treatments-xml/data/03/99/CF/0399CF10C439AA58FF7D6584FF2BFCBC.xml
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<document id="2C37F2036C721AD5204778A8DC40BD37" ID-CLB-Dataset="20908" ID-DOI="10.11646/zootaxa.5093.4.7" ID-GBIF-Dataset="a9b22ea0-f9d5-4817-9169-7dd6372923f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912825" ID-ZooBank="C7E53A34-E925-49C0-9866-390018806954" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354875717" checkinUser="plazi" docAuthor="Rahayu, Dwi Listyo &amp; Pratiwi, Rianta" docDate="2022" docId="0399CF10C439AA58FF7D6584FF2BFCBC" docLanguage="en" docName="zootaxa.5093.4.7.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diogenes matabiru Rahayu &amp; Pratiwi 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="498" masterDocId="FFA0B768C438AA5DFFEA6513FFDCFF85" masterDocTitle="Diogenes matabiru, a new species of hermit crab from Lombok Island, Indonesia (Crustacea, Decapoda, Anomura, Diogenidae)" masterLastPageNumber="500" masterPageNumber="493" pageNumber="494" updateTime="1699304820249" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2633B3DD27600AFFAD499C8079F1B8D0">Diogenes matabiru, a new species of hermit crab from Lombok Island, Indonesia (Crustacea, Decapoda, Anomura, Diogenidae)</mods:title>
</mods:titleInfo>
<mods:name id="CB596767B2E7052929DD2793DBFB97E5" type="personal">
<mods:role id="DE31D00E9D742A40266238A68E2A4926">
<mods:roleTerm id="BC8A8D5A86FC32615251E0E2366FFD8E">Author</mods:roleTerm>
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<mods:namePart id="5C5F18DAF8259D199D9B19A633D8226B">Rahayu, Dwi Listyo</mods:namePart>
<mods:nameIdentifier id="5AA33A32881C560EE92CB187539D57F6" type="ORCID">0000-0003-4342-9045</mods:nameIdentifier>
<mods:affiliation id="73000B3CDAD2090A6B82391125D453FD">Marine Bio-industry Division, National Research and Innovation Agency (BRIN), Dusun Teluk Kodek, Pemenang 83756, Lombok Utara, NTB, Indonesia. dwilistyo @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4342 - 9045</mods:affiliation>
<mods:nameIdentifier id="EF60A0D099349C9B423773705A7D3070" type="email">dwilistyo@yahoo.com</mods:nameIdentifier>
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<mods:name id="F427A60B652845AE23A65C8233D414D9" type="personal">
<mods:role id="6C75593E7366C9CFA60BCB10072740FE">
<mods:roleTerm id="BFEA62CBE57ED0F71BB81318F9933E46">Author</mods:roleTerm>
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<mods:namePart id="69ADEF93369E9B1C964ED17DA26664A8">Pratiwi, Rianta</mods:namePart>
<mods:nameIdentifier id="FEB751DA0DE908D93D113777010DEF21" type="ORCID">0000-0002-5516-3996</mods:nameIdentifier>
<mods:affiliation id="2FAF2CA39171A9EEC4C1C19E40CD930F">Research Center for Oceanography, National Research and Innovation Agency (BRIN), Jl. Pasir Putih 1, Ancol Timur, Jakarta Utara, Indonesia. pratiwiafriadi @ gmail. com; https: // orcid. org / 0000 - 0002 - 5516 - 3996</mods:affiliation>
<mods:nameIdentifier id="D8C8D6F3E5236F728197436918AFECE9" type="email">pratiwiafriadi@gmail.com</mods:nameIdentifier>
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<mods:title id="7F79C8223192D00A93D36D9101DA847E">Zootaxa</mods:title>
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<mods:part id="D241D1BFF1CC2BFCC2827403F0A7481E">
<mods:date id="FAC885754BE39EA4BFD3C5A01AB21791">2022</mods:date>
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<mods:number id="D9A7A6947353E69C05242B39FADFAC1B">2022-01-28</mods:number>
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<subSubSection id="C32A2D8DC439AA5CFF7D6584FE14FF34" box="[151,456,151,178]" pageId="1" pageNumber="494" type="nomenclature">
<paragraph id="8B8F7E06C439AA5CFF7D6584FE14FF34" blockId="1.[151,456,151,178]" box="[151,456,151,178]" pageId="1" pageNumber="494">
<heading id="D0C7C96AC439AA5CFF7D6584FE14FF34" bold="true" box="[151,456,151,178]" fontSize="11" level="1" pageId="1" pageNumber="494" reason="1">
<emphasis id="B944A214C439AA5CFF7D6584FE14FF34" bold="true" box="[151,456,151,178]" pageId="1" pageNumber="494">
<taxonomicName id="4C300585C439AA5CFF7D6584FEA2FF37" ID-CoL="46bca9b0-786a-4687-a553-e50a93ebe3a1" authority="Rahayu &amp; Pratiwi, 2022" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[151,382,151,178]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="matabiru" status="sp. nov.">
<emphasis id="B944A214C439AA5CFF7D6584FEA2FF37" bold="true" box="[151,382,151,178]" italics="true" pageId="1" pageNumber="494">Diogenes matabiru</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2771F6FC439AA5CFE6F6584FE14FF34" box="[389,456,151,177]" pageId="1" pageNumber="494" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A2D8DC439AA5CFF7D65CCFA54FDB8" pageId="1" pageNumber="494" type="materials_examined">
<paragraph id="8B8F7E06C439AA5CFF7D65CCFBBAFE98" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
<emphasis id="B944A214C439AA5CFF7D65CCFEA1FF7C" bold="true" box="[151,381,223,249]" pageId="1" pageNumber="494">Material examined</emphasis>
.
<materialsCitation id="3B58745BC439AA5CFE6465CCFE8EFE98" ID-GBIF-Occurrence="3459547311" collectingDate="2014-09-29" collectionCode="MZB" collectorName="R. Pratiwi" location="Lombok Island" pageId="1" pageNumber="494" specimenCount="1" typeStatus="holotype">
<typeStatus id="548BC0A4C439AA5CFE6465CCFE25FF7C" box="[398,505,223,249]" pageId="1" pageNumber="494" type="holotype">Holotype</typeStatus>
, male,
<quantity id="4CC8D3E3C439AA5CFDA565CCFD74FF7F" box="[591,680,223,250]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="494" unit="mm" value="2.7">2.7 mm</quantity>
(
<collectionCode id="ED21E6C3C439AA5CFD5365CCFD2BFF7C" box="[697,759,223,249]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="494" type="Museum">MZB</collectionCode>
Cru 5343), Jerowaru,
<location id="8EEF28DDC439AA5CFBEA65CCFB6DFF7C" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFBEA65CCFB6DFF7C" box="[1024,1201,223,249]" name="Lombok Island" pageId="1" pageNumber="494">Lombok Island</location>
, coll.
<collectorName id="26C51BD0C439AA5CFB1065F3FAAEFF7C" box="[1274,1394,223,250]" pageId="1" pageNumber="494">R. Pratiwi</collectorName>
,
<date id="FF8E58C6C439AA5CFA6A65CCFE8EFE98" pageId="1" pageNumber="494" value="2014-09-29">
<collectingDate id="EFCAA12EC439AA5CFA6A65CCFE8EFE98" pageId="1" pageNumber="494" value="2014-09-29">29 September 2014</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B58745BC439AA5CFEB66417FBBEFE98" ID-GBIF-Occurrence="3459547307" box="[348,1122,259,286]" collectingDate="2014-09-29" collectionCode="ZRC" collectorName="R. Pratiwi" location="Lombok Island" pageId="1" pageNumber="494" specimenCode="ZRC 2021.0900" specimenCount="1" typeStatus="paratype">
<typeStatus id="548BC0A4C439AA5CFEB66417FE1DFE9B" box="[348,449,260,286]" pageId="1" pageNumber="494" type="paratype">Paratype</typeStatus>
, male,
<quantity id="4CC8D3E3C439AA5CFDFA6410FDBAFE9B" box="[528,614,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="494" unit="mm" value="2.5">2.5 mm</quantity>
(
<specimenCode id="DB96D67DC439AA5CFD9E6410FCECFE98" box="[628,816,259,285]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="494">ZRC 2021.0900</specimenCode>
), same locality as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D6434FB89FE28" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
<emphasis id="B944A214C439AA5CFF2D6434FE0BFEC4" bold="true" box="[199,471,295,321]" pageId="1" pageNumber="494">Comparative material.</emphasis>
<taxonomicName id="4C300585C439AA5CFE37643BFC4CFEC4" authority="Whitelegge, 1897" authorityName="Whitelegge" authorityYear="1897" box="[477,912,295,321]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C439AA5CFE37643BFD1EFEC4" box="[477,706,295,321]" italics="true" pageId="1" pageNumber="494">Diogenes pallescens</emphasis>
<materialsCitation id="3B58745BC439AA5CFD2D6434FC4CFEC4" ID-GBIF-Occurrence="3459547306" box="[711,912,295,321]" collectingDate="2007-06" collectionCode="MZB" collectorName="Cru" location="Lombok" municipality="Kuta" pageId="1" pageNumber="494" specimenCount="1">
<bibRefCitation id="EFA103F7C439AA5CFD2D6434FC4CFEC4" author="Whitelegge, T." box="[711,912,295,321]" pageId="1" pageNumber="494" pagination="127 - 151" refId="ref7789" refString="Whitelegge, T. (1897) The fauna of Funafuti [VI.] The Crustacea. Australian Museum Memoir, 3 (1), 127 - 151, pls. 6 + 7. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1897.492" type="journal article" year="1897">Whitelegge, 1897</bibRefCitation>
</materialsCitation>
</taxonomicName>
. 1 ovigerous female,
<quantity id="4CC8D3E3C439AA5CFB976434FB0EFEC7" box="[1149,1234,295,322]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="494" unit="mm" value="1.8">1.8 mm</quantity>
(
<collectionCode id="ED21E6C3C439AA5CFB356434FAC1FEC4" box="[1247,1309,295,321]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="494" type="Museum">MZB</collectionCode>
<collectorName id="26C51BD0C439AA5CFACE6434FA92FEC4" box="[1316,1358,295,321]" pageId="1" pageNumber="494">Cru</collectorName>
5344),
<collectingMunicipality id="6BEBE47CC439AA5CFF7D645FFF0CFEE3" box="[151,208,332,358]" pageId="1" pageNumber="494">Kuta</collectingMunicipality>
,
<location id="8EEF28DDC439AA5CFF346458FE9CFEE0" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFF346458FE9CFEE0" box="[222,320,331,357]" municipality="Kuta" name="Lombok" pageId="1" pageNumber="494">Lombok</location>
,
<date id="FF8E58C6C439AA5CFEA4645FFE1AFEE0" box="[334,454,331,358]" pageId="1" pageNumber="494" value="2007-06">
<collectingDate id="EFCAA12EC439AA5CFEA4645FFE1AFEE0" box="[334,454,331,358]" pageId="1" pageNumber="494" value="2007-06">June 2007</collectingDate>
</date>
;
<materialsCitation id="3B58745BC439AA5CFE3F6458FF0EFE0C" ID-GBIF-Occurrence="3459547302" collectingDate="2003-03-31" collectionCode="MZB" collectorName="Cru" location="Tanjung Merah" pageId="1" pageNumber="494" specimenCount="1" stateProvince="Sulawesi Utara">
1 ovigerous female,
<quantity id="4CC8D3E3C439AA5CFD2B6458FCC6FEE3" box="[705,794,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="494" unit="mm" value="2.5">2.5 mm</quantity>
(
<collectionCode id="ED21E6C3C439AA5CFCC16458FCB5FEE0" box="[811,873,331,357]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="494" type="Museum">MZB</collectionCode>
<collectorName id="26C51BD0C439AA5CFC996458FC41FEE0" box="[883,925,331,357]" pageId="1" pageNumber="494">Cru</collectorName>
5345),
<location id="8EEF28DDC439AA5CFC136458FB76FEE0" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFC136458FB76FEE0" box="[1017,1194,331,357]" name="Tanjung Merah" pageId="1" pageNumber="494" stateProvince="Sulawesi Utara">Tanjung Merah</location>
,
<collectingRegion id="49F4B0E4C439AA5CFB5D6458FAC3FEE0" box="[1207,1311,331,357]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="494">Sulawesi</collectingRegion>
,
<date id="FF8E58C6C439AA5CFAC76458FF0EFE0C" pageId="1" pageNumber="494" value="2003-03-31">
<collectingDate id="EFCAA12EC439AA5CFAC76458FF0EFE0C" pageId="1" pageNumber="494" value="2003-03-31">31 March 2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B58745BC439AA5CFF35647CFB76FE0C" ID-GBIF-Occurrence="3459547304" box="[223,1194,367,394]" collectingDate="2006-07-24" collectionCode="MZB" collectorName="Cru" location="Lombok" municipality="Medana" pageId="1" pageNumber="494" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
<specimenCount id="9D36B58FC439AA5CFF35647CFEF3FE0C" box="[223,303,367,393]" pageId="1" pageNumber="494" type="male">1 male</specimenCount>
,
<quantity id="4CC8D3E3C439AA5CFED1647CFE49FE0F" box="[315,405,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="494" unit="mm" value="2.4">2.4 mm</quantity>
,
<specimenCount id="9D36B58FC439AA5CFE4B647CFDD6FE0C" box="[417,522,367,393]" pageId="1" pageNumber="494" type="female">1 female</specimenCount>
,
<quantity id="4CC8D3E3C439AA5CFDE4647CFDBEFE0F" box="[526,610,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="494" unit="mm" value="1.3">1.3 mm</quantity>
(
<collectionCode id="ED21E6C3C439AA5CFD98647CFD6CFE0C" box="[626,688,367,393]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="494" type="Museum">MZB</collectionCode>
<collectorName id="26C51BD0C439AA5CFD52647CFD3EFE0C" box="[696,738,367,393]" pageId="1" pageNumber="494">Cru</collectorName>
5346),
<collectingMunicipality id="6BEBE47CC439AA5CFCD1647CFC44FE0C" box="[827,920,367,393]" pageId="1" pageNumber="494">Medana</collectingMunicipality>
,
<location id="8EEF28DDC439AA5CFC4F647CFBDBFE0C" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFC4F647CFBDBFE0C" box="[933,1031,367,393]" municipality="Medana" name="Lombok" pageId="1" pageNumber="494" stateProvince="Sulawesi Utara">Lombok</location>
,
<date id="FF8E58C6C439AA5CFBF9647CFB76FE0C" box="[1043,1194,367,393]" pageId="1" pageNumber="494" value="2006-07-24">
<collectingDate id="EFCAA12EC439AA5CFBF9647CFB76FE0C" box="[1043,1194,367,393]" pageId="1" pageNumber="494" value="2006-07-24">24 July 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B58745BC439AA5CFB5C647CFB8CFE28" ID-GBIF-Occurrence="3459547301" collectingDate="2021-08-20" collectionCode="MZB" collectorName="Cru" location="Lombok" municipality="Teluk Kodek" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
<specimenCount id="9D36B58FC439AA5CFB5C647CFAC7FE0C" box="[1206,1307,367,393]" pageId="1" pageNumber="494" type="female">1 female</specimenCount>
, 2.0 mm, 1 ovigerous female,
<quantity id="4CC8D3E3C439AA5CFE8D6480FE61FE2B" box="[359,445,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="494" unit="mm" value="2.7">2.7 mm</quantity>
(
<collectionCode id="ED21E6C3C439AA5CFE216480FDD5FE28" box="[459,521,403,429]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="494" type="Museum">MZB</collectionCode>
<collectorName id="26C51BD0C439AA5CFDFA6480FDE6FE28" box="[528,570,403,429]" pageId="1" pageNumber="494">Cru</collectorName>
5347),
<collectingMunicipality id="6BEBE47CC439AA5CFD7A6480FCF9FE28" box="[656,805,403,429]" pageId="1" pageNumber="494">Teluk Kodek</collectingMunicipality>
,
<location id="8EEF28DDC439AA5CFCC56480FC4DFE28" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFCC56480FC4DFE28" box="[815,913,403,429]" municipality="Teluk Kodek" name="Lombok" pageId="1" pageNumber="494" stateProvince="Sulawesi Utara">Lombok</location>
,
<date id="FF8E58C6C439AA5CFC766480FB8CFE28" box="[924,1104,403,429]" pageId="1" pageNumber="494" value="2021-08-20">
<collectingDate id="EFCAA12EC439AA5CFC766480FB8CFE28" box="[924,1104,403,429]" pageId="1" pageNumber="494" value="2021-08-20">20 August 2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D64ABFAE3FD9C" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
<taxonomicName id="4C300585C439AA5CFF2D64ABFD9FFE54" authority="Morgan, 1987" authorityName="Morgan" authorityYear="1987" box="[199,579,439,466]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C439AA5CFF2D64ABFE4AFE54" box="[199,406,439,465]" italics="true" pageId="1" pageNumber="494">Diogenes biramus</emphasis>
<bibRefCitation id="EFA103F7C439AA5CFE7564ABFD9FFE54" author="Morgan, G. J." box="[415,579,439,466]" pageId="1" pageNumber="494" pagination="165 - 186" refId="ref7142" refString="Morgan, G. J. (1987) Hermit crabs (Decapoda, Anomura: Coenobitidae, Diogenidae, Paguridae) of Darwin and Port Essington, Northern Australia. The Beagle, Record of the Northern Territory Museum of Art and Sciences, 4 (1), 165 - 186. https: // doi. org / 10.5962 / p. 271277" type="journal article" year="1987">Morgan, 1987</bibRefCitation>
</taxonomicName>
.
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5349),
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, 2.0 mm (
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),
<collectingMunicipality id="6BEBE47CC439AA5CFAC264C8FA4BFE70" box="[1320,1431,475,501]" pageId="1" pageNumber="494">Pulau Osi</collectingMunicipality>
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,
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<collectingDate id="EFCAA12EC439AA5CFEB864ECFE36FD9C" box="[338,490,511,538]" pageId="1" pageNumber="494" value="1989-06-16">16 June 1989</collectingDate>
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;
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<specimenCount id="9D36B58FC439AA5CFE1C64ECFD86FD9C" box="[502,602,511,537]" pageId="1" pageNumber="494" type="female">1 female</specimenCount>
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(
<specimenCode id="DB96D67DC439AA5CFD2364ECFC58FD9C" box="[713,900,511,537]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="494">ZRC 2021.0902</specimenCode>
),
<collectingMunicipality id="6BEBE47CC439AA5CFC7D64ECFC28FD9C" box="[919,1012,511,537]" pageId="1" pageNumber="494">Medana</collectingMunicipality>
,
<location id="8EEF28DDC439AA5CFC1564ECFBBDFD9C" LSID="urn:lsid:plazi:treatment:0399CF10C439AA58FF7D6584FF2BFCBC:8EEF28DDC439AA5CFC1564ECFBBDFD9C" box="[1023,1121,511,537]" municipality="Medana" name="Lombok" pageId="1" pageNumber="494" stateProvince="Maluku Utara">Lombok</location>
,
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<collectingDate id="EFCAA12EC439AA5CFB8664ECFAE7FD9C" box="[1132,1339,511,537]" pageId="1" pageNumber="494" value="2021-09-02">2 September 2021</collectingDate>
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.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D6737FA54FDB8" blockId="1.[151,1437,223,2014]" box="[199,1416,547,574]" pageId="1" pageNumber="494">
<taxonomicName id="4C300585C439AA5CFF2D6737FD84FDB8" authority="Haig &amp; Ball, 1988" authorityName="Haig &amp; Ball" authorityYear="1988" box="[199,600,547,573]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C439AA5CFF2D6737FE5CFDB8" box="[199,384,547,573]" italics="true" pageId="1" pageNumber="494">Diogenes viridis</emphasis>
<materialsCitation id="3B58745BC439AA5CFE6C6730FD84FDB8" ID-GBIF-Occurrence="3459547305" box="[390,600,547,573]" collectingDate="2006-10-04" collectionCode="MNHN" country="Vanuatu" location="Santo" pageId="1" pageNumber="494" specimenCount="1">
<bibRefCitation id="EFA103F7C439AA5CFE6C6730FD84FDB8" author="Haig, J. &amp; Ball, E. E." box="[390,600,547,573]" pageId="1" pageNumber="494" pagination="151 - 196" refId="ref6775" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196. https: // doi. org / 10.3853 / j. 0067 - 1975.40.1988.153" type="journal article" year="1988">Haig &amp; Ball, 1988</bibRefCitation>
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</taxonomicName>
. 1 ovigerous female,
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,
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,
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<collectingDate id="EFCAA12EC439AA5CFB896730FACDFDB8" box="[1123,1297,547,573]" pageId="1" pageNumber="494" value="2006-10-04">4 October 2006</collectingDate>
</date>
(
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).
</paragraph>
</subSubSection>
<subSubSection id="C32A2D8DC439AA5FFF1D6754FC6CFAFC" lastPageId="2" lastPageNumber="495" pageId="1" pageNumber="494" type="description">
<paragraph id="8B8F7E06C439AA5CFF1D6754FC0DFD74" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
<emphasis id="B944A214C439AA5CFF1D6754FE5DFDE4" bold="true" box="[247,385,583,609]" pageId="1" pageNumber="494">Description</emphasis>
. Shield (
<figureCitation id="130B6283C439AA5CFE0D6754FD97FDE4" box="[487,587,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Figs. 1A</figureCitation>
,
<figureCitation id="130B6283C439AA5CFDBD6754FDA7FDE4" box="[599,635,583,609]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="1" pageNumber="494">2A</figureCitation>
) about 1.2 times as long as broad; anterior margin between rostral lobe and lateral projections somewhat concave; anterolateral angle rounded, margin sloping; posterior margin truncate; dorsal surface with scattered spines, prominent spines along lateral margin, and tufts of sparse setae. Rostrum broadly rounded. Lateral projections triangular, exceeding tip of rostral lobe, each with moderately large marginal spine. Dorsal margins of branchiostegites with row of 4 or 5 small spines.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D67E8FC54FC20" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
Ocular peduncle (
<figureCitation id="130B6283C439AA5CFE6567E8FE2FFC90" box="[399,499,763,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Figs. 1A</figureCitation>
,
<figureCitation id="130B6283C439AA5CFE1567E8FDFFFC90" box="[511,547,763,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="1" pageNumber="494">2A</figureCitation>
) slender, cylindrical, approximately as long as shield, inflated proximally, with tufts of very sparse setae dorsoproximally; cornea not dilated, diameter about 0.2 times peduncular length. Ocular acicles broad, subrectangular, basally separated by approximately 0.3 width of 1 acicle, with row of spines on terminal margin, mesialmost spine largest. Intercalary rostral process slender, simple at tip, overreaching tip of mesial spine of ocular acicle, with few long setae; no ventral spine.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D66BCFBA6FC68" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
Antennular peduncle (
<figureCitation id="130B6283C439AA5CFE2366BCFDEDFC4C" box="[457,561,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Figs. 1A</figureCitation>
,
<figureCitation id="130B6283C439AA5CFDAB66BCFDBBFC4C" box="[577,615,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="1" pageNumber="494">2A</figureCitation>
), when fully extended, not reaching cornea. Ultimate and penultimate segments glabrous. Basal segment with 2 or 3 small spines on ventrolateral distal margin
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D66E4FD4CFB6F" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
Antennal peduncle (
<figureCitation id="130B6283C439AA5CFE4266E4FE21FB94" box="[424,509,1015,1041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Fig. 1A</figureCitation>
) reaching 0.5 length of ocular peduncle; fifth segment with 1 long and 1 short setae distally, fourth segment with few short setae distally, third segment short, stout, also with few short setae, second segment with dorsolateral distal angle produced into prominent spine, dorsomesial distal angle unarmed, small spine on dorsodistal margin medially, mesial and lateral margins with few setae, first segments unarmed. Antennal acicle overreaching distal margin of fourth segment, terminating in strong spine; lateral margin with 1 large spine, mesial margin with 3 moderately large spines, and few setae. Antennal flagellum slightly longer than shield, consisting of 1618 articles, each with 1 or 2 simple setae.
</paragraph>
<paragraph id="8B8F7E06C439AA5CFF2D61E0FC52FAD0" blockId="1.[151,1437,223,2014]" pageId="1" pageNumber="494">
Third maxilliped (
<figureCitation id="130B6283C439AA5CFE7961E0FE34FA88" box="[403,488,1267,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Fig. 1B</figureCitation>
) with coxa and basis unarmed; crista dentata on ischium obsolete; merus longer than carpus, both segments with long setae on mesial margin, lateral margin with fewer setae; propodus slightly longer than carpus; dactyl shorter than propodus, both with setae ventrally.
</paragraph>
<paragraph id="8B8F7E06C439AA5FFF2D604CFE59FE98" blockId="1.[151,1437,223,2014]" lastBlockId="2.[151,1437,151,1942]" lastPageId="2" lastPageNumber="495" pageId="1" pageNumber="494">
Left cheliped (
<figureCitation id="130B6283C439AA5CFE86604CFE11FAFC" box="[364,461,1375,1401]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="1" pageNumber="494">Figs. 1C</figureCitation>
,
<figureCitation id="130B6283C439AA5CFE30604CFE22FAFC" box="[474,510,1375,1401]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="1" pageNumber="494">2A</figureCitation>
) larger than right, narrow hiatus between dactyl and fixed finger. Dactyl about 0.8 times as long as upper margin of palm, slightly arched; cutting edge with row of blunt calcareous teeth, 1 large tooth proximally, terminating in small calcareous claw, slightly overlapped by fixed finger; outer surface convex with partial covering of spines, and with long stiff setae; upper margin with 2 rows of moderately large spines decreasing in size distally; inner surface slightly convex, with covering tubercles and tufts of short setae. Fixed finger outer surface slightly convex, armed with moderately large sharp and blunt spines and long stiff setae; lower margin with row of moderately small, subacute spines, forming weak curve with lower, similarly armed margin of palm; cutting edge of fixed finger with row of calcareous teeth, large tooth subproximally, terminating in large calcareous claw; inner surface with scattered small tubercles and few tufts of short setae. Palm slightly shorter than carpus, with row of moderately large spines on upper margin, second row shorter and with smaller spines; outer surface convex, with sparse short setae, longitudinal row of sharp, large spines on midline, extending beyond midlength of palm but not reaching articulation with dactyl; additional longitudinal row of small spines between midline and upper margin merging into scattered spines adjacent to articulation with dactyl; lower half of outer surface with sparse spines forming irregular longitudinal rows; inner surface with few small tubercles and short setae. Carpus as long as merus; upper margin with row of irregularly spaced prominent spines and sparse long setae; outer surface convex with longitudinal row of large spine on midline; lower margin with large spine distally and few small spines medially; inner surface with few tubercles and sparse setae. Merus subtriangular in dorsal view; dorsodistal margin with strong spine; dorsal surface with rows of moderately small spines and tufts of long plumose setae; lateral face tuberculate, ventrolateral margin with row of large spines, distal spines strongest and accompanied by long setae; mesial face with sparse tubercles, ventromesial margin with row of moderately large spines, distal spines strongest and accompanied by plumose setae; ventral surface tuberculate. Ischium unarmed, but with sparse short setae on ventromesial margin.
</paragraph>
<paragraph id="8B8F7E06C43AAA5FFF2D6434FB20FD2C" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
Right cheliped (
<figureCitation id="130B6283C43AAA5FFE976434FE3DFEC4" box="[381,481,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Figs. 1D</figureCitation>
,
<figureCitation id="130B6283C43AAA5FFE046434FDCEFEC4" box="[494,530,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="2" pageNumber="495">2A</figureCitation>
) reaching proximal third of palm of left cheliped; narrow hiatus between dactyl and fixed finger. Dactyl longer than palm; upper margin with row small spines and numerous long setae; outer surface with scattered small spines and long setae; cutting edge with row of low, blunt, calcareous teeth, terminating in moderately large calcareous claw overlapped by fixed finger. Fixed finger with row of tubercles in midline partially obscured by long setae on outer surface; lower margin smooth but with sparse long setae; cutting edge with row of small calcareous teeth, terminating in moderately large calcareous claw. Palm with row of moderately large spines and tufts of long setae on upper margin; outer surface with longitudinal row of small spines in midline and with long setae. Carpus with row of large spines on upper margin; outer face with row of small spines in midline, lower half of surface with sparse tubercles, inner surface with setae distally. Merus triangular, dorsal margin with 1 large spine distally, and with numerous long, simple or plumose setae; ventrolateral margins with irregular row of small spines, ventromesial margin with few small spines. Ischium unarmed, but with few long setae.
</paragraph>
<paragraph id="8B8F7E06C43AAA5FFF2D67A0FC11FB94" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
P2 and P3 (
<figureCitation id="130B6283C43AAA5FFEAC67A0FE42FD48" box="[326,414,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="2" pageNumber="495">Fig. 2A</figureCitation>
) slender. P2 (
<figureCitation id="130B6283C43AAA5FFDDF67A0FD54FD48" box="[565,648,691,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1E</figureCitation>
) with dactyls 1.3 times as long as propodi, somewhat curved ventrally, but not twisted in dorsal view, terminating in moderately small corneous claws; dorsal and ventral margins each with mixture of short and long, plumose and simple setae; mesial surfaces each with row of setae in midline. Propodi 1.4 times as long as carpi, unarmed, each with sparse long and short setae on dorsal and ventral margins; mesial faces each with few setae. Carpi 0.7 times as long as meri; moderately large spine on dorsodistal and dorsoproximal and sparse, long setae; ventral margins unarmed. Meri each with sparse setae on dorsal and ventral margins, otherwise unarmed. Ischia unarmed, each with sparse setae. P3 (
<figureCitation id="130B6283C43AAA5FFD106698FC91FC20" box="[762,845,907,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1F</figureCitation>
) with dactyls and propodi same as in P2; carpi each with prominent dorsodistal spine, smaller spine on subdistal and subproximal dorsally, and sparse setae, lateral faces and ventral margins unarmed; meri and ischia also unarmed but with few setae. P4 semichelate (
<figureCitation id="130B6283C43AAA5FFB3C66C0FAF3FC68" box="[1238,1327,979,1005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1G</figureCitation>
); propodi with tiny dorsodistal spine, carpi with moderately large dorsodistal spine.
</paragraph>
<paragraph id="8B8F7E06C43AAA5FFF2D6108FC56FBF8" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
Telson (
<figureCitation id="130B6283C43AAA5FFECB6108FEABFBB0" box="[289,375,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1H</figureCitation>
) with distinct median cleft, long setae on terminal margin; left posterior lobe longer than right, left terminal margin with row of small spines, stronger spines extending onto lateral margin; right terminal margin also with row of small spines, but not extended onto lateral margin.
</paragraph>
<paragraph id="8B8F7E06C43AAA5FFF2D6195FC6CFAFC" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
<emphasis id="B944A214C43AAA5FFF2D6195FEB4FB25" bold="true" box="[199,360,1158,1184]" pageId="2" pageNumber="495">Colour in life</emphasis>
. Shield cream with large dark brown spot medially, smaller dark brown patches anteriorly and laterally. Ocular peduncle generally white, greenish brown patch proximally, orangish red band on third distal, blue areas proximal to cornea, cornea light brown. Ocular acicle white with brown spot mesially. Antennular peduncles cream with dark brow ring on ultimate and penultimate segments distally. Antennal peduncles light brown, white band on fifth segment distally. Chelipeds cream, greenish brown on dactyl and fixed finger, palm with dark brown patches, spines white. P2 and P3 cream mottled with light brown, dark brown band on each propodi and carpi medially, meri with dark brown medially and subproximally (
<figureCitation id="130B6283C43AAA5FFCA3604CFC78FAFC" box="[841,932,1375,1401]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="2" pageNumber="495">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A2D8DC43AAA5FFF2D6090FE7DFA44" pageId="2" pageNumber="495" type="etymology">
<paragraph id="8B8F7E06C43AAA5FFF2D6090FE7DFA44" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
<emphasis id="B944A214C43AAA5FFF2D6090FE94FA18" bold="true" box="[199,328,1411,1437]" pageId="2" pageNumber="495">Etymology</emphasis>
:
<taxonomicName id="4C300585C43AAA5FFEBC6090FE61FA18" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[342,445,1411,1437]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="matabiru">
<emphasis id="B944A214C43AAA5FFEBC6090FE61FA18" box="[342,445,1411,1437]" italics="true" pageId="2" pageNumber="495">matabiru</emphasis>
</taxonomicName>
means blue eye in Bahasa
<collectingCountry id="F3273E96C43AAA5FFD036090FC85FA18" box="[745,857,1411,1437]" name="Indonesia" pageId="2" pageNumber="495">Indonesia</collectingCountry>
, alluding to the blue colour of ocular peduncles; use as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32A2D8DC43AAA58FF2D60D8FF2BFCBC" lastPageId="5" lastPageNumber="498" pageId="2" pageNumber="495" type="discussion">
<paragraph id="8B8F7E06C43AAA5FFF2D60D8FD0EF813" blockId="2.[151,1437,151,1942]" pageId="2" pageNumber="495">
<emphasis id="B944A214C43AAA5FFF2D60D8FEE8FA60" bold="true" box="[199,308,1483,1509]" pageId="2" pageNumber="495">Remarks</emphasis>
. Within the eight species of the
<taxonomicName id="4C300585C43AAA5FFD7960D8FCF4FA60" authorityName="Whitelegge" authorityYear="1897" box="[659,808,1483,1509]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFD7960D8FCF4FA60" box="[659,808,1483,1509]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
group, four species have the antennular peduncle shorter than the ocular peduncle, namely,
<taxonomicName id="4C300585C43AAA5FFDF760FCFD6AF98C" authorityName="Whitelegge" authorityYear="1897" box="[541,694,1519,1545]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFDF760FCFD6AF98C" box="[541,694,1519,1545]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C300585C43AAA5FFD2F60FCFC99F98C" authorityName="Morgan" authorityYear="1987" box="[709,837,1519,1545]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43AAA5FFD2F60FCFC99F98C" box="[709,837,1519,1545]" italics="true" pageId="2" pageNumber="495">D. biramus</emphasis>
</taxonomicName>
,
<emphasis id="B944A214C43AAA5FFCBE60FCFC1AF98C" box="[852,966,1519,1545]" italics="true" pageId="2" pageNumber="495">
<taxonomicName id="4C300585C43AAA5FFCBE60FCFC1DF98C" authorityName="Haig &amp; Ball" authorityYear="1988" box="[852,961,1519,1545]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="viridis">D. viridis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C300585C43AAA5FFC1560FCFBBDF98D" authorityName="Asakura &amp; Godwin" authorityYear="2006" box="[1023,1121,1519,1544]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="patae">
<emphasis id="B944A214C43AAA5FFC1560FCFBBDF98D" box="[1023,1121,1519,1544]" italics="true" pageId="2" pageNumber="495">D. patae</emphasis>
</taxonomicName>
. Among these four species,
<taxonomicName id="4C300585C43AAA5FFF7D6300FEFEF9A8" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[151,290,1555,1581]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="matabiru">
<emphasis id="B944A214C43AAA5FFF7D6300FEFEF9A8" box="[151,290,1555,1581]" italics="true" pageId="2" pageNumber="495">D. matabiru</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C300585C43AAA5FFDD36300FD0DF9A8" authorityName="Whitelegge" authorityYear="1897" box="[569,721,1555,1581]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFDD36300FD0DF9A8" box="[569,721,1555,1581]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
in having strong spines on the shield and the left cheliped, the ocular peduncle as long as the shield, and P4 with distal spine on the dorsal margin of the carpi. However, the shield of the new species is clearly longer than broad (
<figureCitation id="130B6283C43AAA5FFD5B6348FCD4F9F0" box="[689,776,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1A</figureCitation>
) (as long as broad in
<taxonomicName id="4C300585C43AAA5FFC176348FB49F9F0" authorityName="Whitelegge" authorityYear="1897" box="[1021,1173,1627,1653]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFC176348FB49F9F0" box="[1021,1173,1627,1653]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
), the intercalary rostral process is very slender, reaching well beyond the tip of the longest spine of the ocular acicle (
<figureCitation id="130B6283C43AAA5FFB21636CFAFFF91C" box="[1227,1315,1663,1689]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1A</figureCitation>
) (larger at base to tapering at tip, as long as or only slightly reaching beyond the tip of the ocular acicle in
<taxonomicName id="4C300585C43AAA5FFB1963B0FA50F938" authorityName="Whitelegge" authorityYear="1897" box="[1267,1420,1699,1725]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFB1963B0FA50F938" box="[1267,1420,1699,1725]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
), the left cheliped with outer surface of fixed finger slightly convex, armed with strong spines, outer surface of the palm with row of strong spines on the midline and row of smaller spines between the midline and the upper margin (
<figureCitation id="130B6283C43AAA5FFF4A621CFF2AF8AC" box="[160,246,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1C</figureCitation>
), and the ventral margin of the merus of P2 unarmed (
<figureCitation id="130B6283C43AAA5FFCBF621CFC74F8AC" box="[853,936,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1859,1883]" captionTargetBox="[162,1424,189,1818]" captionTargetId="figure-17@3.[151,1436,181,1835]" captionTargetPageId="3" captionText="FIGURE 1. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="2" pageNumber="495">Fig. 1E</figureCitation>
) (outer surface of the fixed finger somewhat flattened, the palm with scattered tubercles or spines and with longitudinal row of strong spines beginning near the midpoint of proximal margin not reaching midlength, and the ventral margin of the merus of P2 with 1 to 5 spines in
<taxonomicName id="4C300585C43AAA5FFF5E6268FE97F810" authorityName="Whitelegge" authorityYear="1897" box="[180,331,1915,1941]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43AAA5FFF5E6268FE97F810" box="[180,331,1915,1941]" italics="true" pageId="2" pageNumber="495">D. pallescens</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFA103F7C43AAA5FFE896268FD87F810" author="McLaughlin, P. A." box="[355,603,1915,1941]" pageId="2" pageNumber="495" pagination="81 - 94" refId="ref6976" refString="McLaughlin, P. A. (2002) Diogenes pallescens Whitelegge, D. gardineri Alcock and D. serenei Forest (Decapoda: Anomura: Paguroidea: Diogenidae): Distinct species or morphological variant? Raffles Bulletin of Zoology, 50 (1), 81 - 94." type="journal article" year="2002">McLaughlin 2002: 91</bibRefCitation>
, figs.14).
</paragraph>
<caption id="DF4F2E8EC43BAA5EFF7D6250FE59F842" ID-DOI="http://doi.org/10.5281/zenodo.5912827" ID-Zenodo-Dep="5912827" httpUri="https://zenodo.org/record/5912827/files/figure.png" pageId="3" pageNumber="496" startId="3.[151,250,1859,1883]" targetBox="[162,1424,189,1818]" targetPageId="3">
<paragraph id="8B8F7E06C43BAA5EFF7D6250FE59F842" blockId="3.[151,1436,1859,1991]" pageId="3" pageNumber="496">
<emphasis id="B944A214C43BAA5EFF7D6250FED1F8DE" bold="true" box="[151,269,1859,1883]" pageId="3" pageNumber="496">FIGURE 1</emphasis>
.
<taxonomicName id="4C300585C43BAA5EFEF36257FE00F8DE" authority="Rahayu &amp; Pratiwi, 2022" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[281,476,1859,1883]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="matabiru" status="sp. nov.">
<emphasis id="B944A214C43BAA5EFEF36257FE00F8DE" box="[281,476,1859,1883]" italics="true" pageId="3" pageNumber="496">Diogenes matabiru</emphasis>
</taxonomicName>
<emphasis id="B944A214C43BAA5EFE096250FDC0F8DE" bold="true" box="[483,540,1859,1883]" pageId="3" pageNumber="496">
<taxonomicNameLabel id="A2771F6FC43BAA5EFE096250FDC0F8DE" box="[483,540,1859,1883]" pageId="3" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male, 2.7 mm (MZB Cru 5343). A, shield and cephalic appendages; B, chela, carpus and merus of left cheliped, dorsolateral view; C, chela, carpus and merus of right cheliped, dorsolateral view; D, left third maxilliped, mesial view; E, right P2, lateral view; F, left P3, lateral view; G, left P4, lateral view; H, telson. Setae partially omitted. Scale = 1 mm.
</paragraph>
</caption>
<caption id="DF4F2E8EC43CAA59FF7D6080FC61FA76" ID-DOI="http://doi.org/10.5281/zenodo.5912829" ID-Zenodo-Dep="5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="4" pageNumber="497" startId="4.[151,250,1427,1451]" targetBox="[151,1435,181,1402]" targetPageId="4">
<paragraph id="8B8F7E06C43CAA59FF7D6080FC61FA76" blockId="4.[151,1437,1427,1523]" pageId="4" pageNumber="497">
<emphasis id="B944A214C43CAA59FF7D6080FED1FA2E" bold="true" box="[151,269,1427,1451]" pageId="4" pageNumber="497">FIGURE 2</emphasis>
. A.
<taxonomicName id="4C300585C43CAA59FEDC6080FE24FA2E" authority="Rahayu &amp; Pratiwi, 2022" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[310,504,1427,1451]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="matabiru" status="sp. nov.">
<emphasis id="B944A214C43CAA59FEDC6080FE24FA2E" box="[310,504,1427,1451]" italics="true" pageId="4" pageNumber="497">Diogenes matabiru</emphasis>
</taxonomicName>
<emphasis id="B944A214C43CAA59FE146080FDEBFA2E" bold="true" box="[510,567,1427,1451]" pageId="4" pageNumber="497">
<taxonomicNameLabel id="A2771F6FC43CAA59FE146080FDEBFA2E" box="[510,567,1427,1451]" pageId="4" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male, 2.7 mm (MZB Cru 5343); B,
<taxonomicName id="4C300585C43CAA59FBE46080FA44FA2E" authority="Whitelegge, 1897" authorityName="Whitelegge" authorityYear="1897" box="[1038,1432,1427,1451]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43CAA59FBE46080FB01FA2E" box="[1038,1245,1427,1451]" italics="true" pageId="4" pageNumber="497">Diogenes pallescens</emphasis>
<bibRefCitation id="EFA103F7C43CAA59FB086080FA44FA2E" author="Whitelegge, T." box="[1250,1432,1427,1451]" pageId="4" pageNumber="497" pagination="127 - 151" refId="ref7789" refString="Whitelegge, T. (1897) The fauna of Funafuti [VI.] The Crustacea. Australian Museum Memoir, 3 (1), 127 - 151, pls. 6 + 7. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1897.492" type="journal article" year="1897">Whitelegge, 1897</bibRefCitation>
</taxonomicName>
, ovigerous female, 2.7 mm (MZB Cru 5347); C.
<taxonomicName id="4C300585C43CAA59FD6A60A4FC0DFA4A" authority="Morgan, 1987" authorityName="Morgan" authorityYear="1987" box="[640,977,1463,1487]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43CAA59FD6A60A4FCE5FA4A" box="[640,825,1463,1487]" italics="true" pageId="4" pageNumber="497">Diogenes biramus</emphasis>
<bibRefCitation id="EFA103F7C43CAA59FCAA60A4FC0DFA4A" author="Morgan, G. J." box="[832,977,1463,1487]" pageId="4" pageNumber="497" pagination="165 - 186" refId="ref7142" refString="Morgan, G. J. (1987) Hermit crabs (Decapoda, Anomura: Coenobitidae, Diogenidae, Paguridae) of Darwin and Port Essington, Northern Australia. The Beagle, Record of the Northern Territory Museum of Art and Sciences, 4 (1), 165 - 186. https: // doi. org / 10.5962 / p. 271277" type="journal article" year="1987">Morgan, 1987</bibRefCitation>
</taxonomicName>
, female, mm (ZRC 2021.0902); D,
<taxonomicName id="4C300585C43CAA59FAD760A4FE46FA76" authority="Haig &amp; Ball, 1988" authorityName="Haig &amp; Ball" authorityYear="1988" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43CAA59FAD760A4FF04FA76" italics="true" pageId="4" pageNumber="497">Diogenes viridis</emphasis>
<bibRefCitation id="EFA103F7C43CAA59FF3460C8FE46FA76" author="Haig, J. &amp; Ball, E. E." box="[222,410,1499,1523]" pageId="4" pageNumber="497" pagination="151 - 196" refId="ref6775" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196. https: // doi. org / 10.3853 / j. 0067 - 1975.40.1988.153" type="journal article" year="1988">Haig &amp; Ball, 1988</bibRefCitation>
</taxonomicName>
(Panglao, Philippines, 2004; specimen not collected).
</paragraph>
</caption>
<paragraph id="8B8F7E06C43CAA59FF2D6331FEF5F96A" blockId="4.[151,1437,1569,2028]" pageId="4" pageNumber="497">
<taxonomicName id="4C300585C43CAA59FF2D6331FE4EF9BE" authorityName="Morgan" authorityYear="1987" box="[199,402,1569,1595]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43CAA59FF2D6331FE4EF9BE" box="[199,402,1569,1595]" italics="true" pageId="4" pageNumber="497">Diogenes biramus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C300585C43CAA59FE296331FDF7F9BE" authorityName="Haig &amp; Ball" authorityYear="1988" box="[451,555,1569,1595]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43CAA59FE296331FDF7F9BE" box="[451,555,1569,1595]" italics="true" pageId="4" pageNumber="497">D. viridis</emphasis>
</taxonomicName>
differ in having the ocular peduncles much longer than the shield, the intercalary rostral process is short, triangular, reaching only midlength of the ocular acicle, and the terminal margin of the telson lacks the medial cleft (
<bibRefCitation id="EFA103F7C43CAA59FE7F637AFDEDF906" author="Morgan, G. J." box="[405,561,1641,1667]" pageId="4" pageNumber="497" pagination="165 - 186" refId="ref7142" refString="Morgan, G. J. (1987) Hermit crabs (Decapoda, Anomura: Coenobitidae, Diogenidae, Paguridae) of Darwin and Port Essington, Northern Australia. The Beagle, Record of the Northern Territory Museum of Art and Sciences, 4 (1), 165 - 186. https: // doi. org / 10.5962 / p. 271277" type="journal article" year="1987">Morgan 1987</bibRefCitation>
;
<bibRefCitation id="EFA103F7C43CAA59FDD6637AFCD9F906" author="Haig, J. &amp; Ball, E. E." box="[572,773,1641,1667]" pageId="4" pageNumber="497" pagination="151 - 196" refId="ref6775" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196. https: // doi. org / 10.3853 / j. 0067 - 1975.40.1988.153" type="journal article" year="1988">Haig &amp; Ball 1988</bibRefCitation>
) (the ocular peduncles as long as the shield, the intercalary rostral process is long, overreaching well beyond the ocular acicle, and the medial cleft of the telson is present in
<taxonomicName id="4C300585C43CAA59FA6B639DFF22F94E" authority="Rahayu &amp; Pratiwi, 2022" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="matabiru" status="sp. nov.">
<emphasis id="B944A214C43CAA59FA6B639DFF22F94E" italics="true" pageId="4" pageNumber="497">D. matabiru</emphasis>
</taxonomicName>
<emphasis id="B944A214C43CAA59FEEE63A2FE9EF94E" bold="true" box="[260,322,1713,1739]" pageId="4" pageNumber="497">
<taxonomicNameLabel id="A2771F6FC43CAA59FEEE63A2FE9EF94E" box="[260,322,1713,1739]" pageId="4" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C300585C43CAA59FEB363A1FDDBF94E" authorityName="Asakura &amp; Godwin" authorityYear="2006" box="[345,519,1714,1739]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="patae">
<emphasis id="B944A214C43CAA59FEB363A1FDDBF94E" box="[345,519,1714,1739]" italics="true" pageId="4" pageNumber="497">Diogenes patae</emphasis>
</taxonomicName>
is different in having dense setae on the chelipeds, P2 and P3 (
<bibRefCitation id="EFA103F7C43CAA59FB2A63A2FF0FF96A" author="Asakura, A. &amp; Godwin, S." pageId="4" pageNumber="497" pagination="457 - 463" refId="ref6418" refString="Asakura, A. &amp; Godwin, S. (2006) Diogenes patae n. sp., a new species of hermit crab (Crustacea, Decapoda, Anomura, Diogenidae) from American Samoa. Zoosystema, 28 (2), 457 - 463." type="journal article" year="2006">Asakura &amp; Godwin 2006</bibRefCitation>
: fig. 2).
</paragraph>
<paragraph id="8B8F7E06C43CAA58FF2D63EAFEFDFF50" blockId="4.[151,1437,1569,2028]" lastBlockId="5.[151,1437,151,825]" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
The life coloration will easily separate
<taxonomicName id="4C300585C43CAA59FD6863E9FCD0F896" authority="Rahayu &amp; Pratiwi, 2022" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[642,780,1785,1811]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="matabiru" status="sp. nov.">
<emphasis id="B944A214C43CAA59FD6863E9FCD0F896" box="[642,780,1785,1811]" italics="true" pageId="4" pageNumber="497">D. matabiru</emphasis>
</taxonomicName>
<emphasis id="B944A214C43CAA59FCF963EAFC8FF896" bold="true" box="[787,851,1785,1811]" pageId="4" pageNumber="497">
<taxonomicNameLabel id="A2771F6FC43CAA59FCF963EAFC8FF896" box="[787,851,1785,1811]" pageId="4" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C300585C43CAA59FC7063E9FBEEF896" authorityName="Whitelegge" authorityYear="1897" box="[922,1074,1785,1811]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43CAA59FC7063E9FBEEF896" box="[922,1074,1785,1811]" italics="true" pageId="4" pageNumber="497">D. pallescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C300585C43CAA59FBAA63E9FB63F896" authorityName="Morgan" authorityYear="1987" box="[1088,1215,1785,1811]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43CAA59FBAA63E9FB63F896" box="[1088,1215,1785,1811]" italics="true" pageId="4" pageNumber="497">D. biramus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C300585C43CAA59FB1263E9FABFF896" authorityName="Haig &amp; Ball" authorityYear="1988" box="[1272,1379,1785,1811]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43CAA59FB1263E9FABFF896" box="[1272,1379,1785,1811]" italics="true" pageId="4" pageNumber="497">D. viridis</emphasis>
</taxonomicName>
. The ocular peduncle of
<taxonomicName id="4C300585C43CAA59FE9F620DFD93F8B2" authorityName="Rahayu &amp; Pratiwi" authorityYear="2022" box="[373,591,1821,1847]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="matabiru">
<emphasis id="B944A214C43CAA59FE9F620DFD93F8B2" box="[373,591,1821,1847]" italics="true" pageId="4" pageNumber="497">Diogenes matabiru</emphasis>
</taxonomicName>
is white, with greenish brown large patch proximally, orangish red broad band on third distal, and a blue area near cornea, the cheliped palm is cream with dark brown patches on dorsolateral and dorsomesial surface (
<figureCitation id="130B6283C43CAA59FE546276FDC7F8FA" box="[446,539,1893,1919]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="4" pageNumber="497">Fig. 2A</figureCitation>
). In
<taxonomicName id="4C300585C43CAA59FDB86275FD30F8FA" authorityName="Whitelegge" authorityYear="1897" box="[594,748,1893,1919]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="B944A214C43CAA59FDB86275FD30F8FA" box="[594,748,1893,1919]" italics="true" pageId="4" pageNumber="497">D. pallescens</emphasis>
</taxonomicName>
the ocular peduncle is white, light and dark brown patches on proximal half, brown band on distal third, the cheliped palm is cream with dark brown patches proximally on dorsal surface (
<figureCitation id="130B6283C43CAA59FEAC62BEFE43F842" box="[326,415,1965,1991]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="4" pageNumber="497">Fig. 2B</figureCitation>
). In
<taxonomicName id="4C300585C43CAA59FE3F62BDFD89F842" authorityName="Morgan" authorityYear="1987" box="[469,597,1965,1991]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43CAA59FE3F62BDFD89F842" box="[469,597,1965,1991]" italics="true" pageId="4" pageNumber="497">D. biramus</emphasis>
</taxonomicName>
the ocular peduncle is uniformly cream, the cheliped palm is light brown with pinkish brown patches on dorsolateral surface (
<figureCitation id="130B6283C43CAA59FD0262C2FC9DF86E" box="[744,833,2001,2027]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="4" pageNumber="497">Fig. 2C</figureCitation>
), while in
<taxonomicName id="4C300585C43CAA59FC5062C1FBF9F86E" authorityName="Haig &amp; Ball" authorityYear="1988" box="[954,1061,2001,2027]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43CAA59FC5062C1FBF9F86E" box="[954,1061,2001,2027]" italics="true" pageId="4" pageNumber="497">D. viridis</emphasis>
</taxonomicName>
the ocular peduncle is uniformly light pink, the cheliped chela, palm, and carpus white, the merus green (
<figureCitation id="130B6283C43DAA58FC2C6584FBFDFF34" box="[966,1057,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="5" pageNumber="498">Fig. 2D</figureCitation>
). The life colouration of
<taxonomicName id="4C300585C43DAA58FAD16584FA40FF35" authorityName="Asakura &amp; Godwin" authorityYear="2006" box="[1339,1436,151,176]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="patae">
<emphasis id="B944A214C43DAA58FAD16584FA40FF35" box="[1339,1436,151,176]" italics="true" pageId="5" pageNumber="498">D. patae</emphasis>
</taxonomicName>
is unknown.
</paragraph>
<paragraph id="8B8F7E06C43DAA58FF2D65CCFD15FD74" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="498">
<bibRefCitation id="EFA103F7C43DAA58FF2D65CCFE0DFF7C" author="Rahayu, D. L. &amp; Forest, J." box="[199,465,222,249]" pageId="5" pageNumber="498" pagination="383 - 415" refId="ref7560" refString="Rahayu, D. L. &amp; Forest, J. (1995) Le genre Diogenes (Crustacea, Decapoda, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin Museum national d'Histoire naturelle, Paris, 16 (2 - 4), 383 - 415." type="journal article" year="1995">Rahayu &amp; Forest (1995)</bibRefCitation>
discussed the similarity between
<taxonomicName id="4C300585C43DAA58FCD565CCFC67FF7C" authorityName="Morgan" authorityYear="1987" box="[831,955,223,249]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43DAA58FCD565CCFC67FF7C" box="[831,955,223,249]" italics="true" pageId="5" pageNumber="498">D. biramus</emphasis>
</taxonomicName>
and
<emphasis id="B944A214C43DAA58FC0765CCFB87FF7C" box="[1005,1115,223,249]" italics="true" pageId="5" pageNumber="498">
<taxonomicName id="4C300585C43DAA58FC0765CCFB8BFF7C" authorityName="Haig &amp; Ball" authorityYear="1988" box="[1005,1111,223,249]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">D. viridis</taxonomicName>
,
</emphasis>
and suggested the possibility that the latter is a junior synonym of the former. The respective
<typeStatus id="548BC0A4C43DAA58FC606410FC66FE98" box="[906,954,259,285]" pageId="5" pageNumber="498">type</typeStatus>
localities of each species are not distant:
<taxonomicName id="4C300585C43DAA58FF7D6434FEC4FEC4" authorityName="Morgan" authorityYear="1987" box="[151,280,295,321]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43DAA58FF7D6434FEC4FEC4" box="[151,280,295,321]" italics="true" pageId="5" pageNumber="498">D. biramus</emphasis>
</taxonomicName>
was described from Coral Bay, the
<collectingRegion id="49F4B0E4C43DAA58FD526434FC56FEC4" box="[696,906,295,321]" country="Australia" name="Northern Territory" pageId="5" pageNumber="498">Northern Territory</collectingRegion>
,
<collectingCountry id="F3273E96C43DAA58FC7C6434FBDCFEC4" box="[918,1024,295,321]" name="Australia" pageId="5" pageNumber="498">Australia</collectingCountry>
, while
<taxonomicName id="4C300585C43DAA58FBBE6434FB1DFEC4" authorityName="Haig &amp; Ball" authorityYear="1988" box="[1108,1217,295,321]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43DAA58FBBE6434FB1DFEC4" box="[1108,1217,295,321]" italics="true" pageId="5" pageNumber="498">D. viridis</emphasis>
</taxonomicName>
was from
<collectingRegion id="49F4B0E4C43DAA58FAD46434FA44FEC4" box="[1342,1432,295,321]" country="Indonesia" name="Maluku Utara" pageId="5" pageNumber="498">Maluku</collectingRegion>
,
<collectingCountry id="F3273E96C43DAA58FF7D6458FED4FEE0" box="[151,264,331,357]" name="Indonesia" pageId="5" pageNumber="498">Indonesia</collectingCountry>
. However, examination of the specimens from
<collectingCountry id="F3273E96C43DAA58FCC36458FC4AFEE0" box="[809,918,331,357]" name="Indonesia" pageId="5" pageNumber="498">Indonesia</collectingCountry>
and
<collectingCountry id="F3273E96C43DAA58FC3B6458FBF1FEE0" box="[977,1069,331,357]" name="Vanuatu" pageId="5" pageNumber="498">Vanuatu</collectingCountry>
revealed several characters that separate these two species. The most distinctive character is the colouration of P2 and P3. In
<taxonomicName id="4C300585C43DAA58FB2F647CFA99FE0C" authorityName="Morgan" authorityYear="1987" box="[1221,1349,367,393]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43DAA58FB2F647CFA99FE0C" box="[1221,1349,367,393]" italics="true" pageId="5" pageNumber="498">D. biramus</emphasis>
</taxonomicName>
P2 and P3 cream, dactyl with light brown band medially, propodus with dark brown band medially and proximally, carpus and merus mottled with light brown, dark brown band medially (
<figureCitation id="130B6283C43DAA58FC9564A4FC06FE54" box="[895,986,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="5" pageNumber="498">Fig. 2C</figureCitation>
). In
<taxonomicName id="4C300585C43DAA58FBE564A4FBA7FE54" authorityName="Haig &amp; Ball" authorityYear="1988" box="[1039,1147,439,465]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43DAA58FBE564A4FBA7FE54" box="[1039,1147,439,465]" italics="true" pageId="5" pageNumber="498">D. viridis</emphasis>
</taxonomicName>
the dactyls of P2 and P3 are white, propodi, carpi and meri green with small white spots, white band distally (
<figureCitation id="130B6283C43DAA58FB6B64C8FB03FE70" box="[1153,1247,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[151,1435,181,1402]" captionTargetId="figure-24@4.[151,1435,181,1402]" captionTargetPageId="4" captionText="FIGURE 2. A. Diogenes matabiru n. sp., holotype, male, 2.7 mm (MZB Cru 5343); B, Diogenes pallescens Whitelegge, 1897, ovigerous female, 2.7 mm (MZB Cru 5347); C. Diogenes biramus Morgan, 1987, female, mm (ZRC 2021.0902); D, Diogenes viridis Haig &amp; Ball, 1988 (Panglao, Philippines, 2004; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.5912829" httpUri="https://zenodo.org/record/5912829/files/figure.png" pageId="5" pageNumber="498">Fig. 2D</figureCitation>
). Morphological differences of these two species are also remarkable.
<taxonomicName id="4C300585C43DAA58FD1964ECFC1DFD9C" authorityName="Morgan" authorityYear="1987" box="[755,961,511,537]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="biramus">
<emphasis id="B944A214C43DAA58FD1964ECFC1DFD9C" box="[755,961,511,537]" italics="true" pageId="5" pageNumber="498">Diogenes biramus</emphasis>
</taxonomicName>
has the following features: shield slightly longer than broad, left cheliped dactyl much longer than palm, fixed finger slightly flattened and lower margin with row of large spines that continued to lower margin of palm, propodus of P4 without any spine, telson subrectangular, broader than long.
<taxonomicName id="4C300585C43DAA58FE866778FDF9FD00" authorityName="Haig &amp; Ball" authorityYear="1988" box="[364,549,619,645]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B944A214C43DAA58FE866778FDF9FD00" box="[364,549,619,645]" italics="true" pageId="5" pageNumber="498">Diogenes viridis</emphasis>
</taxonomicName>
has the following features: shield as long as broad, left cheliped with dactyl as long as palm, fixed finger flattened, deflexed, lower margin with very small spines continued to lower margin of palm, P4 with distal spine on propodus, often also on carpus, telson longer than broad. The differences mentioned above confirmed the validity of these two species.
</paragraph>
<paragraph id="8B8F7E06C43DAA58FF2D67E9FF2BFCBC" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="498">
<emphasis id="B944A214C43DAA58FF2D67E9FEFFFC91" bold="true" box="[199,291,762,788]" pageId="5" pageNumber="498">Habitat</emphasis>
. The species was collected from muddy sandy substrate in mangrove environment, together with
<taxonomicName id="4C300585C43DAA58FA6B67E8FF2CFCBC" authorityName="Rahayu &amp; Forest" authorityYear="1995" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="tumidus">
<emphasis id="B944A214C43DAA58FA6B67E8FF2CFCBC" italics="true" pageId="5" pageNumber="498">D. tumidus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>