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<mods:title id="A647DC2010E39BEB209A877C2C13274A">Vernonieae (Asteraceae) of southern Africa: A generic disposition of the species and a study of their pollen</mods:title>
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<mods:namePart id="76366E578A3D70A4A96DB71751D34555">Robinson, Harold</mods:namePart>
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<mods:namePart id="A0C60ABE99C880AEC959864D47BBD58C">Skvarla, John J.</mods:namePart>
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<mods:namePart id="9AFB9305188405E61F01030EC747654C">Funk, Vicki A.</mods:namePart>
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<treatment id="8B7752A104CF4E2063DD8A3845A86CB2" ID-GBIF-Taxon="127870825" LSID="urn:lsid:plazi:treatment:8B7752A104CF4E2063DD8A3845A86CB2" httpUri="http://treatment.plazi.org/id/8B7752A104CF4E2063DD8A3845A86CB2" lastPageId="19" lastPageNumber="68" pageId="17" pageNumber="66" scope_family="Asteraceae">
<subSubSection id="09D518DE9FC147BE33FB004B1989C593" pageId="17" pageNumber="66" type="nomenclature">
<paragraph id="E99F9040E20F6E5D4AC84E0317C4696E" pageId="17" pageNumber="66">
<taxonomicName id="2155AA272E1215BBA514532B3ADD891D" ID-CoL="62RH4" LSID="6D197571-5723-58EC-9D9D-0BC173917171" authority="L. f." class="Magnoliopsida" family="Asteraceae" genus="Ethulia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ethulia" order="Asterales" pageId="17" pageNumber="66" phylum="Tracheophyta" rank="genus">Ethulia L.f.</taxonomicName>
<figureCitation id="292AE0D51B4837D2C91E7CBDC3B1B842" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photographs of Erlangea and Ethulia: A-B Erlangea misera S. Moore, and C-E Ethulia conyzoides L. f. subsp. conyzoides, note: Ethulia has no capillary pappus. See Appendix C for citation details." figureDoi="10.3897/phytokeys.60.6734.figure7" httpUri="https://binary.pensoft.net/fig/77479" pageId="17" pageNumber="66">
Figures 7
<normalizedToken id="FE0992E3D2E49E11F9CF56B117198C05" originalValue="CE">C-E</normalizedToken>
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<figureCitation id="9FF222C9677B0FC2D9C1840B482AFF23" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scanning electron micrographs of acetolyzed echinolophate pollen of Erlangea and sublophate pollen of Ethulia. A-C Erlangea misera (Oliv. &amp; Hiern) S. Moore. A Poral view B Near polar view C Grain fragment. D-F Ethulia conzyoides L. f. D Equatorial view, showing comparatively blunt spines E Lateral view F Grain fragment. (D-F Funk 12708 G Petelot 4047 H Lewis 6025; Ethulia views from Robinson and Skvarla (2010 ))." figureDoi="10.3897/phytokeys.60.6734.figure8" httpUri="https://binary.pensoft.net/fig/77480" pageId="17" pageNumber="66">
; 8
<normalizedToken id="E0D555CB4C9926B7748E18635D156541" originalValue="DF">D-F</normalizedToken>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="79737C1D7AB89DE11742BE22E0FB8C08" pageId="18" pageNumber="67" type="reference_group">
<paragraph id="94CD9E93897E30762BD42AE134849323" pageId="18" pageNumber="67">
<taxonomicName id="682C1BEE79239D56717959D026E78B7E" class="Magnoliopsida" family="Asteraceae" genus="Ethulia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ethulia" order="Asterales" pageId="18" pageNumber="67" phylum="Tracheophyta" rank="genus">
<pageBreakToken id="027B3D44190A732DF0342699C59A4A72" pageId="18" pageNumber="67" start="start">Ethulia</pageBreakToken>
</taxonomicName>
L.f. Dec. Prima Pl. Rar. Horti Upsal. 1 (1762); L.f. ex L., Sp. Pl. ed. II: 1171 (1763). - Type:
<taxonomicName id="2E83584FFE736ED0F94B6D4C1F3007C2" authorityName="L. f. subsp. conyzoides" authorityYear="1762" class="Magnoliopsida" family="Asteraceae" genus="Ethulia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ethulia conyzoides" order="Asterales" pageId="18" pageNumber="67" phylum="Tracheophyta" rank="species" species="conyzoides">
<emphasis id="BA77768CC94AF05BD72CCA131D540540" italics="true" pageId="18" pageNumber="67">Ethulia conyzoides</emphasis>
</taxonomicName>
L.f.
</paragraph>
<paragraph id="D44C80BDD63D06451CA804AEC96AB3E6" pageId="18" pageNumber="67">
<taxonomicName id="C09480005E4BE45E0CE0D2BDCEBC4C4A" class="Magnoliopsida" family="Asteraceae" genus="Hoehnelia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoehnelia" order="Asterales" pageId="18" pageNumber="67" phylum="Tracheophyta" rank="genus">Hoehnelia</taxonomicName>
Schweinf. in
<normalizedToken id="ACE55FBE8A07400D0F26D6B979665F0D" originalValue="Höhnel">Hoehnel</normalizedToken>
, Zum Rudolf-See und Stephanie-See 86 (1892). - Type:
<taxonomicName id="F5ABB76B955C075D2050502260ABA010" authorityName="Schweinf" authorityYear="1892" class="Magnoliopsida" family="Asteraceae" genus="Hoehnelia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoehnelia vernonioides" order="Asterales" pageId="18" pageNumber="67" phylum="Tracheophyta" rank="species" species="vernonioides">
<emphasis id="2322227CF0F979670A499D39A3E30B31" italics="true" pageId="18" pageNumber="67">Hoehnelia vernonioides</emphasis>
</taxonomicName>
Schweinf. in
<normalizedToken id="E78286C3DD553EB7C47627B9B8CF10EC" originalValue="Höhnel">Hoehnel</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="5C454C8FBF876D7B35F659D5AD70A978" pageId="18" pageNumber="67" type="resources">
<paragraph id="47611E8236B5BFAE6A9CE4EAA595EC87" pageId="18" pageNumber="67">Resources.</paragraph>
<paragraph id="628BE62A5E712AE0062E42A88CC1D0C3" pageId="18" pageNumber="67">
Treatment of the genus by
<bibRefCitation id="570B7C836745E4D6D494DDA4598B3F66" author="Gilbert, MG" journalOrPublisher="Kew Bulletin" pageId="50" pageNumber="99" pagination="165 - 193" publicationUrl="10.2307/4113733" refId="B11" refString="Gilbert, MG, Jeffrey, C, 1988. A revision of Ethulia (Compositae: Vernonieae). Kew Bulletin 43 (2): 165 - 193, DOI: 10.2307/4113733" title="A revision of Ethulia (Compositae: Vernonieae)." url="10.2307/4113733" volume="43" year="1988">Gilbert and Jeffrey (1988)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8ACF75E5EEF52106BAE75D561B10160" lastPageId="19" lastPageNumber="68" pageId="18" pageNumber="67" type="descriptions">
<paragraph id="289CD5997ECE53F37C4C262CA5B3DAFD" pageId="18" pageNumber="67">Descriptions.</paragraph>
<paragraph id="84C2E85C28F1A029981243B07B93154A" lastPageId="19" lastPageNumber="68" pageId="18" pageNumber="67">
Annual or short-lived perennial herbs, rarely rhizomatous; stems terete and usually striate, with broad solid pith; hairs uniseriate with erect apical cells, with glandular dots. Leaves alternate, sessile or short petiolate; blades thinly
<pageBreakToken id="C46A425284C3FA496D357F23A9BC6E41" pageId="19" pageNumber="68" start="start">herbaceous</pageBreakToken>
, ovate to linear lanceolate, base cuneate or continuous onto stem, margins subentire to serrate or dentate, apex acute to obtuse, surfaces glabrous to densely pubescent; venation pinnate with ascending secondary veins. Inflorescence terminal, corymbiform to rather cymiform, lower bracteoles a reduced foliiform, peduncular bracteoles filiform. Heads rather small, with broadly campanulate involucres; involucral bracts 15-40 in 2-3 usually subequal series; receptacle flat or slightly convex, epaleaceous. Florets 3-100 in a head, strongly exserted; corollas white or pink to purple, with glandular dots on surface, with a narrow cylindrical base, limb narrowly funnelform to narrowly campanulate; lobes lanceolate, without apical hairs; bases of anther thecae rounded, not tailed; apical appendages glabrous; style base without node; branches with sweeping hairs shortly acute. Achenes cylindrical with 2-6 usually paler ribs, sides with glandular dots, rarely with short white setulae; raphids short-oblong; pappus lacking or a coroniform rim. Chromosome number n = 10, 20 (
<bibRefCitation id="E35CB5F6EC6868C5B928F2234306A8B4" author="Pilz, GE" editor="Loeve, A" journalOrPublisher="Taxon" pageId="51" pageNumber="100" pagination="347 - 367" refId="B34" refString="Pilz, GE, 1980. . In: Loeve, A, Ed., IOPB Chromosome number reports LXVII. Taxon 29: 347 - 367" volume="29" volumeTitle="IOPB Chromosome number reports LXVII." year="1980">Pilz 1980</bibRefCitation>
;
<bibRefCitation id="7C52ABD49BB5BE1B732400D8D4E23506" author="Gilbert, MG" journalOrPublisher="Kew Bulletin" pageId="50" pageNumber="99" pagination="165 - 193" publicationUrl="10.2307/4113733" refId="B11" refString="Gilbert, MG, Jeffrey, C, 1988. A revision of Ethulia (Compositae: Vernonieae). Kew Bulletin 43 (2): 165 - 193, DOI: 10.2307/4113733" title="A revision of Ethulia (Compositae: Vernonieae)." url="10.2307/4113733" volume="43" year="1988">Gilbert and Jeffrey 1988</bibRefCitation>
).
</paragraph>
<paragraph id="F3DB59D651CC0F49EF97391FB8B0AB99" pageId="19" pageNumber="68">
Pollen: ca. 35
<normalizedToken id="83885C840EBD2D8A2DFBA02279C8B83D" originalValue="μm">μm</normalizedToken>
in diam. in fluid; tricolporate, sublophate, echinate, spines long; tectum continuous in intercolpi and at poles, distinctly microperforate; columellae below spines firmly attached to footlayer (Fig.
<figureCitation id="46F47C4CB44E2E53E353D799372CA356" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scanning electron micrographs of acetolyzed echinolophate pollen of Erlangea and sublophate pollen of Ethulia. A-C Erlangea misera (Oliv. &amp; Hiern) S. Moore. A Poral view B Near polar view C Grain fragment. D-F Ethulia conzyoides L. f. D Equatorial view, showing comparatively blunt spines E Lateral view F Grain fragment. (D-F Funk 12708 G Petelot 4047 H Lewis 6025; Ethulia views from Robinson and Skvarla (2010 ))." figureDoi="10.3897/phytokeys.60.6734.figure8" httpUri="https://binary.pensoft.net/fig/77480" pageId="19" pageNumber="68">
8
<normalizedToken id="96E6D29A1E63B46184A8B3ACE235D0D4" originalValue="DF">D-F</normalizedToken>
</figureCitation>
).
</paragraph>
<paragraph id="5495A493D65FF084FD77983D992C1212" pageId="19" pageNumber="68">
Notable secondary metabolites: 5-alkylcoumarins (
<bibRefCitation id="FDD833A6939691373E7BE0117D0A244E" author="Bohlmann, F" journalOrPublisher="Plant Systematics and Evolution" pageId="50" pageNumber="99" pagination="3 - 43" publicationUrl="10.1007/978-3-7091-6928-5_2" refId="B5" refString="Bohlmann, F, Jakupovic, J, 1990. Progress in the chemistry of the Vernonieae (Compositae). Plant Systematics and Evolution 4: 3 - 43, DOI: 10.1007/978-3-7091-6928-5_2" title="Progress in the chemistry of the Vernonieae (Compositae)." url="10.1007/978-3-7091-6928-5_2" volume="4" year="1990">Bohlmann and Jakupovic 1990</bibRefCitation>
, as
<taxonomicName id="85BC8202B59D10433B32E4D5E7B11A2E" authorityName="L. f. subsp. conyzoides" authorityYear="1762" class="Magnoliopsida" family="Compositae" genus="Ethulia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ethulia conyzoides" order="Asterales" pageId="19" pageNumber="68" phylum="Tracheophyta" rank="species" species="conyzoides">
<emphasis id="121DA4B0423AA17E40BE8ED6532D496E" italics="true" pageId="19" pageNumber="68">Ethulia conyzoides</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>