="[309,654,890,915]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BF136EC6708F168FE88FC8E70D9249A" blockId="6.[309,654,890,915]" box="[309,654,890,915]" pageId="6" pageNumber="7">
<heading id="D0B981806708F168FE88FC8E70D9249A" box="[309,654,890,915]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="5">
<taxonomicName id="4C4E4D6F6708F168FE88FC8E70D9249A" ID-CoL="4VJZX" authority="(Spix, 1824)" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[309,654,890,915]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FE88FC8E70D9249A" bold="true" box="[309,654,890,915]" pageId="6" pageNumber="7">
<emphasis id="B93AEAFE6708F168FE88FC8E73A9249A" bold="true" box="[309,510,890,915]" italics="true" pageId="6" pageNumber="7">Scinax x‑signatus</emphasis>
(
<bibRefCitation id="EFDF4B1D6708F168FDB7FC8E70D0249A" author="Spix, J. B." box="[522,647,890,915]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix, 1824</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676708F168FF17FC4E714C2F5A" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BF136EC6708F168FF17FC4E739624DA" blockId="6.[170,795,954,2131]" box="[170,449,954,979]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17FC4E73EA24DA" ID-CoL="4ZYM" authority="Spix, 1824" box="[170,445,954,979]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FF17FC4E731324DA" box="[170,324,954,979]" italics="true" pageId="6" pageNumber="7">Hyla x‑signata</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FEF7FC4E73EA24DA" author="Spix, J. B." box="[330,445,954,979]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix, 1824</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF136EC6708F168FF17FC2E73B323FA" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17FC2E73F724FA" ID-CoL="3NCJF" authority="Spix, 1824" authorityName="Spix" authorityYear="1824" box="[170,416,986,1011]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="affinis">
<emphasis id="B93AEAFE6708F168FF17FC2E737324FA" box="[170,292,986,1011]" italics="true" pageId="6" pageNumber="7">Hyla affinis</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FE91FC2E73F724FA" author="Spix, J. B." box="[300,416,986,1011]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix, 1824</bibRefCitation>
</taxonomicName>
– Considered a synonym of 
<taxonomicName id="4C4E4D6F6708F168FD54FC2E7363231A" ID-CoL="4ZYM" authority="Spix, 1824" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FD54FC2E7363231A" italics="true" pageId="6" pageNumber="7">Hyla x‑signata</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFDF4B1D6708F168FEDFFC0E70F4231A" author="Hoogmoed, M. S. &amp; Gruber, U." box="[354,675,1018,1043]" pageId="6" pageNumber="7" pagination="319 - 345" refId="ref21766" refString="Hoogmoed, M. S. &amp; Gruber, U. 1983. Spix andWagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (The Netherlands). Spixiana, 9: 319 - 345." type="journal article" year="1983">Hoogmoed &amp; Gruber (1983)</bibRefCitation>
. 
<bibRefCitation id="EFDF4B1D6708F168FD12FC0E733F233A" author="Sturaro, M. J. &amp; Peloso, P. L. V." pageId="6" pageNumber="7" pagination="9 - 23" refId="ref25030" refString="Sturaro, M. J. &amp; Peloso, P. L. V. 2014. A new species of Scinax Wagler, 1830 (Anura: Hylidae) from the middle Amazon River basin, Brazil. Papeis Avulsos de Zoologia, 54 (2): 9 - 23." type="journal article" year="2014">Sturaro &amp; Peloso (2014)</bibRefCitation>
questioned this association based on the description and figure provided by 
<bibRefCitation id="EFDF4B1D6708F168FD2BFBCE7141235A" author="Spix, J. B." box="[662,790,1082,1107]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix (1824)</bibRefCitation>
. Our study of photographs of the 
<typeStatus id="54F5884E6708F168FDD3FBAE7084237A" box="[622,723,1114,1139]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
(ZSM 2945) indicates that the situation is uncertain. Only a study of the taxonomy of amazonian populations associated with 
<taxonomicName id="4C4E4D6F6708F168FED1FB4E707223DA" ID-CoL="4VJZX" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[364,549,1210,1235]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FED1FB4E707223DA" box="[364,549,1210,1235]" italics="true" pageId="6" pageNumber="7">Scinax x‑signatus</emphasis>
</taxonomicName>
would allow to clarify the status of this nomen.
</paragraph>
<paragraph id="8BF136EC6708F168FF17FB0E73CF213A" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17FB0E73E5221A" ID-CoL="3NCQM" authority="Spix, 1824" authorityName="Spix" authorityYear="1824" box="[170,434,1274,1299]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="coerulea">
<emphasis id="B93AEAFE6708F168FF17FB0E736B221A" box="[170,316,1274,1299]" italics="true" pageId="6" pageNumber="7">Hyla coerulea</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FEFCFB0E73E5221A" author="Spix, J. B." box="[321,434,1274,1299]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix, 1824</bibRefCitation>
</taxonomicName>
– Considered a synonym of 
<taxonomicName id="4C4E4D6F6708F168FD54FB0E7363223A" ID-CoL="3NDFD" authority="Spix, 1824" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signata">
<emphasis id="B93AEAFE6708F168FD54FB0E7363223A" italics="true" pageId="6" pageNumber="7">Hyla x‑signata</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFDF4B1D6708F168FEDFFAEE70F4223A" author="Hoogmoed, M. S. &amp; Gruber, U." box="[354,675,1306,1331]" pageId="6" pageNumber="7" pagination="319 - 345" refId="ref21766" refString="Hoogmoed, M. S. &amp; Gruber, U. 1983. Spix andWagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (The Netherlands). Spixiana, 9: 319 - 345." type="journal article" year="1983">Hoogmoed &amp; Gruber (1983)</bibRefCitation>
. 
<bibRefCitation id="EFDF4B1D6708F168FD12FAEE733F225A" author="Sturaro, M. J. &amp; Peloso, P. L. V." pageId="6" pageNumber="7" pagination="9 - 23" refId="ref25030" refString="Sturaro, M. J. &amp; Peloso, P. L. V. 2014. A new species of Scinax Wagler, 1830 (Anura: Hylidae) from the middle Amazon River basin, Brazil. Papeis Avulsos de Zoologia, 54 (2): 9 - 23." type="journal article" year="2014">Sturaro &amp; Peloso (2014)</bibRefCitation>
questioned this association based on the description and figure provided by 
<bibRefCitation id="EFDF4B1D6708F168FD2BFAAE7141227A" author="Spix, J. B." box="[662,790,1370,1395]" pageId="6" pageNumber="7" refId="ref24980" refString="Spix, J. B. d. 1824. Animalia nova sive species novae Testudinum et Ranarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae Regis, Typis Franc. Seraph. Hubschmanni, Monachii, XXXIX + 53 pp." type="book" year="1824">Spix (1824)</bibRefCitation>
. Our study of photographs of the 
<typeStatus id="54F5884E6708F168FDFCFA8E70FD229A" box="[577,682,1402,1427]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
designated by 
<bibRefCitation id="EFDF4B1D6708F168FEA9FA6E701F22BA" author="Hoogmoed, M. S. &amp; Gruber, U." box="[276,584,1434,1459]" pageId="6" pageNumber="7" pagination="319 - 345" refId="ref21766" refString="Hoogmoed, M. S. &amp; Gruber, U. 1983. Spix andWagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (The Netherlands). Spixiana, 9: 319 - 345." type="journal article" year="1983">Hoogmoed &amp; Gruber (1983)</bibRefCitation>
(ZSM 2710-0-1) indicates that the situation is uncertain. Only a study of the taxonomy of amazonian populations associated with 
<taxonomicName id="4C4E4D6F6708F168FEB4FA0E7394211A" ID-CoL="4VJZX" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[265,451,1530,1555]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FEB4FA0E7394211A" box="[265,451,1530,1555]" italics="true" pageId="6" pageNumber="7">Scinax x‑signatus</emphasis>
</taxonomicName>
would allow to clarify the status of this nomen.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F9CE73BD21BA" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F9CE73E2215A" ID-CoL="6MM7Q" authority="Daudin, 1802" authorityName="Daudin" authorityYear="1802" box="[170,437,1594,1619]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FF17F9CE734C215A" box="[170,283,1594,1619]" italics="true" pageId="6" pageNumber="7">Hyla rubra</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FE9CF9CE73E2215A" author="Daudin, F. M." box="[289,437,1594,1619]" pageId="6" pageNumber="7" refId="ref19306" refString="Daudin, F. M. 1802 &quot; An. XI &quot;. Histoire Naturelle des Rainettes, des Grenouilles et des Crapauds. Levrault, Paris. 108 p." type="book" year="1802">Daudin, 1802</bibRefCitation>
</taxonomicName>
(part) – 
<bibRefCitation id="EFDF4B1D6708F168FDAFF9CE7141215A" author="Dumeril, A. M. C. &amp; Bibron, G." box="[530,790,1594,1619]" pageId="6" pageNumber="7" refId="ref20031" refString="Dumeril, A. M. C. &amp; Bibron, G. 1841. Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Tome 8. Comprenant l'Histoire Generale des Batraciens, et la description des cinquante-duex generes et des cent soixante-trois especes des duex premiers sous-ordres: Les peromeles qui n'ont pas de members, et les anoures qui sont prives de la queue. Librairie Encyclopedique de Roret, Paris." type="book" year="1841">Duméril &amp; Bibron, 1841</bibRefCitation>
. First treatment as a synonym of 
<taxonomicName id="4C4E4D6F6708F168FD83F9AE7350219A" ID-CoL="6MM7Q" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FD83F9AE70E6217A" box="[574,689,1626,1651]" italics="true" pageId="6" pageNumber="7">Hyla rubra</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FD05F9AE7350219A" author="Laurenti, J. N." pageId="6" pageNumber="7" refId="ref22285" refString="Laurenti, J. N. 1768. Specimen Medicum, Exhibens Synopsin Reptilium Emendatum cum Experimentis Circa Venena et Antidota Reptilium Austriacorum. Wien, Joan. Thom. nob. de Trattnern." type="book" year="1768">Laurenti, 1768</bibRefCitation>
</taxonomicName>
(not 
<bibRefCitation id="EFDF4B1D6708F168FEFDF98E7382219A" author="Daudin, F. M." box="[320,469,1658,1683]" pageId="6" pageNumber="7" refId="ref19306" refString="Daudin, F. M. 1802 &quot; An. XI &quot;. Histoire Naturelle des Rainettes, des Grenouilles et des Crapauds. Levrault, Paris. 108 p." type="book" year="1802">Daudin, 1802</bibRefCitation>
; see 
<bibRefCitation id="EFDF4B1D6708F168FDB4F98E70D3219A" author="Leon, J. R." box="[521,644,1658,1683]" pageId="6" pageNumber="7" pagination="505 - 545" refId="ref22320" refString="Leon, J. R. 1969. The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18: 505 - 545." type="journal article" year="1969">León, 1969</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6708F168FD30F98E7141219A" author="Rivero, J. A." box="[653,790,1658,1683]" pageId="6" pageNumber="7" pagination="108 - 119" refId="ref24342" refString="Rivero, J. A. 1969. Sobre la Hyla rubra Laurenti y la Hyla x-signata Spix (Amphibia, Salientia). Memorias de la Sociedad de Historia Natural La Salle, 29: 108 - 119." type="journal article" year="1969">Rivero, 1969</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6708F168FF6DF96E73B721BA" author="Duellman, W. E. &amp; Wiens, J. J." box="[208,480,1690,1715]" pageId="6" pageNumber="7" pagination="1 - 57" refId="ref19931" refString="Duellman, W. E. &amp; Wiens, J. J. 1993. Hylid frogs of the genus Scinax Wagler, 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History, University of Kansas, 153: 1 - 57." type="journal article" year="1993">Duellman &amp; Wiens, 1993</bibRefCitation>
).
</paragraph>
<paragraph id="8BF136EC6708F168FF17F94E73B021FA" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F94E732721DA" authority="(Spix, 1824)" box="[170,368,1722,1747]" class="Amphibia" family="Hylidae" genus="Scytopis" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="xsignatus">
<emphasis id="B93AEAFE6708F168FF17F94E732721DA" box="[170,368,1722,1747]" italics="true" pageId="6" pageNumber="7">Scytopis xsignatus</emphasis>
</taxonomicName>
[sic] – 
<bibRefCitation id="EFDF4B1D6708F168FE78F94E701021DA" author="Cope, E. D." box="[453,583,1722,1747]" pageId="6" pageNumber="7" pagination="147 - 169" refId="ref19137" refString="Cope, E. D. 1870 &quot; 1869 &quot;. Seventh contribution to the herpetology of tropical America. Proceedings of the American Philosophical Society, 11 (81): 147 - 169." type="journal article" year="1870">Cope, 1870</bibRefCitation>
. First combination with 
<taxonomicName id="4C4E4D6F6708F168FEBAF92E73B421FA" authority="Cope, 1862" authorityName="Cope" authorityYear="1862" box="[263,483,1754,1779]" class="Amphibia" family="Hylidae" genus="Scytopis" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FEBAF92E730821FA" box="[263,351,1754,1779]" italics="true" pageId="6" pageNumber="7">Scytopis</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FED8F92E73B421FA" author="Cope, E. D." box="[357,483,1754,1779]" pageId="6" pageNumber="7" pagination="346 - 359" refId="ref19046" refString="Cope, E. D. 1862. Catalogues of the reptiles obtained during the explorations of the Parana, Paraguay, Vermejo and Uruguay Rivers, by Capt. Thos. J. Page, U. S. N.; and of those procured by Lieut. N. Michler, U. S. Top. Eng., Commander of the expedition conducting the survey of the Atrato River. Proceedings of the Academy of Natural Sciences of Philadelphia, 14: 346 - 359." type="journal article" year="1862">Cope, 1862</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F90E707F203A" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F90E73E5201A" ID-CoL="6MM7Q" box="[170,434,1786,1811]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FF17F90E734C201A" box="[170,283,1786,1811]" italics="true" pageId="6" pageNumber="7">Hyla rubra</emphasis>
var. 
<emphasis id="B93AEAFE6708F168FEF3F90E73E5201A" box="[334,434,1786,1811]" italics="true" pageId="6" pageNumber="7">x‑signata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFDF4B1D6708F168FE76F90E7018201A" author="Peters, W. C. H." box="[459,591,1786,1811]" pageId="6" pageNumber="7" pagination="196 - 227" refId="ref23678" refString="Peters, W. C. H. 1872. Uber die von Spix in Brasilien gesammelten batrachier des Konigl. Naturalienkabinets zu Munchen. Monatsberichtederkoniglich Akademie der Wissenschaften zu Berlin, 1872: 196 - 227." type="journal article" year="1872">Peters 1872</bibRefCitation>
. First treatment as a variety of 
<taxonomicName id="4C4E4D6F6708F168FEEDF8EE707F203A" ID-CoL="6MM7Q" authority="Laurenti." authorityName="Laurenti." box="[336,552,1818,1843]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FEEDF8EE7397203A" box="[336,448,1818,1843]" italics="true" pageId="6" pageNumber="7">Hyla rubra</emphasis>
Laurenti.
</taxonomicName>
</paragraph>
<paragraph id="8BF136EC6708F168FF17F8CE70D2207A" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F8CE73DF205A" ID-CoL="6MM7Q" box="[170,392,1850,1875]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FF17F8CE73DF205A" box="[170,392,1850,1875]" italics="true" pageId="6" pageNumber="7">Hyla rubra x‑signata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFDF4B1D6708F168FE1BF8CE7064205A" author="Muller, L." box="[422,563,1850,1875]" pageId="6" pageNumber="7" pagination="257 - 304" refId="ref23064" refString="Muller, L. 1927. Amphibien und Reptilien der Ausbeute Prof. Bresslau's in Brasilien 1913 - 1914. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 40: 257 - 304." type="journal article" year="1927">Müller, 1927</bibRefCitation>
. First treatment as a subspecies of 
<taxonomicName id="4C4E4D6F6708F168FED6F8AE70D6207A" ID-CoL="6MM7Q" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" box="[363,641,1882,1907]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rubra">
<emphasis id="B93AEAFE6708F168FED6F8AE738B207A" box="[363,476,1882,1907]" italics="true" pageId="6" pageNumber="7">Hyla rubra</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FE5CF8AE70D6207A" author="Laurenti, J. N." box="[481,641,1882,1907]" pageId="6" pageNumber="7" refId="ref22285" refString="Laurenti, J. N. 1768. Specimen Medicum, Exhibens Synopsin Reptilium Emendatum cum Experimentis Circa Venena et Antidota Reptilium Austriacorum. Wien, Joan. Thom. nob. de Trattnern." type="book" year="1768">Laurenti, 1768</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F88E730720BA" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F88E73FB209A" ID-CoL="4ZYM" box="[170,428,1914,1939]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FF17F88E73FB209A" box="[170,428,1914,1939]" italics="true" pageId="6" pageNumber="7">Hyla x‑signata x‑signata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFDF4B1D6708F168FE78F88E706F209A" author="Lutz, B." box="[453,568,1914,1939]" pageId="6" pageNumber="7" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
. First use as nominal subspecies.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F84E708920FA" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F84E732E20DA" authorityName="Fitzinger" authorityYear="1843" box="[170,377,1978,2003]" class="Amphibia" family="Hylidae" genus="Ololygon" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signata">
<emphasis id="B93AEAFE6708F168FF17F84E732E20DA" box="[170,377,1978,2003]" italics="true" pageId="6" pageNumber="7">Ololygon x‑signata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFDF4B1D6708F168FE26F84E714120DA" author="Fouquette Jr., M. J. &amp; Delahoussaye, A. J." box="[411,790,1978,2003]" pageId="6" pageNumber="7" pagination="387 - 396" refId="ref20934" refString="Fouquette Jr., M. J. &amp; Delahoussaye, A. J. 1977. Sperm morphology in the Hyla rubra group (Amphibia, Anura, Hylidae), and its bearing on generic status. Journal of Herpetology, 11: 387 - 396." type="journal article" year="1977">Fouquette &amp; Delahoussaye, 1977</bibRefCitation>
. First combination with 
<taxonomicName id="4C4E4D6F6708F168FE72F82E708D20FA" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[463,730,2010,2035]" class="Amphibia" family="Hylidae" genus="Ololygon" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FE72F82E706620FA" box="[463,561,2010,2035]" italics="true" pageId="6" pageNumber="7">Ololygon</emphasis>
Fitzinger, 1843
</taxonomicName>
.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F80E70632F3A" blockId="6.[170,795,954,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FF17F80E730C2F1A" ID-CoL="3NDFD" authorityName="Wagler" authorityYear="1830" box="[170,347,2042,2067]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signata">
<emphasis id="B93AEAFE6708F168FF17F80E730C2F1A" box="[170,347,2042,2067]" italics="true" pageId="6" pageNumber="7">Scinax x‑signata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFDF4B1D6708F168FECAF80E70DC2F1A" author="Duellman, W. E. &amp; Wiens, J. J." box="[375,651,2042,2067]" pageId="6" pageNumber="7" pagination="1 - 23" refId="ref19880" refString="Duellman, W. E. &amp; Wiens, J. J. 1992. The status of the Hylid frog genus Ololygon and the recognition of Scinax Wagler, 1830. Occasional Papers of the Museum of Natural History, University of Kansas, 151: 1 - 23." type="journal article" year="1992">Duellman &amp; Wiens, 1992</bibRefCitation>
. First combination with 
<taxonomicName id="4C4E4D6F6708F168FEE8F7EE70672F3A" ID-CoL="63RG8" authority="Wagler, 1830" authorityName="Wagler" authorityYear="1830" box="[341,560,2074,2099]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE6708F168FEE8F7EE73CC2F3A" box="[341,411,2074,2099]" italics="true" pageId="6" pageNumber="7">Scinax</emphasis>
<bibRefCitation id="EFDF4B1D6708F168FE22F7EE70672F3A" author="Wagler, J." box="[415,560,2074,2099]" pageId="6" pageNumber="7" refId="ref25585" refString="Wagler, J. 1830. Naturliches System der Amphibien: mit vorangehender Classification der Saugethiere und Vogel: ein Beitrag zur vergleichenden Zoologie. Cotta'schen, Munchen." type="book" year="1830">Wagler, 1830</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF136EC6708F168FF17F7CE714C2F5A" blockId="6.[170,795,954,2131]" box="[170,795,2106,2131]" pageId="6" pageNumber="7">
<emphasis id="B93AEAFE6708F168FF17F7CE73362F5A" box="[170,353,2106,2131]" italics="true" pageId="6" pageNumber="7">Scinaxx‑signatus</emphasis>
– 
<bibRefCitation id="EFDF4B1D6708F168FEC8F7CE70312F5A" author="Kohler, J. &amp; Bohme, W." box="[373,614,2106,2131]" pageId="6" pageNumber="7" pagination="133 - 140" refId="ref22066" refString="Kohler, J. &amp; Bohme, W. 1996. Anuran amphibians from the region of Pre-Cambrian rock outcrops (inselbergs) in northeastern Bolivia, with a note on the gender of Scinax Wagler, 1830. Revue Francaise d'Aquariologie, 23: 133 - 140." type="journal article" year="1996">Köhler &amp; Böhme,1996</bibRefCitation>
.Gender change.
</paragraph>
</subSubSection>
<subSection id="E2C12F076708F168FCD8FF2E77CB26DA" pageId="6" pageNumber="7" type="multiple">
<subSubSection id="C35465676708F168FCD8FF2E719C27FA" box="[869,971,218,243]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="8BF136EC6708F168FCD8FF2E719C27FA" blockId="6.[869,971,218,243]" box="[869,971,218,243]" pageId="6" pageNumber="7">
<heading id="D0B981806708F168FCD8FF2E719C27FA" box="[869,971,218,243]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="5">
<emphasis id="B93AEAFE6708F168FCD8FF2E719C27FA" bold="true" box="[869,971,218,243]" pageId="6" pageNumber="7">
<typeStatus id="54F5884E6708F168FCD8FF2E719C27FA" box="[869,971,218,243]" pageId="6" pageNumber="7" type="neotype">Neotype</typeStatus>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676708F168FC36FEEE77CB26DA" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BF136EC6708F168FC36FEEE77CB26DA" blockId="6.[869,1494,282,467]" pageId="6" pageNumber="7">
CFBH 44688, adult male, campus of the Universidade Estadual de Santa Cruz – UESC, Salobrinho, Ilhéus, State of 
<collectingRegion id="498AF80E6708F168FC6CFEAE7646267A" box="[977,1041,346,371]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, 
<collectingCountry id="F359767C6708F168FB9FFEAE7609267A" box="[1058,1118,346,371]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
[
<geoCoordinate id="EE7A502B6708F168FBCCFEAE76A1267A" box="[1137,1270,345,371]" degrees="39" direction="west" minutes="10" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="24" value="-39.173336">39°10′24″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FABAFEAE77D5267A" box="[1287,1410,345,371]" degrees="14" direction="south" minutes="47" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="52" value="-14.797778">14°47′52″S</geoCoordinate>
; about 
<quantity id="4CB69B096708F168FCD8FE8E71FC269A" box="[869,939,378,403]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="m" value="30.0">30 m</quantity>
above sea level (a.s.l.)], collected 
<date id="FFF0102C6708F168FACCFE8E71CB26BA" pageId="6" pageNumber="7" value="2018-04-10">10 April 2018</date>
by G. Novaes-e-Fagundes. 
<uri id="FFDF3AEE6708F168FAA5FE6E77CB26DA" pageId="6" pageNumber="7">
urn:lsid:zoobank. org:act: 
<uuid id="FFE80C396708F168FC12FE4E77CB26DA" box="[943,1436,442,467]" pageId="6" pageNumber="7">F357D8CC-446B-4F2C-9B53-190AFEEFD533</uuid>
</uri>
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C12F076708F168FCD8FE0E76EF23FA" pageId="6" pageNumber="7" type="multiple">
<subSubSection id="C35465676708F168FCD8FE0E7618251A" box="[869,1103,506,531]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="8BF136EC6708F168FCD8FE0E7618251A" blockId="6.[869,1103,506,531]" box="[869,1103,506,531]" pageId="6" pageNumber="7">
<heading id="D0B981806708F168FCD8FE0E7618251A" box="[869,1103,506,531]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="5">
<emphasis id="B93AEAFE6708F168FCD8FE0E7618251A" bold="true" box="[869,1103,506,531]" pageId="6" pageNumber="7">Referred specimens</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676708F168FC36FDCE76EF23FA" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BF136EC6708F168FC36FDCE76EF23FA" blockId="6.[868,1495,570,1267]" pageId="6" pageNumber="7">
Fifteen adults (
<specimenCount id="9D48FD656708F168FB96FDCE76DA255A" box="[1067,1165,570,595]" pageId="6" pageNumber="7" type="male">12 males</specimenCount>
and three females) from eight localities in the State of 
<collectingRegion id="498AF80E6708F168FBD1FDAE76FB257A" box="[1132,1196,602,627]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, 
<collectingCountry id="F359767C6708F168FB08FDAE76A4257A" box="[1205,1267,602,627]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
. CFBH 21071 (male), Povoado Senote, Caetité (
<geoCoordinate id="EE7A502B6708F168FB0EFD8E7760259A" box="[1203,1335,633,659]" degrees="42" direction="west" minutes="28" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="48" value="-42.48">42°28′48″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FAEDFD8E779B259A" box="[1360,1484,633,659]" degrees="14" direction="south" minutes="04" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="55" value="-14.081944">14°04′55″S</geoCoordinate>
); MHNJCH 1014 (male), Floresta Nacional Contendas do Sincorá, Contendas do Sincorá (
<geoCoordinate id="EE7A502B6708F168FAF3FD4E778425DA" box="[1358,1491,697,723]" degrees="41" direction="west" minutes="07" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="04" value="-41.117775">41°07′04″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FCD8FD2E71B625FA" box="[869,993,729,755]" degrees="13" direction="south" minutes="55" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="20" value="-13.922222">13°55′20″S</geoCoordinate>
); MZUESC 20683 (male), Condomínio Parque Universitário, Salobrinho, Ilhéus (
<geoCoordinate id="EE7A502B6708F168FAF3FD0E7784241A" box="[1358,1491,761,787]" degrees="39" direction="west" minutes="10" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="43" value="-39.17861">39°10′43″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FCD8FCEE71B6243A" box="[869,993,793,819]" degrees="14" direction="south" minutes="47" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="44" value="-14.795556">14°47′44″S</geoCoordinate>
); CFBH 44687 (male), Fazenda Lagoa Nova, Irajuba (
<geoCoordinate id="EE7A502B6708F168FC7BFCCE761C245A" box="[966,1099,825,851]" degrees="39" direction="west" minutes="59" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="57" value="-39.99917">39°59′57″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FBE6FCCE7680245A" box="[1115,1239,825,851]" degrees="13" direction="south" minutes="12" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="19" value="-13.205277">13°12′19″S</geoCoordinate>
); CFBH 18797 (male), Fazendas Santo Onofre and Cana Brava, Maracás (approx. 
<geoCoordinate id="EE7A502B6708F168FCD9FC8E765E249A" box="[868,1033,889,915]" degrees="40" direction="west" minutes="25" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="23.58" value="-40.423218">40°25′23.58″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FBA6FC8E76EF249A" box="[1051,1208,889,915]" degrees="13" direction="south" minutes="23" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="29.30" value="-13.391472">13°23′29.30″S</geoCoordinate>
); MHNJCH 1701 (male), near Fazendas Santo Onofre and Cana Brava, Maracás (
<geoCoordinate id="EE7A502B6708F168FCD6FC4E764724DA" box="[875,1040,953,979]" degrees="40" direction="west" minutes="26" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="26.23" value="-40.44062">40°26′26.23″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FB94FC4E769124DA" box="[1065,1222,953,979]" degrees="13" direction="south" minutes="21" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="59.10" value="-13.366417">13°21′59.10″S</geoCoordinate>
); MHNJCH 1698-1700 (males), Assentamento do Cumbe, Maracás (
<geoCoordinate id="EE7A502B6708F168FAF3FC2E778424FA" box="[1358,1491,985,1011]" degrees="40" direction="west" minutes="27" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="38" value="-40.460556">40°27′38″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FCD8FC0E71B6231A" box="[869,993,1017,1043]" degrees="13" direction="south" minutes="26" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="40" value="-13.444445">13°26′40″S</geoCoordinate>
); MZUESC 14890, 14893 (males), and 14891 (female), Companhia de Pesquisa de Recursos Minerais – CPRM, Morro do Chapéu (
<geoCoordinate id="EE7A502B6708F168FB16FBCE7778235A" box="[1195,1327,1081,1107]" degrees="41" direction="west" minutes="09" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="28" value="-41.15778">41°09′28″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FA81FBCE77E4235A" box="[1340,1459,1081,1107]" degrees="11" direction="south" minutes="32" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="56" value="-11.548889">11°32′56″S</geoCoordinate>
or 
<geoCoordinate id="EE7A502B6708F168FCD8FBAE71BF237A" box="[869,1000,1113,1139]" degrees="41" direction="west" minutes="06" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="26" value="-41.10722">41°06′26″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FC46FBAE7620237A" box="[1019,1143,1113,1139]" degrees="11" direction="south" minutes="29" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="34" value="-11.492778">11°29′34″S</geoCoordinate>
); MZUESC 15894 and 17503 (females), Praça Municipal, Potiraguá (
<geoCoordinate id="EE7A502B6708F168FAF3FB8E7784239A" box="[1358,1491,1145,1171]" degrees="39" direction="west" minutes="57" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="30" value="-39.958336">39°57′30″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FCD8FB6E71B623BA" box="[869,993,1177,1203]" degrees="15" direction="south" minutes="37" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="07" value="-15.618611">15°37′07″S</geoCoordinate>
); and UFMG 4787 (male), Route Sebastião Laranjeiras-Candiba, Sebastião Laranjeiras (approx. 
<geoCoordinate id="EE7A502B6708F168FCD8FB2E765E23FA" box="[869,1033,1241,1267]" degrees="42" direction="west" minutes="56" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="21.34" value="-42.939262">42°56′21.34″W</geoCoordinate>
, 
<geoCoordinate id="EE7A502B6708F168FBACFB2E76F923FA" box="[1041,1198,1241,1267]" degrees="14" direction="south" minutes="33" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="13.74" value="-14.553817">14°33′13.74″S</geoCoordinate>
).
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C12F076708F169FCD8FAEE73C525BA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="multiple">
<subSubSection id="C35465676708F168FCD8FAEE7783223A" box="[869,1492,1306,1331]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="8BF136EC6708F168FCD8FAEE7783223A" blockId="6.[869,1492,1306,1331]" box="[869,1492,1306,1331]" pageId="6" pageNumber="7">
<heading id="D0B981806708F168FCD8FAEE7783223A" box="[869,1492,1306,1331]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="5">
<emphasis id="B93AEAFE6708F168FCD8FAEE7783223A" bold="true" box="[869,1492,1306,1331]" pageId="6" pageNumber="7">
Diagnosis (based on 
<typeStatus id="54F5884E6708F168FBE9FAEE76E0223A" box="[1108,1207,1306,1331]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
and referred specimens)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676708F169FC36FAAE73C525BA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BF136EC6708F168FC36FAAE71BB2F1A" blockId="6.[868,1494,1370,2131]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FC36FAAE761F227A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[907,1096,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FC36FAAE761F227A" box="[907,1096,1370,1395]" italics="true" pageId="6" pageNumber="7">Scinax x‑signatus</emphasis>
</taxonomicName>
is a species of 
<emphasis id="B93AEAFE6708F168FB42FAAE771D227A" box="[1279,1354,1370,1395]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FB42FAAE7710227A" authorityName="Wagler" authorityYear="1830" box="[1279,1351,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Scinax</taxonomicName>
,
</emphasis>
as it shares three synapomorphies of this genus: webbing between toes I and II that does not extend beyond the subarticular tubercle of toe I; origin of the m. 
<emphasis id="B93AEAFE6708F168FB5DFA4E778222DA" box="[1248,1493,1466,1491]" italics="true" pageId="6" pageNumber="7">pectoralis abdominalis</emphasis>
through well-defined tendons; and m. 
<emphasis id="B93AEAFE6708F168FAAEFA2E71CA211A" italics="true" pageId="6" pageNumber="7">pectoralis abdominalis</emphasis>
overlapping m. 
<emphasis id="B93AEAFE6708F168FBE3FA0E7749211A" box="[1118,1310,1530,1555]" italics="true" pageId="6" pageNumber="7">obliquus externus</emphasis>
(
<bibRefCitation id="EFDF4B1D6708F168FA93FA0E7785211A" author="da Silva, H. R." box="[1326,1490,1530,1555]" pageId="6" pageNumber="7" refId="ref24934" refString="da Silva, H. R. 1998. Phylogenetic relationships of the family Hylidae with emphasis on the relationships within the subfamily Hylinae (Amphibia: Anura). Ph. D. dissertation, Department of Systematics and Ecology, University of Kansas." type="book" year="1998">da Silva, 1998</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6708F168FCD8F9EE7644213A" author="Faivovich, J." box="[869,1043,1562,1587]" pageId="6" pageNumber="7" pagination="367 - 393" refId="ref20258" refString="Faivovich, J. 2002. A cladistic analysis of Scinax (Anura: Hylidae). Cladistics, 18 (4): 367 - 393." type="journal article" year="2002">Faivovich, 2002</bibRefCitation>
; Faivovich 
<emphasis id="B93AEAFE6708F168FB32F9EE769E213A" box="[1167,1225,1562,1587]" italics="true" pageId="6" pageNumber="7">et al.,</emphasis>
2005). A single synapomorphy is known for the 
<taxonomicName id="4C4E4D6F6708F168FBCBF9CE7692215A" box="[1142,1221,1594,1619]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6708F168FBCBF9CE7692215A" box="[1142,1221,1594,1619]" italics="true" pageId="6" pageNumber="7">S.ruber</emphasis>
</taxonomicName>
Clade: tadpoles with the vent tube above the margin of the lower fin (
<bibRefCitation id="EFDF4B1D6708F168FAD8F9AE71F7219A" author="Faivovich, J." pageId="6" pageNumber="7" pagination="367 - 393" refId="ref20258" refString="Faivovich, J. 2002. A cladistic analysis of Scinax (Anura: Hylidae). Cladistics, 18 (4): 367 - 393." type="journal article" year="2002">Faivovich, 2002</bibRefCitation>
; Faivovich 
<emphasis id="B93AEAFE6708F168FB9CF98E7608219A" box="[1057,1119,1658,1683]" italics="true" pageId="6" pageNumber="7">et al.,</emphasis>
2005). While tadpoles unequivocally associated to 
<taxonomicName id="4C4E4D6F6708F168FBF1F96E768321BA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1100,1236,1690,1715]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FBF1F96E768321BA" box="[1100,1236,1690,1715]" italics="true" pageId="6" pageNumber="7">S. x‑signatus</emphasis>
</taxonomicName>
remain unknown, this species was associated to the 
<taxonomicName id="4C4E4D6F6708F168FB7DF94E774421DA" box="[1216,1299,1722,1747]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6708F168FB7DF94E774421DA" box="[1216,1299,1722,1747]" italics="true" pageId="6" pageNumber="7">S. ruber</emphasis>
</taxonomicName>
Clade by having the unique combination of external vocal sac and presence of pectoral fold [internal vocal sac and pectoral fold absent in most species of the 
<taxonomicName id="4C4E4D6F6708F168FB09F8EE7717203A" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[1204,1344,1818,1843]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="catharinae">
<emphasis id="B93AEAFE6708F168FB09F8EE7717203A" box="[1204,1344,1818,1843]" italics="true" pageId="6" pageNumber="7">S. catharinae</emphasis>
</taxonomicName>
Clade; in few species where the vocal sac is external, the pectoral fold is absent (
<emphasis id="B93AEAFE6708F168FC6BF8AE7751207A" box="[982,1286,1882,1907]" italics="true" pageId="6" pageNumber="7">
e.g., 
<taxonomicName id="4C4E4D6F6708F168FBBBF8AE76CD207A" box="[1030,1178,1882,1907]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="garibaldiae">S. garibaldiae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6708F168FB19F8AE7751207A" baseAuthorityName="Bokermann" baseAuthorityYear="1964" box="[1188,1286,1882,1907]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rizibilis">S. rizibilis</taxonomicName>
</emphasis>
); otherwise, in the two cases where the pectoral fold is present, the vocal sac is internal (
<taxonomicName id="4C4E4D6F6708F168FBB9F86E760520BA" baseAuthorityName="Cruz &amp; Peixoto" baseAuthorityYear="1983" box="[1028,1106,1946,1971]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="agilis">
<emphasis id="B93AEAFE6708F168FBB9F86E760520BA" box="[1028,1106,1946,1971]" italics="true" pageId="6" pageNumber="7">S.agilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F6708F168FB38F86E771020BA" authorityName="Lourenco, Luna &amp; Pombal" authorityYear="2014" box="[1157,1351,1946,1971]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="melanodactylus">
<emphasis id="B93AEAFE6708F168FB38F86E771020BA" box="[1157,1351,1946,1971]" italics="true" pageId="6" pageNumber="7">S.melanodactylus</emphasis>
</taxonomicName>
); J.Faivovich &amp; K. Araujo-Vieira, 
<emphasis id="B93AEAFE6708F168FB8AF84E76F720DA" box="[1079,1184,1978,2003]" italics="true" pageId="6" pageNumber="7">pers. obs.;</emphasis>
see also 
<bibRefCitation id="EFDF4B1D6708F168FAB6F84E778520DA" author="Bokermann, W. C. A." box="[1291,1490,1978,2003]" pageId="6" pageNumber="7" pagination="67 - 76" refId="ref18186" refString="Bokermann, W. C. A. 1964. Dos nuevas especies de Hyla de Minas Gerais y notas sobre Hyla alvarengai (Amphibia, Salientia, Hylidae). Neotropica, 10 (32): 67 - 76." type="journal article" year="1964">Bokermann, 1964</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6708F168FCD8F82E760E20FA" author="Cruz, C. D. &amp; Peixoto, O. L." box="[869,1113,2010,2035]" pageId="6" pageNumber="7" pagination="721 - 724" refId="ref19212" refString="Cruz, C. D. &amp; Peixoto, O. L. 1982. Uma nova especie de Hyla do Estado do Espirito Santo, Brasil (Amphibia, Anura, Hylidae). Revista Brasileira de Biologia, 42 (4): 721 - 724." type="journal article" year="1982">Cruz &amp; Peixoto, 1982</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6708F168FBDBF82E774020FA" author="Faivovich, J." box="[1126,1303,2010,2035]" pageId="6" pageNumber="7" pagination="367 - 393" refId="ref20258" refString="Faivovich, J. 2002. A cladistic analysis of Scinax (Anura: Hylidae). Cladistics, 18 (4): 367 - 393." type="journal article" year="2002">Faivovich, 2002</bibRefCitation>
; Lourenço 
<emphasis id="B93AEAFE6708F168FA25F82E778220FA" box="[1432,1493,2010,2035]" italics="true" pageId="6" pageNumber="7">et al.,</emphasis>
2014, 2019].
</paragraph>
<paragraph id="8BF136EC6708F169FC36F7EE73C525BA" blockId="6.[868,1494,1370,2131]" lastBlockId="7.[94,719,218,691]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D6F6708F168FC36F7EE76122F3A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[907,1093,2074,2099]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6708F168FC36F7EE76122F3A" box="[907,1093,2074,2099]" italics="true" pageId="6" pageNumber="7">Scinax x‑signatus</emphasis>
</taxonomicName>
can be differentiated from all other species of the 
<taxonomicName id="4C4E4D6F6708F168FBB7F7CE760B2F5A" box="[1034,1116,2106,2131]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6708F168FBB7F7CE760B2F5A" box="[1034,1116,2106,2131]" italics="true" pageId="6" pageNumber="7">S. ruber</emphasis>
</taxonomicName>
Clade by the combination of the following characters: (1) male SVL 34.5-38.4 mm, 
<emphasis id="B93AEAFE6709F169FD3EFF2E70C627FA" box="[643,657,218,243]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= 13; (2) snout rounded in dorsal view and profile; (3) pointed tubercles on lower jaw absent; (4) vocal sac subgular, weakly bilobate; (5) spicule-shaped papillary epidermal projections on the nuptial pads and pectoral region present in males; (6) pectoral glands present in males; (7) dorsal color pattern with large irregular dark blotches, commonly with dark X-shaped mark composed of one or two pairs of inverted parenthesis-like blotches; (8) hidden surfaces of thighs dark with irregular pale blotches, yellow in living specimens; (9) iris yellowish golden or bronze with a median black streak; (10) physiological chlorosis absent; and (11) advertisement call composed of a single, multipulsed note, 0.11- 
<geoCoordinate id="EE7A502B6709F169FE55FD8E707D259A" box="[488,554,634,659]" degrees="0.25" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="555" value="-0.25">0.25 s</geoCoordinate>
duration, 6-14 pulses/note, 52-64 pulses/s.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C12F076709F163FFE3FD2E73DE265A" lastPageId="13" lastPageNumber="14" pageId="7" pageNumber="8" type="multiple">
<subSubSection id="C35465676709F169FFE3FD2E709925FA" box="[94,718,730,755]" pageId="7" pageNumber="8" type="multiple">
<paragraph id="8BF136EC6709F169FFE3FD2E709925FA" blockId="7.[94,718,730,755]" box="[94,718,730,755]" pageId="7" pageNumber="8">
<heading id="D0B981806709F169FFE3FD2E709925FA" box="[94,718,730,755]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="8" reason="5">
<emphasis id="B93AEAFE6709F169FFE3FD2E709925FA" bold="true" box="[94,718,730,755]" pageId="7" pageNumber="8">
Comparisons with other species of 
<taxonomicName id="4C4E4D6F6709F169FE4BFD2E70D225FA" box="[502,645,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6709F169FE4BFD2E70D225FA" bold="true" box="[502,645,730,755]" italics="true" pageId="7" pageNumber="8">Scinax ruber</emphasis>
</taxonomicName>
Clade
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676709F166FF39FCEE714C23BA" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BF136EC6709F169FF39FCEE730323FA" blockId="7.[93,720,794,2131]" pageId="7" pageNumber="8">
The SVL in males of 
<taxonomicName id="4C4E4D6F6709F169FED5FCEE7075243A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[360,546,794,819]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FED5FCEE7075243A" box="[360,546,794,819]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
(34.5-38.4 mm, 
<emphasis id="B93AEAFE6709F169FFDDFCCE7239245A" box="[96,110,826,851]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= 13) distinguishes it from the larger species 
<taxonomicName id="4C4E4D6F6709F169FDD0FCCE72C1247A" authorityName="De la Riva" authorityYear="1993" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="castroviejoi">
<emphasis id="B93AEAFE6709F169FDD0FCCE72C1247A" italics="true" pageId="7" pageNumber="8">S. castroviejoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F6709F169FF76FCAE736C247A" baseAuthorityName="Bokermann" baseAuthorityYear="1968" box="[203,315,858,883]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="eurydice">
<emphasis id="B93AEAFE6709F169FF76FCAE736C247A" box="[203,315,858,883]" italics="true" pageId="7" pageNumber="8">S.eurydice</emphasis>
</taxonomicName>
(SVL males 44.0-52.0 mm; 
<bibRefCitation id="EFDF4B1D6709F169FDE3FCAE72C0249A" author="De la Riva, I." pageId="7" pageNumber="8" pagination="41 - 46" refId="ref19370" refString="De la Riva, I. 1993. A new species of Scinax (Anura, Hylidae) from Argentina and Bolivia. Journal of Herpetology, 27: 41 - 46." type="journal article" year="1993">De la Riva, 1993</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FF19FC8E7339249A" author="Bokermann, W. C. A." box="[164,366,890,915]" pageId="7" pageNumber="8" pagination="25 - 31" refId="ref18261" refString="Bokermann, W. C. A. 1968. Three new Hyla from the Plateau of Maracas, central Bahia, Brazil. Journal of Herpetology, 1: 25 - 31." type="journal article" year="1968">Bokermann, 1968</bibRefCitation>
), and from the smaller species 
<emphasis id="B93AEAFE6709F169FFE3FC6E73CB233A" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FFE3FC6E72E624BA" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[94,177,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altae">S. altae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF02FC6E737824BA" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[191,303,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="auratus">S. auratus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE81FC6E738724BA" authorityName="Drummond, Baeta &amp; Pires" authorityYear="2007" box="[316,464,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabralensis">S. cabralensis</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE60FC6E703724BA" baseAuthorityName="Lutz" baseAuthorityYear="1968" box="[477,608,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caldarum">S. caldarum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDD3FC6E72F924DA" baseAuthorityName="Duellman" baseAuthorityYear="1972" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cruentomma">S. cruentomma</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF00FC4E734924DA" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[189,286,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="danae">S. danae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE91FC4E73CD24DA" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[300,410,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="exiguus">S. exiguus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE15FC4E702D24DA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[424,634,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscomarginatus">S. fuscomarginatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FD35FC4E72D324FA" authorityName="Nunes &amp; Pombal" authorityYear="2010" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="juncae">S. juncae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF2DFC2E736124FA" baseAuthorityName="Pyburn" baseAuthorityYear="1993" box="[144,310,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="karenanneae">S. karenanneae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEFFFC2E73F924FA" authorityName="Pyburn" authorityYear="1992" box="[322,430,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lindsayi">S. lindsayi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE04FC2E706E24FA" baseAuthorityName="Bokermann" baseAuthorityYear="1964" box="[441,569,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="madeirae">S. madeirae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDF9FC2E709D24FA" baseAuthorityName="Cardoso &amp; Sazima" baseAuthorityYear="1980" box="[580,714,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="maracaya">S. maracaya</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FFDCFC0E7347231A" authorityName="Ferrao, Fraga, Moravec, Kaefer &amp; Lima" authorityYear="2018" box="[97,272,1018,1043]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruberoculatus">S. ruberoculatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEA6FC0E73C6231A" authorityName="Araujo-Vieira, Brandao &amp; Faria" authorityYear="2015" box="[283,401,1018,1043]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rupestris">S. rupestris</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE20FC0E7047231A" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[413,528,1018,1043]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="staufferi">S. staufferi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDA6FC0E709B231A" authorityName="Ferrao, Moravec, Kaefer, Fraga &amp; Lima" authorityYear="2018" box="[539,716,1018,1043]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="strussmannae">S. strussmannae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FFE3FBEE7357233A" authorityName="Nunes, Kwet &amp; Pombal" authorityYear="2012" box="[94,256,1050,1075]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tymbamirim">S. tymbamirim</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEB1FBEE73CF233A" authorityName="Brusquetti, Jansen, Barrio-Amoros, Segalla &amp; Haddad" authorityYear="2014" box="[268,408,1050,1075]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="villasboasi">S. villasboasi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FE64FBEE701D233A" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[473,586,1050,1075]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="wandae">
<emphasis id="B93AEAFE6709F169FE64FBEE701D233A" box="[473,586,1050,1075]" italics="true" pageId="7" pageNumber="8">S. wandae</emphasis>
</taxonomicName>
(SVL males 15.7-29.0 mm; 
<bibRefCitation id="EFDF4B1D6709F169FEB5FBCE7328235A" author="Lutz, B." box="[264,383,1082,1107]" pageId="7" pageNumber="8" pagination="1 - 18" refId="ref22735" refString="Lutz, B. 1968. New brazilian forms of Hyla. Pearce-Sellards Series Texas Memorial Museum, 10: 1 - 18." type="journal article" year="1968">Lutz, 1968</bibRefCitation>
, 
<bibRefCitation id="EFDF4B1D6709F169FE31FBCE7392235A" author="Lutz, B." box="[396,453,1082,1107]" pageId="7" pageNumber="8" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FE6CFBCE70D2235A" author="Duellman, W. E." box="[465,645,1082,1107]" pageId="7" pageNumber="8" pagination="1 - 753" refId="ref19657" refString="Duellman, W. E. 1970. The hylid frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1: 1 - 753." type="journal article" year="1970">Duellman, 1970</bibRefCitation>
, 
<bibRefCitation id="EFDF4B1D6709F169FD2FFBCE709C235A" author="Duellman, W. E." box="[658,715,1082,1107]" pageId="7" pageNumber="8" pagination="864 - 870" refId="ref19849" refString="Duellman, W. E. 1986. Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986: 864 - 870." type="journal article" year="1986">1986</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FFE3FBAE73CE237A" author="Pyburn, W. F. &amp; Fouquette, M. J." box="[94,409,1114,1139]" pageId="7" pageNumber="8" pagination="97 - 101" refId="ref24172" refString="Pyburn, W. F. &amp; Fouquette, M. J. 1971. A new striped treefrog from Central Colombia. Journal of Herpetology, 5: 97 - 101." type="journal article" year="1971">Pyburn &amp; Fouquette, 1971</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FE17FBAE709C237A" author="Cardoso, A. J. &amp; Sazima, I." box="[426,715,1114,1139]" pageId="7" pageNumber="8" pagination="75 - 79" refId="ref18559" refString="Cardoso, A. J. &amp; Sazima, I. 1980. Nova especie de Hyla do sudeste brasileiro (Amphibia, Anura, Hylidae). Revista Brasileira de Biologia, 40 (1): 75 - 79." type="journal article" year="1980">Cardoso &amp; Sazima, 1980</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FFE3FB8E733E239A" author="Duellman, W. E. &amp; Wiens, J. J." box="[94,361,1146,1171]" pageId="7" pageNumber="8" pagination="1 - 57" refId="ref19931" refString="Duellman, W. E. &amp; Wiens, J. J. 1993. Hylid frogs of the genus Scinax Wagler, 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History, University of Kansas, 153: 1 - 57." type="journal article" year="1993">Duellman &amp; Wiens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FECDFB8E7053239A" author="Pyburn, W. F." box="[368,516,1146,1171]" pageId="7" pageNumber="8" pagination="405 - 411" refId="ref24083" refString="Pyburn, W. F. 1992. A new tree frog of the genus Scinax from the Vaupes River of northwestern Brazil. The Texas Journal of Science, 44 (4): 405 - 411." type="journal article" year="1992">Pyburn, 1992</bibRefCitation>
, 
<bibRefCitation id="EFDF4B1D6709F169FDB6FB8E7013239A" author="Pyburn, W. F." box="[523,580,1146,1171]" pageId="7" pageNumber="8" pagination="46 - 50" refId="ref24122" refString="Pyburn, W. F. 1993. A new species of dimorphic tree frog, genus Hyla (Amphibia: Anura: Hylidae), from the Vaupes River of Colombia. Proceedings of the Biological Society of Washington, 106 (1): 46 - 50." type="journal article" year="1993">1993</bibRefCitation>
; Drummond 
<emphasis id="B93AEAFE6709F169FFE0FB6E72CE23BA" box="[93,153,1178,1203]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2007; 
<bibRefCitation id="EFDF4B1D6709F169FF54FB6E73A723BA" author="Nunes, I. &amp; Pombal Jr., J. P." box="[233,496,1178,1203]" pageId="7" pageNumber="8" pagination="347 - 353" refId="ref23378" refString="Nunes, I. &amp; Pombal Jr., J. P. 2010. A new Scinax Wagler (Amphibia, Anura, Hylidae) from the Atlantic Rain Forest remains of southern State of Bahia, North-eastern Brazil. Amphibia-Reptilia, 31 (3): 347 - 353." type="journal article" year="2010">Nunes &amp; Pombal, 2010</bibRefCitation>
, 
<bibRefCitation id="EFDF4B1D6709F169FE41FB6E706223BA" author="Nunes, I. &amp; Pombal Jr., J. P." box="[508,565,1178,1203]" pageId="7" pageNumber="8" pagination="80 - 88" refId="ref23430" refString="Nunes, I. &amp; Pombal Jr., J. P. 2011. A new snouted treefrog of the speciose genus Scinax Wagler (Anura, Hylidae) from northeastern Brazil. Herpetologica, 67 (1): 80 - 88." type="journal article" year="2011">2011</bibRefCitation>
; Nunes 
<emphasis id="B93AEAFE6709F169FD2EFB6E709823BA" box="[659,719,1178,1203]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2012; Brusquetti 
<emphasis id="B93AEAFE6709F169FE9EFB4E733723DA" box="[291,352,1210,1235]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2014; Araujo-Vieira 
<emphasis id="B93AEAFE6709F169FDF6FB4E70DF23DA" box="[587,648,1210,1235]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2015; Ferrão 
<emphasis id="B93AEAFE6709F169FF16FB2E72B423FA" box="[171,227,1242,1267]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018a, b).
</paragraph>
<paragraph id="8BF136EC6709F169FF39FB0E7381201A" blockId="7.[93,720,794,2131]" pageId="7" pageNumber="8">
The snout rounded in dorsal view and profile differentiates 
<taxonomicName id="4C4E4D6F6709F169FF02FAEE732D223A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[191,378,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FF02FAEE732D223A" box="[191,378,1306,1331]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
from 
<emphasis id="B93AEAFE6709F169FE7DFAEE701D225A" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FE7DFAEE705D223A" baseAuthorityName="B.Lutz" baseAuthorityYear="1973" box="[448,522,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="alter">S. alter</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDA8FAEE70D5223A" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[533,642,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="auratus">S. auratus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FD30FAEE72C1225A" authorityName="Nunes &amp; Pombal" authorityYear="2011" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cretatus">S. cretatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF1CFACE730F225A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[161,344,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="crospedospilus">S. crospedospilus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEDEFACE738D225A" authorityName="Nunes, Kwet &amp; Pombal" authorityYear="2012" box="[355,474,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="imbegue">S. imbegue</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE58FACE7011225A" authorityName="Nunes &amp; Pombal" authorityYear="2010" box="[485,582,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="juncae">S. juncae</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FD38FACE72ED227A" authorityName="Nunes, Kwet &amp; Pombal" authorityYear="2012" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tymbamirim">
<emphasis id="B93AEAFE6709F169FD38FACE72ED227A" italics="true" pageId="7" pageNumber="8">S. tymbamirim</emphasis>
</taxonomicName>
(sub-elliptical with a pointed tip in dorsal view and slightly acute in profile), 
<taxonomicName id="4C4E4D6F6709F169FE24FA8E7070229A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[409,551,1402,1427]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscovarius">
<emphasis id="B93AEAFE6709F169FE24FA8E7070229A" box="[409,551,1402,1427]" italics="true" pageId="7" pageNumber="8">S.fuscovarius</emphasis>
</taxonomicName>
(roundly acute in dorsal view and protruding in profile), 
<emphasis id="B93AEAFE6709F169FDFBFA6E70C022DA" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FDFBFA6E709D22BA" baseAuthorityName="Lutz" baseAuthorityYear="1968" box="[582,714,1434,1459]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caldarum">S. caldarum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FFE3FA4E729822DA" authorityName="Pugliese, Pombal &amp; Sazima" authorityYear="2004" box="[94,207,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="curicica">S. curicica</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF63FA4E731E22DA" baseAuthorityName="B.Lutz" baseAuthorityYear="1951" box="[222,329,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="duartei">S. duartei</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEE5FA4E73B522DA" baseAuthorityName="Cardoso &amp; Sazima" baseAuthorityYear="1980" box="[344,482,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="maracaya">S. maracaya</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE4CFA4E70C322DA" authorityName="Conte, Araujo-Vieira, Crivellari &amp; Berneck" authorityYear="2016" box="[497,660,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rossaferesae">S. rossaferesae</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FFE3FA2E729F22FA" authorityName="Nunes, Carvalho &amp; Pereira" authorityYear="2010" box="[94,200,1498,1523]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tigrinus">
<emphasis id="B93AEAFE6709F169FFE3FA2E729F22FA" box="[94,200,1498,1523]" italics="true" pageId="7" pageNumber="8">S. tigrinus</emphasis>
</taxonomicName>
(sub-elliptical or subovoid in dorsal view and slightly acute in profile), 
<taxonomicName id="4C4E4D6F6709F169FED0FA0E7052211A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[365,517,1530,1555]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="squalirostris">
<emphasis id="B93AEAFE6709F169FED0FA0E7052211A" box="[365,517,1530,1555]" italics="true" pageId="7" pageNumber="8">S. squalirostris</emphasis>
</taxonomicName>
(pointed in dorsal view and acute in profile), and species of the 
<taxonomicName id="4C4E4D6F6709F169FDE4F9EE7098213A" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[601,719,1562,1587]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B93AEAFE6709F169FDE4F9EE7098213A" box="[601,719,1562,1587]" italics="true" pageId="7" pageNumber="8">S. rostratus</emphasis>
</taxonomicName>
group (elongate pointed in dorsal view and acute with or without a fleshy proboscis in profile). Furthermore, the absence of pointed tubercles on the lower jaw differentiates 
<taxonomicName id="4C4E4D6F6709F169FF2EF96E734E21BA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[147,281,1690,1715]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FF2EF96E734E21BA" box="[147,281,1690,1715]" italics="true" pageId="7" pageNumber="8">S. x‑signatus</emphasis>
</taxonomicName>
from almost all species of the 
<taxonomicName id="4C4E4D6F6709F169FDCCF96E722921DA" baseAuthorityName="Peters" baseAuthorityYear="1863" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B93AEAFE6709F169FDCCF96E722921DA" italics="true" pageId="7" pageNumber="8">S. rostratus</emphasis>
</taxonomicName>
Group; exceptions are 
<taxonomicName id="4C4E4D6F6709F169FECAF94E73BA21DA" baseAuthorityName="Pyburn" baseAuthorityYear="1973" box="[375,493,1722,1747]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="B93AEAFE6709F169FECAF94E73BA21DA" box="[375,493,1722,1747]" italics="true" pageId="7" pageNumber="8">S.kennedyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F6709F169FD9FF94E70C021DA" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[546,663,1722,1747]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B93AEAFE6709F169FD9FF94E70C021DA" box="[546,663,1722,1747]" italics="true" pageId="7" pageNumber="8">S.rostratus</emphasis>
</taxonomicName>
(
<emphasis id="B93AEAFE6709F169FD19F94E709821DA" box="[676,719,1722,1747]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FFE3F92E734821FA" author="Duellman, W. E." box="[94,287,1754,1779]" pageId="7" pageNumber="8" pagination="177 - 192" refId="ref19730" refString="Duellman, W. E. 1972 a. South American Frogs of the Hyla rostrata group (Amphibia, Anura, Hylidae). Zoologische Mededelingen, 47 (14): 177 - 192." type="journal article" year="1972">
Duellman, 1972 
<bibRefCitation id="EFDF4B1D6709F169FEAFF92E734821FA" author="Bokermann, W. C. A. &amp; Sazima, I." box="[274,287,1754,1779]" pageId="7" pageNumber="8" pagination="521 - 528" refId="ref18334" refString="Bokermann, W. C. A. &amp; Sazima, I. 1973. Anfibios da Serra do Cipo, Minas Gerais, Brasil. 1: Duas especies novas de Hyla (Anura, Hylidae). Revista Brasileira de Biologia, 33 (3): 521 - 528." type="journal article" year="1973">a</bibRefCitation>
</bibRefCitation>
, 1973; 
<bibRefCitation id="EFDF4B1D6709F169FED2F92E705021FA" author="Pyburn, W. F." box="[367,519,1754,1779]" pageId="7" pageNumber="8" pagination="297 - 301" refId="ref24055" refString="Pyburn, W. F. 1973. A new hylid frog from the Llanos of Colombia. Journal of Herpetology, 7: 297 - 301." type="journal article" year="1973">Pyburn, 1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FDAFF92E72C0201A" author="Lescure, J. &amp; Marty, C." pageId="7" pageNumber="8" refId="ref22401" refString="Lescure, J. &amp; Marty, C. 2000. Atlas des amphibiens de Guyane. Collection Patrimoines Naturels." type="book" year="2000">Lescure &amp; Marty, 2000</bibRefCitation>
; 
<collectingRegion id="498AF80E6709F169FF1DF90E7281201A" box="[160,214,1786,1811]" country="Peru" name="Lima" pageId="7" pageNumber="8">Lima</collectingRegion>
<emphasis id="B93AEAFE6709F169FF61F90E7342201A" box="[220,277,1786,1811]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2005; this study).
</paragraph>
<paragraph id="8BF136EC6709F169FF39F8EE70E42F5A" blockId="7.[93,720,794,2131]" pageId="7" pageNumber="8">
The presence of a weakly bilobate subgular vocal sac in 
<taxonomicName id="4C4E4D6F6709F169FFE3F8CE7347205A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[94,272,1850,1875]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FFE3F8CE7347205A" box="[94,272,1850,1875]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
distinguishes it from 
<taxonomicName id="4C4E4D6F6709F169FE49F8CE70F2205A" authorityName="Bokermann" authorityYear="1968" box="[500,677,1850,1875]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="camposseabrai">
<emphasis id="B93AEAFE6709F169FE49F8CE70F2205A" box="[500,677,1850,1875]" italics="true" pageId="7" pageNumber="8">S.camposseabrai</emphasis>
</taxonomicName>
(bilobate subgular vocal sac; see also 
<bibRefCitation id="EFDF4B1D6709F169FE65F8AE72C0209A" author="Caramaschi, U. &amp; Cardoso, M. C. S." pageId="7" pageNumber="8" pagination="549 - 552" refId="ref18520" refString="Caramaschi, U. &amp; Cardoso, M. C. S. 2006. Taxonomicstatusof Hylacamposseabrai Bokermann, 1968 (Anura: Hylidae). Journal of Herpetology, 40 (4): 549 - 552." type="journal article" year="2006">Caramaschi &amp; Cardoso, 2006</bibRefCitation>
: fig. 1) and from the remaining species of the 
<taxonomicName id="4C4E4D6F6709F169FD3CF88E7098209A" box="[641,719,1914,1939]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6709F169FD3CF88E7098209A" box="[641,719,1914,1939]" italics="true" pageId="7" pageNumber="8">S. ruber</emphasis>
</taxonomicName>
Clade with single subgular vocal sac;exceptions are 
<emphasis id="B93AEAFE6709F169FDC8F86E737D2F1A" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FDC8F86E72C820DA" baseAuthorityName="Cope" baseAuthorityYear="1862" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acuminatus">S.acuminatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF15F84E72AF20DA" baseAuthorityName="Werner" baseAuthorityYear="1903" box="[168,248,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="dolloi">S.dolloi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEBDF84E732720DA" baseAuthorityName="Cope" baseAuthorityYear="1874" box="[256,368,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="funereus">S.funereus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FEC5F84E705520DA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[376,514,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscovarius">S.fuscovarius</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDB7F84E700420DA" baseAuthorityName="Barbour" baseAuthorityYear="1909" box="[522,595,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hayii">S.hayii</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDE6F84E72C520FA" baseAuthorityName="Pyburn" baseAuthorityYear="1993" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="karenanneae">S.karenanneae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF27F82E737B20FA" authorityName="Junca, Napoli, Nunes, Merces &amp; Abreu" authorityYear="2015" box="[154,300,2010,2035]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="montivagus">S.montivagus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE89F82E732920FA" authorityName="Ferrao, Moravec, Fraga, Pinheiro de Almeida, Kaefer &amp; Lima" authorityYear="2017" box="[308,382,2010,2035]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="onca">S.onca</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE3AF82E73B620FA" authorityName="Duellman &amp; Wiens" authorityYear="1993" box="[391,481,2010,2035]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="oreites">S.oreites</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FE54F82E703F20FA" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1937" box="[489,616,2010,2035]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachycrus">S.pachycrus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FDCDF82E72202F1A" authorityName="Pombal, Haddad &amp; Kasahara" authorityYear="1995" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="perereca">S.perereca</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FF3CF80E73712F1A" authorityName="Ferrao, Fraga, Moravec, Kaefer &amp; Lima" authorityYear="2018" box="[129,294,2042,2067]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruberoculatus">S. ruberoculatus</taxonomicName>
,
</emphasis>
and 
<emphasis id="B93AEAFE6709F169FEE2F80E739C2F1A" box="[351,459,2042,2067]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FEE2F80E73912F1A" authorityName="Ron, Duellman, Caminer &amp; Pazmino" authorityYear="2018" box="[351,454,2042,2067]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tsachila">S.tsachila</taxonomicName>
,
</emphasis>
that have a weakly bilobate subgular vocal sac (
<emphasis id="B93AEAFE6709F169FED5F7EE73C72F3A" box="[360,400,2074,2099]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FE2BF7EE73AD2F3A" author="Cei, J. M." box="[406,506,2074,2099]" pageId="7" pageNumber="8" refId="ref18752" refString="Cei, J. M. 1980. Amphibians of Argentina. Monitore Zoologico Italiano, Nuova Serie, Monographia, ixii + 609 p." type="book" year="1980">Cei, 1980</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FDB9F7EE72C02F5A" author="Duellman, W. E. &amp; Wiens, J. J." pageId="7" pageNumber="8" pagination="1 - 57" refId="ref19931" refString="Duellman, W. E. &amp; Wiens, J. J. 1993. Hylid frogs of the genus Scinax Wagler, 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History, University of Kansas, 153: 1 - 57." type="journal article" year="1993">Duellman &amp; Wiens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FF23F7CE737A2F5A" author="Pyburn, W. F." box="[158,301,2106,2131]" pageId="7" pageNumber="8" pagination="46 - 50" refId="ref24122" refString="Pyburn, W. F. 1993. A new species of dimorphic tree frog, genus Hyla (Amphibia: Anura: Hylidae), from the Vaupes River of Colombia. Proceedings of the Biological Society of Washington, 106 (1): 46 - 50." type="journal article" year="1993">Pyburn, 1993</bibRefCitation>
;Ferrão 
<emphasis id="B93AEAFE6709F169FEC6F7CE73E72F5A" box="[379,432,2106,2131]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2017, 2018a; this study).
</paragraph>
<paragraph id="8BF136EC6709F169FC82FF2E76D6251A" blockId="7.[793,1419,218,2131]" pageId="7" pageNumber="8">
The presence of spicule-shaped papillary epidermal projections on the nuptial pad and pectoral region in males differentiates 
<taxonomicName id="4C4E4D6F6709F169FB9FFEEE7688263A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1058,1247,282,307]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FB9FFEEE7688263A" box="[1058,1247,282,307]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
from all other species of the 
<taxonomicName id="4C4E4D6F6709F169FC7AFECE764B265A" box="[967,1052,314,339]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6709F169FC7AFECE764B265A" box="[967,1052,314,339]" italics="true" pageId="7" pageNumber="8">S. ruber</emphasis>
</taxonomicName>
Clade, except for 
<taxonomicName id="4C4E4D6F6709F169FB48FECE77DE265A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[1269,1417,314,339]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscovarius">
<emphasis id="B93AEAFE6709F169FB48FECE77DE265A" box="[1269,1417,314,339]" italics="true" pageId="7" pageNumber="8">S. fuscovarius</emphasis>
</taxonomicName>
(see also Luna 
<emphasis id="B93AEAFE6709F169FC7EFEAE71A9267A" box="[963,1022,346,371]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018: fig. 10A, C). The presence of pectoral glands in males differentiates 
<taxonomicName id="4C4E4D6F6709F169FB74FE8E7719269A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1225,1358,378,403]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FB74FE8E7719269A" box="[1225,1358,378,403]" italics="true" pageId="7" pageNumber="8">S. x‑signatus</emphasis>
</taxonomicName>
from most species of the 
<taxonomicName id="4C4E4D6F6709F169FC44FE6E761D26BA" box="[1017,1098,410,435]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B93AEAFE6709F169FC44FE6E761D26BA" box="[1017,1098,410,435]" italics="true" pageId="7" pageNumber="8">S. ruber</emphasis>
</taxonomicName>
Clade, except for 
<emphasis id="B93AEAFE6709F169FAACFE6E762526DA" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FAACFE6E77D126BA" baseAuthorityName="Cope" baseAuthorityYear="1874" box="[1297,1414,410,435]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="funereus">S. funereus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FE4E71FD26DA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[794,938,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscovarius">S.fuscovarius</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC0FFE4E764E26DA" baseAuthorityName="Cope" baseAuthorityYear="1862" box="[946,1049,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nasicus">S.nasicus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB9CFE4E763A26DA" authorityName="Ferrao, Moravec, Fraga, Pinheiro de Almeida, Kaefer &amp; Lima" authorityYear="2017" box="[1057,1133,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="onca">S.onca</taxonomicName>
,
</emphasis>
and 
<emphasis id="B93AEAFE6709F169FB18FE4E775426DA" box="[1189,1283,442,467]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FB18FE4E76A826DA" authorityName="Cochran" authorityYear="1952" box="[1189,1279,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="similis">S.similis</taxonomicName>
,
</emphasis>
and species of the 
<taxonomicName id="4C4E4D6F6709F169FCD9FE2E71B926FA" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[868,1006,474,499]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="uruguayus">
<emphasis id="B93AEAFE6709F169FCD9FE2E71B926FA" box="[868,1006,474,499]" italics="true" pageId="7" pageNumber="8">S. uruguayus</emphasis>
</taxonomicName>
Group (
<emphasis id="B93AEAFE6709F169FBF5FE2E762426FA" box="[1096,1139,474,499]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FBC4FE2E77D126FA" author="Muller, L. &amp; Hellmich, W." box="[1145,1414,474,499]" pageId="7" pageNumber="8" refId="ref23096" refString="Muller, L. &amp; Hellmich, W. 1936. Wissenschaftliche Ergebnisse der Deutschen Gran Chaco-Expedition. Amphibien und Reptilien. I. Amphibia, Chelonia, Loricata. Stuttgart, Strecker und Schroder." type="book" year="1936">Müller &amp; Hellmich, 1936</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FCA6FE0E71D9251A" author="Lutz, B." box="[795,910,506,531]" pageId="7" pageNumber="8" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FC2AFE0E71A9251A" author="Cei, J. M." box="[919,1022,506,531]" pageId="7" pageNumber="8" refId="ref18752" refString="Cei, J. M. 1980. Amphibians of Argentina. Monitore Zoologico Italiano, Nuova Serie, Monographia, ixii + 609 p." type="book" year="1980">Cei, 1980</bibRefCitation>
; this study).
</paragraph>
<paragraph id="8BF136EC6709F169FC82FDEE71C4237A" blockId="7.[793,1419,218,2131]" pageId="7" pageNumber="8">
The dorsal pattern with large irregular dark blotches, commonly with dark X-shaped marks composed of one or two pairs of inverted parenthesis-like blotches, distinguishes 
<taxonomicName id="4C4E4D6F6709F169FCC8FD8E7679259A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[885,1070,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FCC8FD8E7679259A" box="[885,1070,634,659]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
from 
<emphasis id="B93AEAFE6709F169FBCCFD8E7694243A" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FBCCFD8E76E8259A" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[1137,1215,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altae">S. altae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB74FD8E7746259A" baseAuthorityName="B.Lutz" baseAuthorityYear="1973" box="[1225,1297,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="alter">S. alter</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FAA6FD8E77D1259A" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[1307,1414,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="auratus">S. auratus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FD6E71FA25BA" baseAuthorityName="Goin" baseAuthorityYear="1966" box="[794,941,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="boesemani">S. boesemani</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC07FD6E766925BA" baseAuthorityName="Lutz" baseAuthorityYear="1968" box="[954,1086,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caldarum">S. caldarum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBF6FD6E76E825BA" authorityName="Nunes &amp; Pombal" authorityYear="2011" box="[1099,1215,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cretatus">S. cretatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB76FD6E77D125BA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[1227,1414,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="crospedospilus">S. crospedospilus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FD4E71DD25DA" authorityName="Pugliese, Pombal &amp; Sazima" authorityYear="2004" box="[794,906,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="curicica">S. curicica</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC2AFD4E767F25DA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[919,1064,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuspidatus">S. cuspidatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB8BFD4E76F625DA" baseAuthorityName="B.Lutz" baseAuthorityYear="1951" box="[1078,1185,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="duartei">S. duartei</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB12FD4E774925DA" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[1199,1310,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="exiguus">S. exiguus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FA91FD4E71CF25FA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscomarginatus">S. fuscomarginatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC1EFD2E764B25FA" authorityName="Nunes, Kwet &amp; Pombal" authorityYear="2012" box="[931,1052,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="imbegue">S. imbegue</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB95FD2E76DC25FA" authorityName="Nunes &amp; Pombal" authorityYear="2010" box="[1064,1163,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="juncae">S. juncae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB2AFD2E774025FA" baseAuthorityName="Bokermann" baseAuthorityYear="1964" box="[1175,1303,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="madeirae">S. madeirae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FA9EFD2E77D225FA" authorityName="Duellman &amp; Wiens" authorityYear="1993" box="[1315,1413,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="oreites">S. oreites</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FD0E71F6241A" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1937" box="[794,929,762,787]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachycrus">S. pachycrus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC10FD0E7628241A" baseAuthorityName="Fowler" baseAuthorityYear="1913" box="[941,1151,762,787]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="quinquefasciatus">S. quinquefasciatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB37FD0E7689241A" box="[1162,1246,762,787]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruber">S. ruber</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB54FD0E77D2241A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[1257,1413,762,787]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="squalirostris">S. squalirostris</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FCEE71C7243A" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[794,912,794,819]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="staufferi">S. staufferi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC20FCEE7659243A" authorityName="Ron, Duellman, Caminer &amp; Pazmino" authorityYear="2018" box="[925,1038,794,819]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tsachila">S. tsachila</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBA1FCEE76E8243A" authorityName="Nunes, Kwet &amp; Pombal" authorityYear="2012" box="[1052,1215,794,819]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tymbamirim">S. tymbamirim</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FAB8FCEE717D245A" authorityName="Brusquetti, Jansen, Barrio-Amoros, Segalla &amp; Haddad" authorityYear="2014" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="villasboasi">
<emphasis id="B93AEAFE6709F169FAB8FCEE717D245A" italics="true" pageId="7" pageNumber="8">S. villasboasi</emphasis>
</taxonomicName>
(variable number of dorsal and/or lateral stripes; 
<emphasis id="B93AEAFE6709F169FAE2FCCE77DD245A" box="[1375,1418,826,851]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FCA7FCAE7187247A" author="Duellman, W. E." box="[794,976,858,883]" pageId="7" pageNumber="8" pagination="1 - 753" refId="ref19657" refString="Duellman, W. E. 1970. The hylid frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1: 1 - 753." type="journal article" year="1970">Duellman, 1970</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FC62FCAE760F247A" author="Lutz, B." box="[991,1112,858,883]" pageId="7" pageNumber="8" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FBDAFCAE77D1247A" author="Duellman, W. E. &amp; Wiens, J. J." box="[1127,1414,858,883]" pageId="7" pageNumber="8" pagination="1 - 57" refId="ref19931" refString="Duellman, W. E. &amp; Wiens, J. J. 1993. Hylid frogs of the genus Scinax Wagler, 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History, University of Kansas, 153: 1 - 57." type="journal article" year="1993">Duellman &amp; Wiens, 1993</bibRefCitation>
; Pugliese 
<emphasis id="B93AEAFE6709F169FCC3FC8E71EF249A" box="[894,952,890,915]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2004; Nunes 
<emphasis id="B93AEAFE6709F169FBEFFC8E76DB249A" box="[1106,1164,890,915]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2012; Brusquetti 
<emphasis id="B93AEAFE6709F169FAECFC8E77DC249A" box="[1361,1419,890,915]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2014; Ron 
<emphasis id="B93AEAFE6709F169FC2EFC6E719A24BA" box="[915,973,922,947]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018; this study), and 
<emphasis id="B93AEAFE6709F169FB71FC6E71F524FA" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FB71FC6E774A24BA" baseAuthorityName="Fouquette &amp; Pyburn" baseAuthorityYear="1972" box="[1228,1309,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="blairi">S. blairi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FA95FC6E710724DA" authorityName="Drummond, Baeta &amp; Pires" authorityYear="2007" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabralensis">S. cabralensis</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCE2FC4E71AC24DA" baseAuthorityName="De la Riva" baseAuthorityYear="1990" box="[863,1019,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="chiquitanus">S. chiquitanus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBB7FC4E763B24DA" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[1034,1132,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="danae">S. danae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBC1FC4E775224DA" authorityName="Moravec, Tuanama, Perez-Pena &amp; Lehr" authorityYear="2009" box="[1148,1285,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="iquitorum">S. iquitorum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FAA9FC4E77D124DA" authorityName="Pyburn" authorityYear="1992" box="[1300,1414,954,979]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lindsayi">S. lindsayi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FC2E71C924FA" baseAuthorityName="Cardoso &amp; Sazima" baseAuthorityYear="1980" box="[794,926,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="maracaya">S. maracaya</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FC66FC2E76DD24FA" authorityName="Ferrao, Moravec, Kaefer, Fraga &amp; Lima" authorityYear="2018" box="[987,1162,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B93AEAFE6709F169FC66FC2E76DD24FA" box="[987,1162,986,1011]" italics="true" pageId="7" pageNumber="8">S. strussmannae</emphasis>
</taxonomicName>
(scattered or homogeneously distributed spots and/or irregular blotches; 
<emphasis id="B93AEAFE6709F169FADDFC0E77DC231A" box="[1376,1419,1018,1043]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FCA7FBEE7613233A" author="Fouquette Jr., M. J. &amp; Pyburn, W. F." box="[794,1092,1050,1075]" pageId="7" pageNumber="8" pagination="176 - 181" refId="ref20983" refString="Fouquette Jr., M. J. &amp;. Pyburn, W. F. 1972. A new Colombian treefrog of the Hyla rubra complex. Herpetologica, 28: 176 - 181." type="journal article" year="1972">Fouquette &amp; Pyburn, 1972</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FBF2FBEE7708233A" author="Cardoso, A. J. &amp; Sazima, I." box="[1103,1375,1050,1075]" pageId="7" pageNumber="8" pagination="75 - 79" refId="ref18559" refString="Cardoso, A. J. &amp; Sazima, I. 1980. Nova especie de Hyla do sudeste brasileiro (Amphibia, Anura, Hylidae). Revista Brasileira de Biologia, 40 (1): 75 - 79." type="journal article" year="1980">Cardoso &amp; Sazima, 1980</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FAD7FBEE71F0235A" author="De la Riva, I." pageId="7" pageNumber="8" pagination="81 - 86" refId="ref19337" refString="De la Riva, I. 1990. Una especie nueva de Ololygon (Anura: Hylidae) procedente de Bolivia. Revista Espanola de Herpetologia, 4: 81 - 86." type="journal article" year="1990">De la Riva, 1990</bibRefCitation>
; Drummond 
<emphasis id="B93AEAFE6709F169FB88FBCE763A235A" box="[1077,1133,1082,1107]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2007; Ferrão 
<emphasis id="B93AEAFE6709F169FABDFBCE776E235A" box="[1280,1337,1082,1107]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018b; this study).
</paragraph>
<paragraph id="8BF136EC6709F169FC82FB8E771B203A" blockId="7.[793,1419,218,2131]" pageId="7" pageNumber="8">
The hidden surfaces of thighs dark colored with light irregular pale blotches, yellow in living specimens differentiate 
<taxonomicName id="4C4E4D6F6709F169FC10FB4E763D23DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[941,1130,1210,1235]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FC10FB4E763D23DA" box="[941,1130,1210,1235]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
from 
<emphasis id="B93AEAFE6709F169FB0BFB4E7674229A" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FB0BFB4E775F23DA" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[1206,1288,1210,1235]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altae">S. altae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FAABFB4E77D123DA" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[1302,1414,1210,1235]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="auratus">S. auratus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FB2E719A23FA" baseAuthorityName="Rivero" baseAuthorityYear="1961" box="[794,973,1242,1267]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="baumgardneri">S. baumgardneri</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC6AFB2E763023FA" baseAuthorityName="Goin" baseAuthorityYear="1966" box="[983,1127,1242,1267]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="boesemani">S. boesemani</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBCCFB2E76B723FA" authorityName="Nunes &amp; Pombal" authorityYear="2011" box="[1137,1248,1242,1267]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cretatus">S. cretatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB57FB2E716E221A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="crospedospilus">S. crospedospilus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCF9FB0E71BE221A" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[836,1001,1274,1299]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cruentomma">S. cruentomma</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC4EFB0E76D6221A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[1011,1153,1274,1299]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuspidatus">S. cuspidatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB36FB0E76BE221A" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[1163,1257,1274,1299]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="danae">S. danae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB4EFB0E77D2221A" baseAuthorityName="Cope" baseAuthorityYear="1876" box="[1267,1413,1274,1299]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="elaeochroa">S. elaeochroa</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA6FAEE71DC223A" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[795,907,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="exiguus">S. exiguus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC24FAEE763B223A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[921,1132,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fuscomarginatus">S. fuscomarginatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBC7FAEE76A7223A" authorityName="Duellman &amp; Wiens" authorityYear="1993" box="[1146,1264,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ictericus">S. ictericus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB43FAEE77D0223A" authorityName="Moravec, Tuanama, Perez-Pena &amp; Lehr" authorityYear="2009" box="[1278,1415,1306,1331]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="iquitorum">S. iquitorum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FACE71CC225A" baseAuthorityName="Bokermann" baseAuthorityYear="1964" box="[794,923,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="madeirae">S. madeirae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC15FACE7664225A" authorityName="Barrio-Amoros, Orellana &amp; Chacon-Ortiz" authorityYear="2004" box="[936,1075,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="manriquei">S. manriquei</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB82FACE769F225A" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1937" box="[1087,1224,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pachycrus">S. pachycrus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB68FACE77D2225A" authorityName="Ferrao, Fraga, Moravec, Kaefer &amp; Lima" authorityYear="2018" box="[1237,1413,1338,1363]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruberoculatus">S. ruberoculatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FAAE71C6227A" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[794,913,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="staufferi">S. staufferi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC1DFAAE7602227A" authorityName="Ferrao, Moravec, Kaefer, Fraga &amp; Lima" authorityYear="2018" box="[928,1109,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="strussmannae">S. strussmannae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FBD9FAAE7754227A" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[1124,1283,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="squalirostris">S. squalirostris</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FAAEFAAE77D2227A" authorityName="Ron, Duellman, Caminer &amp; Pazmino" authorityYear="2018" box="[1299,1413,1370,1395]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tsachila">S. tsachila</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FCA7FA8E71F3229A" authorityName="Brusquetti, Jansen, Barrio-Amoros, Segalla &amp; Haddad" authorityYear="2014" box="[794,932,1402,1427]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="villasboasi">S. villasboasi</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FC13FA8E7648229A" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[942,1055,1402,1427]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="wandae">S. wandae</taxonomicName>
,
</emphasis>
and species of the 
<taxonomicName id="4C4E4D6F6709F169FABCFA8E77DC229A" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[1281,1419,1402,1427]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="uruguayus">
<emphasis id="B93AEAFE6709F169FABCFA8E77DC229A" box="[1281,1419,1402,1427]" italics="true" pageId="7" pageNumber="8">S. uruguayus</emphasis>
</taxonomicName>
Group (hidden surfaces of thighs uniform, light or dark colored; 
<emphasis id="B93AEAFE6709F169FCC0FA4E71FF22DA" box="[893,936,1466,1491]" italics="true" pageId="7" pageNumber="8">e.g.,</emphasis>
<bibRefCitation id="EFDF4B1D6709F169FC0FFA4E766822DA" author="Rivero, J. A." box="[946,1087,1466,1491]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref24320" refString="Rivero, J. A. 1961. SalientiaofVenezuela. BulletinoftheMuseumofComparative Zoology, Harvard College, 126: 1 - 207." type="journal article" year="1961">Rivero, 1961</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FBF1FA4E775622DA" author="Duellman, W. E." box="[1100,1281,1466,1491]" pageId="7" pageNumber="8" pagination="1 - 753" refId="ref19657" refString="Duellman, W. E. 1970. The hylid frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1: 1 - 753." type="journal article" year="1970">Duellman, 1970</bibRefCitation>
, 
<bibRefCitation id="EFDF4B1D6709F169FAB3FA4E771022DA" author="Duellman, W. E." box="[1294,1351,1466,1491]" pageId="7" pageNumber="8" pagination="864 - 870" refId="ref19849" refString="Duellman, W. E. 1986. Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986: 864 - 870." type="journal article" year="1986">1986</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FAE8FA4E710422FA" author="Lutz, B." pageId="7" pageNumber="8" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FCE3FA2E764222FA" author="De la Riva, I." box="[862,1045,1498,1523]" pageId="7" pageNumber="8" pagination="81 - 86" refId="ref19337" refString="De la Riva, I. 1990. Una especie nueva de Ololygon (Anura: Hylidae) procedente de Bolivia. Revista Espanola de Herpetologia, 4: 81 - 86." type="journal article" year="1990">De la Riva, 1990</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FB9DFA2E776322FA" author="Duellman, W. E. &amp; Wiens, J. J." box="[1056,1332,1498,1523]" pageId="7" pageNumber="8" pagination="1 - 57" refId="ref19931" refString="Duellman, W. E. &amp; Wiens, J. J. 1993. Hylid frogs of the genus Scinax Wagler, 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History, University of Kansas, 153: 1 - 57." type="journal article" year="1993">Duellman &amp; Wiens, 1993</bibRefCitation>
; Barrio-Amorós 
<emphasis id="B93AEAFE6709F169FCC7FA0E71E2211A" box="[890,949,1530,1555]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2004; 
<bibRefCitation id="EFDF4B1D6709F169FBB9FA0E775E211A" author="Nunes, I. &amp; Pombal Jr., J. P." box="[1028,1289,1530,1555]" pageId="7" pageNumber="8" pagination="80 - 88" refId="ref23430" refString="Nunes, I. &amp; Pombal Jr., J. P. 2011. A new snouted treefrog of the speciose genus Scinax Wagler (Anura, Hylidae) from northeastern Brazil. Herpetologica, 67 (1): 80 - 88." type="journal article" year="2011">Nunes &amp; Pombal, 2011</bibRefCitation>
; Brusquetti 
<emphasis id="B93AEAFE6709F169FCA7F9EE7104213A" box="[794,851,1562,1587]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2014; Ferrão 
<emphasis id="B93AEAFE6709F169FC56F9EE7673213A" box="[1003,1060,1562,1587]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018a, b; Ron 
<emphasis id="B93AEAFE6709F169FB75F9EE7755213A" box="[1224,1282,1562,1587]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018; Baldo 
<emphasis id="B93AEAFE6709F169FCA6F9CE7102215A" box="[795,853,1594,1619]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2019; this study), 
<emphasis id="B93AEAFE6709F169FB9DF9CE76A4215A" box="[1056,1267,1594,1619]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FB9DF9CE76C1215A" baseAuthorityName="Cope" baseAuthorityYear="1874" box="[1056,1174,1594,1619]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="funereus">S. funereus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F169FB1DF9CE76B9215A" authorityName="Ferrao, Moravec, Fraga, Pinheiro de Almeida, Kaefer &amp; Lima" authorityYear="2017" box="[1184,1262,1594,1619]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="onca">S. onca</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6709F169FA91F9CE7111217A" authorityName="Moravec, Tuanama, Perez-Pena &amp; Lehr" authorityYear="2009" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="iquitorum">
<emphasis id="B93AEAFE6709F169FA91F9CE7111217A" italics="true" pageId="7" pageNumber="8">S. iquitorum</emphasis>
</taxonomicName>
(hidden surfaces of thighs with horizontal or irregular dark blotches; 
<bibRefCitation id="EFDF4B1D6709F169FC4AF98E76FC219A" author="Duellman, W. E." box="[1015,1195,1658,1683]" pageId="7" pageNumber="8" pagination="1 - 21" refId="ref19691" refString="Duellman, W. E. 1971. A taxonomic review of South American hylid frogs, genus Phrynohyas. Occasional Papers of the Museum of Natural History, University of Kansas, 4: 1 - 21." type="journal article" year="1971">Duellman, 1971</bibRefCitation>
; Ferrão 
<emphasis id="B93AEAFE6709F169FAB5F98E7713219A" box="[1288,1348,1658,1683]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2017; Moravec 
<emphasis id="B93AEAFE6709F169FC3DF96E71EE21BA" box="[896,953,1690,1715]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2009; this study), and from species of the 
<taxonomicName id="4C4E4D6F6709F169FCA6F94E71C121DA" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[795,918,1722,1747]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B93AEAFE6709F169FCA6F94E71C121DA" box="[795,918,1722,1747]" italics="true" pageId="7" pageNumber="8">S. rostratus</emphasis>
</taxonomicName>
Group (hidden surfaces of thighs uniform light or marked with bold dark and light mottling or broad vertical bars; 
<bibRefCitation id="EFDF4B1D6709F169FC4BF90E76E3201A" author="Duellman, W. E." box="[1014,1204,1786,1811]" pageId="7" pageNumber="8" pagination="177 - 192" refId="ref19730" refString="Duellman, W. E. 1972 a. South American Frogs of the Hyla rostrata group (Amphibia, Anura, Hylidae). Zoologische Mededelingen, 47 (14): 177 - 192." type="journal article" year="1972">
Duellman, 1972 
<bibRefCitation id="EFDF4B1D6709F169FB1AF90E76E3201A" author="Bokermann, W. C. A. &amp; Sazima, I." box="[1191,1204,1786,1811]" pageId="7" pageNumber="8" pagination="521 - 528" refId="ref18334" refString="Bokermann, W. C. A. &amp; Sazima, I. 1973. Anfibios da Serra do Cipo, Minas Gerais, Brasil. 1: Duas especies novas de Hyla (Anura, Hylidae). Revista Brasileira de Biologia, 33 (3): 521 - 528." type="journal article" year="1973">a</bibRefCitation>
</bibRefCitation>
, 1973; 
<bibRefCitation id="EFDF4B1D6709F169FABDF90E77D1201A" author="Henle, K." box="[1280,1414,1786,1811]" pageId="7" pageNumber="8" pagination="76 - 82" refId="ref21505" refString="Henle, K. 1991. Ololygon pedromedinae sp. nov., ein neuer Knickzehenlaubfrosch (Hylidae) aus Peru. Salamandra, 27 (1 - 2): 76 - 82." type="journal article" year="1991">Henle, 1991</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6709F169FCA6F8EE7658203A" author="Lescure, J. &amp; Marty, C." box="[795,1039,1818,1843]" pageId="7" pageNumber="8" refId="ref22401" refString="Lescure, J. &amp; Marty, C. 2000. Atlas des amphibiens de Guyane. Collection Patrimoines Naturels." type="book" year="2000">Lescure &amp; Marty, 2000</bibRefCitation>
; 
<collectingRegion id="498AF80E6709F169FBA5F8EE7619203A" box="[1048,1102,1818,1843]" country="Peru" name="Lima" pageId="7" pageNumber="8">Lima</collectingRegion>
<emphasis id="B93AEAFE6709F169FBEEF8EE76DC203A" box="[1107,1163,1818,1843]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2005; this study).
</paragraph>
<paragraph id="8BF136EC6709F166FC82F8CE70D426BA" blockId="7.[793,1419,218,2131]" lastBlockId="8.[168,796,218,1203]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
The yellowish golden or bronze iris, with a median black streak, distinguishes 
<taxonomicName id="4C4E4D6F6709F169FBF4F8AE7753207A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1097,1284,1882,1907]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FBF4F8AE7753207A" box="[1097,1284,1882,1907]" italics="true" pageId="7" pageNumber="8">Scinax x‑signatus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C4E4D6F6709F169FAF4F8AE71D4209A" baseAuthorityName="Duellman" baseAuthorityYear="1972" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cruentomma">
<emphasis id="B93AEAFE6709F169FAF4F8AE71D4209A" italics="true" pageId="7" pageNumber="8">S. cruentomma</emphasis>
</taxonomicName>
(silvery bronze iris, with a median red streak; 
<bibRefCitation id="EFDF4B1D6709F169FCA7F86E764E20BA" author="Duellman, W. E." box="[794,1049,1946,1971]" pageId="7" pageNumber="8" pagination="265 - 271" refId="ref19767" refString="Duellman, W. E. 1972 b. A New Species of Hyla from Amazonian Ecuador. Copeia, 1972: 265 - 271." type="journal article" year="1972">
Duellman 
<emphasis id="B93AEAFE6709F169FC33F86E719F20BA" box="[910,968,1946,1971]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
1972b
</bibRefCitation>
), 
<taxonomicName id="4C4E4D6F6709F169FB96F86E768E20BA" authorityName="Ferrao, Fraga, Moravec, Kaefer &amp; Lima" authorityYear="2018" box="[1067,1241,1946,1971]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ruberoculatus">
<emphasis id="B93AEAFE6709F169FB96F86E768E20BA" box="[1067,1241,1946,1971]" italics="true" pageId="7" pageNumber="8">S. ruberoculatus</emphasis>
</taxonomicName>
(bicolored, reddish upper half and grey lower half; Ferrão 
<emphasis id="B93AEAFE6709F169FB47F84E776420DA" box="[1274,1331,1978,2003]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2018a), and species of the 
<taxonomicName id="4C4E4D6F6709F169FC4FF82E762A20FA" baseAuthorityName="Schmidt" baseAuthorityYear="1944" box="[1010,1149,2010,2035]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="uruguayus">
<emphasis id="B93AEAFE6709F169FC4FF82E762A20FA" box="[1010,1149,2010,2035]" italics="true" pageId="7" pageNumber="8">S. uruguayus</emphasis>
</taxonomicName>
Group (bicolored, golden upper half and dark brown to black lower half; Baldo 
<emphasis id="B93AEAFE6709F169FCA6F7EE710F2F3A" box="[795,856,2074,2099]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
2019). The absence of physiological chlorosis in 
<taxonomicName id="4C4E4D6F6709F169FCA7F7CE71C82F5A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[794,927,2106,2131]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6709F169FCA7F7CE71C82F5A" box="[794,927,2106,2131]" italics="true" pageId="7" pageNumber="8">S. x‑signatus</emphasis>
</taxonomicName>
distinguishes it from 
<emphasis id="B93AEAFE6709F166FB33F7CE73E0263A" italics="true" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D6F6709F169FB33F7CE774A2F5A" baseAuthorityName="Goin" baseAuthorityYear="1966" box="[1166,1309,2106,2131]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="boesemani">S. boesemani</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6709F166FA98F7CE729E27FA" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">S. caprarius</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FF6EFF2E732E27FA" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[211,377,218,243]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cruentomma">S. cruentomma</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FE3FFF2E705827FA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[386,527,218,243]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cuspidatus">S. cuspidatus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FDA4FF2E70FD27FA" baseAuthorityName="Cope" baseAuthorityYear="1876" box="[537,682,218,243]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="elaeochroa">S. elaeochroa</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FD08FF2E7294261A" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="funereus">S. funereus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FF73FF0E7317261A" authorityName="Duellman &amp; Wiens" authorityYear="1993" box="[206,320,250,275]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ictericus">S. ictericus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FEF6FF0E7387261A" authorityName="Moravec, Tuanama, Perez-Pena &amp; Lehr" authorityYear="2009" box="[331,464,250,275]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="iquitorum">S. iquitorum</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FE66FF0E70D5261A" baseAuthorityName="Pyburn" baseAuthorityYear="1993" box="[475,642,250,275]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="karenanneae">S. karenanneae</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FD30FF0E7141261A" authorityName="Barrio-Amoros, Orellana &amp; Chacon-Ortiz" authorityYear="2004" box="[653,790,250,275]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="manriquei">S. manriquei</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FF11FEEE72AD263A" authorityName="Ferrao, Moravec, Fraga, Pinheiro de Almeida, Kaefer &amp; Lima" authorityYear="2017" box="[172,250,282,307]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="onca">S. onca</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FEB9FEEE73E4263A" authorityName="Ferrao, Moravec, Kaefer, Fraga &amp; Lima" authorityYear="2018" box="[260,435,282,307]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="strussmannae">S. strussmannae</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6706F166FE53FEEE700E263A" authorityName="Ron, Duellman, Caminer &amp; Pazmino" authorityYear="2018" box="[494,601,282,307]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="tsachila">
<emphasis id="B93AEAFE6706F166FE53FEEE700E263A" box="[494,601,282,307]" italics="true" pageId="8" pageNumber="9">S. tsachila</emphasis>
</taxonomicName>
(present in these species; 
<bibRefCitation id="EFDF4B1D6706F166FEB4FECE73D1265A" author="Leon, J. R." box="[265,390,314,339]" pageId="8" pageNumber="9" pagination="505 - 545" refId="ref22320" refString="Leon, J. R. 1969. The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18: 505 - 545." type="journal article" year="1969">León, 1969</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6706F166FE2DFECE7052265A" author="Lutz, B." box="[400,517,314,339]" pageId="8" pageNumber="9" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6706F166FDADFECE70F1265A" author="Pyburn, W. F." box="[528,678,314,339]" pageId="8" pageNumber="9" pagination="46 - 50" refId="ref24122" refString="Pyburn, W. F. 1993. A new species of dimorphic tree frog, genus Hyla (Amphibia: Anura: Hylidae), from the Vaupes River of Colombia. Proceedings of the Biological Society of Washington, 106 (1): 46 - 50." type="journal article" year="1993">Pyburn, 1993</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6706F166FD0CFECE72B2267A" author="La Marca, E." pageId="8" pageNumber="9" pagination="1 - 9" refId="ref22206" refString="La Marca, E. 2004. Descripcion de dos nuevos anfibios del Piedemonte Andino de Venezuela. Herpetotropicos, 1 (1): 1 - 9." type="journal article" year="2004">La Marca, 2004</bibRefCitation>
;Moravec 
<emphasis id="B93AEAFE6706F166FEF1FEAE73D4267A" box="[332,387,346,371]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2009;Cole 
<emphasis id="B93AEAFE6706F166FE41FEAE7064267A" box="[508,563,346,371]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2013; 
<bibRefCitation id="EFDF4B1D6706F166FDC5FEAE731D269A" author="Melo-Sampaio, P. R. &amp; Souza, M. B." pageId="8" pageNumber="9" pagination="1 - 7" refId="ref22967" refString="Melo-Sampaio, P. R. &amp; Souza, M. B. 2015. New and noteworthy distributional records of treefrogs (Anura) from southwestern Amazonia. Check List, 11: 1 - 7." type="journal article" year="2015">Melo-Sampaio &amp; Souza, 2015</bibRefCitation>
; Ferrão 
<emphasis id="B93AEAFE6706F166FE18FE8E73B7269A" box="[421,480,378,403]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2017, 2018b; 
<bibRefCitation id="EFDF4B1D6706F166FDC2FE8E72B426BA" author="Acosta-Galvis, A. R." pageId="8" pageNumber="9" pagination="131 - 159" refId="ref17424" refString="Acosta-Galvis, A. R. 2018. Una nueva rana de huesos verdes del genero Scinax (Anura: Hylidae) asociada a los bosques subandinos de la cuenca del rio Magdalena, Colombia. Biota Colombiana, 19 (Supp. 1): 131 - 159." type="journal article" year="2018">Acosta-Galvis, 2018</bibRefCitation>
; Ron 
<emphasis id="B93AEAFE6706F166FEA3FE6E730126BA" box="[286,342,410,435]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2018;Taboada 
<emphasis id="B93AEAFE6706F166FDBDFE6E706F26BA" box="[512,568,410,435]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2020).
</paragraph>
<paragraph id="8BF136EC6706F166FF6DFE4E714C23BA" blockId="8.[168,796,218,1203]" pageId="8" pageNumber="9">
The advertisement call composed of a single multipulsed short note (0.11- 
<geoCoordinate id="EE7A502B6706F166FE7DFE2E705226FA" box="[448,517,474,499]" degrees="0.25" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-0.25">0.25 s</geoCoordinate>
), with 6-14 pulses/note, and pulse rate of 52-64 pulses/s differentiates 
<taxonomicName id="4C4E4D6F6706F166FD69FE0E734F253A" baseAuthorityName="Spix" baseAuthorityYear="1824" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6706F166FD69FE0E734F253A" italics="true" pageId="8" pageNumber="9">Scinax x‑signatus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C4E4D6F6706F166FEDBFDEE73AA253A" authorityName="De la Riva" authorityYear="1993" box="[358,509,538,563]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="castroviejoi">
<emphasis id="B93AEAFE6706F166FEDBFDEE73AA253A" box="[358,509,538,563]" italics="true" pageId="8" pageNumber="9">S. castroviejoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F6706F166FDFDFDEE70E0253A" baseAuthorityName="Bokermann" baseAuthorityYear="1968" box="[576,695,538,563]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="eurydice">
<emphasis id="B93AEAFE6706F166FDFDFDEE70E0253A" box="[576,695,538,563]" italics="true" pageId="8" pageNumber="9">S. eurydice</emphasis>
</taxonomicName>
(two or three multipulsed notes; 
<bibRefCitation id="EFDF4B1D6706F166FE00FDCE703C255A" author="De la Riva, I." box="[445,619,570,595]" pageId="8" pageNumber="9" pagination="41 - 46" refId="ref19370" refString="De la Riva, I. 1993. A new species of Scinax (Anura, Hylidae) from Argentina and Bolivia. Journal of Herpetology, 27: 41 - 46." type="journal article" year="1993">De la Riva, 1993</bibRefCitation>
; De la Riva 
<emphasis id="B93AEAFE6706F166FD5CFDCE714E255A" box="[737,793,570,595]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
1994; Pombal 
<emphasis id="B93AEAFE6706F166FEF1FDAE73D0257A" box="[332,391,602,627]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
1995a; Magrini 
<emphasis id="B93AEAFE6706F166FD82FDAE702D257A" box="[575,634,602,627]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2011; Mângia 
<emphasis id="B93AEAFE6706F166FF17FD8E72B3259A" box="[170,228,634,659]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2017), 
<emphasis id="B93AEAFE6706F166FE84FD8E73AA259A" box="[313,509,634,659]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D6F6706F166FE84FD8E73D5259A" baseAuthorityName="B.Lutz" baseAuthorityYear="1973" box="[313,386,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="alter">S. alter</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FE30FD8E73AD259A" authorityName="Pugliese, Pombal &amp; Sazima" authorityYear="2004" box="[397,506,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="curicica">S. curicica</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6706F166FD8AFD8E70FB259A" authorityName="Pombal, Haddad &amp; Kasahara" authorityYear="1995" box="[567,684,634,659]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="perereca">
<emphasis id="B93AEAFE6706F166FD8AFD8E70FB259A" box="[567,684,634,659]" italics="true" pageId="8" pageNumber="9">S. perereca</emphasis>
</taxonomicName>
(note duration 0.28- 
<geoCoordinate id="EE7A502B6706F166FE96FD6E733725BA" box="[299,352,666,691]" degrees="4.5" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="5555" value="-4.5">4.5 s</geoCoordinate>
and 21-152 pulses/note; Pombal 
<emphasis id="B93AEAFE6706F166FD62FD6E714D25BA" box="[735,794,666,691]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
1995a, b; Pugliese 
<emphasis id="B93AEAFE6706F166FEC3FD4E73ED25DA" box="[382,442,698,723]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2004), 
<emphasis id="B93AEAFE6706F166FDB3FD4E737B25FA" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D6F6706F166FDB3FD4E70E325DA" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[526,692,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cruentomma">S. cruentomma</taxonomicName>
, 
<taxonomicName id="4C4E4D6F6706F166FD02FD4E737F25FA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="fuscomarginatus">S. fuscomarginatus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C4E4D6F6706F166FEC5FD2E707825FA" authorityName="Ferrao, Moravec, Kaefer, Fraga &amp; Lima" authorityYear="2018" box="[376,559,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B93AEAFE6706F166FEC5FD2E707825FA" box="[376,559,730,755]" italics="true" pageId="8" pageNumber="9">S. strussmannae</emphasis>
</taxonomicName>
(17-90 pulses/note and 113-272 pulses/s; De la Riva 
<emphasis id="B93AEAFE6706F166FD9CFD0E700B241A" box="[545,604,762,787]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
1994; 
<bibRefCitation id="EFDF4B1D6706F166FD15FD0E72A5243A" author="Duellman, W. E." pageId="8" pageNumber="9" pagination="265 - 271" refId="ref19767" refString="Duellman, W. E. 1972 b. A New Species of Hyla from Amazonian Ecuador. Copeia, 1972: 265 - 271." type="journal article" year="1972">Duellman, 1972b</bibRefCitation>
; Brusquetti 
<emphasis id="B93AEAFE6706F166FECCFCEE73FE243A" box="[369,425,794,819]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2014; Carvalho 
<emphasis id="B93AEAFE6706F166FDE8FCEE70DA243A" box="[597,653,794,819]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2015;Ferrão 
<emphasis id="B93AEAFE6706F166FF14FCCE72B0245A" box="[169,231,826,851]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2018b), 
<taxonomicName id="4C4E4D6F6706F166FEE9FCCE7395245A" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[340,450,826,851]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B93AEAFE6706F166FEE9FCCE7395245A" box="[340,450,826,851]" italics="true" pageId="8" pageNumber="9">S. exiguus</emphasis>
</taxonomicName>
(23-90 pulses/note; Carvalho 
<emphasis id="B93AEAFE6706F166FF17FCAE72B2247A" box="[170,229,858,883]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2017), 
<taxonomicName id="4C4E4D6F6706F166FE87FCAE73EE247A" baseAuthorityName="Bokermann" baseAuthorityYear="1964" box="[314,441,858,883]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="madeirae">
<emphasis id="B93AEAFE6706F166FE87FCAE73EE247A" box="[314,441,858,883]" italics="true" pageId="8" pageNumber="9">S. madeirae</emphasis>
</taxonomicName>
(note duration 0.72- 
<geoCoordinate id="EE7A502B6706F166FD18FCAE70B0247A" box="[677,743,858,883]" degrees="1.16" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-1.16">1.16 s</geoCoordinate>
and 104-145 pulses/s; Brusquetti 
<emphasis id="B93AEAFE6706F166FDBEFC8E7014249A" box="[515,579,890,915]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2014), 
<taxonomicName id="4C4E4D6F6706F166FD1EFC8E714D249A" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[675,794,890,915]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B93AEAFE6706F166FD1EFC8E714D249A" box="[675,794,890,915]" italics="true" pageId="8" pageNumber="9">S. staufferi</emphasis>
</taxonomicName>
(100-130 pulses/s; 
<bibRefCitation id="EFDF4B1D6706F166FEC5FC6E73A124BA" author="Leon, J. R." box="[376,502,922,947]" pageId="8" pageNumber="9" pagination="505 - 545" refId="ref22320" refString="Leon, J. R. 1969. The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18: 505 - 545." type="journal article" year="1969">León, 1969</bibRefCitation>
), 
<taxonomicName id="4C4E4D6F6706F166FDB8FC6E702324BA" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[517,628,922,947]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="wandae">
<emphasis id="B93AEAFE6706F166FDB8FC6E702324BA" box="[517,628,922,947]" italics="true" pageId="8" pageNumber="9">S. wandae</emphasis>
</taxonomicName>
(note duration 0.44- 
<geoCoordinate id="EE7A502B6706F166FF5CFC4E737724DA" box="[225,288,954,979]" degrees="0.69" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-0.69">0.69 s</geoCoordinate>
and 70-108 pulses/note; 
<bibRefCitation id="EFDF4B1D6706F166FD8BFC4E72B424FA" author="Pyburn, W. F. &amp; Fouquette, M. J." pageId="8" pageNumber="9" pagination="97 - 101" refId="ref24172" refString="Pyburn, W. F. &amp; Fouquette, M. J. 1971. A new striped treefrog from Central Colombia. Journal of Herpetology, 5: 97 - 101." type="journal article" year="1971">Pyburn &amp; Fouquette, 1971</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6706F166FF4FFC2E73FF24FA" author="Duellman, W. E." box="[242,424,986,1011]" pageId="8" pageNumber="9" pagination="864 - 870" refId="ref19849" refString="Duellman, W. E. 1986. Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986: 864 - 870." type="journal article" year="1986">Duellman, 1986</bibRefCitation>
; Pombal 
<emphasis id="B93AEAFE6706F166FDAAFC2E700324FA" box="[535,596,986,1011]" italics="true" pageId="8" pageNumber="9">et al.,</emphasis>
2011), and from some large species of the 
<taxonomicName id="4C4E4D6F6706F166FE57FC0E7031231A" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[490,614,1018,1043]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B93AEAFE6706F166FE57FC0E7031231A" box="[490,614,1018,1043]" italics="true" pageId="8" pageNumber="9">S. rostratus</emphasis>
</taxonomicName>
Group: 
<taxonomicName id="4C4E4D6F6706F166FD76FC0E72A4233A" baseAuthorityName="Cope" baseAuthorityYear="1887" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B93AEAFE6706F166FD76FC0E72A4233A" italics="true" pageId="8" pageNumber="9">S. boulengeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F6706F166FE90FBEE7384233A" baseAuthorityName="Brongersma" baseAuthorityYear="1933" box="[301,467,1050,1075]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B93AEAFE6706F166FE90FBEE7384233A" box="[301,467,1050,1075]" italics="true" pageId="8" pageNumber="9">S. proboscideus</emphasis>
</taxonomicName>
(80-230 pulses/s; 
<bibRefCitation id="EFDF4B1D6706F166FD27FBEE7141233A" author="Leon, J. R." box="[666,790,1050,1075]" pageId="8" pageNumber="9" pagination="505 - 545" refId="ref22320" refString="Leon, J. R. 1969. The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18: 505 - 545." type="journal article" year="1969">León, 1969</bibRefCitation>
; 
<bibRefCitation id="EFDF4B1D6706F166FF16FBCE7338235A" author="Duellman, W. E." box="[171,367,1082,1107]" pageId="8" pageNumber="9" pagination="177 - 192" refId="ref19730" refString="Duellman, W. E. 1972 a. South American Frogs of the Hyla rostrata group (Amphibia, Anura, Hylidae). Zoologische Mededelingen, 47 (14): 177 - 192." type="journal article" year="1972">Duellman, 1972a</bibRefCitation>
), 
<taxonomicName id="4C4E4D6F6706F166FE3DFBCE7391235A" box="[384,454,1082,1107]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="jolyi">
<emphasis id="B93AEAFE6706F166FE3DFBCE7391235A" box="[384,454,1082,1107]" italics="true" pageId="8" pageNumber="9">S. jolyi</emphasis>
</taxonomicName>
(note duration 
<geoCoordinate id="EE7A502B6706F166FDC3FBCE70E4235A" box="[638,691,1082,1107]" degrees="2.5" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="5555" value="-2.5">2.5 s</geoCoordinate>
and 180 pulses/note; 
<bibRefCitation id="EFDF4B1D6706F166FE85FBAE707C237A" author="Lescure, J. &amp; Marty, C." box="[312,555,1114,1139]" pageId="8" pageNumber="9" refId="ref22401" refString="Lescure, J. &amp; Marty, C. 2000. Atlas des amphibiens de Guyane. Collection Patrimoines Naturels." type="book" year="2000">Lescure &amp; Marty,2000</bibRefCitation>
), 
<taxonomicName id="4C4E4D6F6706F166FD85FBAE70F9237A" baseAuthorityName="Pyburn" baseAuthorityYear="1973" box="[568,686,1114,1139]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="B93AEAFE6706F166FD85FBAE70F9237A" box="[568,686,1114,1139]" italics="true" pageId="8" pageNumber="9">S.kennedyi</emphasis>
</taxonomicName>
(note duration 0.66- 
<geoCoordinate id="EE7A502B6706F166FE9AFB8E730D239A" box="[295,346,1146,1171]" degrees="2.9" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="5555" value="-2.9">2.9 s</geoCoordinate>
; 
<bibRefCitation id="EFDF4B1D6706F166FEDCFB8E73A0239A" author="Pyburn, W. F." box="[353,503,1146,1171]" pageId="8" pageNumber="9" pagination="297 - 301" refId="ref24055" refString="Pyburn, W. F. 1973. A new hylid frog from the Llanos of Colombia. Journal of Herpetology, 7: 297 - 301." type="journal article" year="1973">Pyburn, 1973</bibRefCitation>
),and 
<taxonomicName id="4C4E4D6F6706F166FD8FFB8E70F9239A" baseAuthorityName="Duellman" baseAuthorityYear="1973" box="[562,686,1146,1171]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sugillatus">
<emphasis id="B93AEAFE6706F166FD8FFB8E70F9239A" box="[562,686,1146,1171]" italics="true" pageId="8" pageNumber="9">S.sugillatus</emphasis>
</taxonomicName>
(note duration 0.28- 
<geoCoordinate id="EE7A502B6706F166FE95FB6E733123BA" box="[296,358,1178,1203]" degrees="0.60" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-0.6">0.60 s</geoCoordinate>
and 110-140 pulses/s; 
<bibRefCitation id="EFDF4B1D6706F166FDE2FB6E714623BA" author="Duellman, W. E." box="[607,785,1178,1203]" pageId="8" pageNumber="9" pagination="219 - 227" refId="ref19793" refString="Duellman, W. E. 1973. Descriptions of new hylid frogs from Colombia and Ecuador. Herpetologica, 29: 219 - 227." type="journal article" year="1973">Duellman, 1973</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35465676706F166FF17FB2E73B023FA" box="[170,487,1242,1267]" pageId="8" pageNumber="9" type="multiple">
<paragraph id="8BF136EC6706F166FF17FB2E73B023FA" blockId="8.[170,487,1242,1267]" box="[170,487,1242,1267]" pageId="8" pageNumber="9">
<emphasis id="B93AEAFE6706F166FF17FB2E73B023FA" bold="true" box="[170,487,1242,1267]" pageId="8" pageNumber="9">
Description of the 
<typeStatus id="54F5884E6706F166FE39FB2E73B023FA" box="[388,487,1242,1267]" pageId="8" pageNumber="9" type="neotype">neotype</typeStatus>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35465676706F163FF6DFAEE73DE265A" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF136EC6706F166FF6DFAEE77EC241A" blockId="8.[170,794,1306,1427]" lastBlockId="8.[868,1495,218,1427]" pageId="8" pageNumber="9">
Head as wide as long, HL 35.1% and HW 33.2% of SVL (
<figureCitation id="13752A696706F166FF0DFACE72B9225A" box="[176,238,1338,1363]" captionStart="Figure 2" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,1452,2076]" captionTargetId="figure-704@8.[170,1493,1452,2076]" captionTargetPageId="8" captionText="Figure 2. Neotype of Scinax x‑signatus(CFBH 44688,male).(A) Dorsal view.(B)Ventral view.Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4614065" httpUri="https://zenodo.org/record/4614065/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). Snout rounded in dorsal view and profile, with a low protuberance on the tip (
<figureCitation id="13752A696706F166FE44FAAE7032227A" box="[505,613,1370,1395]" captionStart="Figure 3" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[132,1379,204,2044]" captionTargetId="figure-1@9.[132,1379,204,2044]" captionTargetPageId="9" captionText="Figure 3. Neotype of Scinax x‑signatus (CFBH 44688, male). (A) Head, profile. (B) Head, dorsal view. (C) Right hand, ventral view. (D) Right foot, ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4614067" httpUri="https://zenodo.org/record/4614067/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, B</figureCitation>
). Nostrils dorsolateral, elliptical, protruded; IND 39.7% of IOD. 
<emphasis id="B93AEAFE6706F166FD7CFA8E71ED27FA" box="[705,954,218,1427]" italics="true" pageId="8" pageNumber="9">Canthus rostralis</emphasis>
marked, convex. Loreal region slightly concave. Eyes large, protuberant, ED 94.7% of IOD and 92.3% of END. Pupil horizontal, subelliptical. Tympanum rounded, separated from eye by a distance almost half TD; TD 75.0% of ED. Tympanic annulus rounded, with the posterior upper portion hidden by the supratympanic fold. Supratympanic fold evident, from the posterior upper portion of the tympanum to the insertion of the forearm. Vocal sac subgular, weakly bilobate, externally evident by the loose skin, not occupying space between head and body, and ventrally not reaching the pectoral fold (
<figureCitation id="13752A696706F166FC23FDCE71BC255A" box="[926,1003,570,595]" captionStart="Figure 2" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,1452,2076]" captionTargetId="figure-704@8.[170,1493,1452,2076]" captionTargetPageId="8" captionText="Figure 2. Neotype of Scinax x‑signatus(CFBH 44688,male).(A) Dorsal view.(B)Ventral view.Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4614065" httpUri="https://zenodo.org/record/4614065/files/figure.png" pageId="8" pageNumber="9">Fig. 2B</figureCitation>
). Pectoral fold present, with pre- and postaxillar elements. Vocal slits present, nearly parallel to the mandible, originating laterally to the tongue and running towards the corner of the mouth. Tongue ovoid, free laterally and posteriorly, slightly notched posteriorly. Vomerine teeth in two slightly separated convex series, bearing five (right) and four (left) teeth. Choanae oval.
</paragraph>
<paragraph id="8BF136EC6706F164FC36FCEE73CF23FA" blockId="8.[868,1495,218,1427]" lastBlockId="10.[169,1496,801,2131]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
Axillary membrane absent. Upper arm more slender than forearm. A series of small, flat, ulnar tubercles on the forearm. Fingers short and slender, fringed (
<figureCitation id="13752A696706F166FAC1FCAE779C247A" box="[1404,1483,858,883]" captionStart="Figure 3" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[132,1379,204,2044]" captionTargetId="figure-1@9.[132,1379,204,2044]" captionTargetPageId="9" captionText="Figure 3. Neotype of Scinax x‑signatus (CFBH 44688, male). (A) Head, profile. (B) Head, dorsal view. (C) Right hand, ventral view. (D) Right foot, ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4614067" httpUri="https://zenodo.org/record/4614067/files/figure.png" pageId="8" pageNumber="9">Fig. 3C</figureCitation>
). Relative finger length II&lt;III≈V&lt;IV. Discs elliptical, wider than long, 3FD 59.2% of TD; disc of Finger II smaller than others. Subarticular tubercles single, conical on fingers II and III; rounded and quadrangular on fingers IV and V. Supernumerary tubercles small, single, rounded. Inner metacarpal tubercle single, elliptical; outer metacarpal tubercle flat, nearly triangular, bilobate. Webbing absent between fingers II and III; basal between fingers III, IV, and V. Slightly thickened, light-colored nuptial pad, covering Metacarpal II dorsomedially, and ventrally extending from the base of inner metacarpal tubercle, obscuring its outer margin, to the subarticular tubercle (
<figureCitation id="13752A696706F166FCD1FB0E718B221A" box="[876,988,1274,1299]" captionStart="Figure 4" captionStartId="10.[170,229,801,826]" captionTargetBox="[170,1493,204,771]" captionTargetId="figure-698@10.[170,1493,204,771]" captionTargetPageId="10" captionText="Figure 4. Neotype of Scinax x‑signatus(CFBH 44688,male).Nuptial pad of the righthand.(A) Dorsal view.(B)Ventral view.Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4614069" httpUri="https://zenodo.org/record/4614069/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, B</figureCitation>
). Glandular acini on inner margin of fingers II-III; on Finger II from the distal margin of nuptial pad to the disc. Spicule-shaped papillary epidermal projections on the nuptial pad, margins of fingers II-III (
<figureCitation id="13752A696706F166FAE1FAAE779C227A" box="[1372,1483,1370,1395]" captionStart="Figure 4" captionStartId="10.[170,229,801,826]" captionTargetBox="[170,1493,204,771]" captionTargetId="figure-698@10.[170,1493,204,771]" captionTargetPageId="10" captionText="Figure 4. Neotype of Scinax x‑signatus(CFBH 44688,male).Nuptial pad of the righthand.(A) Dorsal view.(B)Ventral view.Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4614069" httpUri="https://zenodo.org/record/4614069/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, B</figureCitation>
), and dorsum of fingers II-V. Hindlimbs robust; TL 49.0% of SVL, FL 39.8% of SVL. Toes slender, fringed (Fig. 3D). Relative toe length I&lt;II&lt;III≈V&lt;IV. Discs elliptical, wider than long, slightly smaller than discs of fingers, 4TD = 3FD. Subarticular tubercles single, conical, rounded; supernumerary tubercles small, single, rounded. Inner metatarsal tubercle single, elliptical; outer metatarsal tubercle single, slightly marked, two thirds smaller than inner tubercle.Webbing formula I 2-2⁺ II 1¹ 
<superScript id="7C3B9BA46704F164FECDFBAD73222361" attach="none" box="[368,373,1113,1128]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
⁴-2⁺ III 1-2¹ 
<superScript id="7C3B9BA46704F164FE55FBAD73BA2361" attach="none" box="[488,493,1113,1128]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
³ IV 2⁺-1 V. Fringe on lateral margin of Toe V extends along the margin of the sole by a poorly developed ridge that reaches the distal portion of the metatarsus.Ventrolateral margin of tarsus smooth; heel slightly granular.
</paragraph>
<caption id="DF3166646706F166FF17F7CE76122F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614065" ID-Zenodo-Dep="4614065" box="[170,1093,2106,2131]" httpUri="https://zenodo.org/record/4614065/files/figure.png" pageId="8" pageNumber="9" startId="8.[170,229,2106,2131]" targetBox="[170,1493,1452,2076]" targetPageId="8">
<paragraph id="8BF136EC6706F166FF17F7CE76122F5A" blockId="8.[170,1093,2106,2131]" box="[170,1093,2106,2131]" pageId="8" pageNumber="9">
<emphasis id="B93AEAFE6706F166FF17F7CE730F2F5A" bold="true" box="[170,344,2106,2131]" pageId="8" pageNumber="9">Figure 2. Neotype of</emphasis>
<taxonomicName id="4C4E4D6F6706F166FEE0F7CE73BF2F5A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[349,488,2106,2131]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6706F166FEE0F7CE73BF2F5A" box="[349,488,2106,2131]" italics="true" pageId="8" pageNumber="9">Scinax x‑signatus</emphasis>
</taxonomicName>
(CFBH 44688,male).(A) Dorsal view.(B)Ventral view.Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="DF3166646707F167FFE3F7EE72A82F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614067" ID-Zenodo-Dep="4614067" httpUri="https://zenodo.org/record/4614067/files/figure.png" pageId="9" pageNumber="10" startId="9.[94,153,2074,2099]" targetBox="[132,1379,204,2044]" targetPageId="9">
<paragraph id="8BF136EC6707F167FFE3F7EE72A82F5A" blockId="9.[94,1418,2074,2131]" pageId="9" pageNumber="10">
<emphasis id="B93AEAFE6707F167FFE3F7EE73452F3A" bold="true" box="[94,274,2074,2099]" pageId="9" pageNumber="10">Figure 3. Neotype of</emphasis>
<taxonomicName id="4C4E4D6F6707F167FEA6F7EE73F02F3A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[283,423,2074,2099]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6707F167FEA6F7EE73F02F3A" box="[283,423,2074,2099]" italics="true" pageId="9" pageNumber="10">Scinax x‑signatus</emphasis>
</taxonomicName>
(CFBH 44688, male). (A) Head, profile. (B) Head, dorsal view. (C) Right hand, ventral view. (D) Right foot, ventral view. Scale bars = 2 mm.
</paragraph>
</caption>
<caption id="DF3166646704F164FF17FCD5776F2433" ID-DOI="http://doi.org/10.5281/zenodo.4614069" ID-Zenodo-Dep="4614069" box="[170,1336,801,826]" httpUri="https://zenodo.org/record/4614069/files/figure.png" pageId="10" pageNumber="11" startId="10.[170,229,801,826]" targetBox="[170,1493,204,771]" targetPageId="10">
<paragraph id="8BF136EC6704F164FF17FCD5776F2433" blockId="10.[169,1496,801,2131]" box="[170,1336,801,826]" pageId="10" pageNumber="11">
<emphasis id="B93AEAFE6704F164FF17FCD5730F2433" bold="true" box="[170,344,801,826]" pageId="10" pageNumber="11">Figure 4. Neotype of</emphasis>
<taxonomicName id="4C4E4D6F6704F164FEE0FCD573BF2433" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[349,488,801,826]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6704F164FEE0FCD573BF2433" box="[349,488,801,826]" italics="true" pageId="10" pageNumber="11">Scinax x‑signatus</emphasis>
</taxonomicName>
<emphasis id="B93AEAFE6704F164FE56FCD5776F2433" bold="true" box="[491,1336,801,826]" pageId="10" pageNumber="11">(CFBH 44688,male).Nuptial pad of the right hand.(A) Dorsal view.(B)Ventral view.Scale bars = 1 mm.</emphasis>
</paragraph>
</caption>
<paragraph id="8BF136EC6704F164FF6DFB0E70A6213A" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">Cloacal opening directed posteriorly at upper level of thighs. Skin on dorsum of head, upper eyelid, trunk, and limbs smooth, with scattered granules. Posterior corner of eyes, around tympanum and forearm insertion, supratympanic fold, and flanks granular. Vocal sac, hidden surfaces of limbs, and inguinal region smooth; other ventral surfaces and subcloacal area densely covered with rounded, flat granules. Pectoral region and inner margin of upper- and forelimbs with glandular acini, covered with spicule-shaped papillary epidermal projections.</paragraph>
<paragraph id="8BF136EC6704F164FF17F9AE7782233A" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">
<emphasis id="B93AEAFE6704F164FF17F9AE73F9217A" bold="true" box="[170,430,1626,1651]" pageId="10" pageNumber="11">Measurements (mm):</emphasis>
SVL 36.7; HL 12.9; HW 12.2; IND 2.5; IOD 3.8; ED 3.6; END 3.9; TD 2.7; FL 14.6; TL 18.0; 3FD 1.6; 4TD 1.6. 
<emphasis id="B93AEAFE6704F164FF17F92E732221FA" bold="true" box="[170,373,1754,1779]" pageId="10" pageNumber="11">Coloration in life:</emphasis>
The description is based on the freshly euthanized specimen (
<figureCitation id="13752A696704F164FE00F90E73AD201A" box="[445,506,1786,1811]" captionStart="Hindlimbs" captionStartId="10.[208,322,858,883]" captionTargetBox="[94,1417,204,1300]" captionTargetId="figure-698@10.[170,1493,204,771]" captionTargetPageId="10" captionText="Hindlimbs robust; TL 49.0% of SVL, FL 39.8% of flanks. Flanks light with irregular, dark brown blotches. SVL. Toes slender, fringed (Fig. 3D). Relative toe length Dorsal surfaces of discs, fingers, toes, forearms, and tar- I&lt;II&lt;III≈V&lt;IV. Discs elliptical, wider than long, slightly sus brownish gray with transverse, brown bars; upper smaller than discs of fingers, 4TD = 3FD. Subarticular arms uniform; shanks and thighs with large dark brown tubercles single, conical, rounded; supernumerary tu- blotches.Toe webbing covered by brown melanophores. bercles small, single, rounded. Inner metatarsal tubercle Iris grayish bronze with thin black reticulations, thin yelsingle, elliptical; outer metatarsal tubercle single, slightly low halo bordering the pupil, and a median black streak. marked, two thirds smaller than inner tubercle.Webbing Soles and palms light brown; glandular pectoral reformula I 2-2⁺ II 1¹/⁴-2⁺ III 1-2¹/³ IV 2⁺-1 V. Fringe on lateral gion yellowish white; other ventral areas creamy white, margin of Toe V extends along the margin of the sole by immaculate,but margins of gular region, around forearm a poorly developed ridge that reaches the distal portion insertion and knees, tarsus, and shanks finely spotted of the metatarsus.Ventrolateral margin of tarsus smooth; with brown (Fig. 5B). Inguinal region yellow, with irregheel slightly granular. ular dark blotches. Hidden surfaces of thighs and shanks Cloacal opening directed posteriorly at upper level of brown, with small to medium-size, rounded or irregular thighs. Skin on dorsum of head, upper eyelid, trunk, and yellow blotches (Fig. 5C).White bones. limbs smooth, with scattered granules. Posterior corner of eyes, around tympanum and forearm insertion, su- Coloration in preservative: Paler than in the fresh specpratympanic fold, and flanks granular. Vocal sac, hidden imen.The coloration on the glandular pectoral region, insurfaces of limbs, and inguinal region smooth; other ven- guinal region, and hidden surfaces of thighs and shanks tral surfaces and subcloacal area densely covered with faded to light beige or cream white. rounded, flat granules. Pectoral region and inner margin of upper- and forelimbs with glandular acini, covered Variation with spicule-shaped papillary epidermal projections. See Table 1 for measurements of the available speci- Measurements (mm): SVL 36.7; HL 12.9; HW 12.2; mens. Vomerine teeth vary from 5 to 8. In some individ- IND 2.5; IOD 3.8; ED 3.6; END 3.9; TD 2.7; FL 14.6; TL 18.0; uals, the medial constriction of the vocal sac is barely 3FD 1.6; 4TD 1.6. evident externally. Snout rounded in dorsal view and" figureDoi="http://doi.org/10.5281/zenodo.4614071" httpUri="https://zenodo.org/record/4614071/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
). Dorsal color dark brown, with two pairs of large, irregular, black blotches on the suprascapular and sacral regions, and scattered, small, round or irregular, light blotches; interocular region with an inverted triangle-shaped, black marking (
<figureCitation id="13752A696704F164FD07F88E7147209A" box="[698,784,1914,1939]" captionStart="Hindlimbs" captionStartId="10.[208,322,858,883]" captionTargetBox="[94,1417,204,1300]" captionTargetId="figure-698@10.[170,1493,204,771]" captionTargetPageId="10" captionText="Hindlimbs robust; TL 49.0% of SVL, FL 39.8% of flanks. Flanks light with irregular, dark brown blotches. SVL. Toes slender, fringed (Fig. 3D). Relative toe length Dorsal surfaces of discs, fingers, toes, forearms, and tar- I&lt;II&lt;III≈V&lt;IV. Discs elliptical, wider than long, slightly sus brownish gray with transverse, brown bars; upper smaller than discs of fingers, 4TD = 3FD. Subarticular arms uniform; shanks and thighs with large dark brown tubercles single, conical, rounded; supernumerary tu- blotches.Toe webbing covered by brown melanophores. bercles small, single, rounded. Inner metatarsal tubercle Iris grayish bronze with thin black reticulations, thin yelsingle, elliptical; outer metatarsal tubercle single, slightly low halo bordering the pupil, and a median black streak. marked, two thirds smaller than inner tubercle.Webbing Soles and palms light brown; glandular pectoral reformula I 2-2⁺ II 1¹/⁴-2⁺ III 1-2¹/³ IV 2⁺-1 V. Fringe on lateral gion yellowish white; other ventral areas creamy white, margin of Toe V extends along the margin of the sole by immaculate,but margins of gular region, around forearm a poorly developed ridge that reaches the distal portion insertion and knees, tarsus, and shanks finely spotted of the metatarsus.Ventrolateral margin of tarsus smooth; with brown (Fig. 5B). Inguinal region yellow, with irregheel slightly granular. ular dark blotches. Hidden surfaces of thighs and shanks Cloacal opening directed posteriorly at upper level of brown, with small to medium-size, rounded or irregular thighs. Skin on dorsum of head, upper eyelid, trunk, and yellow blotches (Fig. 5C).White bones. limbs smooth, with scattered granules. Posterior corner of eyes, around tympanum and forearm insertion, su- Coloration in preservative: Paler than in the fresh specpratympanic fold, and flanks granular. Vocal sac, hidden imen.The coloration on the glandular pectoral region, insurfaces of limbs, and inguinal region smooth; other ven- guinal region, and hidden surfaces of thighs and shanks tral surfaces and subcloacal area densely covered with faded to light beige or cream white. rounded, flat granules. Pectoral region and inner margin of upper- and forelimbs with glandular acini, covered Variation with spicule-shaped papillary epidermal projections. See Table 1 for measurements of the available speci- Measurements (mm): SVL 36.7; HL 12.9; HW 12.2; mens. Vomerine teeth vary from 5 to 8. In some individ- IND 2.5; IOD 3.8; ED 3.6; END 3.9; TD 2.7; FL 14.6; TL 18.0; uals, the medial constriction of the vocal sac is barely 3FD 1.6; 4TD 1.6. evident externally. Snout rounded in dorsal view and" figureDoi="http://doi.org/10.5281/zenodo.4614071" httpUri="https://zenodo.org/record/4614071/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Upper lip light with diffuse brown blotches anteriorly, and a white stripe on the infraorbital region extending to posterior margin of the tympanum. Loreal region brown with small, irregular, black dots; dark brown canthal line. Post-orbital dark brown line from anterior corner of the eyes, upper margin of tympanum, to the middle of the flanks. Flanks light with irregular, dark brown blotches. Dorsal surfaces of discs, fingers, toes, forearms, and tarsus brownish gray with transverse, brown bars; upper arms uniform; shanks and thighs with large dark brown blotches.Toe webbing covered by brown melanophores. Iris grayish bronze with thin black reticulations, thin yellow halo bordering the pupil, and a median black streak.
</paragraph>
<paragraph id="8BF136EC6704F164FC36FBCE7758223A" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">Soles and palms light brown; glandular pectoral region yellowish white; other ventral areas creamy white, immaculate,but margins of gular region, around forearm insertion and knees, tarsus, and shanks finely spotted with brown (Fig. 5B). Inguinal region yellow, with irregular dark blotches. Hidden surfaces of thighs and shanks brown, with small to medium-size, rounded or irregular yellow blotches (Fig. 5C).White bones.</paragraph>
<paragraph id="8BF136EC6704F164FCD8FAAE76A522DA" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">
<emphasis id="B93AEAFE6704F164FCD8FAAE76C8227A" bold="true" box="[869,1183,1370,1395]" pageId="10" pageNumber="11">Coloration in preservative:</emphasis>
Paler than in the fresh specimen.The coloration on the glandular pectoral region, inguinal region, and hidden surfaces of thighs and shanks faded to light beige or cream white.
</paragraph>
<paragraph id="8BF136EC6704F164FCD8FA0E7795201A" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">
<emphasis id="B93AEAFE6704F164FCD8FA0E7185211A" bold="true" box="[869,978,1530,1555]" pageId="10" pageNumber="11">Variation</emphasis>
See Table 1 for measurements of the available specimens. Vomerine teeth vary from 5 to 8. In some individuals, the medial constriction of the vocal sac is barely evident externally. Snout rounded in dorsal view and profile, with or without a low protuberance on the tip. Toe webbing formula varies as follows: I (2⁺-2⁻) – (2¹ 
<superScript id="7C3B9BA46704F164FA1DF92D77F221E1" attach="none" box="[1440,1445,1753,1768]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
²-2⁻) II (1¹ 
<superScript id="7C3B9BA46704F164FC2EF90D71CF2001" attach="none" box="[915,920,1785,1800]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
²-1) – (2¹ 
<superScript id="7C3B9BA46704F164FC4EF90D71AF2001" attach="none" box="[1011,1016,1785,1800]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
²-2⁻) III (1¹ 
<superScript id="7C3B9BA46704F164FBDFF90D76302001" attach="none" box="[1122,1127,1785,1800]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
²-1) – (2¹ 
<superScript id="7C3B9BA46704F164FB7FF90D76902001" attach="none" box="[1218,1223,1785,1800]" fontSize="6" pageId="10" pageNumber="11">/</superScript>
²-2) IV (2⁺-2⁻) – (1⁺-1) V.
</paragraph>
<paragraph id="8BF136EC6704F164FC36F8EE71B320DA" blockId="10.[169,1496,801,2131]" pageId="10" pageNumber="11">
Dorsal skin similar to that of 
<typeStatus id="54F5884E6704F164FB71F8EE777C203A" box="[1228,1323,1818,1843]" pageId="10" pageNumber="11" type="neotype">neotype</typeStatus>
, with scattered or uniformly distributed granules. Ulnar and tarsal tubercles protuberant or inconspicuous. Three or four, low, rounded tubercles can be present next to the tibio-tarsal articulation; the distal one is elongated in some individuals.
</paragraph>
<paragraph id="8BF136EC6704F165FC36F82E702E21FA" blockId="10.[169,1496,801,2131]" lastBlockId="11.[94,721,1402,2131]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
In freshly euthanized specimens, dorsal color varies from beige or gray to dark brown, with large irregular brown to black blotches, and small white blotches (
<figureCitation id="13752A696704F164FCD1F7CE71FD2F5A" box="[876,938,2106,2131]" captionStart="Figure 6" captionStartId="12.[170,229,2074,2099]" captionTargetBox="[170,1493,204,2044]" captionTargetId="figure-1@12.[170,1493,204,2044]" captionTargetPageId="12" captionText="Figure 6.Variationin the color pattern of freshly euthanized specimens of Scinax x‑signatus.(A), (D), (G),and (I) CFBH44687 (male).(B),(E),and (K) MHNJCH 1701 (male). (C),(F),and (L) MHNJCH 1698 (male),(H) and (J) MZUESC 20683 (male)." figureDoi="http://doi.org/10.5281/zenodo.4614073" httpUri="https://zenodo.org/record/4614073/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
). In males, pectoral region light yellow and abdomen yellowish beige; in females, cream white. Inguinal region light yellow or yellow, with irregular dark blotches; hidden surfaces of thighs, shanks, and tarsi brown to black, with small to large, rounded or irregular yellow blotches. Flanks light yellow or cream white; axillae yellow in some individuals. Iris yellowish golden or bronze, with many thin, dark reticulations, and a median black streak. In life, overall coloration similar to fleshly euthanized specimens. Still, dark and light tones are more intense and brighter, especially yellow coloration on inguinal region and hidden surfaces of hindlimbs (
<figureCitation id="13752A696705F165FDE0F94E70CB21DA" box="[605,668,1722,1747]" captionStart="Figure7" captionStartId="13.[94,153,2074,2099]" captionTargetBox="[94,1417,797,2044]" captionTargetId="figure-301@13.[94,1417,797,2044]" captionTargetPageId="13" captionText="Figure7.LivingspecimensofScinaxx‑signatus.(A)CFBH44687(male).(B)MZUESC15894(female).(C)MZUESC20683(male).(D)MHNJCH1701(male).(E-F)MZUESC 17503 (female).Notice the coloration inlife in the flank,inguinal region,and posterior surfaceof thigh in (F).Photos B,E,and F courtesyof CarlosAugusto S.Costa." figureDoi="http://doi.org/10.5281/zenodo.4614075" httpUri="https://zenodo.org/record/4614075/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
). Iris iridescent yellowish golden or bronze (
<figureCitation id="13752A696705F165FDB1F92E701221FA" box="[524,581,1754,1779]" captionStart="Figure7" captionStartId="13.[94,153,2074,2099]" captionTargetBox="[94,1417,797,2044]" captionTargetId="figure-301@13.[94,1417,797,2044]" captionTargetPageId="13" captionText="Figure7.LivingspecimensofScinaxx‑signatus.(A)CFBH44687(male).(B)MZUESC15894(female).(C)MZUESC20683(male).(D)MHNJCH1701(male).(E-F)MZUESC 17503 (female).Notice the coloration inlife in the flank,inguinal region,and posterior surfaceof thigh in (F).Photos B,E,and F courtesyof CarlosAugusto S.Costa." figureDoi="http://doi.org/10.5281/zenodo.4614075" httpUri="https://zenodo.org/record/4614075/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
A-E).
</paragraph>
<caption id="DF3166646705F165FFE2FAC677DD2242" ID-DOI="http://doi.org/10.5281/zenodo.4614071" ID-Zenodo-Dep="4614071" box="[95,1418,1330,1355]" httpUri="https://zenodo.org/record/4614071/files/figure.png" pageId="11" pageNumber="12" startId="11.[95,154,1330,1355]" targetBox="[94,1417,204,1300]" targetPageId="11">
<paragraph id="8BF136EC6705F165FFE2FAC677DD2242" blockId="11.[95,1418,1330,1355]" box="[95,1418,1330,1355]" pageId="11" pageNumber="12">
<emphasis id="B93AEAFE6705F165FFE2FAC6735E2242" bold="true" box="[95,265,1330,1355]" pageId="11" pageNumber="12">Figure5.Neotype of</emphasis>
<taxonomicName id="4C4E4D6F6705F165FEB6FAC673C32242" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[267,404,1330,1355]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6705F165FEB6FAC673C32242" box="[267,404,1330,1355]" italics="true" pageId="11" pageNumber="12">Scinax x‑signatus</emphasis>
</taxonomicName>
(CFBH 44688,male).Freshly euthanized individual.(A)Dorsal view.(B)Ventral view.(C)Dorsolateral view.Scale bars=10mm.
</paragraph>
</caption>
<paragraph id="8BF136EC6705F165FF39F90E776E21DA" blockId="11.[94,721,1402,2131]" lastBlockId="11.[793,1419,1402,2131]" pageId="11" pageNumber="12">
In preservative, dorsal pattern varies from light beige or light gray to dark brown, with large, irregular dark blotches, usually with X-shaped marks composed of one or two pairs of inverted parentheses-like blotches, with or without light blotches (
<figureCitation id="13752A696705F165FD8BF88E702F209A" box="[566,632,1914,1939]" captionStart="Figure 8" captionStartId="14.[170,229,1978,2003]" captionTargetBox="[236,1427,204,1948]" captionTargetId="figure-1@14.[236,1427,204,1948]" captionTargetPageId="14" captionText="Figure 8. Variation in the dorsal color pattern of preserved Scinax x‑signatus.(A) MHNJCH 1700 (SVL 36.9 mm, male). (B) MHNJCH 1698 (SVL 35.0 mm, male). (C) MHNJCH 1699 (SVL 37.8 mm, male). (D) MHNJCH 1701 (SVL 35.0 mm, male). (E) CFBH 44687 (SVL 37.2 mm, male). (F) MZUESC 20683 (SVL 34.5 mm, male). (G) UFMG 4787 (SVL 38.0 mm, male). (H) CFBH 18797 (SVL 37.0 mm, male). (I) MZUESC 14890 (SVL 35.5 mm, male). (J) MZUESC 14893 (SVL 37.1 mm, male). (K)CFBH21071 (SVL36.8 mm,male).(L)MHNJCH1014 (SVL 38.4mm,male).(M)MZUESC17503 (SVL44.7 mm,female).(N) MZUESC14891 (SVL 38.2mm,female). (O) MZUESC 15894 (SVL 32.9 mm,female).Scale bars= 10mm." figureDoi="http://doi.org/10.5281/zenodo.4614077" httpUri="https://zenodo.org/record/4614077/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
). Small, dark brown, irregular blotches on the dorsum of all individuals. Interocular marking can be inverted triangle, T-shaped, or W-shaped, sometimes fragmented. Upper lip light with diffuse brown blotches, sometimes with a white infraorbital stripe that extends to the tympanum. Dark canthal line present in all specimens. Post-orbital line varies in extension, reaching posteriorly level of forearm insertion or middle of flanks. Dark blotches on flanks and inguinal region rounded or irregular. Small to large, rounded or irregular light blotches on hidden surfaces of hindlimbs. Ventral surfaces from cream white to light beige, finely or conspicuously covered with brown spots in some individuals. Dark coloration predominates on dorsum of specimen MHNJCH 1014 (
<figureCitation id="13752A696705F165FB71F9AE7741217A" box="[1228,1302,1626,1651]" captionStart="Figure 8" captionStartId="14.[170,229,1978,2003]" captionTargetBox="[236,1427,204,1948]" captionTargetId="figure-1@14.[236,1427,204,1948]" captionTargetPageId="14" captionText="Figure 8. Variation in the dorsal color pattern of preserved Scinax x‑signatus.(A) MHNJCH 1700 (SVL 36.9 mm, male). (B) MHNJCH 1698 (SVL 35.0 mm, male). (C) MHNJCH 1699 (SVL 37.8 mm, male). (D) MHNJCH 1701 (SVL 35.0 mm, male). (E) CFBH 44687 (SVL 37.2 mm, male). (F) MZUESC 20683 (SVL 34.5 mm, male). (G) UFMG 4787 (SVL 38.0 mm, male). (H) CFBH 18797 (SVL 37.0 mm, male). (I) MZUESC 14890 (SVL 35.5 mm, male). (J) MZUESC 14893 (SVL 37.1 mm, male). (K)CFBH21071 (SVL36.8 mm,male).(L)MHNJCH1014 (SVL 38.4mm,male).(M)MZUESC17503 (SVL44.7 mm,female).(N) MZUESC14891 (SVL 38.2mm,female). (O) MZUESC 15894 (SVL 32.9 mm,female).Scale bars= 10mm." figureDoi="http://doi.org/10.5281/zenodo.4614077" httpUri="https://zenodo.org/record/4614077/files/figure.png" pageId="11" pageNumber="12">Fig. 8L</figureCitation>
); however, this resulted from the fixation process in 10% formalin. In life, this specimen showed the X-shaped mark and other dorsal blotches common to the other specimens.
</paragraph>
<paragraph id="8BF136EC6705F163FC82F92E73DE265A" blockId="11.[793,1419,1402,2131]" lastBlockId="13.[93,719,218,339]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
The occurrence of glandular tissue in the pectoral region has been considered a secondary sexually dimorphic character occurring in males (
<bibRefCitation id="EFDF4B1D6705F165FB6EF8EE771A203A" author="Lutz, B." box="[1235,1357,1818,1843]" pageId="11" pageNumber="12" refId="ref22759" refString="Lutz, B. 1973. Brazilian Species of Hyla. University of Texas Press, Austin &amp; London. 260 p." type="book" year="1973">Lutz, 1973</bibRefCitation>
), and this is corroborated here (
<figureCitation id="13752A696705F165FB8AF8CE76D1205A" box="[1079,1158,1850,1875]" captionStart="Figure 9" captionStartId="15.[94,153,2042,2067]" captionTargetBox="[142,1370,204,2012]" captionTargetId="figure-1@15.[142,1370,204,2012]" captionTargetPageId="15" captionText="Figure 9. Pectoral glands and spicule-shaped papillary epidermal projections in males of Scinax x‑signatus.(A) Pectoral glands of CFBH 44687;notice the whitish cream glandular area (acini) in the pectoral region. Right hand of MZUESC 14893 and MHNJCH 1014, respectively.(B) Dorsal view. (C) Ventral view.White arrows indicate the spicule-shaped projections on nuptial pads and inner margin of Finger III. Scale bars = 1 mm (upper) and 2 mm (lower)." figureDoi="http://doi.org/10.5281/zenodo.4614079" httpUri="https://zenodo.org/record/4614079/files/figure.png" pageId="11" pageNumber="12">Fig. 9A</figureCitation>
). This is also the case of the spicule-shaped papillary epidermal projections on the nuptial pad (
<figureCitation id="13752A696705F165FC6AF88E7615209A" box="[983,1090,1914,1939]" captionStart="Figure 9" captionStartId="15.[94,153,2042,2067]" captionTargetBox="[142,1370,204,2012]" captionTargetId="figure-1@15.[142,1370,204,2012]" captionTargetPageId="15" captionText="Figure 9. Pectoral glands and spicule-shaped papillary epidermal projections in males of Scinax x‑signatus.(A) Pectoral glands of CFBH 44687;notice the whitish cream glandular area (acini) in the pectoral region. Right hand of MZUESC 14893 and MHNJCH 1014, respectively.(B) Dorsal view. (C) Ventral view.White arrows indicate the spicule-shaped projections on nuptial pads and inner margin of Finger III. Scale bars = 1 mm (upper) and 2 mm (lower)." figureDoi="http://doi.org/10.5281/zenodo.4614079" httpUri="https://zenodo.org/record/4614079/files/figure.png" pageId="11" pageNumber="12">Fig. 9B, C</figureCitation>
), inner margin of upper- and forearms, and pectoral region. The glandular areas (acini) on the inner margins of upper- and forearms, and fingers II-III (excluding the nuptial pad) are absent in some specimens (
<emphasis id="B93AEAFE6705F165FC22F80E719D2F1A" box="[927,970,2042,2067]" italics="true" pageId="11" pageNumber="12">e.g.,</emphasis>
MHNJCH 1014, 1698-1700); when present, spicule-shaped papillary epidermal projections also occur on these areas. Spicule-shaped projections can be present, scattered, and apparently not associated to acini on fingers II-V in some individuals. Although our sample of females is small (three individuals), females tend to be larger than males (
<tableCitation id="C6CC03576703F163FE97FECE732D265A" box="[298,378,314,339]" captionStart="Table 1" captionStartId="1.[96,146,1594,1619]" captionTargetPageId="1" captionText="Table 1. Measurements (in mm) of Scinax x‑signatus (including the neotype).See Materialsand methods for the abbreviations of measurements." httpUri="http://table.plazi.org/id/DF316664670FF16FFFDDF9CE70E1217A" pageId="13" pageNumber="14" tableUuid="DF316664670FF16FFFDDF9CE70E1217A">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</subSection>
<caption id="DF3166646702F162FF17F7EE71622F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614073" ID-Zenodo-Dep="4614073" httpUri="https://zenodo.org/record/4614073/files/figure.png" pageId="12" pageNumber="13" startId="12.[170,229,2074,2099]" targetBox="[170,1493,204,2044]" targetPageId="12">
<paragraph id="8BF136EC6702F162FF17F7EE71622F5A" blockId="12.[170,1493,2074,2131]" pageId="12" pageNumber="13">
<emphasis id="B93AEAFE6702F162FF17F7EE71532F3A" bold="true" box="[170,772,2074,2099]" pageId="12" pageNumber="13">Figure 6.Variation in the color pattern of freshly euthanized specimens of</emphasis>
<emphasis id="B93AEAFE6702F162FCB5F7EE71CF2F3A" box="[776,920,2074,2099]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C4E4D6F6702F162FCB5F7EE71C22F3A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[776,917,2074,2099]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="x-signatus">Scinax x‑signatus</taxonomicName>
.
</emphasis>
(A), (D), (G),and (I) CFBH44687 (male).(B),(E),and (K) MHNJCH 1701 (male). (C),(F),and (L) MHNJCH 1698 (male),(H) and (J) MZUESC 20683 (male).
</paragraph>
</caption>
<subSection id="E2C12F076703F17FFFE3FE8E70FC267A" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="multiple">
<subSubSection id="C35465676703F163FFE3FE8E736A269A" box="[94,317,378,403]" pageId="13" pageNumber="14" type="multiple">
<paragraph id="8BF136EC6703F163FFE3FE8E736A269A" blockId="13.[94,317,378,403]" box="[94,317,378,403]" pageId="13" pageNumber="14">
<heading id="D0B981806703F163FFE3FE8E736A269A" box="[94,317,378,403]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="14" reason="5">
<emphasis id="B93AEAFE6703F163FFE3FE8E736A269A" bold="true" box="[94,317,378,403]" pageId="13" pageNumber="14">Advertisement call</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35465676703F17FFF39FE4E70FC267A" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="description">
<paragraph id="8BF136EC6703F163FF39FE4E707825FA" blockId="13.[93,720,442,755]" pageId="13" pageNumber="14">
The advertisement call of 
<taxonomicName id="4C4E4D6F6703F163FE23FE4E700226DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[414,597,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE6703F163FE23FE4E700226DA" box="[414,597,442,467]" italics="true" pageId="13" pageNumber="14">Scinax x‑signatus</emphasis>
</taxonomicName>
consists of a single multipulsed note,emitted at a highly variable repetition rate (2-105 notes/minute; 
<tableCitation id="C6CC03576703F163FE75FE0E704F251A" box="[456,536,506,531]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
; 
<figureCitation id="13752A696703F163FD9FFE0E702B251A" box="[546,636,506,531]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10A</figureCitation>
), which is likely affected by conspecific chorus density, since the longer intervals between notes (up to 
<geoCoordinate id="EE7A502B6703F163FE46FDCE706C255A" box="[507,571,570,595]" degrees="25.3" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="5555" value="-25.3">25.3 s</geoCoordinate>
; 
<tableCitation id="C6CC03576703F163FDFCFDCE70D9255A" box="[577,654,570,595]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
) were observed in the recording of the 
<typeStatus id="54F5884E6703F163FE7FFDAE7077257A" box="[450,544,602,627]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
, which was calling alone with no nearby conspecific. Notwithstanding, much longer intervals are the exception, and the notes are repeated at faster rates (29-105 notes/minute; 
<tableCitation id="C6CC03576703F163FDC8FD4E709325DA" box="[629,708,698,723]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
), but never composing a stereotyped series.
</paragraph>
<paragraph id="8BF136EC6703F163FC82FF2E71F726BA" blockId="13.[793,1419,218,755]" pageId="13" pageNumber="14">
Note duration is 0.11- 
<geoCoordinate id="EE7A502B6703F163FB90FF2E763C27FA" box="[1069,1131,218,243]" degrees="0.25" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="555" value="-0.25">0.25 s</geoCoordinate>
(
<tableCitation id="C6CC03576703F163FBCBFF2E769D27FA" box="[1142,1226,218,243]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
; 
<figureCitation id="13752A696703F163FB6FFF2E774F27FA" box="[1234,1304,218,243]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
A-C); each note is composed of 6-14 pulses with modulating amplitude increasing from the first third, reaching the maximum amplitude around the middle of the note,and gradually decaying towards the last pulse (
<figureCitation id="13752A696703F163FB7BFEAE7769267A" box="[1222,1342,346,371]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C</figureCitation>
). Pulse rate is 52-64 pulses/s and pulse duration 
<geoCoordinate id="EE7A502B6703F163FB4BFE8E7765269A" box="[1270,1330,378,403]" degrees="0.013" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="0.013">0.013</geoCoordinate>
<geoCoordinate id="EE7A502B6703F163FA8EFE8E7720269A" box="[1331,1399,378,403]" degrees="0.018" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="-0.018">
- 
<geoCoordinate id="EE7A502B6703F163FA86FE8E7720269A" box="[1339,1399,378,403]" degrees="0.018" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="-0.018">0.018</geoCoordinate>
</geoCoordinate>
s (
<figureCitation id="13752A696703F163FC9DFE6E71C226BA" box="[800,917,410,435]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C</figureCitation>
).
</paragraph>
<paragraph id="8BF136EC6703F17FFC82FE4E70FC267A" blockId="13.[793,1419,218,755]" lastBlockId="17.[94,719,218,371]" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
Calls have a broadband spectrum (BW90% 2250-2799 Hz; 
<tableCitation id="C6CC03576703F163FC02FE2E764526FA" box="[959,1042,474,499]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
; 
<figureCitation id="13752A696703F163FBA1FE2E762E26FA" box="[1052,1145,474,499]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10C</figureCitation>
). The power spectrum is usually biphasic, with two main emphasized frequency bands (
<figureCitation id="13752A696703F163FCD0FDEE719A253A" box="[877,973,538,563]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10C</figureCitation>
). The lower band (
<emphasis id="B93AEAFE6703F163FB1DFDEE7694253A" box="[1184,1219,538,563]" italics="true" pageId="13" pageNumber="14">i.e.,</emphasis>
the low-frequency band) comprises most of the power of the spectrum, with the dominant frequency ranging between 904-1359 Hz (
<figureCitation id="13752A696703F163FC04FD8E764D259A" box="[953,1050,634,659]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="13" pageNumber="14">Fig. 10C</figureCitation>
), and also including part of the freq5% (861-991 Hz; 
<tableCitation id="C6CC03576703F163FBB5FD6E760B25BA" box="[1032,1116,666,691]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
). The upper band (
<emphasis id="B93AEAFE6703F163FA85FD6E770C25BA" box="[1336,1371,666,691]" italics="true" pageId="13" pageNumber="14">i.e.,</emphasis>
the high-frequency band) has less power than the first one, surrounding the freq95% (3188-3704 Hz; 
<tableCitation id="C6CC03576703F163FB4FFD2E771125FA" box="[1266,1350,730,755]" captionStart="Table2" captionStartId="2.[93,143,510,535]" captionTargetPageId="2" captionText="Table2.Parametersof the advertisementcall ofScinax x‑signatusfrom four localitiesinthe State ofBahia,Brazil.Values areexpressedas minimum-maximum(mean ± SD).Abbreviations:N= numberof notes/numberof intervals/number of pulses;Freq = Frequency;SD = standard deviation;and SVL = snout-vent length.*Data reported by Novaes &amp; Zina (2016)." httpUri="http://table.plazi.org/id/DF316664670CF16CFFE0FE0A760F253E" pageId="13" pageNumber="14" tableUuid="DF316664670CF16CFFE0FE0A760F253E">Table 2</tableCitation>
), with its peak frequency between 2885-3618 Hz (
<figureCitation id="13752A69671FF17FFDDFFF2E709227FA" box="[610,709,218,243]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="17" pageNumber="18">Fig. 10C</figureCitation>
). Between the two power spectrum bands, there is a low-power“valley” (or gap) around 2.0-2.5 kHz (
<figureCitation id="13752A69671FF17FFDD5FEEE7092263A" box="[616,709,282,307]" captionStart="Figure 10" captionStartId="16.[170,229,1978,2003]" captionTargetBox="[220,1442,208,1944]" captionTargetId="figure-1@16.[217,1445,204,1948]" captionTargetPageId="16" captionText="Figure 10. Advertisement call of the neotype of Scinax x‑signatus (CFBH 44688,male).(A) Waveformshowing seven notes;the dashed squarehighlights two notes showedin (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlightsthe note showedin (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the powerspectrum highlights the dominant frequency inthe low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right)." figureDoi="http://doi.org/10.5281/zenodo.4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="17" pageNumber="18">Fig. 10C</figureCitation>
). The dominant frequency does not alternate between the lower and upper bands, remaining in the lower band.
</paragraph>
</subSubSection>
</subSection>
<caption id="DF3166646703F163FFE3F7EE77292F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614075" ID-Zenodo-Dep="4614075" httpUri="https://zenodo.org/record/4614075/files/figure.png" pageId="13" pageNumber="14" startId="13.[94,153,2074,2099]" targetBox="[94,1417,797,2044]" targetPageId="13">
<paragraph id="8BF136EC6703F163FFE3F7EE77292F5A" blockId="13.[93,1417,2074,2131]" pageId="13" pageNumber="14">
<emphasis id="B93AEAFE6703F163FFE3F7EE731B2F3A" bold="true" box="[94,332,2074,2099]" pageId="13" pageNumber="14">Figure7.Living specimens of</emphasis>
<emphasis id="B93AEAFE6703F163FEF0F7EE738B2F3A" box="[333,476,2074,2099]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C4E4D6F6703F163FEF0F7EE738F2F3A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[333,472,2074,2099]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="x-signatus">Scinax x‑signatus</taxonomicName>
.
</emphasis>
(A)CFBH44687(male).(B)MZUESC15894(female).(C)MZUESC20683(male).(D)MHNJCH1701(male).(E-F)MZUESC 17503 (female).Notice the coloration in life in the flank,inguinal region,and posterior surface of thigh in (F).Photos B,E,and F courtesy of Carlos Augusto S.Costa.
</paragraph>
</caption>
<caption id="DF3166646700F160FF17F84E70E52F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614077" ID-Zenodo-Dep="4614077" httpUri="https://zenodo.org/record/4614077/files/figure.png" pageId="14" pageNumber="15" startId="14.[170,229,1978,2003]" targetBox="[236,1427,204,1948]" targetPageId="14">
<paragraph id="8BF136EC6700F160FF17F84E70E52F5A" blockId="14.[169,1496,1978,2131]" pageId="14" pageNumber="15">
<emphasis id="B93AEAFE6700F160FF17F84E70C220DA" bold="true" box="[170,661,1978,2003]" pageId="14" pageNumber="15">Figure 8. Variation in the dorsal color pattern of preserved</emphasis>
<emphasis id="B93AEAFE6700F160FD20F84E716720DA" box="[669,816,1978,2003]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C4E4D6F6700F160FD20F84E717B20DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[669,812,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="x-signatus">Scinax x‑signatus</taxonomicName>
.
</emphasis>
(A) MHNJCH 1700 (SVL 36.9 mm, male). (B) MHNJCH 1698 (SVL 35.0 mm, male). (C) MHNJCH 1699 (SVL 37.8 mm, male). (D) MHNJCH 1701 (SVL 35.0 mm, male). (E) CFBH 44687 (SVL 37.2 mm, male). (F) MZUESC 20683 (SVL 34.5 mm, male). (G) UFMG 4787 (SVL 38.0 mm, male). (H) CFBH 18797 (SVL 37.0 mm, male). (I) MZUESC 14890 (SVL 35.5 mm, male). (J) MZUESC 14893 (SVL 37.1 mm, male). (K)CFBH21071 (SVL36.8 mm,male).(L)MHNJCH1014 (SVL 38.4mm,male).(M)MZUESC17503 (SVL44.7 mm,female).(N) MZUESC14891 (SVL 38.2mm,female). (O) MZUESC 15894 (SVL 32.9 mm,female).Scale bars= 10mm.
</paragraph>
</caption>
<caption id="DF3166646701F161FFE3F80E76C22F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614079" ID-Zenodo-Dep="4614079" httpUri="https://zenodo.org/record/4614079/files/figure.png" pageId="15" pageNumber="16" startId="15.[94,153,2042,2067]" targetBox="[142,1370,204,2012]" targetPageId="15">
<paragraph id="8BF136EC6701F161FFE3F80E76C22F5A" blockId="15.[94,1417,2042,2131]" pageId="15" pageNumber="16">
<emphasis id="B93AEAFE6701F161FFE3F80E716D2F1A" bold="true" box="[94,826,2042,2067]" pageId="15" pageNumber="16">Figure 9. Pectoral glands and spicule-shaped papillary epidermal projections in males of</emphasis>
<emphasis id="B93AEAFE6701F161FCFDF80E71862F1A" box="[832,977,2042,2067]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D6F6701F161FCFDF80E719A2F1A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[832,973,2042,2067]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="x-signatus">Scinax x‑signatus</taxonomicName>
.
</emphasis>
(A) Pectoral glands of CFBH 44687;notice the whitish cream glandular area (acini) in the pectoral region. Right hand of MZUESC 14893 and MHNJCH 1014, respectively.(B) Dorsal view. (C) Ventral view.White arrows indicate the spicule-shaped projections on nuptial pads and inner margin of Finger III. Scale bars = 1 mm (upper) and 2 mm (lower).
</paragraph>
</caption>
<caption id="DF316664671EF17EFF17F84E70002F5A" ID-DOI="http://doi.org/10.5281/zenodo.4614083" ID-Zenodo-Dep="4614083" httpUri="https://zenodo.org/record/4614083/files/figure.png" pageId="16" pageNumber="17" startId="16.[170,229,1978,2003]" targetBox="[220,1442,208,1944]" targetPageId="16">
<paragraph id="8BF136EC671EF17EFF17F84E70002F5A" blockId="16.[170,1493,1978,2131]" pageId="16" pageNumber="17">
<emphasis id="B93AEAFE671EF17EFF17F84E706620DA" bold="true" box="[170,561,1978,2003]" pageId="16" pageNumber="17">Figure 10. Advertisement call of the neotype of</emphasis>
<taxonomicName id="4C4E4D6F671EF17EFD88F84E709720DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[565,704,1978,2003]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671EF17EFD88F84E709720DA" box="[565,704,1978,2003]" italics="true" pageId="16" pageNumber="17">Scinax x‑signatus</emphasis>
</taxonomicName>
(CFBH 44688,male).(A) Waveform showing seven notes;the dashed square highlights two notes showed in (B). (B)Waveform (on top) and spectrogram (on bottom) showing two consecutive notes;dashed square highlights the note showed in (C).(C)Waveform (on top), spectrogram (in the middle), and power spectrum (on bottom) of a single note; notice the amplitude modulation in the intermediate pulses,with four crescent peaks;the arrows and dashed vertical lines in the power spectrum highlights the dominant frequency in the low-frequency band(on the left) and the peak frequency of the high-frequency band (on the right).
</paragraph>
</caption>
<subSection id="E2C12F07671FF17FFFE3FE6E72AF247A" pageId="17" pageNumber="18" type="multiple">
<subSubSection id="C3546567671FF17FFFE3FE6E701326BA" box="[94,580,410,435]" pageId="17" pageNumber="18" type="multiple">
<paragraph id="8BF136EC671FF17FFFE3FE6E701326BA" blockId="17.[94,580,410,435]" box="[94,580,410,435]" pageId="17" pageNumber="18">
<heading id="D0B98180671FF17FFFE3FE6E701326BA" box="[94,580,410,435]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="5">
<emphasis id="B93AEAFE671FF17FFFE3FE6E701326BA" bold="true" box="[94,580,410,435]" pageId="17" pageNumber="18">Notes on calling site and calling behavior</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3546567671FF17FFF39FE2E72AF247A" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BF136EC671FF17FFF39FE2E72AF247A" blockId="17.[94,720,474,883]" pageId="17" pageNumber="18">
Males of 
<taxonomicName id="4C4E4D6F671FF17FFF57FE2E73F326FA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[234,420,474,499]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFF57FE2E73F326FA" box="[234,420,474,499]" italics="true" pageId="17" pageNumber="18">Scinax x‑signatus</emphasis>
</taxonomicName>
call near lentic water bodies, either natural or artificial (such as pools and tanks). They usually call from the ground, either uncovered or hidden among the vegetation. Less often, they call perched at low heights (below 1.5 m; rarely above that height) on the vegetation inside or at the margins of water bodies. Other species of 
<taxonomicName id="4C4E4D6F671FF17FFE28FD6E738C25BA" authorityName="Wagler" authorityYear="1830" box="[405,475,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE671FF17FFE28FD6E738C25BA" box="[405,475,666,691]" italics="true" pageId="17" pageNumber="18">Scinax</emphasis>
</taxonomicName>
found calling syntopically with 
<taxonomicName id="4C4E4D6F671FF17FFF68FD4E730D25DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[213,346,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFF68FD4E730D25DA" box="[213,346,698,723]" italics="true" pageId="17" pageNumber="18">S. x‑signatus</emphasis>
</taxonomicName>
are 
<taxonomicName id="4C4E4D6F671FF17FFE34FD4E739825DA" authorityName="Wagler" authorityYear="1830" box="[393,463,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE671FF17FFE34FD4E739825DA" box="[393,463,698,723]" italics="true" pageId="17" pageNumber="18">Scinax</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C4E4D6F671FF17FFE44FD4E700025DA" baseAuthorityName="Barbour" baseAuthorityYear="1909" box="[505,599,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hayii">
aff. 
<emphasis id="B93AEAFE671FF17FFD9CFD4E700025DA" box="[545,599,698,723]" italics="true" pageId="17" pageNumber="18">hayii</emphasis>
</taxonomicName>
<emphasis id="B93AEAFE671FF17FFDEBFD4E728025FA" italics="true" pageId="17" pageNumber="18">
, 
<taxonomicName id="4C4E4D6F671FF17FFDDDFD4E709C25DA" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[608,715,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="auratus">S. auratus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F671FF17FFFE2FD2E728425FA" baseAuthorityName="Bokermann" baseAuthorityYear="1968" box="[95,211,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="eurydice">S. eurydice</taxonomicName>
,
</emphasis>
and 
<emphasis id="B93AEAFE671FF17FFEAFFD2E703725FA" box="[274,608,730,755]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C4E4D6F671FF17FFEAFFD2E73CD25FA" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1937" box="[274,410,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pachycrus">S. pachycrus</taxonomicName>
. 
<taxonomicName id="4C4E4D6F671FF17FFE19FD2E703725FA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[420,608,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">Scinax x‑signatus</taxonomicName>
</emphasis>
seems to tolerate some degree of light and sound disturbance; as we recorded the 
<typeStatus id="54F5884E671FF17FFEA2FCEE732B243A" box="[287,380,794,819]" pageId="17" pageNumber="18" type="neotype">neotype</typeStatus>
inside the University Campus, with plenty of artificial light and crowd voices as background noise.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C12F07671FF17CFEABFC4E7028213A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="multiple">
<subSubSection id="C3546567671FF17FFEABFC4E704124DA" box="[278,534,954,979]" pageId="17" pageNumber="18" type="multiple">
<paragraph id="8BF136EC671FF17FFEABFC4E704124DA" blockId="17.[278,534,954,979]" box="[278,534,954,979]" pageId="17" pageNumber="18">
<heading id="D0B98180671FF17FFEABFC4E704124DA" box="[278,534,954,979]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="5">
<emphasis id="B93AEAFE671FF17FFEABFC4E704124DA" bold="true" box="[278,534,954,979]" pageId="17" pageNumber="18">Phylogenetic analysis</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3546567671FF17CFF39FC0E7028213A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BF136EC671FF17FFF39FC0E7657259A" blockId="17.[94,719,1018,1075]" lastBlockId="17.[793,1420,218,1075]" pageId="17" pageNumber="18">
The two most parsimonious trees (length 468) recovered all specimens considered in the literature as 
<taxonomicName id="4C4E4D6F671FF17FFD34FBEE71DF27FA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[649,904,218,1075]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFD34FBEE71DF27FA" box="[649,904,218,1075]" italics="true" pageId="17" pageNumber="18">Scinax x‑signatus</emphasis>
</taxonomicName>
closely related with the 
<typeStatus id="54F5884E671FF17FFB1EFF2E775727FA" box="[1187,1280,218,243]" pageId="17" pageNumber="18" type="neotype">neotype</typeStatus>
and our referred specimens (100% jackknife; 
<figureCitation id="13752A69671FF17FFB2AFF0E76B7261A" box="[1175,1248,250,275]" captionStart="Figure 11" captionStartId="17.[95,154,2042,2067]" captionTargetBox="[97,1416,1090,2012]" captionTargetId="figure-432@17.[94,1417,1086,2012]" captionTargetPageId="17" captionText="Figure 11.Oneofthe two most parsimonioustrees(length468 steps)obtained from the analysisof the 16S rRNAmitochondrial geneinTNTwithgaps as fifthstate. Values above or below nodes are jackknife support values. Nodes without values indicate &lt;50% jackknife support;black dot indicates a node that collapses in the strict consensus;anasterisk (*) indicates groups with 100% jackknife support." figureDoi="http://doi.org/10.5281/zenodo.4614086" httpUri="https://zenodo.org/record/4614086/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
). These include specimens from 
<collectingRegion id="498AF80E671FF17FFC59FEEE7673263A" box="[996,1060,282,307]" country="Brazil" name="Bahia" pageId="17" pageNumber="18">Bahia</collectingRegion>
, 
<collectingRegion id="498AF80E671FF17FFB8AFEEE762E263A" box="[1079,1145,282,307]" country="Brazil" name="Ceara" pageId="17" pageNumber="18">Ceará</collectingRegion>
, and 
<collectingRegion id="498AF80E671FF17FFB78FEEE7704263A" box="[1221,1363,282,307]" country="Brazil" name="Pernambuco" pageId="17" pageNumber="18">Pernambuco</collectingRegion>
(NE 
<collectingCountry id="F359767C671FF17FFCA7FECE710F265A" box="[794,856,314,339]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
) and 
<collectingRegion id="498AF80E671FF17FFC2CFECE71B7265A" box="[913,992,314,339]" country="Brazil" name="Amapa" pageId="17" pageNumber="18">Amapá</collectingRegion>
(N 
<collectingCountry id="F359767C671FF17FFBBEFECE7614265A" box="[1027,1091,314,339]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
),and from 
<collectingCountry id="F359767C671FF17FFB06FECE770D265A" box="[1211,1370,314,339]" name="French Guiana" pageId="17" pageNumber="18">French Guiana</collectingCountry>
and 
<collectingCountry id="F359767C671FF17FFCA4FEAE71C3267A" box="[793,916,346,371]" name="Martinique" pageId="17" pageNumber="18">Martinique</collectingCountry>
(
<figureCitation id="13752A69671FF17FFC1FFEAE71B9267A" box="[930,1006,346,371]" captionStart="Figure 12" captionStartId="18.[170,229,1103,1128]" captionTargetBox="[173,1490,208,1072]" captionTargetId="figure-327@18.[170,1493,204,1073]" captionTargetPageId="18" captionText="Figure 12. Distribution map of Scinax x‑signatusshowing localities of specimens with sequences of the 16S rRNA mitochondrial gene included in this study.Gray shadow on the lefthighlights the putativedistribution ofS.x‑signatus taken from IUCNRed ListofThreatened Species (Rodrigues et al., 2010).Note that althoughit includes Paraguay,Bolivia, and Peru,we are not aware of published records assigned to S.x‑signatus from thesecountries.The dashed square on the left highlights the area on the right showing the distribution of S.x‑signatus in the State of Bahia, Brazil.Black triangle:Januária (Scinax sp.).Red star:Ilhéus (neotype of S.x‑signatus). Red dots.1 = Potiraguá.2 = Una.3 = Sebastião Laranjeiras.4 = Caetité.5 = Maracás.6 = Irajuba.7 = Morro do Chapéu.8 = Parque Nacional doCatimbau. 9 = Sanharó.10 = Fernando de Noronha.11 = Ubajara.12 = Macapá.13 = Montravel.14 = Korou.15 = Ile Royale.16= Diamant.17 = Unknown.Abbreviations. Countries: BRA = Brazil. BOL = Bolivia. COL = Colombia.GUF = French Guiana. GUY = Guyana. PER = Peru. PRY = Paraguay. SUR = Surinam.VEN = Venezuela. Brazilian states: AC = Acre.AL = Alagoas.AM = Amazonas.AP = Amapá.BA = Bahia.CE = Ceará. ES = Espírito Santo.GO = Goiás.MA = Maranhão.MG = Minas Gerais.MS = Mato Grosso do Sul.MT = Mato Grosso.PA = Pará.PB = Paraíba.PE = Pernambuco.PI = Piauí.PR= Paraná.RJ = Rio de Janeiro.RN = Rio Grande do Norte.RO = Rondônia.RR = Roraima.RS= Rio Grande do Sul.SC= SantaCatarina.SE = Sergipe.SP = São Paulo.TO =Tocantins." figureDoi="http://doi.org/10.5281/zenodo.4614088" httpUri="https://zenodo.org/record/4614088/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
). The selected fragment of the mitochondrial ribosomal gene 16S rRNA showed uncorrected pairwise distances of 0.2-1.7% among the 28 individuals of 
<taxonomicName id="4C4E4D6F671FF17FFC84FE4E719626DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[825,961,442,467]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFC84FE4E719626DA" box="[825,961,442,467]" italics="true" pageId="17" pageNumber="18">S. x‑signatus</emphasis>
</taxonomicName>
(
<tableCitation id="C6CC0357671FF17FFC6DFE4E767026DA" box="[976,1063,442,467]" captionStart="Table3" captionStartId="19.[93,143,507,532]" captionTargetPageId="19" captionText="Table3. Uncorrectedpairwise distances amongsequences of the mitochondrial 16S rRNAgene ofScinax x‑signatus(including the neotype)from localities in the Brazilian statesof Amapá (AP),Bahia (BA),Ceará (CE),and Pernambuco(PE), and alsofrom French Guiana (GUF) and Martinique (MTQ);plus one related lineage from Minas Gerais (MG),Brazil (BRA).Values are given as percentage.The neotype is underlined.See Material and Methods and Appendix 2 for details." httpUri="http://table.plazi.org/id/DF316664671DF17DFFE0FE0F7593253D" pageId="17" pageNumber="18" tableUuid="DF316664671DF17DFFE0FE0F7593253D">Table 3</tableCitation>
). The maximum value (1.7%) is between specimens from Kourou and Ile Royale (
<collectingCountry id="F359767C671FF17FFA81FE2E7139251A" name="French Guiana" pageId="17" pageNumber="18">French Guiana</collectingCountry>
), and those from Ubajara (
<collectingRegion id="498AF80E671FF17FFB03FE0E7756251A" box="[1214,1281,506,531]" country="Brazil" name="Ceara" pageId="17" pageNumber="18">Ceará</collectingRegion>
, NE 
<collectingCountry id="F359767C671FF17FFAFCFE0E77D6251A" box="[1345,1409,506,531]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
), Fernando de Noronha, and Sanharó (
<collectingRegion id="498AF80E671FF17FFB76FDEE770A253A" box="[1227,1373,538,563]" country="Brazil" name="Pernambuco" pageId="17" pageNumber="18">Pernambuco</collectingRegion>
, NE 
<collectingCountry id="F359767C671FF17FFCA7FDCE710D255A" box="[794,858,570,595]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
); the geographic distances between these points are approx. 
<quantity id="4CB69B09671FF17FFC23FDAE7652257A" box="[926,1029,602,627]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="km" value="1.7">1,700 km</quantity>
(see distances between points 8-9 and 
<quantity id="4CB69B09671FF17FFCF6FD8E71F2259A" box="[843,933,634,659]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="17" pageNumber="18" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">14-15 in</quantity>
<figureCitation id="13752A69671FF17FFC16FD8E71A2259A" box="[939,1013,634,659]" captionStart="Figure 12" captionStartId="18.[170,229,1103,1128]" captionTargetBox="[173,1490,208,1072]" captionTargetId="figure-327@18.[170,1493,204,1073]" captionTargetPageId="18" captionText="Figure 12. Distribution map of Scinax x‑signatusshowing localities of specimens with sequences of the 16S rRNA mitochondrial gene included in this study.Gray shadow on the lefthighlights the putativedistribution ofS.x‑signatus taken from IUCNRed ListofThreatened Species (Rodrigues et al., 2010).Note that althoughit includes Paraguay,Bolivia, and Peru,we are not aware of published records assigned to S.x‑signatus from thesecountries.The dashed square on the left highlights the area on the right showing the distribution of S.x‑signatus in the State of Bahia, Brazil.Black triangle:Januária (Scinax sp.).Red star:Ilhéus (neotype of S.x‑signatus). Red dots.1 = Potiraguá.2 = Una.3 = Sebastião Laranjeiras.4 = Caetité.5 = Maracás.6 = Irajuba.7 = Morro do Chapéu.8 = Parque Nacional doCatimbau. 9 = Sanharó.10 = Fernando de Noronha.11 = Ubajara.12 = Macapá.13 = Montravel.14 = Korou.15 = Ile Royale.16= Diamant.17 = Unknown.Abbreviations. Countries: BRA = Brazil. BOL = Bolivia. COL = Colombia.GUF = French Guiana. GUY = Guyana. PER = Peru. PRY = Paraguay. SUR = Surinam.VEN = Venezuela. Brazilian states: AC = Acre.AL = Alagoas.AM = Amazonas.AP = Amapá.BA = Bahia.CE = Ceará. ES = Espírito Santo.GO = Goiás.MA = Maranhão.MG = Minas Gerais.MS = Mato Grosso do Sul.MT = Mato Grosso.PA = Pará.PB = Paraíba.PE = Pernambuco.PI = Piauí.PR= Paraná.RJ = Rio de Janeiro.RN = Rio Grande do Norte.RO = Rondônia.RR = Roraima.RS= Rio Grande do Sul.SC= SantaCatarina.SE = Sergipe.SP = São Paulo.TO =Tocantins." figureDoi="http://doi.org/10.5281/zenodo.4614088" httpUri="https://zenodo.org/record/4614088/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BF136EC671FF17CFC82FD6E7028213A" blockId="17.[793,1420,218,1075]" lastBlockId="18.[170,795,1466,1587]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<taxonomicName id="4C4E4D6F671FF17FFC82FD6E765425BA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[831,1027,666,691]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFC82FD6E765425BA" box="[831,1027,666,691]" italics="true" pageId="17" pageNumber="18">Scinax x‑signatus</emphasis>
</taxonomicName>
is moderately supported (85% jackknife) as sister taxon of 
<taxonomicName id="4C4E4D6F671FF17FFBE3FD4E76F325DA" authorityName="Wagler" authorityYear="1830" box="[1118,1188,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE671FF17FFBE3FD4E76F325DA" box="[1118,1188,698,723]" italics="true" pageId="17" pageNumber="18">Scinax</emphasis>
</taxonomicName>
sp. (as 
<taxonomicName id="4C4E4D6F671FF17FFABDFD4E77DE25DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1280,1417,698,723]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFABDFD4E77DE25DA" box="[1280,1417,698,723]" italics="true" pageId="17" pageNumber="18">S. x‑signatus</emphasis>
</taxonomicName>
“ 
<taxonomicName id="4C4E4D6F671FF17FFC9CFD2E713D25FA" authorityName="Wagler" authorityYear="1830" box="[801,874,730,755]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">Scinax</taxonomicName>
_64” in 
<bibRefCitation id="EFDF4B1D671FF17FFC00FD2E76DD25FA" author="Vacher, J. - P." box="[957,1162,730,755]" pageId="17" pageNumber="18" pagination="1 - 11" refId="ref25345" refString="Vacher, J. - P.; Chave, J.; Ficetola, F.; Sommeria-Klein, G.; Tao, S.; Thebaud, C.; Blanc, M.; Camacho, A.; Cassimiro, J.; Colston, T. J.; Dewynter, M.; Ernst, R.; Gaucher, P.; Gomes, J. O.; Jairam, R.; Kok, P. J. R.; Dias Lima, J.; Martinez, Q.; Marty, C.; Noonan, B. P.; Nunes, P. M. S.; Ouboter P.; Recoder, R.; Rodrigues, M. T.; Snyder, A.; Souza, S. M. &amp; Fouquet, A. 2020. Large-scale DNA-based survey of frogs in Amazonia suggests a vast underestimation of species richness and endemism. Journal of Biogeography, 2020, 00: 1 - 11." type="journal article" year="2020">
Vacher 
<emphasis id="B93AEAFE671FF17FFBACFD2E761C25FA" box="[1041,1099,730,755]" italics="true" pageId="17" pageNumber="18">et al.,</emphasis>
2020
</bibRefCitation>
) from Parque Nacional Cavernas do Peruaçu, Januária, N 
<collectingRegion id="498AF80E671FF17FFB15FD0E776A241A" box="[1192,1341,762,787]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="18">Minas Gerais</collectingRegion>
, 
<collectingCountry id="F359767C671FF17FFAF7FD0E77DF241A" box="[1354,1416,762,787]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
. Uncorrected pairwise distances between 
<taxonomicName id="4C4E4D6F671FF17FFB43FCEE77DE243A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1278,1417,794,819]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFB43FCEE77DE243A" box="[1278,1417,794,819]" italics="true" pageId="17" pageNumber="18">S. x‑signatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D6F671FF17FFCF4FCCE71D8245A" authorityName="Wagler" authorityYear="1830" box="[841,911,826,851]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE671FF17FFCF4FCCE71D8245A" box="[841,911,826,851]" italics="true" pageId="17" pageNumber="18">Scinax</emphasis>
</taxonomicName>
sp.are 6.9-10.2%, with a sequence divergence of 8.0% between one specimen of 
<taxonomicName id="4C4E4D6F671FF17FFB11FCAE7763247A" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[1196,1332,858,883]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="x-signatus">
<emphasis id="B93AEAFE671FF17FFB11FCAE7763247A" box="[1196,1332,858,883]" italics="true" pageId="17" pageNumber="18">S. x‑signatus</emphasis>
</taxonomicName>
(UFMG 4787) from Sebastião Laranjeiras (
<collectingRegion id="498AF80E671FF17FFB24FC8E768C249A" box="[1177,1243,890,915]" country="Brazil" name="Bahia" pageId="17" pageNumber="18">Bahia</collectingRegion>
, NE 
<collectingCountry id="F359767C671FF17FFAB0FC8E771C249A" box="[1293,1355,890,915]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
) only distant approx. 
<quantity id="4CB69B09671FF17FFC7AFC6E764E24BA" box="[967,1049,922,947]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="km" value="170.0">170 km</quantity>
ENE from the locality of this candidate species in N 
<collectingRegion id="498AF80E671FF17FFC52FC4E762B24DA" box="[1007,1148,954,979]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="18">Minas Gerais</collectingRegion>
(see 
<figureCitation id="13752A69671FF17FFB09FC4E76A824DA" box="[1204,1279,954,979]" captionStart="Figure 12" captionStartId="18.[170,229,1103,1128]" captionTargetBox="[173,1490,208,1072]" captionTargetId="figure-327@18.[170,1493,204,1073]" captionTargetPageId="18" captionText="Figure 12. Distribution map of Scinax x‑signatusshowing localities of specimens with sequences of the 16S rRNA mitochondrial gene included in this study.Gray shadow on the lefthighlights the putativedistribution ofS.x‑signatus taken from IUCNRed ListofThreatened Species (Rodrigues et al., 2010).Note that althoughit includes Paraguay,Bolivia, and Peru,we are not aware of published records assigned to S.x‑signatus from thesecountries.The dashed square on the left highlights the area on the right showing the distribution of S.x‑signatus in the State of Bahia, Brazil.Black triangle:Januária (Scinax sp.).Red star:Ilhéus (neotype of S.x‑signatus). Red dots.1 = Potiraguá.2 = Una.3 = Sebastião Laranjeiras.4 = Caetité.5 = Maracás.6 = Irajuba.7 = Morro do Chapéu.8 = Parque Nacional doCatimbau. 9 = Sanharó.10 = Fernando de Noronha.11 = Ubajara.12 = Macapá.13 = Montravel.14 = Korou.15 = Ile Royale.16= Diamant.17 = Unknown.Abbreviations. Countries: BRA = Brazil. BOL = Bolivia. COL = Colombia.GUF = French Guiana. GUY = Guyana. PER = Peru. PRY = Paraguay. SUR = Surinam.VEN = Venezuela. Brazilian states: AC = Acre.AL = Alagoas.AM = Amazonas.AP = Amapá.BA = Bahia.CE = Ceará. ES = Espírito Santo.GO = Goiás.MA = Maranhão.MG = Minas Gerais.MS = Mato Grosso do Sul.MT = Mato Grosso.PA = Pará.PB = Paraíba.PE = Pernambuco.PI = Piauí.PR= Paraná.RJ = Rio de Janeiro.RN = Rio Grande do Norte.RO = Rondônia.RR = Roraima.RS= Rio Grande do Sul.SC= SantaCatarina.SE = Sergipe.SP = São Paulo.TO =Tocantins." figureDoi="http://doi.org/10.5281/zenodo.4614088" httpUri="https://zenodo.org/record/4614088/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
). The voucher specimen of 
<taxonomicName id="4C4E4D6F671FF17FFC7CFC2E765024FA" authorityName="Wagler" authorityYear="1830" box="[961,1031,986,1011]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B93AEAFE671FF17FFC7CFC2E765024FA" box="[961,1031,986,1011]" italics="true" pageId="17" pageNumber="18">Scinax</emphasis>
</taxonomicName>
sp. (MTJ0578) was not available for morphological study, and therefore we are not aware of any diagnostic characters for this candidate species. The clade 
<emphasis id="B93AEAFE671CF17CFF4DFA4E73B722DA" box="[240,480,1466,1491]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E4D6F671CF17CFF4DFA4E732F22DA" baseAuthorityName="Spix" baseAuthorityYear="1824" box="[240,376,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="x-signatus">S. x‑signatus</taxonomicName>
+ 
<taxonomicName id="4C4E4D6F671CF17CFE27FA4E73B722DA" authorityName="Wagler" authorityYear="1830" box="[410,480,1466,1491]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">Scinax</taxonomicName>
</emphasis>
sp. is well-supported (99% jackknife) as sister taxon of 
<emphasis id="B93AEAFE671CF17CFE58FA2E702C22FA" box="[485,635,1498,1523]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E4D6F671CF17CFE58FA2E702022FA" baseAuthorityName="A.Lutz" baseAuthorityYear="1925" box="[485,631,1498,1523]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fuscovarius">S. fuscovarius</taxonomicName>
,
</emphasis>
followed by a poorly supported clade (&lt;50% jackknife) composed of 
<emphasis id="B93AEAFE671CF17CFF17F9EE73BD213A" box="[170,490,1562,1587]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E4D6F671CF17CFF17F9EE734B213A" baseAuthorityName="Bokermann" baseAuthorityYear="1968" box="[170,284,1562,1587]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="eurydice">S. eurydice</taxonomicName>
, 
<taxonomicName id="4C4E4D6F671CF17CFE98F9EE73DA213A" baseAuthorityName="Cope" baseAuthorityYear="1862" box="[293,397,1562,1587]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="nasicus">S. nasicus</taxonomicName>
, 
<taxonomicName id="4C4E4D6F671CF17CFE2BF9EE73B0213A" box="[406,487,1562,1587]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="ruber">S. ruber</taxonomicName>
,
</emphasis>
and 
<emphasis id="B93AEAFE671CF17CFD9DF9EE7028213A" box="[544,639,1562,1587]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E4D6F671CF17CFD9DF9EE702C213A" authorityName="Cochran" authorityYear="1952" box="[544,635,1562,1587]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="similis">S. similis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>