229 lines
26 KiB
XML
229 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="71E81728B37D5C0397B5B5BEFD3A3F0D" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole nitidobruna Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="4676573B-D010-4733-BB04-D4F3AB90022E" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Salata, Sebastian</mods:namePart>
|
|
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
|
|
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
|
|
<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Fisher, Brian L.</mods:namePart>
|
|
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
|
|
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2020</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>949</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>185</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
|
|
<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="165090175" LSID="urn:lsid:zoobank.org:act:4676573B-D010-4733-BB04-D4F3AB90022E" httpUri="http://treatment.plazi.org/id/71E81728B37D5C0397B5B5BEFD3A3F0D" lastPageNumber="1" pageId="0" pageNumber="1">
|
|
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName LSID="71E81728-B37D-5C03-97B5-B5BEFD3A3F0D" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna" status="sp. nov.">Pheidole nitidobruna</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">Figs 48A-F</figureCitation>
|
|
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64F</figureCitation>
|
|
<figureCitation captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Distribution. Pheidole mahamavo sp. nov. (A). Pheidole mainty sp. nov. (B). Pheidole mamiratra sp. nov. (C). Pheidole manantenina sp. nov. (D). Pheidole masoandro sp. nov. (E). Pheidole mavohavoana sp. nov. (F). Pheidole midongy sp. nov. (G). Pheidole mikros sp. nov. (H). Pheidole mivory sp. nov. (I). Pheidole nitidobruna sp. nov. (J). Pheidole parvula sp. nov. (K). Pheidole parvulogibba sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure66" httpUri="https://binary.pensoft.net/fig/432235" pageId="0" pageNumber="1">, 66J</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="type material">
|
|
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
|
|
Madagascar. • 1 major worker; Antsiranana; Makirovana forest;
|
|
<geoCoordinate degrees="14.16044" direction="south" orientation="latitude" precision="1" value="-14.16044">-14.16044</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="49.95216" direction="east" orientation="longitude" precision="1" value="49.95216">49.95216</geoCoordinate>
|
|
; alt. 550 m; 1 May 2011; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF26855; CASENT0923273 (CASC).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
|
|
• 1w.; same data as for holotype, CASENT0245024 (CASC).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0245024) and full-face view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0923273).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Minute species.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
|
HL <0.9 mm and WL <0.8 mm; head in full-face view elongate, not widening posteriorly with posterior sides slightly convex, in lateral view sub-oval and short with convex dorsal and ventral sides; body brown; sides of head with moderately sparse, moderately long, suberect to erect pilosity; entire head distinctly sculptured, medial part of frons with thick and longitudinal rugae and smooth to indistinctly rugulate interspaces, occipital lobes with dense and irregular rugae with indistinctly rugoreticulate interspaces; scape, when laid back, exceeding the midlength of head by one-fifth of its length; lateral sides of promesonotum with smooth notches; inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, lower and wider than inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity; base of first gastral tergite shagreened.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
HL <0.45 mm and WL <0.5 mm, scape, when laid back, exceeding the posterior head margin by one-fifth of its length; propodeal spines reduced to small tubercles; head relatively rectangular; body yellowish brown; head with frons and vertex smooth; mesosoma smooth.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="description">
|
|
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
|
Measurements (
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 1): HL: 0.9; HW: 0.79; SL: 0.53; EL: 0.1; WL: 0.79; PSL: 0.11; MTL: 0.46; PNW: 0.44; PTW: 0.11; PPW: 0.28; CI: 113.1; SI: 66.7; PSLI: 12.7; PPI: 37.9; PNI: 55.8; MTI: 57.4.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
|
In full-face view elongate, not widening posteriorly, with anterior and posterior sides slightly convex (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48B</figureCitation>
|
|
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately sparse, moderately long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, longitudinal, and moderately dense rugae and smooth to indistinctly rugulate interspaces, rugae directed slightly outward on posteromedial part; anterolateral sides with thick, sparse, and longitudinal rugae with smooth to indistinctly rugulate interspaces, posterolateral sides with dense, thick, and irregular rugae with distinctly rugulae interspaces. Occipital lobes with dense, irregular rugae and indistinctly rugoreticulate interspaces. Area posterolateral from eyes with weaker sculpture, rugoreticulate. Gena with relatively sparse, thick, and longitudinal rugae and smooth interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48B, D</figureCitation>
|
|
). Inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, lower and wider than inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
|
|
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64F</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
|
In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines short, with wide base and acute apex; humeral area produced (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D</figureCitation>
|
|
). Surface shiny; promesonotum with sparse, thick to moderately thick, irregular rugae and foveolate interspaces, lateral sides with smooth notches; anepisternum, katepisternum, and propodeum with denser rugofoveolae. Pilosity moderately sparse, moderately long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
|
Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thin apex, in rear view node dorsoventrally concave; pilosity moderately sparse and erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
|
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view trapezoid, lateral margins medially with two distinct dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
|
Shiny and smooth with shagreened base; pilosity moderately dense, moderately short, and erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
|
Brown with yellow legs (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48D, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 1): HL: 0.44; HW: 0.38; SL: 0.42; EL: 0.08; WL: 0.5; PSL: 0.03; MTL: 0.31; PNW: 0.25; PTW: 0.06; PPW: 0.08; CI: 117.2; SI: 110.3; PSLI: 7.4; PPI: 75.4; PNI: 67.0; MTI: 82.3.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
|
Cephalic margin indistinctly relatively straight (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48A</figureCitation>
|
|
). Pilosity relatively sparse, moderately long, decumbent to suberect. Sculpture shiny and smooth; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48A, C</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
|
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove indistinct; propodeal spines reduced to small tubercles (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48C</figureCitation>
|
|
). Sculpture shiny and smooth. Pilosity moderately sparse, moderately long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
|
Yellowish brown, legs and antenna yellow (Fig.
|
|
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Pheidole nitidobruna sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0245024) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923273)." figureDoi="10.3897/zookeys.949.51269.figure48" httpUri="https://binary.pensoft.net/fig/432217" pageId="0" pageNumber="1">48C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Latin for smooth and brown in reference to smooth sculpture and dark body colouration.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The species was collected at 550 m in elevation, in rainforest. Nest was located in a rotten log.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="comments">
|
|
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. belongs to the group of species characterised by small body size (major workers: HL <1.05 mm, WL <0.9 mm and minor workers HL <0.5 mm, WL <0.6 mm), major workers with head elongate and not widening posteriorly, and minor workers with predominantly smooth and relatively rectangular head and yellow to brown body colouration. The group includes four species:
|
|
<taxonomicName lsidName="P. flavominuta" pageId="0" pageNumber="1" rank="species" species="flavominuta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. flavominuta</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.,
|
|
<taxonomicName lsidName="P. nitidobruna" pageId="0" pageNumber="1" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.,
|
|
<taxonomicName lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., and
|
|
<taxonomicName lsidName="P. beanka" pageId="0" pageNumber="1" rank="species" species="beanka">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. beanka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is known only from Makirovana forest in Antsiranana and is parapatric with
|
|
<taxonomicName lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., which is recorded from the area between Andapa and Antisiranana. Major workers of
|
|
<taxonomicName lsidName="P. nitidobruna" pageId="0" pageNumber="1" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be separated from
|
|
<taxonomicName lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by smooth to indistinctly rugulate interspaces on frons and occipital lobes and presence of smooth notches on lateral sides of promesonotum. Minor workers of
|
|
<taxonomicName lsidName="P. nitidobruna" pageId="0" pageNumber="1" rank="species" species="nitidobruna">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. nitidobruna</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be separated from
|
|
<taxonomicName lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. based on entirely smooth vertex and frons and presence of propodeal spines reduced to small tubercles.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |