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<document id="4C99F8A8A69C29BEE3634E262053F80F" ID-DOI="10.5281/zenodo.172627" ID-GBIF-Dataset="7968031c-5c71-43b2-b6af-63b183cbc82c" ID-ISSN="1175­5326" ID-Zenodo-Dep="172627" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459759352879" checkinUser="plazi" docAuthor="Bruce, A. J." docDate="2006" docId="727A87E6FF837B34D535FA80FC86FC2F" docLanguage="en" docName="zt01224p022.pdf" docOrigin="Zootaxa 1224" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Periclimenaeus zanzibaricus Bruce 1969" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="8E43FF9EFF8D7B27D43DFFACFFFDFF99" masterDocTitle="Periclimenaeus nielbrucei sp. nov. (Crustacea: Decapoda: Pontoniinae), a new sponge associate from the Capricorn Islands, Queensland, with notes on related Periclimenaeus species" masterLastPageNumber="22" masterPageNumber="1" pageNumber="15" updateTime="1698214615013" updateUser="plazi">
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<mods:title id="E8D5340D2BE3876C5EC2BD08A2C965BF">Periclimenaeus nielbrucei sp. nov. (Crustacea: Decapoda: Pontoniinae), a new sponge associate from the Capricorn Islands, Queensland, with notes on related Periclimenaeus species</mods:title>
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<mods:namePart id="3B93E93A62FE3C1E9E7DC7DCD76227A4">Bruce, A. J.</mods:namePart>
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<mods:date id="0A3FF0C6407294D3D42E17D63733FD33">2006</mods:date>
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<treatment id="727A87E6FF837B34D535FA80FC86FC2F" ID-DOI="http://doi.org/10.5281/zenodo.6258734" ID-GBIF-Taxon="119344468" ID-Zenodo-Dep="6258734" LSID="urn:lsid:plazi:treatment:727A87E6FF837B34D535FA80FC86FC2F" httpUri="http://treatment.plazi.org/id/727A87E6FF837B34D535FA80FC86FC2F" lastPageId="19" lastPageNumber="20" pageId="14" pageNumber="15">
<subSubSection id="B2C9657BFF837B29D535FA80FE80FAF0" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="FA6C36F0FF837B29D535FA80FD03FADF" blockId="14.[264,766,1324,1385]" box="[264,766,1324,1350]" pageId="14" pageNumber="15">
<heading id="A124819CFF837B29D535FA80FD03FADF" bold="true" box="[264,766,1324,1350]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="3DD34D73FF837B29D535FA80FD03FADF" ID-CoL="3dc90072-c9e6-4a4b-bd09-744858ec4593" authority="Bruce, 1969" authorityName="Bruce" authorityYear="1969" box="[264,766,1324,1350]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF837B29D535FA80FD03FADF" bold="true" box="[264,766,1324,1350]" pageId="14" pageNumber="15">
<emphasis id="C8A7EAE2FF837B29D535FA80FD9FFADF" bold="true" box="[264,610,1324,1350]" italics="true" pageId="14" pageNumber="15">Periclimenaeus zanzibaricus</emphasis>
<bibRefCitation id="9E424B01FF837B29D654FA80FD03FADF" author="Bruce" box="[617,766,1324,1350]" pageId="14" pageNumber="21" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologisches Mededelingen, Leiden, 43 (20), 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FA6C36F0FF837B29D535FAE3FE80FAF0" blockId="14.[264,766,1324,1385]" box="[264,381,1359,1385]" pageId="14" pageNumber="15">
(
<figureCitation id="62E82A75FF837B29D52CFAE3FE88FAF0" box="[273,373,1359,1385]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="16.[312,416,1387,1411]" captionStartId-1="18.[312,416,1712,1736]" captionTargetBox-0="[320,1249,335,1373]" captionTargetBox-1="[321,1268,551,1691]" captionTargetId-0="figure@16.[318,1268,296,1373]" captionTargetId-1="figure@18.[318,1268,490,1729]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionText-0="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." captionText-1="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri-0="https://zenodo.org/record/172635/files/figure.png" httpUri-1="https://zenodo.org/record/172636/files/figure.png" pageId="14" pageNumber="15">Fig. 8–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2C9657BFF837B29D535FA33FBE3FA2C" box="[264,1054,1438,1462]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="FA6C36F0FF837B29D535FA33FBE3FA2C" blockId="14.[264,1054,1438,1462]" box="[264,1054,1438,1462]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="DAC311DEFF837B29D535FA33FBE3FA2C" box="[264,1054,1438,1462]" pageId="14" pageNumber="15">
<taxonomicName id="3DD34D73FF837B29D535FA33FD01FA2C" ID-CoL="3dc90072-c9e6-4a4b-bd09-744858ec4593" authority="Bruce, 1969: 174" authorityName="Bruce" authorityPageNumber="174" authorityYear="1969" box="[264,764,1438,1462]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF837B29D535FA33FDC6FA2F" box="[264,571,1439,1462]" italics="true" pageId="14" pageNumber="15">Periclimenaeus zanzibaricus</emphasis>
<treatmentCitation id="7B7210E1FF837B29D67FFA32FD01FA2C" author="Bruce" box="[578,764,1438,1461]" page="174" pageId="14" pageNumber="15" year="1969">
<bibRefCitation id="9E424B01FF837B29D67FFA32FD35FA2C" author="Bruce" box="[578,712,1438,1461]" pageId="14" pageNumber="21" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologisches Mededelingen, Leiden, 43 (20), 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
: 174
</treatmentCitation>
</taxonomicName>
–175. — 
<treatmentCitation id="7B7210E1FF837B29D763FA32FBE7FA2C" author="Bruce" box="[862,1050,1438,1461]" page="474" pageId="14" pageNumber="15" year="1976">
<bibRefCitation id="9E424B01FF837B29D763FA32FC19FA2C" author="Bruce" box="[862,996,1438,1461]" pageId="14" pageNumber="21" refString="Bruce, A. J. (1976) A Synopsis of the Pontoniinid Shrimp Fauna of central East Africa. Journal of the Marine Biological Association of India, 16 (2) (1974), 462 - 490." type="journal article" year="1976">Bruce, 1976</bibRefCitation>
: 474
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2C9657BFF837B29D535FA4BFE8BF880" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="FA6C36F0FF837B29D535FA4BFE1BF999" blockId="14.[264,1323,1511,1817]" box="[264,486,1511,1536]" pageId="14" pageNumber="15">
<emphasis id="C8A7EAE2FF837B29D535FA4BFE1BF999" box="[264,486,1511,1536]" italics="true" pageId="14" pageNumber="15">Material examined</emphasis>
</paragraph>
<paragraph id="FA6C36F0FF837B29D505F9A3FE8BF880" blockId="14.[264,1323,1511,1817]" pageId="14" pageNumber="15">
(1) 1 ov. Ψ 
<emphasis id="C8A7EAE2FF837B29D5F3F9A2FDC4F9B1" bold="true" box="[462,569,1550,1576]" pageId="14" pageNumber="15">allotype,</emphasis>
<collectionCode id="9CC2AE35FF837B29D679F9A3FD61F9B0" LSID="urn:lsid:biocol.org:col:34992" box="[580,668,1551,1577]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15">RMNH</collectionCode>
D.51673; 4, 6 ovig. Ψ, 1 Ψ, 
<specimenCount id="ECD5FD79FF837B29D01AF9A3FB8DF9B0" box="[1063,1136,1551,1577]" pageId="14" pageNumber="15" type="generic">1 juv.</specimenCount>
, Uroa, Unguja, 
<collectingCountry id="82C47660FF837B29D535F99BFE8FF9C8" box="[264,370,1591,1617]" name="Tanzania" pageId="14" pageNumber="15">Zanzibar</collectingCountry>
, 
<date id="8E6D1030FF837B29D542F99BFDC5F9C8" box="[383,568,1591,1617]" pageId="14" pageNumber="15" value="1960-03-21">21 March 1960</date>
, AJB 289A, AJB 289/1­12, unidentified sponge, shore pools at neap tide, 
<typeStatus id="25688852FF837B29D5BDF9F3FE0EF9E0" box="[384,499,1631,1657]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, 
<collectionCode id="9CC2AE35FF837B29D5C3F9F3FDABF9E0" LSID="urn:lsid:biocol.org:col:34992" box="[510,598,1631,1657]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15">RMNH</collectionCode>
D. 51674; 1 spm, 
<typeStatus id="25688852FF837B29D713F9F3FC6BF9E0" box="[814,918,1631,1657]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
, 
<collectionCode id="9CC2AE35FF837B29D79DF9F3FC05F9E0" box="[928,1016,1631,1657]" pageId="14" pageNumber="15">BMNH</collectionCode>
2006. 407–408. (2) 1 ɗ, 1 ovig. Ψ, 2 juv, CLs 1.8, 2.5, 1.5, 
<quantity id="3D2B9B15FF837B29D6A3F92BFD03F938" box="[670,766,1671,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="mm" value="1.5">1.5 mm</quantity>
, stn AJB/59, Myora, North Stradbroke Island, Queensland, 
<date id="8E6D1030FF837B29D599F903FDC5F950" box="[420,568,1711,1737]" pageId="14" pageNumber="15" value="1969-07-01">1 July 1969</date>
, QM W27989. (3) 1 ɗ, 1 ov. Ψ, CLs 1.9, 
<quantity id="3D2B9B15FF837B29D07AF903FB55F950" box="[1095,1192,1711,1737]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="15" unit="mm" value="3.3">3.3 mm</quantity>
, 
<collectingCountry id="82C47660FF837B29D08AF903FAD7F950" box="[1207,1322,1711,1737]" name="Kenya" pageId="14" pageNumber="15">Mombasa</collectingCountry>
Island, 
<collectingCountry id="82C47660FF837B29D55DF97BFE4DF968" box="[352,432,1751,1777]" name="Kenya" pageId="14" pageNumber="15">Kenya</collectingCountry>
, 
<date id="8E6D1030FF837B29D583F97BFD5EF968" box="[446,675,1751,1777]" pageId="14" pageNumber="15" value="1973-12-11">11 December 1973</date>
, AJB 2086, seaward reef, reef pools, at 
<collectionCode id="9CC2AE35FF837B29D0B4F97BFB35F968" box="[1161,1224,1751,1777]" pageId="14" pageNumber="15">LWS</collectionCode>
, 
<collectionCode id="9CC2AE35FF837B29D0EBF97BFAD6F968" LSID="urn:lsid:biocol.org:col:34871" box="[1238,1323,1751,1777]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">USNM</collectionCode>
1086753.
</paragraph>
</subSubSection>
<subSubSection id="B2C9657BFF827B28D535FEB0FCA1FCA7" pageId="15" pageNumber="16" type="discussion">
<paragraph id="FA6C36F0FF827B28D535FEB0FE93FEAC" blockId="15.[264,1323,284,830]" box="[264,366,284,309]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D535FEB0FE93FEAC" box="[264,366,284,309]" italics="true" pageId="15" pageNumber="16">Remarks</emphasis>
</paragraph>
<paragraph id="FA6C36F0FF827B28D505FEE8FCA1FCA7" blockId="15.[264,1323,284,830]" pageId="15" pageNumber="16">
<bibRefCitation id="9E424B01FF827B28D505FEE8FE28FEC7" author="Bruce" box="[312,469,324,350]" pageId="15" pageNumber="21" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologisches Mededelingen, Leiden, 43 (20), 253 - 278." type="journal article" year="1969">Bruce (1969)</bibRefCitation>
provided only an un­illustrated preliminary description of this species, which has not been further reported since that date. The 
<typeStatus id="25688852FF827B28D7FDFEC0FBDBFE1F" box="[960,1062,364,390]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
was deposited in the Rijksmuseun van Natuurlijke Historie collection. Further specimens collected simultaneously with the 
<typeStatus id="25688852FF827B28D609FE10FD9AFE4F" box="[564,615,444,470]" pageId="15" pageNumber="16">type</typeStatus>
and other material has now been studied. 
<taxonomicName id="3DD34D73FF827B28D049FE10FE61FE64" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF827B28D049FE10FE61FE64" italics="true" pageId="15" pageNumber="16">Periclimenaeus zanzibaricus</emphasis>
</taxonomicName>
was initially considered most closely related to 
<taxonomicName id="3DD34D73FF827B28D7CCFE48FADBFE67" authority="Nobili 1904" authorityName="Nobili" authorityYear="1904" box="[1009,1318,484,510]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rhodope">
<emphasis id="C8A7EAE2FF827B28D7CCFE48FB8EFE64" box="[1009,1139,484,509]" italics="true" pageId="15" pageNumber="16">P. rhodope</emphasis>
(
<bibRefCitation id="9E424B01FF827B28D0BAFE48FAE3FE67" author="Nobili" box="[1159,1310,484,510]" pageId="15" pageNumber="22" refString="Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Museum d'Histoire naturelle, Paris, 10, 228 - 238." type="journal article" year="1904">Nobili 1904</bibRefCitation>
)
</taxonomicName>
, which at that time was known with certainty only from Nobili’s description and figures, the species referred to 
<taxonomicName id="3DD34D73FF827B28D626FD98FD66FDD4" box="[539,667,564,589]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rhodope">
<emphasis id="C8A7EAE2FF827B28D626FD98FD66FDD4" box="[539,667,564,589]" italics="true" pageId="15" pageNumber="16">P. rhodope</emphasis>
</taxonomicName>
by 
<bibRefCitation id="9E424B01FF827B28D6F1FD98FC75FDD7" author="Holthuis" box="[716,904,564,590]" pageId="15" pageNumber="22" refString="Holthuis, L. B. (1952) The Decapoda of the Siboga Expedition. Part XI. The Palaemonidae collected by the Siboga and Snellius Expeditions with remarks on other species. II. Subfamily Pontoniinae. Siboga Expedition Monograph, 39 a 10, 1 - 252." type="journal article" year="1952">Holthuis (1952)</bibRefCitation>
having been re­named 
<taxonomicName id="3DD34D73FF827B28D098FD98FE66FDEF" authority="Bruce, 1969" authorityName="Bruce" authorityYear="1969" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="C8A7EAE2FF827B28D098FD98FAD7FDD4" box="[1189,1322,564,589]" italics="true" pageId="15" pageNumber="16">P. holthuisi</emphasis>
<bibRefCitation id="9E424B01FF827B28D535FDF0FE66FDEF" author="Bruce" box="[264,411,604,630]" pageId="15" pageNumber="21" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologisches Mededelingen, Leiden, 43 (20), 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
</taxonomicName>
(see above). Further details of 
<taxonomicName id="3DD34D73FF827B28D722FDF0FC5DFDEC" box="[799,928,604,629]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rhodope">
<emphasis id="C8A7EAE2FF827B28D722FDF0FC5DFDEC" box="[799,928,604,629]" italics="true" pageId="15" pageNumber="16">P. rhodope</emphasis>
</taxonomicName>
were provided by 
<bibRefCitation id="9E424B01FF827B28D0B6FDF0FAD4FDEF" author="Bruce" box="[1163,1321,604,630]" pageId="15" pageNumber="21" refString="Bruce, A. J. (1974) Observations upon some specimens of the genus Periclimenaeus Borradaile (Decapoda Natantia, Pontoniinae) originally described by G. Nobili. Bulletin du Museum national d'Histoire naturelle, Paris, (3), no. 258, Zool. 180, 1557 - 1583." type="journal article" year="1974">Bruce (1974)</bibRefCitation>
after the re­examination of Nobili’s 
<typeStatus id="25688852FF827B28D687FD28FD11FD07" box="[698,748,644,670]" pageId="15" pageNumber="16">type</typeStatus>
material held in the collections of the Muséum National d’Histoire Naturelle. The preliminary description of 
<taxonomicName id="3DD34D73FF827B28D034FD00FB3DFD5C" box="[1033,1216,684,709]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF827B28D034FD00FB3DFD5C" box="[1033,1216,684,709]" italics="true" pageId="15" pageNumber="16">P. zanzibaricus</emphasis>
</taxonomicName>
may be amplified by the following details from dissected specimens QM W27989 (289/7 and 289/ 8). The allotype male of 
<taxonomicName id="3DD34D73FF827B28D66EFD50FCF1FC8C" box="[595,780,764,789]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF827B28D66EFD50FCF1FC8C" box="[595,780,764,789]" italics="true" pageId="15" pageNumber="16">P. zanzibaricus</emphasis>
</taxonomicName>
is held in the collection of the Nationaal Natuurhistorisch Museum, Leiden, Crust.D.25615.
</paragraph>
</subSubSection>
<subSubSection id="B2C9657BFF827B34D535FCD8FC86FC2F" lastPageId="19" lastPageNumber="20" pageId="15" pageNumber="16" type="description">
<paragraph id="FA6C36F0FF827B28D535FCD8FE6FFC14" blockId="15.[264,1324,884,1903]" box="[264,402,884,909]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D535FCD8FE6FFC14" box="[264,402,884,909]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
</paragraph>
<paragraph id="FA6C36F0FF827B28D505FC30FC8FFBE5" blockId="15.[264,1324,884,1903]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505FC30FE66FC2C" box="[312,411,924,949]" italics="true" pageId="15" pageNumber="16">Rostrum</emphasis>
(
<figureCitation id="62E82A75FF827B28D596FC30FE12FC2F" box="[427,495,924,950]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
B) slender, about 5.5.times longer than central depth, about 0. 4 of CL, reaching to distal margin of intermediate antennular segment, tip slightly up­turned, dentition 6­7/1, (one small specimen with 5/1, one with 6/0, but tip appears damaged), dorsal teeth all anterior to orbital margin, broadly acute, with 2–3 interdental setae, ventral margin straight, ventral tooth well developed, small, acute, projecting horizontally. Supraorbital tubercle well developed, strongly acute.
</paragraph>
<paragraph id="FA6C36F0FF827B28D505FB26FCBAFB6D" blockId="15.[264,1324,884,1903]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505FB26FE56FB3A" box="[312,427,1162,1187]" italics="true" pageId="15" pageNumber="16">Carapace</emphasis>
(
<figureCitation id="62E82A75FF827B28D5FCFB26FDF7FB3D" box="[449,522,1162,1188]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
A) with antennal spine robust, very acute, slightly postmarginal. Inferior orbital angle (
<figureCitation id="62E82A75FF827B28D629FB1EFDA6FB55" box="[532,603,1202,1228]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
C) well developed, broadly rounded, reflected below lateral orbital rim, with small rounded distomedial lobe.
</paragraph>
<paragraph id="FA6C36F0FF827B28D505FAADFC6CFA82" blockId="15.[264,1324,884,1903]" box="[312,913,1281,1307]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505FAADFDB5FA83" box="[312,584,1281,1306]" italics="true" pageId="15" pageNumber="16">First abdominal tergite</emphasis>
without anteromedian lobe.
</paragraph>
<paragraph id="FA6C36F0FF827B28D505FA85FE32F9A8" blockId="15.[264,1324,884,1903]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505FA85FE7CFADB" box="[312,385,1321,1346]" italics="true" pageId="15" pageNumber="16">Telson</emphasis>
(
<figureCitation id="62E82A75FF827B28D5AEFA85FE2AFADA" box="[403,471,1321,1347]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
K) about 0.6 of CL, 2.5 times longer than anterior width, lateral margins convex, posteriorly convergent, with large submarginal dorsal spines at 0.13 and 0.22 of length, anterior spines 0.2 of telson length, anterior spines 1.18 times posterior spine length, posterior margin rounded with small acute median point, posterior spines (
<figureCitation id="62E82A75FF827B28D0F7FA0CFAF0FA23" box="[1226,1293,1440,1466]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
L) well developed, lateral spines subdorsal, about 0.33 of intermediate spine length, intermediate spine robust, about 0.25 of telson length, submedian spines more slender, densely setulose.
</paragraph>
<paragraph id="FA6C36F0FF827B28D505F9ECFD4EF949" blockId="15.[264,1324,884,1903]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505F9ECFDCFF9C0" box="[312,562,1600,1625]" italics="true" pageId="15" pageNumber="16">Antennular peduncle</emphasis>
(
<figureCitation id="62E82A75FF827B28D67AF993FD6DF9C0" box="[583,656,1599,1625]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
D) with proximal segment of the peduncle with acute stylocerite reaching to about 0.5 of segment length, with small ventromedial tooth at similar level; anterolateral lobe (
<figureCitation id="62E82A75FF827B28D6ABF923FD22F930" box="[662,735,1679,1705]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
E) acutely produced as triangular process rather than rounded lobe with lateral tooth.
</paragraph>
<paragraph id="FA6C36F0FF827B28D505F972FC36F8F6" blockId="15.[264,1324,884,1903]" pageId="15" pageNumber="16">
<emphasis id="C8A7EAE2FF827B28D505F972FE67F96E" box="[312,410,1758,1783]" italics="true" pageId="15" pageNumber="16">Antenna</emphasis>
(
<figureCitation id="62E82A75FF827B28D591F972FE0FF961" box="[428,498,1758,1784]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
F) with subcylindrical carpocerite, about 2.5 times longer than wide, reaching to about 7.2 of scaphocerite length; scaphocerite (
<figureCitation id="62E82A75FF827B28D787F8AAFC00F8B9" box="[954,1021,1798,1824]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
G) about 4.3 times longer than wide, broadly rounded distally, with very robust distolateral tooth at 0.85 of length, about 0.16 of scaphocerite length and far exceeding lamella.
</paragraph>
<caption id="AEAC6678FF9D7B37D505FAC7FE26F997" httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="16" pageNumber="17" targetBox="[320,1249,335,1373]" targetPageId="16">
<paragraph id="FA6C36F0FF9D7B37D505FAC7FE26F997" blockId="16.[264,1325,1387,1550]" pageId="16" pageNumber="17">
<emphasis id="C8A7EAE2FF9D7B37D505FAC7FE47FA1A" bold="true" box="[312,442,1387,1411]" pageId="16" pageNumber="17">FIGURE 8.</emphasis>
<taxonomicName id="3DD34D73FF9D7B37D5FCFAC0FC7DFA1A" authority="Bruce, 1969" authorityName="Bruce" authorityYear="1969" box="[449,896,1388,1411]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF9D7B37D5FCFAC0FD09FA1A" box="[449,756,1388,1411]" italics="true" pageId="16" pageNumber="17">Periclimenaeus zanzibaricus</emphasis>
Bruce, 1969
</taxonomicName>
, ovigerous female paratype, 289/7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod.
</paragraph>
</caption>
<paragraph id="FA6C36F0FF9D7B37D505F9EDFE65F91A" blockId="16.[264,1323,1601,1868]" pageId="16" pageNumber="17">
<emphasis id="C8A7EAE2FF9D7B37D505F9EDFE99F9C3" box="[312,356,1601,1626]" italics="true" pageId="16" pageNumber="17">Eye</emphasis>
(
<figureCitation id="62E82A75FF9D7B37D548F9EDFE47F9C2" box="[373,442,1601,1627]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
H) with hemispherical cornea, well pigmented, with diameter about 0.8 of stalk length.
</paragraph>
<paragraph id="FA6C36F0FF9D7B37D505F93DFE88F965" blockId="16.[264,1323,1601,1868]" pageId="16" pageNumber="17">
<emphasis id="C8A7EAE2FF9D7B37D505F93DFE3CF933" box="[312,449,1681,1706]" italics="true" pageId="16" pageNumber="17">Mouthparts</emphasis>
similar to those of 
<taxonomicName id="3DD34D73FF9D7B37D688F93DFCB1F933" box="[693,844,1681,1706]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nielbrucei">
<emphasis id="C8A7EAE2FF9D7B37D688F93DFCB1F933" box="[693,844,1681,1706]" italics="true" pageId="16" pageNumber="17">P. nielbrucei</emphasis>
</taxonomicName>
, but incisor process (
<figureCitation id="62E82A75FF9D7B37D068F93DFB60F932" box="[1109,1181,1681,1707]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="16" pageNumber="17">
Fig. 
<date id="8E6D1030FF9D7B37D0B2F93DFB60F932" box="[1167,1181,1681,1707]" pageId="16" pageNumber="17">8</date>
</figureCitation>
I) obliquely rounded distally with about 12 long acute teeth, largest laterally and of decreasing size medially.
</paragraph>
<paragraph id="FA6C36F0FF9D7B36D505F8A6FBC7FE67" blockId="16.[264,1323,1601,1868]" lastBlockId="17.[264,1325,284,1870]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="C8A7EAE2FF9D7B37D505F8A6FE13F8BA" box="[312,494,1802,1827]" italics="true" pageId="16" pageNumber="17">First pereiopod</emphasis>
(
<figureCitation id="62E82A75FF9D7B37D5C3F8A6FDB9F8BD" box="[510,580,1802,1828]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
A) slender, reaching to exceed carpocerite by carpus and chela, chela (
<figureCitation id="62E82A75FF9D7B37D566F89EFE5BF8D5" box="[347,422,1842,1868]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
B) with palm about 3.0 times longer than wide, oval in section, slightly swollen proximally, with fingers (
<figureCitation id="62E82A75FF9C7B36D690FEB0FD05FEAF" box="[685,760,284,310]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
C) about 0.45 of palm length, broad, distally rounded, with entire lateral cutting edges, tips with articulated ungues, flanked by small teeth; with numerous strong setae; carpus about 0.85 of merus length,.6.5 times longer than distal width, tapering uniformly proximally; merus 1.1 times carpus length, 8.5 times longer than maximal width; ischium slightly shorter than chela, about 2.6 times longer than width; basis and coxa normal, coxa without distoventral process.
</paragraph>
<paragraph id="FA6C36F0FF9C7B36D505FDA0FB8AFDBF" blockId="17.[264,1325,284,1870]" box="[312,1143,524,550]" pageId="17" pageNumber="18">
<emphasis id="C8A7EAE2FF9C7B36D505FDA0FDE8FDBC" box="[312,533,524,549]" italics="true" pageId="17" pageNumber="18">Second pereiopods</emphasis>
well developed, rather similar and slightly unequal.
</paragraph>
<paragraph id="FA6C36F0FF9C7B36D505FD98FCE1FC44" blockId="17.[264,1325,284,1870]" pageId="17" pageNumber="18">
<emphasis id="C8A7EAE2FF9C7B36D505FD98FDA1FDD4" box="[312,604,564,589]" italics="true" pageId="17" pageNumber="18">Major second pereiopod</emphasis>
with chela (
<figureCitation id="62E82A75FF9C7B36D6C8FD98FCC0FDD7" box="[757,829,564,590]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
D) about 1.10 of CL; with palm of chela subcylindrical, oval in section, twice as long as deep, dorsal and ventral margins subparallel, with strongly marked rounded shoulder dorsally over base of dactylus; fingers (
<figureCitation id="62E82A75FF9C7B36D52CFD00FEAAFD5F" box="[273,343,684,710]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
E,F) 0.5 of palm length, dactylus about 0.6 of palm length, slender, about 3.0 times longer than maximal depth, compressed, distally up­turned, dorsal margin smoothly convex, with stout acute tip distally, cutting edge broadly convex with inconspicuous low thickening centrally over one third of length, distal third entire, not sharp, non­denticulate, fixed finger about 1.7 times longer than basal width, 0.85 of dactylus length, deeply grooved with small shallow fossa with thickened ventral flange proximally, medial margin with large acute tooth proximally, fingers with numerous long simple setae; carpus, merus, and proximal segments similar to 
<emphasis id="C8A7EAE2FF9C7B36D6ABFC68FCE1FC44" box="[662,796,964,989]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="3DD34D73FF9C7B36D6ABFC68FCE5FC44" box="[662,792,964,989]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rhodope">P. rhodope</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="FA6C36F0FF9C7B36D505FC40FC30FAF4" blockId="17.[264,1325,284,1870]" pageId="17" pageNumber="18">
<emphasis id="C8A7EAE2FF9C7B36D505FC40FDAAFB9C" box="[312,599,1004,1029]" italics="true" pageId="17" pageNumber="18">Minor second pereiopod</emphasis>
chela (
<figureCitation id="62E82A75FF9C7B36D691FC40FD0CFB9F" box="[684,753,1004,1030]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
H) similar, about 0.8 of major chela length; with palm of chela subcylindrical, oval in section, twice as long as deep, dorsal and ventral margins sub­parallel, with strongly marked rounded shoulder dorsally over base of dactylus, dactylus (
<figureCitation id="62E82A75FF9C7B36D5D7FBC8FDD2FBE7" box="[490,559,1124,1150]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">
Fig. 
<date id="8E6D1030FF9C7B36D61DFBC8FDD2FBE7" box="[544,559,1124,1150]" pageId="17" pageNumber="18">9</date>
</figureCitation>
I) about 0.58 of palm length, slender, about 3.0 times longer than maximal depth depth, compressed, distally in­turned, dorsal margin smoothly convex, with stout acute tip distally, cutting edge broadly convex with entire, sharp, nondenticulate, thickening posteriorly, fixed finger about 1.8 times longer than basal width, 0.95 of dactylus length, grooved with small depression with thickened edge proximally, medial margin with low acute tooth proximally, fingers with numerous long simple setae; carpus, merus, and proximal segments similar to 
<emphasis id="C8A7EAE2FF9C7B36D77AFAF8FC30FAF4" box="[839,973,1364,1389]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="3DD34D73FF9C7B36D77AFAF8FC34FAF4" box="[839,969,1364,1389]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rhodope">P. rhodope</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="FA6C36F0FF9C7B35D505FAD0FE94FE4F" blockId="17.[264,1325,284,1870]" lastBlockId="18.[264,1325,284,470]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="C8A7EAE2FF9C7B36D505FAD0FDB0FA0C" box="[312,589,1404,1429]" italics="true" pageId="17" pageNumber="18">Ambulatory pereiopods</emphasis>
are generally similar to those of 
<emphasis id="C8A7EAE2FF9C7B36D7DDFAD0FB96FA0C" box="[992,1131,1404,1429]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="3DD34D73FF9C7B36D7DDFAD0FB99FA0C" box="[992,1124,1404,1429]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rhodope">P. rhodope</taxonomicName>
:
</emphasis>
third pereiopod with dactylus (
<figureCitation id="62E82A75FF9C7B36D58AFA08FE00FA27" box="[439,509,1444,1470]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
J) about 0.3 of propod length, unguis (
<figureCitation id="62E82A75FF9C7B36D7FBFA08FBF1FA27" box="[966,1036,1444,1470]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
K) well demarcated, 3.0 times longer than basal width, 0.45 of dorsal corpus length, moderately curved, concave ventral margin with 5 small acute teeth, corpus 2.1 times basal width, dorsal margin feebly convex, ventral margin with strong, broad acute distoventral accessory tooth, about 0.33 of unguis length, ventral margin feebly convex with 10 small denticles, central denticles largest, most acute, distally directed; propod (
<figureCitation id="62E82A75FF9C7B36D70EF9C0FC8FF91F" box="[819,882,1644,1670]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="17" pageNumber="18">Fig.9</figureCitation>
J) about 0.4 of CL, 5.0 times longer than proximal depth, tapering slightly distally, with 2 strong distoventral spines, 7 equally spaced ventral spines, most proximal ventral spine with 2 smaller adjacent accessory spines; proximal segments as in 
<emphasis id="C8A7EAE2FF9C7B36D6A8F948FCE3F964" box="[661,798,1764,1789]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="3DD34D73FF9C7B36D6A8F948FCE7F964" box="[661,794,1764,1789]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rhodope">P. rhodope</taxonomicName>
.
</emphasis>
Fourth and fifth pereiopods similar; fourth propod with pair of distoventral and 4 ventral spines, 1 accessory spine; fifth propod with numerous setae distally, 1 distoventral spine, 3 ventral spines; dactyls similar to third, slightly more slender, denticulation slightly reduced, fifth dactyl not elongate. Pleopods and uropods without special features, as in 
<taxonomicName id="3DD34D73FF9F7B35D739FEE8FC7FFEC4" box="[772,898,324,349]" class="Malacostraca" family="Palaemonidae" genus="Periclimeneus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rhodope">
<emphasis id="C8A7EAE2FF9F7B35D739FEE8FC7FFEC4" box="[772,898,324,349]" italics="true" pageId="18" pageNumber="19">P. rhodope</emphasis>
</taxonomicName>
, exopod of uropod with small acute distolateral tooth (
<figureCitation id="62E82A75FF9F7B35D5D5FEC0FDCCFE1F" box="[488,561,364,390]" captionStart="FIGURE 8" captionStartId="16.[312,416,1387,1411]" captionTargetBox="[320,1249,335,1373]" captionTargetId="figure@16.[318,1268,296,1373]" captionTargetPageId="16" captionText="FIGURE 8. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female paratype, 289 / 7. A, carapace and rostrum. B, rostrum. C, left inferior orbital angle, dorsal. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, same, scaphocerite. H, eye, dorsal. I. mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, distolateral exopod." httpUri="https://zenodo.org/record/172635/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
M) with strong articulated spine separated by small gap. Ova numerous, of normal size, length about 
<quantity id="3D2B9B15FF9F7B35D6E6FE38FCC4FE37" box="[731,825,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
. Largest male, CL 
<quantity id="3D2B9B15FF9F7B35D025FE38FB88FE37" box="[1048,1141,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="19" unit="mm" value="2.7">2.7 mm</quantity>
, largest female, 
<quantity id="3D2B9B15FF9F7B35D535FE10FE98FE4F" box="[264,357,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="3.3">3.3 mm</quantity>
.
</paragraph>
<caption id="AEAC6678FF9F7B35D505F91CFB17F8B6" httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="18" pageNumber="19" targetBox="[321,1268,551,1691]" targetPageId="18">
<paragraph id="FA6C36F0FF9F7B35D505F91CFB17F8B6" blockId="18.[264,1325,1712,1839]" pageId="18" pageNumber="19">
<emphasis id="C8A7EAE2FF9F7B35D505F91CFE46F951" bold="true" box="[312,443,1712,1736]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
<taxonomicName id="3DD34D73FF9F7B35D5FEF91DFC7AF95E" authority="Bruce, 1969" authorityName="Bruce" authorityYear="1969" box="[451,903,1712,1736]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF9F7B35D5FEF91DFD05F951" box="[451,760,1713,1736]" italics="true" pageId="18" pageNumber="19">Periclimenaeus zanzibaricus</emphasis>
Bruce, 1969
</taxonomicName>
, ovigerous female, paratype, 289/7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle.
</paragraph>
</caption>
<paragraph id="FA6C36F0FF9E7B34D505FEB0FC08FE67" blockId="19.[264,1323,284,950]" pageId="19" pageNumber="20">
The adult Myora specimens (QM W. 27989) also have a rostral dentition of 7/1 and exactly resemble the 
<collectingCountry id="82C47660FF9E7B34D630FEE8FD88FEC7" box="[525,629,324,350]" name="Tanzania" pageId="19" pageNumber="20">Zanzibar</collectingCountry>
specimens except that the male major second pereiopod dactyl shows an elongated thickening, corresponding to the molar process, extending along almost the whole length of the dactylar cutting edge (
<figureCitation id="62E82A75FF9E7B34D7EBFE38FBE0FE37" box="[982,1053,404,430]" captionStart="FIGURE 9" captionStartId="18.[312,416,1712,1736]" captionTargetBox="[321,1268,551,1691]" captionTargetId="figure@18.[318,1268,490,1729]" captionTargetPageId="18" captionText="FIGURE 9. Periclimenaeus zanzibaricus Bruce, 1969, ovigerous female, paratype, 289 / 7. A, first pereiopod. B, same, chela. C, same, fingers. D, antennule. E, same, proximal segment, distolateral angle. F, antenna. G, scaphocerite. H, eye. I, mandible, incisor process. J, third pereiopod, dactyl. K, telson. L, same, posterior spines. M, uropod, exopod, distolateral angle." httpUri="https://zenodo.org/record/172636/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
L). This feature is less marked in the female, which more closely resembles the Indian Ocean material. The inferior orbital angles in the two lots of specimens are identical.
</paragraph>
<paragraph id="FA6C36F0FF9E7B34D505FDA0FC86FC2F" blockId="19.[264,1323,284,950]" pageId="19" pageNumber="20">
Although not mentioned in the original diagnosis of the genus (
<bibRefCitation id="9E424B01FF9E7B34D01BFDA0FB0CFDBF" author="Borradaile" box="[1062,1265,524,550]" pageId="19" pageNumber="21" refString="Borradaile, L. A. (1915) Notes on Carides. Annals and Magazine of Natural History, (8) 15, 205 - 213." type="journal article" year="1915">Borradaile, 1915</bibRefCitation>
), the most characteristic feature of the genus 
<taxonomicName id="3DD34D73FF9E7B34D6C7FD98FC4DFDD4" box="[762,944,564,589]" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="C8A7EAE2FF9E7B34D6C7FD98FC4DFDD4" box="[762,944,564,589]" italics="true" pageId="19" pageNumber="20">Periclimenaeus</emphasis>
</taxonomicName>
is “The dactylus of the larger second pereiopod with a molar tooth, socket in the fixed finger” (
<bibRefCitation id="9E424B01FF9E7B34D7C0FDF0FB4DFDEF" author="Holthuis" box="[1021,1200,604,630]" pageId="19" pageNumber="22" refString="Holthuis, L. B. (1993) The recent genera of the caridean and stenopodidean shrimps (Crustacea, Decapoda) with an appendix on the order Amphionidacea. Nationaal Natuurhistorisch Museum, Leiden, 1 - 328." type="book chapter" year="1993">Holthuis, 1993</bibRefCitation>
), although it may be noted that a remarkably similar sound producing mechanism is also present in some species of 
<taxonomicName id="3DD34D73FF9E7B34D5F4FD00FD20FD5F" authority="Stimpson" authorityName="Stimpson" box="[457,733,684,710]" class="Malacostraca" family="Palaemonidae" genus="Coralliocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="C8A7EAE2FF9E7B34D5F4FD00FD9AFD5C" box="[457,615,684,709]" italics="true" pageId="19" pageNumber="20">Coralliocaris</emphasis>
Stimpson
</taxonomicName>
and is well known in several alpheid genera. In 
<taxonomicName id="3DD34D73FF9E7B34D128FD00FE61FD74" class="Malacostraca" family="Palaemonidae" genus="Periclimenaeus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="C8A7EAE2FF9E7B34D128FD00FE61FD74" italics="true" pageId="19" pageNumber="20">P. zanzibaricus</emphasis>
</taxonomicName>
this mechanism is particularly poorly developed. The shapes of the fingers of the major and minor chela are also remarkably similar. This may illustrate the path by which the mechanism has evolved, from a simple thickening of the posterior part of the dactylar cutting edge opposing a longitudinal groove on that of the fixed finger. The molar process may have developed from this thickening, rather than a pre­existing tooth in this situation, as found in many other pontoniine shrimps.
</paragraph>
</subSubSection>
</treatment>
</document>