treatments-xml/data/03/9F/87/039F87C4D8490B2E9D63FF4DFD5DFA1C.xml

<document id="56589640A175A9B1EE7A0591AF4AC59B" ID-DOI="10.5281/zenodo.12170426" ID-ISSN="2386-3803" ID-Zenodo-Dep="12170426" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718821400646" checkinUser="felipe" docAuthor="Altaba, Cristian R." docDate="2022" docId="039F87C4D8490B2E9D63FF4DFD5DFA1C" docLanguage="en" docName="Nemus.12.168-186.pdf" docOrigin="Nemus 12" docStyle="DocumentStyle:C40A3BA05B19340D21880D691A721036.1:Nemus.2022-.journal_article" docStyleId="C40A3BA05B19340D21880D691A721036" docStyleName="Nemus.2022-.journal_article" docStyleVersion="1" docTitle="Allognathus graellsianus" docType="treatment" docVersion="1" lastPageNumber="171" masterDocId="FFA6FFBCD84A0B2D9D12FF91FF9AFFA6" masterDocTitle="Nomenclature of Helicidae (Gastropoda: Pulmonata) endemic to the Balearics" masterLastPageNumber="186" masterPageNumber="168" pageNumber="171" updateTime="1721071674411" updateUser="juliana">
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<mods:title id="2F0F1AA09315BA74A778C5C4608BB25E">Nomenclature of Helicidae (Gastropoda: Pulmonata) endemic to the Balearics</mods:title>
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<mods:namePart id="D18F1E54C69DE2ADA6BF86EDE509B2B5">Altaba, Cristian R.</mods:namePart>
<mods:affiliation id="C54805D7C30327703F9C86943F9313F3">Department of the Environment and Territory, Government of the Balearic Islands, 07009 Palma, Balearic Islands, Spain. &amp; Research Group on Human Evolution and Cogniton (EVOCOG), University of the Balearic Islands, 07072 Palma, Balearic Islands, Spain. cristianr. altaba @ uib. cat</mods:affiliation>
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<subSubSection id="C32C6559D8490B2E9D63FF4DFD82FF56" box="[113,536,220,240]" pageId="3" pageNumber="171" type="nomenclature">
<paragraph id="8B8936D2D8490B2E9D63FF4DFD82FF56" blockId="3.[113,625,220,1659]" box="[113,536,220,240]" pageId="3" pageNumber="171">
<heading id="D0C181BED8490B2E9D63FF4DFD82FF56" bold="true" box="[113,536,220,240]" fontSize="9" level="1" pageId="3" pageNumber="171" reason="1">
<emphasis id="B942EAC0D8490B2E9D63FF4DFD82FF56" bold="true" box="[113,536,220,240]" pageId="3" pageNumber="171">
<taxonomicName id="4C364D51D8490B2E9D63FF4DFEC1FF56" ID-CoL="BWP9" authority="(L.Pfeiffer, 1848)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1848" box="[113,347,220,240]" class="Gastropoda" family="Helicidae" genus="Allognathus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsianus">
<emphasis id="B942EAC0D8490B2E9D63FF4DFEC1FF56" bold="true" box="[113,347,220,240]" italics="true" pageId="3" pageNumber="171">Allognathus graellsianus</emphasis>
</taxonomicName>
is the correct name
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C6559D8490B2E9D9AFF6DFD5DFA1C" pageId="3" pageNumber="171" type="description">
<paragraph id="8B8936D2D8490B2E9D9AFF6DFEFBFCD0" blockId="3.[113,625,220,1659]" pageId="3" pageNumber="171">
In his pioneering “Catálogo de los moluscos terrestres y de agua dulce de 
<collectingCountry id="F3217642D8490B2E9C76FE8DFE32FE97" box="[356,424,284,305]" name="Spain" pageId="3" pageNumber="171">España</collectingCountry>
”, Mariano de la Paz 
<bibRefCitation id="EFA74B23D8490B2E9D63FEADFF6CFEF4" author="Graells, M. P." box="[113,246,316,338]" pageId="3" pageNumber="171" refId="ref14580" refString="Graells, M. P. 1846. Catalogo de los moluscos terrestres y de agua dulce observados en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais. vi [+ i] + 23 [+ 2] pp., lam. [1]. Calleja. Madrid y Lima." type="journal volume" year="1846">Graells (1846)</bibRefCitation>
ably described and figured a species endemic to the northern Serra de Tramuntana mountain range in Mallorca, which he called 
<taxonomicName id="4C364D51D8490B2E9CFAFEEEFF70FE14" authority="Graells, 1846" authorityName="Graells" authorityYear="1846" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="B942EAC0D8490B2E9CFAFEEEFDEAFE33" box="[488,624,383,405]" italics="true" pageId="3" pageNumber="171">Helix grateloupi</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9D63FE0DFF70FE14" author="Graells, M. P." box="[113,234,412,434]" pageId="3" pageNumber="171" refId="ref14580" refString="Graells, M. P. 1846. Catalogo de los moluscos terrestres y de agua dulce observados en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais. vi [+ i] + 23 [+ 2] pp., lam. [1]. Calleja. Madrid y Lima." type="journal volume" year="1846">Graells, 1846</bibRefCitation>
</taxonomicName>
. However, this species name had already been used: 
<taxonomicName id="4C364D51D8490B2E9DF0FE2EFE61FE74" authority="Pfeiffer, 1842" authorityName="Pfeiffer" authorityYear="1842" box="[226,507,445,469]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupii">
<emphasis id="B942EAC0D8490B2E9DF0FE2EFEE8FE73" box="[226,370,447,469]" italics="true" pageId="3" pageNumber="171">Helix grateloupii</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9C6EFE2CFE61FE74" author="Pfeiffer" box="[380,507,445,467]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
</taxonomicName>
was validly proposed as a synonym (through bibliographic indication; 
<bibRefCitation id="EFA74B23D8490B2E9DA4FE6CFEC8FDB5" author="Pfeiffer" box="[182,338,509,531]" firstAuthor="Pfeiffer" page="99" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer, 1842: 99</bibRefCitation>
) of a species from 
<collectingCountry id="F3217642D8490B2E9F13FE6CFDF5FDB5" box="[513,623,509,531]" name="Madagascar" pageId="3" pageNumber="171">Madagascar</collectingCountry>
(now included in the family 
<taxonomicName id="4C364D51D8490B2E9C9AFD8CFD95FD95" authorityName="Godwin-Austen" authorityYear="1883" box="[392,527,541,563]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
) that had been described as 
<taxonomicName id="4C364D51D8490B2E9C3FFDD1FDA4FDF5" authority="Grateloup, 1840" authorityName="Grateloup" authorityYear="1840" box="[301,574,574,598]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="rufescens">
<emphasis id="B942EAC0D8490B2E9C3FFDD1FE03FDF0" box="[301,409,576,598]" italics="true" pageId="3" pageNumber="171">H. rufescens</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9CB6FDAFFDA4FDF5" author="Grateloup, J. P. S. de" box="[420,574,574,596]" pageId="3" pageNumber="171" pagination="394 - 455" refId="ref14682" refString="Grateloup, J. P. S. de. 1840. Description de plusieurs Coquilles nouvelles ou peu connues de Mollusques exotiques vivants. Actes de la Societe Linneenne de Bordeaux, 11 [1839]: 394 - 455, pls. I-IV." type="journal article" year="1840">Grateloup, 1840</bibRefCitation>
</taxonomicName>
. The latter was preoccupied earliest by 
<taxonomicName id="4C364D51D8490B2E9CA3FDF1FF07FD35" authority="Pennant, 1777" authorityName="Pennant" authorityYear="1777" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="rufescens">
<emphasis id="B942EAC0D8490B2E9CA3FDF1FD8DFDD0" box="[433,535,608,630]" italics="true" pageId="3" pageNumber="171">H. rufescens</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9F0EFDCFFF07FD35" author="Pennant" firstAuthor="Pennant" pageId="3" pageNumber="171" refId="ref16396" refString="Pennant, T. 1777. British zoology. Vol. IV. Crustacea. Mollusca. Testacea. viii + [1 - 10], 1 - 154, [1 - 2], pls. I-XCIII. B. White. London." type="book" year="1777">Pennant, 1777</bibRefCitation>
</taxonomicName>
; this is obviously a juvenile of 
<taxonomicName id="4C364D51D8490B2E9CD6FD10FE88FD11" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Helicidae" genus="Arianta" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="arbustorum">
<emphasis id="B942EAC0D8490B2E9CD6FD10FDF5FD31" box="[452,623,641,663]" italics="true" pageId="3" pageNumber="171">Arianta arbustorum</emphasis>
(
<bibRefCitation id="EFA74B23D8490B2E9D6AFD0FFE97FD11" author="Linnaeus" box="[120,269,670,695]" firstAuthor="Linnaeus" pageId="3" pageNumber="171" refId="ref15243" refString="Linnaeus, C. 1758. Systema naturae per regna tria naturae, secundum" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
. 
<taxonomicName id="4C364D51D8490B2E9C31FD30FE2CFD11" authorityName="Pfeiffer" authorityYear="1842" box="[291,438,673,695]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupii">
<emphasis id="B942EAC0D8490B2E9C31FD30FE2CFD11" box="[291,438,673,695]" italics="true" pageId="3" pageNumber="171">Helix grateloupii</emphasis>
</taxonomicName>
was subsequently validated (article 11.6.1 of the Code) by 
<bibRefCitation id="EFA74B23D8490B2E9CFFFD2EFF15FD53" author="Pfeiffer" firstAuthor="Pfeiffer" page="13" pageId="3" pageNumber="171" refId="ref16467" refString="Pfeiffer, L. 1846. Symbolae ad historiam heliceorum. Sectio tertia. 100 pp. T. Fischer, Cassellum [Kassel]." type="book" year="1846">Pfeiffer (1846: 13)</bibRefCitation>
, and later on described in detail (
<bibRefCitation id="EFA74B23D8490B2E9CD5FD4EFDFCFD53" author="Pfeiffer" box="[455,614,735,757]" firstAuthor="Pfeiffer" page="69" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer, 1848: 69</bibRefCitation>
). Thus, the combination 
<taxonomicName id="4C364D51D8490B2E9C73FC90FF39FC93" authority="(Grateloup, 1840)" baseAuthorityName="Grateloup" baseAuthorityYear="1840" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="rufescens">
<emphasis id="B942EAC0D8490B2E9C73FC90FE63FCB1" box="[353,505,769,791]" italics="true" pageId="3" pageNumber="171">Kalidos rufescens</emphasis>
(
<bibRefCitation id="EFA74B23D8490B2E9F1FFD6EFF07FC93" author="Grateloup, J. P. S. de" pageId="3" pageNumber="171" pagination="394 - 455" refId="ref14682" refString="Grateloup, J. P. S. de. 1840. Description de plusieurs Coquilles nouvelles ou peu connues de Mollusques exotiques vivants. Actes de la Societe Linneenne de Bordeaux, 11 [1839]: 394 - 455, pls. I-IV." type="journal article" year="1840">Grateloup, 1840</bibRefCitation>
)
</taxonomicName>
to designate the Malagasy species (
<bibRefCitation id="EFA74B23D8490B2E9CFFFC8EFF63FCF3" author="Fischer-Piette, E. &amp; Bedoucha, J. &amp; Salvat, F." pageId="3" pageNumber="171" pagination="1 - 50" refId="ref13767" refString="Fischer-Piette, E., Bedoucha, J. &amp; Salvat, F. 1966. Mollusques terrestres de Madagascar. Ariophantidae. Memoires du Museum national d'Histoire naturelle, serie A Zoologie, 40 (1): 1 - 50, pls. I-IV." type="journal article" year="1966">Fischer-Piette et al., 1966: 26</bibRefCitation>
, 
<bibRefCitation id="EFA74B23D8490B2E9C11FCD1FEC1FCFE" author="Fischer-Piette, E. &amp; Blanc, C. P. &amp; Blanc, F. &amp; Salvat, F." box="[259,347,832,856]" pageId="3" pageNumber="171" pagination="1 - 552" refId="ref13814" refString="Fischer-Piette, E., Blanc, C. P., Blanc, F. &amp; Salvat, F. 1994. Gasteropodes terrestres pulmones. Faune de Madagascar, 83: 1 - 552." type="journal article" year="1994">1994: 254</bibRefCitation>
) is incorrect, as it must be 
<taxonomicName id="4C364D51D8490B2E9F4DFCD3FEC7FCD0" authority="(Pfeiffer, 1842)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupii">
<emphasis id="B942EAC0D8490B2E9F4DFCD3FF57FCDE" italics="true" pageId="3" pageNumber="171">K. grateloupii</emphasis>
(
<bibRefCitation id="EFA74B23D8490B2E9DCAFCF1FECDFCD0" author="Pfeiffer" box="[216,343,864,886]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8936D2D8490B2E9D9AFC11FE9AFA3C" blockId="3.[113,625,220,1659]" pageId="3" pageNumber="171">
When discovering the conflict of 
<taxonomicName id="4C364D51D8490B2E9CA4FC13FEB4FC10" authority=", Ludwig Pfeiffer (1848: 437)" authorityName="Ludwig Pfeiffer" authorityPageNumber="437" authorityYear="1848" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="B942EAC0D8490B2E9CA4FC13FDB9FC3E" box="[438,547,898,920]" italics="true" pageId="3" pageNumber="171">H.grateloupi</emphasis>
, Ludwig 
<bibRefCitation id="EFA74B23D8490B2E9D63FC31FEB4FC10" author="Pfeiffer" box="[113,302,928,950]" firstAuthor="Pfeiffer" page="437" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer (1848: 437)</bibRefCitation>
</taxonomicName>
proposed 
<taxonomicName id="4C364D51D8490B2E9C8CFC32FF06FC70" authority="Pfeiffer, 1848" authorityName="Pfeiffer" authorityYear="1848" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsiana">
<emphasis id="B942EAC0D8490B2E9C8CFC32FD83FC1F" box="[414,537,931,953]" italics="true" pageId="3" pageNumber="171">H. graellsiana</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9F37FC31FF06FC70" author="Pfeiffer" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer, 1848</bibRefCitation>
</taxonomicName>
for the species discovered by Graells. During the second half of the 19th century some authors used the name initially proposed by Graells (
<bibRefCitation id="EFA74B23D8490B2E9CD2FB90FDF6FBB1" author="Bourguignat, J. R." box="[448,620,1025,1047]" pageId="3" pageNumber="171" refId="ref11914" refString="Bourguignat, J. R. 1864. Malacologie de l'Algerie, ou histoire naturelle des animaux mollusques terrestres et fluviatiles recueillis jusqu'a ce jour dans nos possessions du nord de l'Afrique. Tome second. 380 + [27] + 4 pp, 27 pls. Challamel Aine and Bastide. Paris and Alger." type="book" year="1864">Bourguignat, 1864</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9D63FBB0FF7DFB90" author="Kobelt, W." box="[113,231,1057,1079]" pageId="3" pageNumber="171" refId="ref15657" refString="Kobelt, W. 1871. Catalog der im europaischen Faunengebiet lebenden Binnenconchylien. Mit besonderer Berucksichtigung der in Rossmassler's Sammlung enthaltenen Arten. xvi + 150 pp. T. Fischer. Cassel." type="book" year="1871">Kobelt, 1871</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9DE1FBB0FEFCFB91" author="Paetel" box="[243,358,1057,1079]" firstAuthor="Paetel" pageId="3" pageNumber="171" refId="ref16141" refString="Paetel, F. 1889. Catalog der Conchylien-Sammlung von Fr. Paetel." type="book" year="1889">Paetel, 1889</bibRefCitation>
), while others chose to use 
<taxonomicName id="4C364D51D8490B2E9D63FBD5FF54FBFC" authorityName="Pfeiffer" authorityYear="1848" box="[113,206,1092,1114]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsiana">
<emphasis id="B942EAC0D8490B2E9D63FBD5FF54FBFC" box="[113,206,1092,1114]" italics="true" pageId="3" pageNumber="171">graellsiana</emphasis>
</taxonomicName>
(for example, 
<bibRefCitation id="EFA74B23D8490B2E9C48FBD3FDAFFBF1" author="Reeve &amp; Sowerby" box="[346,565,1089,1111]" firstAuthor="Reeve" pageId="3" pageNumber="171" refId="ref16872" refString="Reeve, L. &amp; Sowerby, G. B. 1851 - 1854. Conchologia iconica: or, illus- trations of the shells of molluscous animals. Vol. VII containing a" type="book" year="1851">Reeve &amp; Sowerby, 1851</bibRefCitation>
–1854; 
<bibRefCitation id="EFA74B23D8490B2E9D63FBF3FF6CFBDC" author="Hidalgo, J. G." box="[113,246,1122,1146]" pageId="3" pageNumber="171" refId="ref15009" refString="Hidalgo, J. G. 1875. Catalogo iconografico y descriptivo de los moluscos terrestres de Espana, Portugal y las Baleares. iv + 224 pp., 24 lams. S. Martinez. Madrid." type="book" year="1875">Hidalgo, 1875</bibRefCitation>
). Among the latter is Henry Augustus Pilsbry (
<bibRefCitation id="EFA74B23D8490B2E9DD6FB13FE39FB3E" author="Tryon, G. W. &amp; Pilsbry, H. A." box="[196,419,1154,1176]" pageId="3" pageNumber="171" pagination="1 - 69" refId="ref18441" refString="Tryon, G. W. &amp; Pilsbry, H. A. 1888. Manual of Conchology; structures and systematic. Second series: Pulmonata, 4. Helicidae. Vol. II. 296 pp., pls. 1 - 69. Tryon Jr. Philadelphia." type="book chapter" year="1888">Tryon &amp; Pilsbry, 1888</bibRefCitation>
), who designated 
<taxonomicName id="4C364D51D8490B2E9F4FFB15FED0FB1E" authority="Graells, 1846" authorityName="Graells" authorityYear="1846" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="B942EAC0D8490B2E9F4FFB15FF52FB1C" italics="true" pageId="3" pageNumber="171">H. grateloupi</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9DC3FB33FED0FB1E" author="Graells, M. P." box="[209,330,1186,1208]" pageId="3" pageNumber="171" refId="ref14580" refString="Graells, M. P. 1846. Catalogo de los moluscos terrestres y de agua dulce observados en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais. vi [+ i] + 23 [+ 2] pp., lam. [1]. Calleja. Madrid y Lima." type="journal volume" year="1846">Graells, 1846</bibRefCitation>
</taxonomicName>
as the 
<typeStatus id="548D8870D8490B2E9C85FB32FE5AFB1E" box="[407,448,1187,1208]" pageId="3" pageNumber="171">type</typeStatus>
of his new genus 
<taxonomicName id="4C364D51D8490B2E9D63FB54FE85FB7E" authority="Pilsbry" authorityName="Pilsbry" authorityYear="1888" box="[113,287,1218,1243]" class="Gastropoda" family="Helicidae" genus="Allognathus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0D8490B2E9D63FB54FF4DFB7D" box="[113,215,1221,1243]" italics="true" pageId="3" pageNumber="171">Allognathus</emphasis>
Pilsbry
</taxonomicName>
in 
<bibRefCitation id="EFA74B23D8490B2E9C2DFB52FD94FB71" author="Tryon, G. W. &amp; Pilsbry, H. A." box="[319,526,1218,1240]" pageId="3" pageNumber="171" pagination="1 - 69" refId="ref18441" refString="Tryon, G. W. &amp; Pilsbry, H. A. 1888. Manual of Conchology; structures and systematic. Second series: Pulmonata, 4. Helicidae. Vol. II. 296 pp., pls. 1 - 69. Tryon Jr. Philadelphia." type="book chapter" year="1888">Tryon &amp; Pilsbry, 1888</bibRefCitation>
. However, he soon corrected this, stating 
<taxonomicName id="4C364D51D8490B2E9C81FB74FD92FB5D" authorityName="Pfeiffer" authorityYear="1848" box="[403,520,1253,1275]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsiana">
<emphasis id="B942EAC0D8490B2E9C81FB74FD92FB5D" box="[403,520,1253,1275]" italics="true" pageId="3" pageNumber="171">H. graellsiana</emphasis>
</taxonomicName>
is the 
<typeStatus id="548D8870D8490B2E9F54FB72FDF5FB5E" box="[582,623,1251,1272]" pageId="3" pageNumber="171">type</typeStatus>
(Tryon &amp; 
<bibRefCitation id="EFA74B23D8490B2E9DDCFA92FED3FABF" author="Pilsbry, H. A." box="[206,329,1283,1305]" pageId="3" pageNumber="171" pagination="1 - 71" refId="ref18492" refString="Tryon G. W. &amp; Pilsbry, H. A. 1894 - 1895. Manual of Conchology; structural and systematic. Second series: Pulmonata, IX. (Helicidae, Vol. 7). Guide to the study of helices. xlviii + 366 + 126 pp., pls. 1 - 71. The Academy of Natural Sciences. Philadelphia." type="book chapter" year="1894">Pilsbry, 1894</bibRefCitation>
). Throughout the 20th century (
<bibRefCitation id="EFA74B23D8490B2E9D6AFAB2FF7EFA9F" author="Hesse, P." box="[120,228,1315,1337]" pageId="3" pageNumber="171" pagination="631 - 660" refId="ref14887" refString="Hesse, P. 1915 - 1920. Iconographie der Land- &amp; Susswasser-Mollusken mit vorzuglicher Berucksichtigung der europaischen noch nicht abgebildeten Arten von E. A. Rossmassler. Neue Folge. Dreiundzwanzigster Band. [5] + 262 pp., Taf. 631 - 660. Kreidel. Wiesbaden." type="book chapter" year="1915">Hesse, 1915</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9DFFFAB2FEC6FA9F" author="Gasull, L." box="[237,348,1315,1337]" pageId="3" pageNumber="171" pagination="3 - 80" refId="ref14030" refString="Gasull, L. 1964. Algunos moluscos terrestres y de agua dulce de Baleares. Boletin de la Sociedad de Historia Natural de Baleares, 9 [1963]: 3 - 80." type="journal article" year="1964">Gasull, 1964</bibRefCitation>
, 
<bibRefCitation id="EFA74B23D8490B2E9C77FAB2FE0BFA9F" author="Gasull, L." box="[357,401,1315,1337]" pageId="3" pageNumber="171" pagination="149 - 156" refId="ref14100" refString="Gasull, L. 1967. La insularidad de las Islas Baleares desde el punto de vista de la malacologia terrestre. Boletin de la Sociedad de Historia Natural de Baleares, 12 [1966]: 149 - 156." type="journal article" year="1967">1967</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9C88FAB2FF07FAFE" author="Breure, A. S. H. &amp; Gittenberger, E." pageId="3" pageNumber="171" pagination="307 - 312" refId="ref12108" refString="Breure, A. S. H. &amp; Gittenberger, E. 1982. The rock-scraping radula, a striking case of convergence (Mollusca). Netherlands Journal of Zoology, 32 (3): 307 - 312." type="journal article" year="1982">Breure &amp; Gittenberger, 1982</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9DB8FAD2FEBBFAFF" author="Altaba, C. R." box="[170,289,1347,1369]" pageId="3" pageNumber="171" pagination="375 - 416" refId="ref10556" refString="Altaba, C. R. 1991. Mol. luscs. In Altaba C. R. &amp; Ros J. (Eds.): Historia natural dels Paisos Catalans, vol. 8: Invertebrats no artropodes: 375 - 416; 427 - 470. Enciclopedia Catalana. Barcelona." type="book chapter" year="1991">Altaba, 1991</bibRefCitation>
), the species was correctly known (and legally protected since 2008) as 
<taxonomicName id="4C364D51D8490B2E9CFCFAF7FF66FA3C" authority="(Pfeiffer, 1848)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Helicidae" genus="Allognathus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsianus">
<emphasis id="B942EAC0D8490B2E9CFCFAF7FDEAFADA" box="[494,624,1382,1404]" italics="true" pageId="3" pageNumber="171">A. graellsianus</emphasis>
(
<bibRefCitation id="EFA74B23D8490B2E9D65FA15FF6CFA3C" author="Pfeiffer" box="[119,246,1412,1434]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer, 1848</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8936D2D8490B2E9D9AFA35FC5AFE14" blockId="3.[113,625,220,1659]" lastBlockId="3.[661,1173,154,1466]" pageId="3" pageNumber="171">
At the onset of the 21st century, and attending only to the priority principle (but ignoring the species from 
<collectingCountry id="F3217642D8490B2E9D63FA75FF78FA5C" box="[113,226,1508,1530]" name="Madagascar" pageId="3" pageNumber="171">Madagascar</collectingCountry>
), 
<bibRefCitation id="EFA74B23D8490B2E9DE9FA75FE07FA5C" author="Schileyko, A. A." box="[251,413,1508,1530]" pageId="3" pageNumber="171" pagination="1 - 142" refId="ref17943" refString="Schileyko, A. A. 2006. Treatise on Recent terrestrial pulmonate molluscs. Part 13: Helicidae, Pleurodontidae, Polygyridae, Ammonitellidae, Oreohelicidae, Thysanophoridae. Ruthenica, Supplement 2 (13): 1 - 142." type="journal article" year="2006">Schileyko (2006)</bibRefCitation>
proposed to return to the name coined by Graells; this was followed by several authors (
<bibRefCitation id="EFA74B23D8490B2E9C00F9B4FE79F99C" author="Welter-Schultes, F. W." box="[274,483,1573,1595]" pageId="3" pageNumber="171" refId="ref18638" refString="Welter-Schultes, F. W. 2012. European non-marine molluscs, a guide for species identification. A 1 - A 3 + 679 pp. + Q 1 - Q 78. Planet Poster Editions. Gottingen." type="book" year="2012">Welter-Schultes, 2012</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9CFCF9B4FF07F9FC" author="Chueca, L. J. &amp; Fores, M. &amp; Gomez-Moliner, B. J." pageId="3" pageNumber="171" pagination="63 - 74" refId="ref12451" refString="Chueca, L. J., Fores, M. &amp; Gomez-Moliner, B. J. 2013. Consideraciones nomenclaturales sobre las especies del genero AllognOthus (Gastropoda: Pulmonata: Helicidae) y estudio anatomico de AllognOthus hispOnicus tOnitiOnus. Iberus, 31 (1): 63 - 74." type="journal article" year="2013">Chueca et al., 2013</bibRefCitation>
, 
<bibRefCitation id="EFA74B23D8490B2E9DBBF9D4FE22F9FD" author="Altaba, C. R. &amp; Rios Jimenez, F." box="[169,440,1605,1627]" pageId="3" pageNumber="171" pagination="35 - 44" refId="ref10960" refString="Altaba, C. R. &amp; Rios Jimenez, F. 2021. An enigmatic rock-dwelling land snail from southernmost Iberia (Mollusca, Gastropoda, Helicidae). Folia Conchyliologica, 60: 35 - 44." type="journal article" year="2021">Altaba &amp; Ríos Jiménez, 2021</bibRefCitation>
). However, 
<bibRefCitation id="EFA74B23D8490B2E9F3BF9D4FE9FF9DD" author="Welter-Schultes, F. W." pageId="3" pageNumber="171" refId="ref18638" refString="Welter-Schultes, F. W. 2012. European non-marine molluscs, a guide for species identification. A 1 - A 3 + 679 pp. + Q 1 - Q 78. Planet Poster Editions. Gottingen." type="book" year="2012">Welter-Schultes (2012)</bibRefCitation>
, justifies his choice considering that 
<taxonomicName id="4C364D51D8490B2E9F87FF0CFC11FF09" authority="Pfeiffer, 1842" authorityName="Pfeiffer" authorityYear="1842" box="[661,907,154,179]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="grateloupii">
<emphasis id="B942EAC0D8490B2E9F87FF0CFC93FF15" box="[661,777,157,179]" italics="true" pageId="3" pageNumber="171">H. grateloupii</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9E02FF0BFC11FF09" author="Pfeiffer" box="[784,907,154,176]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
</taxonomicName>
might not have been intentionally proposed as a new name (this contradicts the original text), or alternatively that it could have been proposed in synonymy (which is not the case, nor would it affect the availability and date of a name once used first in 1848); it is clear that both Graells and Pfeiffer were never confusing the Malagasy and Mallorcan species. After such oscillating history, the coin eventually falls heads up for Pfeiffer.
</paragraph>
<paragraph id="8B8936D2D8490B2E9FBEFE2DFC3BFD72" blockId="3.[661,1173,154,1466]" pageId="3" pageNumber="171">
Another problematic conclusion is that of 
<bibRefCitation id="EFA74B23D8490B2E9920FE2DFD4AFE55" author="Beckmann, K. H." pageId="3" pageNumber="171" refId="ref11421" refString="Beckmann, K. H. 2007. Die land- uns Susswassermollusken der Balearischen Inseln. 255 pp. Conchbooks. Hackenheim." type="book" year="2007">Beckmann (2007)</bibRefCitation>
, who in his posthumous book on the land and freshwater mollusks of the Balearics maintains the remarkable stand that the name erected by 
<bibRefCitation id="EFA74B23D8490B2E9946FD8CFD54FDF5" author="Graells, M. P." pageId="3" pageNumber="171" refId="ref14580" refString="Graells, M. P. 1846. Catalogo de los moluscos terrestres y de agua dulce observados en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais. vi [+ i] + 23 [+ 2] pp., lam. [1]. Calleja. Madrid y Lima." type="journal volume" year="1846">Graells (1846)</bibRefCitation>
would be a 
<emphasis id="B942EAC0D8490B2E9E2EFDD1FC2DFDF0" box="[828,951,576,598]" italics="true" pageId="3" pageNumber="171">nomen nudum</emphasis>
. 
<bibRefCitation id="EFA74B23D8490B2E9ED3FDACFBFBFDF5" author="Beckmann, K. H." box="[961,1121,573,595]" pageId="3" pageNumber="171" refId="ref11421" refString="Beckmann, K. H. 2007. Die land- uns Susswassermollusken der Balearischen Inseln. 255 pp. Conchbooks. Hackenheim." type="book" year="2007">Beckmann (2007)</bibRefCitation>
cited Fig. 
<quantity id="4CCE9B37D8490B2E9FACFDCFFD6CFDD2" box="[702,758,606,628]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="3" pageNumber="171" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7-8 in</quantity>
the booklet by Graells; with that alone, he should have reached the conclusion that such a name accompanied by a figure has an indication (in the sense of Article 12.2.7 of the Code).
</paragraph>
<paragraph id="8B8936D2D8490B2E9FBEFD4FFC2EFCF3" blockId="3.[661,1173,154,1466]" pageId="3" pageNumber="171">
Another mistake is the spelling “ 
<taxonomicName id="4C364D51D8490B2E9EC9FD70FBB4FD51" box="[987,1070,737,759]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="graellsi">
<emphasis id="B942EAC0D8490B2E9EC9FD70FBB4FD51" box="[987,1070,737,759]" italics="true" pageId="3" pageNumber="171">H. graellsi</emphasis>
</taxonomicName>
” proposed by 
<bibRefCitation id="EFA74B23D8490B2E9FAAFD6EFC38FCB1" author="Bourguignat, J. R." box="[696,930,767,791]" pageId="3" pageNumber="171" refId="ref11914" refString="Bourguignat, J. R. 1864. Malacologie de l'Algerie, ou histoire naturelle des animaux mollusques terrestres et fluviatiles recueillis jusqu'a ce jour dans nos possessions du nord de l'Afrique. Tome second. 380 + [27] + 4 pp, 27 pls. Challamel Aine and Bastide. Paris and Alger." type="book" year="1864">Bourguignat (1864: 356)</bibRefCitation>
. Nobody seems to have followed him. This constitutes an incorrect subsequent spelling (Art. 33.3 of the Code).
</paragraph>
<paragraph id="8B8936D2D8490B2E9FBEFCCEFD5DFA1C" blockId="3.[661,1173,154,1466]" pageId="3" pageNumber="171">
There is another synonym for this species: 
<taxonomicName id="4C364D51D8490B2E997AFCF3FCA6FC30" authority="Ferussac" authorityName="Ferussac" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="tessellata">
<emphasis id="B942EAC0D8490B2E997AFCF3FD7FFC3E" italics="true" pageId="3" pageNumber="171">Helix tessellata</emphasis>
Férussac
</taxonomicName>
in Férussac &amp; Deshayes, 1851. Based on an explicit reference to the Mallorcan species (Férussac in Deshayes, 1819–1851: 232), it is a junior subjective synonym. Besides, this name is preoccupied by 
<taxonomicName id="4C364D51D8490B2E9FA0FB92FC04FBB0" authority="Pfeiffer, 1842" authorityName="Pfeiffer" authorityYear="1842" box="[690,926,1025,1049]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="tessellata">
<emphasis id="B942EAC0D8490B2E9FA0FB92FC83FBBF" box="[690,793,1027,1049]" italics="true" pageId="3" pageNumber="171">H. tessellata</emphasis>
<bibRefCitation id="EFA74B23D8490B2E9E0DFB90FC04FBB0" author="Pfeiffer" box="[799,926,1025,1047]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
</taxonomicName>
. The latter has been attributed to the mention by Anton (1839: 36, nº 1359), but this author only wrote the name without giving any description; 
<bibRefCitation id="EFA74B23D8490B2E9E03FBF0FC58FBD1" author="Pfeiffer" box="[785,962,1121,1143]" firstAuthor="Pfeiffer" page="40" pageId="3" pageNumber="171" refId="ref16443" refString="Pfeiffer, 1842. Symbolae ad Historiam heliceorum. Sectio altera. 147 pp. T. Fischer. Cassellum [Kassel]." type="book" year="1842">Pfeiffer (1842: 40)</bibRefCitation>
describes it adequately, mentioning a letter from Anton and attributing the name to Mühlfeld. This species is now included in the (sub)genus 
<taxonomicName id="4C364D51D8490B2E9E34FB55FC57FB7E" authority="Albers" authorityName="Albers" box="[806,973,1218,1242]" class="Gastropoda" family="Charopidae" genus="Stephanoda" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0D8490B2E9E34FB55FC11FB7C" box="[806,907,1220,1242]" italics="true" pageId="3" pageNumber="171">Stephanoda</emphasis>
Albers
</taxonomicName>
in 
<bibRefCitation id="EFA74B23D8490B2E9EF8FB53FD5AFB5E" author="Albers, J. C. &amp; Martens, E. von" pageId="3" pageNumber="171" refId="ref10327" refString="Albers, J. C. &amp; Martens, E. von 1860. Die Heliceen nach naturlicher Verwandtschaf systematisch geordnet von Joh. Christian Albers. Zweite Ausgabe nach dem hinterlassenen Manuskript besorgt von Eduard von Martens. xviii + 359 pp. W. Engelmann. Leipzig." type="book" year="1860">Albers &amp; Martens, 1860</bibRefCitation>
(
<taxonomicName id="4C364D51D8490B2E9FD9FB73FCC8FB5E" authorityName="Pilsbry" authorityYear="1895" box="[715,850,1250,1272]" class="Gastropoda" family="Endodontidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="family">Endodontidae</taxonomicName>
); it is endemic to the island of Juan Fernández in the southeastern Pacific (Anton, 1839: 36; 
<bibRefCitation id="EFA74B23D8490B2E9FA7FAB2FCA9FA9E" author="Pfeiffer" box="[693,819,1315,1337]" firstAuthor="Pfeiffer" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer, 1848</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9E29FAB2FB8DFA9F" author="Albers, J. C. &amp; Martens, E. von" box="[827,1047,1315,1337]" pageId="3" pageNumber="171" refId="ref10327" refString="Albers, J. C. &amp; Martens, E. von 1860. Die Heliceen nach naturlicher Verwandtschaf systematisch geordnet von Joh. Christian Albers. Zweite Ausgabe nach dem hinterlassenen Manuskript besorgt von Eduard von Martens. xviii + 359 pp. W. Engelmann. Leipzig." type="book" year="1860">Albers &amp; Martens, 1860</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E990DFAB2FB0AFA9E" author="Solem, A." box="[1055,1168,1315,1337]" pageId="3" pageNumber="171" refId="ref18221" refString="Solem, A. 1978. Endodontid land snails from Pacific islands (Mollusca: Pulmonata; Sigmurethra). Part II Families Punctidae and Charopidae, Zoogeography. 336 pp. Field Museum of Natural History. Chicago." type="book" year="1978">Solem, 1978</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9F87FAD2FCF4FAFF" author="Stuardo, J. &amp; Vega, R." box="[661,878,1347,1369]" pageId="3" pageNumber="171" pagination="125 - 146" refId="ref18261" refString="Stuardo, J. &amp; Vega, R. 1985. Synopsis of the Land Mollusca of Chile. With Remarks on Distribution. Studies on Neotropical Fauna and Environment, 20 (3): 125 - 146." type="journal article" year="1985">Stuardo &amp; Vega, 1985</bibRefCitation>
; 
<bibRefCitation id="EFA74B23D8490B2E9E69FAD2FBB8FAFF" author="Valdovinos, C." box="[891,1058,1347,1369]" pageId="3" pageNumber="171" pagination="111 - 164" refId="ref18560" refString="Valdovinos, C. 1999. Biodiversidad de moluscos chilenos: Base de datos taxonomica y distribucional. Gayana Zoologia, 63 (2): 111 - 164." type="journal article" year="1999">Valdovinos, 1999</bibRefCitation>
). The invalidity of the name 
<taxonomicName id="4C364D51D8490B2E9E40FAF7FB8DFADF" authority="Ferussac" authorityName="Ferussac" box="[850,1047,1379,1404]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="171" phylum="Mollusca" rank="species" species="tessellata">
<emphasis id="B942EAC0D8490B2E9E40FAF7FC27FADA" box="[850,957,1382,1404]" italics="true" pageId="3" pageNumber="171">H. tessellata</emphasis>
Férussac
</taxonomicName>
was already detected by 
<bibRefCitation id="EFA74B23D8490B2E9E1FFA12FC5EFA3F" author="Pfeiffer" box="[781,964,1411,1433]" firstAuthor="Pfeiffer" page="117" pageId="3" pageNumber="171" refId="ref16493" refString="Pfeiffer, L. 1848. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et" type="book" year="1848">Pfeiffer (1848: 117)</bibRefCitation>
, so it has never been used.
</paragraph>
</subSubSection>
</treatment>
</document>