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<document id="5B199AAFD34A9870B405E55E2A9969AF" ID-CLB-Dataset="29397" ID-DOI="10.11646/zootaxa.4462.3.1" ID-GBIF-Dataset="ee3428d1-9e3b-495d-99a5-e61ea679b5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441611" ID-ZooBank="75D121A7-F718-4F69-9E33-E82AB371403A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538403914940" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2018" docId="03C887DB60648C03FF67D2CC1D51FC4A" docLanguage="en" docName="zootaxa.4462.3.1.pdf" docOrigin="Zootaxa 4462 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asiagomphus acco Asahina 1996" docType="treatment" docVersion="7" lastPageNumber="303" masterDocId="FFF1FFA360658C01FFF0D13A1956FF82" masterDocTitle="The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)" masterLastPageNumber="330" masterPageNumber="301" pageNumber="302" updateTime="1698561160019" updateUser="ExternalLinkService">
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<mods:title id="825E33D316795090C5B4593D9CEB9D5F">The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)</mods:title>
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<mods:date id="DC1787DF45F81BC24D5A78C2067ACCBA">2018</mods:date>
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<treatment id="03C887DB60648C03FF67D2CC1D51FC4A" ID-DOI="http://doi.org/10.5281/zenodo.5974761" ID-GBIF-Taxon="148424962" ID-Zenodo-Dep="5974761" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A" httpUri="http://treatment.plazi.org/id/03C887DB60648C03FF67D2CC1D51FC4A" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<subSubSection id="C37B654660648C00FF67D2CC18AAFBB0" pageId="1" pageNumber="302" type="nomenclature">
<paragraph id="8BDE36CD60648C00FF67D2CC1B67FB92" blockId="1.[151,561,1014,1074]" box="[151,561,1014,1040]" pageId="1" pageNumber="302">
<heading id="D09681A160648C00FF67D2CC1B67FB92" bold="true" box="[151,561,1014,1040]" fontSize="11" level="1" pageId="1" pageNumber="302" reason="1">
<taxonomicName id="4C614D4E60648C00FF67D2CC1B67FB92" ID-CoL="H9S6" authority="Asahina, 1996" authorityName="Asahina" authorityYear="1996" box="[151,561,1014,1040]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60648C00FF67D2CC1B67FB92" bold="true" box="[151,561,1014,1040]" pageId="1" pageNumber="302">
<emphasis id="B915EADF60648C00FF67D2CC182CFB92" bold="true" box="[151,378,1014,1040]" italics="true" pageId="1" pageNumber="302">Asiagomphus acco</emphasis>
<bibRefCitation id="EFF04B3C60648C00FE71D2CC1B67FB92" box="[385,561,1014,1040]" pageId="1" pageNumber="302" refId="ref17031" refString="Asahina, S. (1996) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 3. Aeschnidae [sic], Corduliidae and Libellulidae. Bulletin of the National Science Museum, Tokyo, Series A, 22, 69 - 80." type="journal article">Asahina, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDE36CD60648C00FF67D52318AAFBB0" blockId="1.[151,561,1014,1074]" box="[151,508,1049,1074]" pageId="1" pageNumber="302">
(
<figureCitation id="135A2A4860648C00FF6FD52319BAFBB0" box="[159,236,1049,1074]" captionStart="FIGURE 1" captionStartId="3.[151,250,1992,2014]" captionTargetBox="[204,1391,193,1976]" captionTargetId="figure@3.[204,1391,193,1976]" captionTargetPageId="3" captionText="FIGURE 1. Asiagomphus acco copula, Xuan Son NP, 30-V-2015." httpUri="https://zenodo.org/record/1441613/files/figure.png" pageId="1" pageNumber="302">Figs. 1</figureCitation>
,
<figureCitation id="135A2A4860648C00FF0AD520185CFBB0" box="[250,266,1050,1074]" captionStart="FIGURE 2" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1437,193,1221]" captionTargetId="figure@5.[151,1437,193,1221]" captionTargetPageId="5" captionText="FIGURE 2. Asiagomphus acco alive in hand. [ac] male, [ef] female. (a) Ba Be NP, 21-VII-2013; (b) Quang Nam Prov., 21- VI-2016; (c) Tam Dao NP, 18-V-2014; (d) Yen Bai Prov., 20-VI-2015; (e) Xuan Son NP, 22-V-2015; (f) Lang Son Prov., 2-V- 2015." httpUri="https://zenodo.org/record/1441615/files/figure.png" pageId="1" pageNumber="302">2</figureCitation>
,
<figureCitation id="135A2A4860648C00FEE6D5231819FBB0" box="[278,335,1049,1074]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (ab) Asiagomphus acco, Tam Dao NP, 18-V-2014; (cd) A. kosterini sp. nov. holotype; (ef) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="1" pageNumber="302">3ab</figureCitation>
,
<figureCitation id="135A2A4860648C00FEABD523182FFBB0" box="[347,377,1049,1074]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="1" pageNumber="302">5h</figureCitation>
,
<figureCitation id="135A2A4860648C00FE75D52318F5FBB0" box="[389,419,1049,1074]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="1" pageNumber="302">6h</figureCitation>
,
<figureCitation id="135A2A4860648C00FE5FD52318A3FBB0" box="[431,501,1049,1074]" captionStart="FIGURE 13" captionStartId="22.[151,250,1766,1788]" captionTargetBox="[151,1437,481,1745]" captionTargetId="figure@22.[151,1437,481,1745]" captionTargetPageId="22" captionText="FIGURE 13. Asiagomphus spp. males in nature. (a) A. acco, Xuan Son NP, 8-VI-2014; (b) A. acco, Pia Oac NR, 8-VII-2014; (c) A. kosterini, Lam Dong Prov., 17-IV-2016; (d) A. pacificus, Cao Bang Prov., 19-V-2015; (e) A. reinhardti, Gia Lai Prov., 20- VI-2016; (f) A. reinhardti, Gia Lai Prov., 13-V-2017." httpUri="https://zenodo.org/record/1441637/files/figure.png" pageId="1" pageNumber="302">13ab</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37B654660648C00FF67D55B1D7FFB51" pageId="1" pageNumber="302" type="reference_group">
<paragraph id="8BDE36CD60648C00FF67D55B1A05FB14" blockId="1.[151,1435,1121,1235]" pageId="1" pageNumber="302">
<taxonomicName id="4C614D4E60648C00FF67D55B1BD3FBF5" ID-CoL="H9S6" authority=": Asahina (1996)" authorityName="Asahina" authorityYear="1996" box="[151,645,1121,1143]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="302" phylum="Arthropoda" rank="subSpecies" species="xanthenatus" subSpecies="acco">
<emphasis id="B915EADF60648C00FF67D55B1881FBF5" box="[151,471,1121,1143]" italics="true" pageId="1" pageNumber="302">Asiagomphus xanthenatus acco</emphasis>
:
<bibRefCitation id="EFF04B3C60648C00FE15D55B1BD3FBF5" box="[485,645,1121,1143]" pageId="1" pageNumber="302" refId="ref17031" refString="Asahina, S. (1996) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 3. Aeschnidae [sic], Corduliidae and Libellulidae. Bulletin of the National Science Museum, Tokyo, Series A, 22, 69 - 80." type="journal article">Asahina (1996)</bibRefCitation>
</taxonomicName>
, pp. 2224, figs. 29 [type, northern Vietnam];
<bibRefCitation id="EFF04B3C60648C00FB9CD55B1C57FBF5" box="[1132,1281,1121,1143]" pageId="1" pageNumber="302" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson (2005)</bibRefCitation>
, p. 132, fig. 15 [records from Guangxi, China and discussion of specific status].
</paragraph>
<paragraph id="8BDE36CD60648C00FF67D5A41D7FFB51" blockId="1.[151,1435,1121,1235]" pageId="1" pageNumber="302">
<taxonomicName id="4C614D4E60648C00FF67D5A41B59FB36" ID-CoL="H9S6" authority=": Do et al. (2011)" authorityName="Do et al." authorityYear="2011" box="[151,527,1182,1205]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60648C00FF67D5A41801FB37" box="[151,343,1182,1205]" italics="true" pageId="1" pageNumber="302">Asiagomphus acco</emphasis>
:
<bibRefCitation id="EFF04B3C60648C00FE97D5A51B59FB36" box="[359,527,1182,1205]" pageId="1" pageNumber="302" refId="ref17122" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Phan, Q. T. (2011) Anisoptera of Cuc Phuong National Park, North Vietnam. International Dragonfly Fund - Report, Newsletter of the International Dragonfly Fund, 33, 1 - 18." type="journal article">
Do
<emphasis id="B915EADF60648C00FE7ED59A1895FB36" box="[398,451,1182,1205]" italics="true" pageId="1" pageNumber="302">et al.</emphasis>
(2011)
</bibRefCitation>
</taxonomicName>
, pp. 810, fig. 8; von
<bibRefCitation id="EFF04B3C60648C00FD0AD5A41AA3FB36" box="[762,1013,1182,1205]" pageId="1" pageNumber="302" refId="ref17651" refString="von Ellenrieder, N., Hauser, M., Gaimari, S. D. &amp; Thai, H. P. (2015) First records of Macromia katae (Macromiidae) and Indothemis carnatica (Libellulidae) from Vietnam (Insecta: Odonata). Check List, 11, 1 - 13. https: // doi. org / 10.15560 / 11.1.1514" type="journal article">
Ellenrieder
<emphasis id="B915EADF60648C00FC83D59A1AFEFB36" box="[883,936,1182,1205]" italics="true" pageId="1" pageNumber="302">et al.</emphasis>
(2015)
</bibRefCitation>
, p. 5 [species list];
<bibRefCitation id="EFF04B3C60648C00FB33D5A41C0DFB36" box="[1219,1371,1182,1204]" pageId="1" pageNumber="302" refId="ref17367" refString="Karube, H. (2014) Vietnamese Odonata collected in 1992 - 2003 surveys. V. Gomphidae. Tombo, 56, 77 - 90." type="journal article">Karube (2014)</bibRefCitation>
, p. 69 [species list];
<bibRefCitation id="EFF04B3C60648C00FEA4D5871BD9FB51" box="[340,655,1213,1235]" pageId="1" pageNumber="302" refId="ref17961" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A list of Lao dragonflies. Privately published by the first author, Koriyama, 110 pp." type="book">Yokoi &amp; Souphanthong (2014)</bibRefCitation>
, p. 84 [distribution map], p. 37 (Plt. 14).
</paragraph>
</subSubSection>
<subSubSection id="C37B654660648C03FF67D5C51980FF56" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302" type="materials_examined">
<paragraph id="8BDE36CD60648C00FF67D5C51997FAE2" blockId="1.[151,1437,1279,2025]" pageId="1" pageNumber="302">
<materialsCitation id="3B093C9060648C00FF67D5C51C0BFA9A" ID-GBIF-Occurrence="1913892983" box="[151,1373,1279,1304]" collectingDate="2014-05-18" collectorName="Tam Dao NP" country="Vietnam" location="Vinh Phuc" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<emphasis id="B915EADF60648C00FF67D5C5182CFA9A" bold="true" box="[151,378,1279,1304]" pageId="1" pageNumber="302">Material examined</emphasis>
:
<specimenCount id="9D67FD4460648C00FE7AD43A18E1FA95" box="[394,439,1280,1304]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
, Tay Thien,
<collectorName id="2694531B60648C00FDB6D43A1B8DFA95" box="[582,731,1280,1303]" pageId="1" pageNumber="302">Tam Dao NP</collectorName>
,
<collectingRegion id="49A5F82F60648C00FD16D5C51A0BFA9A" box="[742,861,1279,1304]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="302">Vinh Phuc</collectingRegion>
Prov, northern
<collectingCountry id="F376765D60648C00FBFED5C51D26FA9A" box="[1038,1136,1279,1304]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB8CD43A1DA1FA9A" box="[1148,1271,1280,1304]" pageId="1" pageNumber="302" value="2014-05-18">
<date id="FFDF100D60648C00FB8CD43A1DA1FA9A" box="[1148,1271,1280,1304]" pageId="1" pageNumber="302" value="2014-05-18">18-V-2014</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FA9AD43A1A55FABF" ID-GBIF-Occurrence="1913892951" collectingDate="2015-06-20" country="Vietnam" location="Yen Bai" pageId="1" pageNumber="302" specimenCount="1" specimenCount-female="1" stateProvince="Yen Bai">
<specimenCount id="9D67FD4460648C00FA9AD43A1CC3FA95" box="[1386,1429,1280,1304]" pageId="1" pageNumber="302" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49A5F82F60648C00FF67D41F19A4FABF" box="[151,242,1316,1341]" country="Vietnam" name="Yen Bai" pageId="1" pageNumber="302">Yen Bai</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FE59D41E1B5DFABF" box="[425,523,1316,1341]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FDE6D41E1BCBFABF" box="[534,669,1316,1341]" pageId="1" pageNumber="302" value="2015-06-20">
<date id="FFDF100D60648C00FDE6D41E1BCBFABF" box="[534,669,1316,1341]" pageId="1" pageNumber="302" value="2015-06-20">20-VI-2015</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FCFFD41E1997FAE2" ID-GBIF-Occurrence="1913892958" collectingDate="2016-06-21" country="Vietnam" location="Quang Nam" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Quang Nam">
<specimenCount id="9D67FD4460648C00FCFFD41E1A6AFABE" box="[783,828,1316,1340]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49A5F82F60648C00FCB9D41E1A87FABE" box="[841,977,1316,1340]" country="Vietnam" name="Quang Nam" pageId="1" pageNumber="302">Quang Nam</collectingRegion>
Prov., central
<collectingCountry id="F376765D60648C00FB86D41E1D8EFABF" box="[1142,1240,1316,1341]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB14D41E1C3DFABF" box="[1252,1387,1316,1341]" pageId="1" pageNumber="302" value="2016-06-21">
<date id="FFDF100D60648C00FB14D41E1C3DFABF" box="[1252,1387,1316,1341]" pageId="1" pageNumber="302" value="2016-06-21">21-VI-2016</date>
</collectingDate>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="8BDE36CD60648C03FF37D4561980FF56" blockId="1.[151,1437,1279,2025]" lastBlockId="2.[151,1437,151,968]" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<materialsCitation id="3B093C9060648C00FF37D4561871FA2A" ID-GBIF-Occurrence="1913892970" collectingDate="2013-07-21" country="Vietnam" location="Ba Be" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan">
<emphasis id="B915EADF60648C00FF37D4561A19FA07" bold="true" box="[199,847,1388,1413]" pageId="1" pageNumber="302">Additional records based on field observations by TK</emphasis>
:
<specimenCount id="9D67FD4460648C00FC90D4561ADBFA06" box="[864,909,1388,1412]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FC6BD4571AB7FA06" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FC6BD4571AB7FA06" box="[923,993,1389,1412]" country="Vietnam" name="Ba Be" pageId="1" pageNumber="302" stateProvince="Bac Kan">Ba Be</location>
NP,
<collectingRegion id="49A5F82F60648C00FBEAD4571D2BFA06" box="[1050,1149,1389,1412]" country="Vietnam" name="Bac Kan" pageId="1" pageNumber="302">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FAC6D4561CCEFA07" box="[1334,1432,1388,1413]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FF67D4AA1871FA2A" box="[151,295,1424,1448]" pageId="1" pageNumber="302" value="2013-07-21">
<date id="FFDF100D60648C00FF67D4AA1871FA2A" box="[151,295,1424,1448]" pageId="1" pageNumber="302" value="2013-07-21">21-VII-2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FEC4D4B51DEAFA2A" ID-GBIF-Occurrence="1913892967" box="[308,1212,1423,1448]" collectingDate="2014-05-18" country="Vietnam" location="Tam Dao" municipality="Tay Thien" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Vinh Phuc">
6 ind.,
<collectingMunicipality id="6BBAACB760648C00FE74D4AA18AFFA2A" box="[388,505,1423,1448]" pageId="1" pageNumber="302">Tay Thien</collectingMunicipality>
,
<location id="8EBE601660648C00FDF5D4AA1B38FA25" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FDF5D4AA1B38FA25" box="[517,622,1424,1447]" country="Vietnam" municipality="Tay Thien" name="Tam Dao" pageId="1" pageNumber="302" stateProvince="Vinh Phuc">Tam Dao</location>
NP,
<collectingRegion id="49A5F82F60648C00FD55D4B51A4DFA2A" box="[677,795,1423,1448]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="302">Vinh Phuc</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FC23D4B51D63FA2A" box="[979,1077,1423,1448]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FBB1D4AA1DEAFA2A" box="[1089,1212,1424,1448]" pageId="1" pageNumber="302" value="2014-05-18">
<date id="FFDF100D60648C00FBB1D4AA1DEAFA2A" box="[1089,1212,1424,1448]" pageId="1" pageNumber="302" value="2014-05-18">18-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FB39D4AA1B71FA4F" ID-GBIF-Occurrence="1913892942" collectingDate="2014-05-25" country="Vietnam" location="Ba Vi" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi">
<specimenCount id="9D67FD4460648C00FB39D4AA1DA0FA25" box="[1225,1270,1424,1448]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FAF2D4AA1C13FA2A" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FAF2D4AA1C13FA2A" box="[1282,1349,1423,1448]" country="Vietnam" name="Ba Vi" pageId="1" pageNumber="302" stateProvince="Ha Noi">Ba Vi</location>
NP,
<collectingRegion id="49A5F82F60648C00FA8BD4AA1992FA4F" country="Vietnam" name="Ha Noi" pageId="1" pageNumber="302">Ha Noi</collectingRegion>
, northern
<collectingCountry id="F376765D60648C00FECBD48E18CBFA4F" box="[315,413,1460,1485]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE5BD48E1B71FA4F" box="[427,551,1460,1485]" pageId="1" pageNumber="302" value="2014-05-25">
<date id="FFDF100D60648C00FE5BD48E1B71FA4F" box="[427,551,1460,1485]" pageId="1" pageNumber="302" value="2014-05-25">25-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FDC5D48E1C12FA4E" ID-GBIF-Occurrence="1913892977" box="[565,1348,1460,1485]" collectingDate="2014-05-31" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
3 ind.,
<location id="8EBE601660648C00FD79D48F1BAAFA4E" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FD79D48F1BAAFA4E" box="[649,764,1460,1484]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FCC7D48E1ACCFA4F" box="[823,922,1460,1485]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FBA8D48E1DECFA4F" box="[1112,1210,1460,1485]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB38D48E1C12FA4E" box="[1224,1348,1460,1484]" pageId="1" pageNumber="302" value="2014-05-31">
<date id="FFDF100D60648C00FB38D48E1C12FA4E" box="[1224,1348,1460,1484]" pageId="1" pageNumber="302" value="2014-05-31">31-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FAA2D48E1A37FA72" ID-GBIF-Occurrence="1913892953" collectingDate="2014-06-01" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
2 ind.,
<location id="8EBE601660648C00FF67D4E2185AFA72" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FF67D4E2185AFA72" box="[151,268,1496,1520]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FEBCD4ED18E6FA72" box="[332,432,1495,1520]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FD85D4ED1B81FA72" box="[629,727,1495,1520]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FD17D4E21A37FA72" box="[743,865,1496,1520]" pageId="1" pageNumber="302" value="2014-06-01">
<date id="FFDF100D60648C00FD17D4E21A37FA72" box="[743,865,1496,1520]" pageId="1" pageNumber="302" value="2014-06-01">1-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FC81D4E2182BF996" ID-GBIF-Occurrence="1913892954" collectingDate="2014-06-08" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
4 ind.,
<location id="8EBE601660648C00FC3AD4E21D69FA72" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FC3AD4E21D69FA72" box="[970,1087,1496,1520]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FB8FD4ED1DB2FA72" box="[1151,1252,1495,1520]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FF67D4C619AFF997" box="[151,249,1532,1557]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FEF3D4C6182BF996" box="[259,381,1532,1556]" pageId="1" pageNumber="302" value="2014-06-08">
<date id="FFDF100D60648C00FEF3D4C6182BF996" box="[259,381,1532,1556]" pageId="1" pageNumber="302" value="2014-06-08">8-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FE78D4C61DD8F997" ID-GBIF-Occurrence="1913892987" box="[392,1166,1532,1557]" collectingDate="2014-06-15" country="Vietnam" location="Tam Dao" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Vinh Phuc">
2 ind.,
<location id="8EBE601660648C00FE25D4C71B6AF996" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FE25D4C71B6AF996" box="[469,572,1533,1556]" country="Vietnam" name="Tam Dao" pageId="1" pageNumber="302" stateProvince="Vinh Phuc">Tam Dao</location>
NP,
<collectingRegion id="49A5F82F60648C00FD80D4C61BB0F997" box="[624,742,1532,1557]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="302">Vinh Phuc</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FC69D4C61AADF997" box="[921,1019,1532,1557]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FBF6D4C61DD8F997" box="[1030,1166,1532,1557]" pageId="1" pageNumber="302" value="2014-06-15">
<date id="FFDF100D60648C00FBF6D4C61DD8F997" box="[1030,1166,1532,1557]" pageId="1" pageNumber="302" value="2014-06-15">15-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FB69D4C618ACF9BA" ID-GBIF-Occurrence="1913892938" collectingDate="2014-06-23" country="Vietnam" location="Cao Bang" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Cao Bang">
1 ind.,
<collectingRegion id="49A5F82F60648C00FB16D4C61C02F997" box="[1254,1364,1532,1557]" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="302">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FEF1D7251835F9BA" box="[257,355,1567,1592]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE83D71A18ACF9BA" box="[371,506,1568,1592]" pageId="1" pageNumber="302" value="2014-06-23">
<date id="FFDF100D60648C00FE83D71A18ACF9BA" box="[371,506,1568,1592]" pageId="1" pageNumber="302" value="2014-06-23">23-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FDFAD71A1DD5F9BA" ID-GBIF-Occurrence="1913892986" box="[522,1155,1567,1592]" collectingDate="2014-06-25" country="Vietnam" location="Bac Kan" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Bac Kan">
1 ind.,
<collectingRegion id="49A5F82F60648C00FD91D71A1B90F9B5" box="[609,710,1568,1591]" country="Vietnam" name="Bac Kan" pageId="1" pageNumber="302">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FC7AD7251ABAF9BA" box="[906,1004,1567,1592]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FC0BD7251DD5F9BA" box="[1019,1155,1567,1592]" pageId="1" pageNumber="302" value="2014-06-25">
<date id="FFDF100D60648C00FC0BD7251DD5F9BA" box="[1019,1155,1567,1592]" pageId="1" pageNumber="302" value="2014-06-25">25-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FB63D71A18A3F9DF" ID-GBIF-Occurrence="1913892949" collectingDate="2014-06-26" country="Vietnam" location="Bac Kan" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Bac Kan">
1 ind.,
<collectingRegion id="49A5F82F60648C00FB1AD71A1C06F9B5" box="[1258,1360,1568,1591]" country="Vietnam" name="Bac Kan" pageId="1" pageNumber="302">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FF0FD77E1837F9DF" box="[255,353,1604,1629]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE9ED77E18A3F9DF" box="[366,501,1604,1629]" pageId="1" pageNumber="302" value="2014-06-26">
<date id="FFDF100D60648C00FE9ED77E18A3F9DF" box="[366,501,1604,1629]" pageId="1" pageNumber="302" value="2014-06-26">26-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FDF3D77E1C46F9DE" ID-GBIF-Occurrence="1913892965" box="[515,1296,1604,1629]" collectingDate="2014-07-04" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
2 ind.,
<location id="8EBE601660648C00FDA5D77F1B91F9DE" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FDA5D77F1B91F9DE" box="[597,711,1604,1628]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FCF0D77E1A34F9DF" box="[768,866,1604,1629]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FBEED77E1DD6F9DF" box="[1054,1152,1604,1629]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB7CD77E1C46F9DE" box="[1164,1296,1604,1628]" pageId="1" pageNumber="302" value="2014-07-04">
<date id="FFDF100D60648C00FB7CD77E1C46F9DE" box="[1164,1296,1604,1628]" pageId="1" pageNumber="302" value="2014-07-04">4-VII-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FAEDD77E1B29F902" ID-GBIF-Occurrence="1913892959" collectingDate="2014-07-08" country="Vietnam" location="Cao Bang" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Cao Bang">
2 ind.,
<collectingRegion id="49A5F82F60648C00FA9FD77E1984F902" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="302">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FE7ED75D18A6F902" box="[398,496,1639,1664]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE0DD7521B29F902" box="[509,639,1640,1664]" pageId="1" pageNumber="302" value="2014-07-08">
<date id="FFDF100D60648C00FE0DD7521B29F902" box="[509,639,1640,1664]" pageId="1" pageNumber="302" value="2014-07-08">8-VII-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FD7ED7521DA9F902" ID-GBIF-Occurrence="1913892957" box="[654,1279,1639,1664]" collectingDate="2014-07-09" country="Vietnam" location="Cao Bang" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Cao Bang">
1 ind.,
<collectingRegion id="49A5F82F60648C00FD10D7521A07F902" box="[736,849,1640,1664]" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="302">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FBFDD75D1D39F902" box="[1037,1135,1639,1664]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB8CD7521DA9F902" box="[1148,1279,1640,1664]" pageId="1" pageNumber="302" value="2014-07-09">
<date id="FFDF100D60648C00FB8CD7521DA9F902" box="[1148,1279,1640,1664]" pageId="1" pageNumber="302" value="2014-07-09">9-VII-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FAFDD75D1BACF927" ID-GBIF-Occurrence="1913892943" collectingDate="2015-04-18" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
5 ind.,
<location id="8EBE601660648C00FA90D7521995F926" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FA90D7521995F926" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FF07D7B61800F927" box="[247,342,1676,1701]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FDF9D7B61B3DF927" box="[521,619,1676,1701]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FD86D7B61BACF927" box="[630,762,1676,1701]" pageId="1" pageNumber="302" value="2015-04-18">
<date id="FFDF100D60648C00FD86D7B61BACF927" box="[630,762,1676,1701]" pageId="1" pageNumber="302" value="2015-04-18">18-IV-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FCF5D7B61C1DF927" ID-GBIF-Occurrence="1913892939" box="[773,1355,1676,1701]" collectingDate="2015-05-02" country="Vietnam" location="Lang Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Lang Son">
2 ind.,
<collectingRegion id="49A5F82F60648C00FCA2D7B71AE8F926" box="[850,958,1676,1701]" country="Vietnam" name="Lang Son" pageId="1" pageNumber="302">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FB81D7B61D85F927" box="[1137,1235,1676,1701]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB2DD7B61C1DF927" box="[1245,1355,1676,1701]" pageId="1" pageNumber="302" value="2015-05-02">
<date id="FFDF100D60648C00FB2DD7B61C1DF927" box="[1245,1355,1676,1701]" pageId="1" pageNumber="302" value="2015-05-02">2-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FAA6D7B61B39F94A" ID-GBIF-Occurrence="1913892940" collectingDate="2015-05-09" country="Vietnam" location="Ha Tinh" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Ha Tinh">
1 ind.,
<collectingRegion id="49A5F82F60648C00FF67D78A19A4F94A" box="[151,242,1711,1736]" country="Vietnam" name="Ha Tinh" pageId="1" pageNumber="302">Ha Tinh</collectingRegion>
Prov., central
<collectingCountry id="F376765D60648C00FE64D79518A0F94A" box="[404,502,1711,1736]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FDF1D7951B39F94A" box="[513,623,1711,1736]" pageId="1" pageNumber="302" value="2015-05-09">
<date id="FFDF100D60648C00FDF1D7951B39F94A" box="[513,623,1711,1736]" pageId="1" pageNumber="302" value="2015-05-09">9-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FD8AD78A1CD5F94A" ID-GBIF-Occurrence="1913892982" box="[634,1411,1711,1736]" collectingDate="2015-05-22" country="Vietnam" location="Cuc Phuong" pageId="1" pageNumber="302" specimenCount="1" specimenCount-female="1" stateProvince="Ninh Binh">
<specimenCount id="9D67FD4460648C00FD8AD78A1BF2F945" box="[634,676,1712,1736]" pageId="1" pageNumber="302" type="female">1 ♀</specimenCount>
,
<location id="8EBE601660648C00FD42D78A1A6DF94A" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FD42D78A1A6DF94A" box="[690,827,1711,1736]" country="Vietnam" name="Cuc Phuong" pageId="1" pageNumber="302" stateProvince="Ninh Binh">Cuc Phuong</location>
NP,
<collectingRegion id="49A5F82F60648C00FC81D7951AB0F94A" box="[881,998,1711,1736]" country="Vietnam" name="Ninh Binh" pageId="1" pageNumber="302">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FB6AD7951DAAF94A" box="[1178,1276,1711,1736]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FAF7D7951CD5F94A" box="[1287,1411,1711,1736]" pageId="1" pageNumber="302" value="2015-05-22">
<date id="FFDF100D60648C00FAF7D7951CD5F94A" box="[1287,1411,1711,1736]" pageId="1" pageNumber="302" value="2015-05-22">22-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FA7ED78A1AC8F96F" ID-GBIF-Occurrence="1913892948" collectingDate="2015-05-30" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
7 ind.,
<location id="8EBE601660648C00FF24D7EF181FF96E" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FF24D7EF181FF96E" box="[212,329,1748,1772]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FE78D7EE18BBF96F" box="[392,493,1748,1773]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FD41D7EE1A45F96F" box="[689,787,1748,1773]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FCD3D7EE1AC8F96F" box="[803,926,1748,1773]" pageId="1" pageNumber="302" value="2015-05-30">
<date id="FFDF100D60648C00FCD3D7EE1AC8F96F" box="[803,926,1748,1773]" pageId="1" pageNumber="302" value="2015-05-30">30-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FC5FD7EE18A0F892" ID-GBIF-Occurrence="1913892946" collectingDate="2015-06-16" country="Vietnam" location="Cuc Phuong" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Ninh Binh">
1 ind.,
<location id="8EBE601660648C00FBF6D7EE1DC3F96F" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FBF6D7EE1DC3F96F" box="[1030,1173,1748,1773]" country="Vietnam" name="Cuc Phuong" pageId="1" pageNumber="302" stateProvince="Ninh Binh">Cuc Phuong</location>
NP,
<collectingRegion id="49A5F82F60648C00FB25D7EE1C19F96F" box="[1237,1359,1748,1773]" country="Vietnam" name="Ninh Binh" pageId="1" pageNumber="302">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FF0FD7CD1837F892" box="[255,353,1783,1808]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE9ED7CD18A0F892" box="[366,502,1783,1808]" pageId="1" pageNumber="302" value="2015-06-16">
<date id="FFDF100D60648C00FE9ED7CD18A0F892" box="[366,502,1783,1808]" pageId="1" pageNumber="302" value="2015-06-16">16-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FDF4D7C21D2CF892" ID-GBIF-Occurrence="1913892968" box="[516,1146,1783,1808]" collectingDate="2015-06-28" country="Vietnam" location="Cao Bang" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Cao Bang">
2 ind.,
<collectingRegion id="49A5F82F60648C00FDA6D7C21B91F892" box="[598,711,1784,1808]" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="302">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FC73D7CD1AB3F892" box="[899,997,1783,1808]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FC03D7CD1D2CF892" box="[1011,1146,1783,1808]" pageId="1" pageNumber="302" value="2015-06-28">
<date id="FFDF100D60648C00FC03D7CD1D2CF892" box="[1011,1146,1783,1808]" pageId="1" pageNumber="302" value="2015-06-28">28-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FB78D7C21B27F8B7" ID-GBIF-Occurrence="1913892960" collectingDate="2015-06-29" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho">
<specimenCount id="9D67FD4460648C00FB78D7C21DE0F88D" box="[1160,1206,1784,1808]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FB35D7C21C60F892" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FB35D7C21C60F892" box="[1221,1334,1784,1808]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FA81D7CD1992F8B7" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FE8BD626188BF8B7" box="[379,477,1820,1845]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE19D6261B27F8B7" box="[489,625,1820,1845]" pageId="1" pageNumber="302" value="2015-06-29">
<date id="FFDF100D60648C00FE19D6261B27F8B7" box="[489,625,1820,1845]" pageId="1" pageNumber="302" value="2015-06-29">29-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FD8DD6261CD4F8B7" ID-GBIF-Occurrence="1913892971" box="[637,1410,1820,1845]" collectingDate="2015-07-19" country="Vietnam" location="Tam Dao" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc">
<specimenCount id="9D67FD4460648C00FD8DD6261BFCF8B6" box="[637,682,1820,1844]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FD46D6271A48F8B6" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FD46D6271A48F8B6" box="[694,798,1821,1844]" country="Vietnam" name="Tam Dao" pageId="1" pageNumber="302" stateProvince="Vinh Phuc">Tam Dao</location>
NP,
<collectingRegion id="49A5F82F60648C00FCA6D6261A9AF8B7" box="[854,972,1820,1845]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="302">Vinh Phuc</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FB73D6261DB3F8B7" box="[1155,1253,1820,1845]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB01D6261CD4F8B7" box="[1265,1410,1820,1845]" pageId="1" pageNumber="302" value="2015-07-19">
<date id="FFDF100D60648C00FB01D6261CD4F8B7" box="[1265,1410,1820,1845]" pageId="1" pageNumber="302" value="2015-07-19">19-VII-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FA7ED6261A3EF8DA" ID-GBIF-Occurrence="1913892952" collectingDate="2016-04-23" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho">
<specimenCount id="9D67FD4460648C00FA7ED62619F8F8D5" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FF4AD67A187CF8DA" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FF4AD67A187CF8DA" box="[186,298,1856,1880]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FE90D6051896F8DA" box="[352,448,1855,1880]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FD86D6051B8EF8DA" box="[630,728,1855,1880]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FD13D6051A3EF8DA" box="[739,872,1855,1880]" pageId="1" pageNumber="302" value="2016-04-23">
<date id="FFDF100D60648C00FD13D6051A3EF8DA" box="[739,872,1855,1880]" pageId="1" pageNumber="302" value="2016-04-23">23-IV-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FC83D67A19B8F8FF" ID-GBIF-Occurrence="1913892941" collectingDate="2016-05-03" country="Vietnam" location="Thua Thien - Hue Prov." pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D67FD4460648C00FC83D67A1AF6F8D5" box="[883,928,1856,1880]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FC5CD6051DEFF8D5" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FC5CD6051DEFF8D5" box="[940,1209,1855,1880]" country="Vietnam" name="Thua Thien - Hue Prov." pageId="1" pageNumber="302">Thua Thien - Hue Prov.</location>
, central
<collectingCountry id="F376765D60648C00FAE8D6051C2CF8DA" box="[1304,1402,1855,1880]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FA75D67A19B8F8FF" pageId="1" pageNumber="302" value="2016-05-03">
<date id="FFDF100D60648C00FA75D67A19B8F8FF" pageId="1" pageNumber="302" value="2016-05-03">3- V-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FF0DD65E1D53F8FF" ID-GBIF-Occurrence="1913892973" box="[253,1029,1892,1917]" collectingDate="2016-05-29" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho">
<specimenCount id="9D67FD4460648C00FF0DD65E1878F8FE" box="[253,302,1892,1916]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FECED65F18E4F8FE" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FECED65F18E4F8FE" box="[318,434,1892,1916]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FE01D65E1B03F8FF" box="[497,597,1892,1917]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FCE7D65E1A2FF8FF" box="[791,889,1892,1917]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FC79D65E1D53F8FF" box="[905,1029,1892,1917]" pageId="1" pageNumber="302" value="2016-05-29">
<date id="FFDF100D60648C00FC79D65E1D53F8FF" box="[905,1029,1892,1917]" pageId="1" pageNumber="302" value="2016-05-29">29-V-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FBE4D65E18DDF822" ID-GBIF-Occurrence="1913892947" collectingDate="2016-06-21" country="Vietnam" location="Quang Nam" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Quang Nam">
6 ind.,
<collectingRegion id="49A5F82F60648C00FB9BD65E1DAEF8FE" box="[1131,1272,1892,1916]" country="Vietnam" name="Quang Nam" pageId="1" pageNumber="302">Quang Nam</collectingRegion>
Prov., central
<collectingCountry id="F376765D60648C00FF67D6BD19AFF822" box="[151,249,1927,1952]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FEF4D6BD18DDF822" box="[260,395,1927,1952]" pageId="1" pageNumber="302" value="2016-06-21">
<date id="FFDF100D60648C00FEF4D6BD18DDF822" box="[260,395,1927,1952]" pageId="1" pageNumber="302" value="2016-06-21">21-VI-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FE67D6B21D79F822" ID-GBIF-Occurrence="1913892972" box="[407,1071,1927,1952]" collectingDate="2016-06-21" country="Vietnam" location="Thua Thien - Hue Prov." pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D67FD4460648C00FE67D6B21895F822" box="[407,451,1928,1952]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FE3FD6BD1B8AF822" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FE3FD6BD1B8AF822" box="[463,732,1927,1952]" country="Vietnam" name="Thua Thien - Hue Prov." pageId="1" pageNumber="302">Thua Thien - Hue Prov.</location>
, central
<collectingCountry id="F376765D60648C00FCCAD6BD1ACAF822" box="[826,924,1927,1952]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FC58D6BD1D79F822" box="[936,1071,1927,1952]" pageId="1" pageNumber="302" value="2016-06-21">
<date id="FFDF100D60648C00FC58D6BD1D79F822" box="[936,1071,1927,1952]" pageId="1" pageNumber="302" value="2016-06-21">21-VI-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FBCBD6B21B65F846" ID-GBIF-Occurrence="1913892950" collectingDate="2017-05-21" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
10 ind.,
<location id="8EBE601660648C00FB67D6B31C51F822" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FB67D6B31C51F822" box="[1175,1287,1928,1952]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FACCD6BD1CCAF822" box="[1340,1436,1927,1952]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FEB9D69618FDF847" box="[329,427,1964,1989]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FE48D6961B65F846" box="[440,563,1964,1988]" pageId="1" pageNumber="302" value="2017-05-21">
<date id="FFDF100D60648C00FE48D6961B65F846" box="[440,563,1964,1988]" pageId="1" pageNumber="302" value="2017-05-21">21-V-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FDB1D6961C10F846" ID-GBIF-Occurrence="1913892966" box="[577,1350,1964,1989]" collectingDate="2017-05-27" country="Vietnam" location="Xuan Son" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Phu Tho">
3 ind.,
<location id="8EBE601660648C00FD63D6971A53F846" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FD63D6971A53F846" box="[659,773,1964,1988]" country="Vietnam" name="Xuan Son" pageId="1" pageNumber="302" stateProvince="Phu Tho">Xuan Son</location>
NP,
<collectingRegion id="49A5F82F60648C00FCCED6961AF6F847" box="[830,928,1964,1989]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="302">Phu Tho</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FBABD6961DEBF847" box="[1115,1213,1964,1989]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FB3AD6961C10F846" box="[1226,1350,1964,1988]" pageId="1" pageNumber="302" value="2017-05-27">
<date id="FFDF100D60648C00FB3AD6961C10F846" box="[1226,1350,1964,1988]" pageId="1" pageNumber="302" value="2017-05-27">27-V-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C00FAA3D6961BFAF86B" ID-GBIF-Occurrence="1913892955" collectingDate="2017-06-05" country="Vietnam" location="Quang Binh" pageId="1" pageNumber="302" specimenCount="1" stateProvince="Quang Binh">
5 ind.,
<collectingRegion id="49A5F82F60648C00FF67D6EA1876F86B" box="[151,288,2000,2025]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="302">Quang Binh</collectingRegion>
Prov., central
<collectingCountry id="F376765D60648C00FE35D6EA1B71F86B" box="[453,551,2000,2025]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C00FDC3D6EA1BFAF86B" box="[563,684,2000,2025]" pageId="1" pageNumber="302" value="2017-06-05">
<date id="FFDF100D60648C00FDC3D6EA1BFAF86B" box="[563,684,2000,2025]" pageId="1" pageNumber="302" value="2017-06-05">5-VI-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060648C03FD49D6EA1985FF32" ID-GBIF-Occurrence="1913892976" collectingDate="2017-06-11" country="Vietnam" lastPageId="2" lastPageNumber="303" location="Cuc Phuong" pageId="1" pageNumber="302" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh">
<specimenCount id="9D67FD4460648C00FD49D6EA1BB0F86A" box="[697,742,2000,2024]" pageId="1" pageNumber="302" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660648C00FD03D6EA1A2BF86B" LSID="urn:lsid:plazi:treatment:03C887DB60648C03FF67D2CC1D51FC4A:8EBE601660648C00FD03D6EA1A2BF86B" box="[755,893,2000,2025]" country="Vietnam" name="Cuc Phuong" pageId="1" pageNumber="302" stateProvince="Ninh Binh">Cuc Phuong</location>
NP,
<collectingRegion id="49A5F82F60648C00FC45D6EA1D7DF86B" box="[949,1067,2000,2025]" country="Vietnam" name="Ninh Binh" pageId="1" pageNumber="302">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60648C00FB13D6EA1C13F86B" box="[1251,1349,2000,2025]" name="Vietnam" pageId="1" pageNumber="302">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560648C03FAA1D6EA1985FF32" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302" value="2017-06-11">
<date id="FFDF100D60648C03FAA1D6EA1985FF32" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302" value="2017-06-11">11-VI- 2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FF10D1A21A11FF32" ID-GBIF-Occurrence="1913892980" box="[224,839,151,176]" collectingDate="2017-07-09" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Bac Kan">
4 ind.,
<collectingRegion id="49A5F82F60678C03FEC4D1A218CEFF2D" box="[308,408,152,175]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FDA5D1AD1BE1FF32" box="[597,695,151,176]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD34D1A21A11FF32" box="[708,839,152,176]" pageId="2" pageNumber="303" value="2017-07-09">
<date id="FFDF100D60678C03FD34D1A21A11FF32" box="[708,839,152,176]" pageId="2" pageNumber="303" value="2017-07-09">9-VII-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FCA5D1A21985FF56" ID-GBIF-Occurrence="1913892964" collectingDate="2017-07-16" country="Vietnam" location="Cao Bang" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Cao Bang">
2 ind.,
<collectingRegion id="49A5F82F60678C03FC58D1A21D4CFF32" box="[936,1050,152,176]" country="Vietnam" name="Cao Bang" pageId="2" pageNumber="303">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FB27D1AD1C6FFF32" box="[1239,1337,151,176]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FAB7D1AD1985FF56" pageId="2" pageNumber="303" value="2017-07-16">
<date id="FFDF100D60678C03FAB7D1AD1985FF56" pageId="2" pageNumber="303" value="2017-07-16">16-VII- 2017</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B654660678C03FF37D1E51D51FC4A" pageId="2" pageNumber="303" type="discussion">
<paragraph id="8BDE36CD60678C03FF37D1E519A1FD07" blockId="2.[151,1437,151,968]" pageId="2" pageNumber="303">
<emphasis id="B915EADF60678C03FF37D1E5181AFF7A" bold="true" box="[199,332,223,248]" pageId="2" pageNumber="303">Discussion.</emphasis>
Males of
<taxonomicName id="4C614D4E60678C03FE4ED1E51BC4FF7A" box="[446,658,223,248]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60678C03FE4ED1E51BC4FF7A" box="[446,658,223,248]" italics="true" pageId="2" pageNumber="303">Asiagomphus acco</emphasis>
</taxonomicName>
are easily identified by their thorax pattern with a dorsal stripe and a single lateral stripe (over the mesepimeron), but without antehumeral stripe or second lateral stripe over the metepisternum, and with all yellow metepimeron. S12 are largely yellow, but S38 are black, with sometimes a small antero-dorsal spot on S7. S9 has a large yellow postero-dorsal spot. The pattern is very consistent (
<figureCitation id="135A2A4860678C03FAB6D0761CC3FEE6" box="[1350,1429,332,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1992,2014]" captionTargetBox="[204,1391,193,1976]" captionTargetId="figure@3.[204,1391,193,1976]" captionTargetPageId="3" captionText="FIGURE 1. Asiagomphus acco copula, Xuan Son NP, 30-V-2015." httpUri="https://zenodo.org/record/1441613/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
,
<figureCitation id="135A2A4860678C03FF67D04A1986FE0A" box="[151,208,368,392]" captionStart="FIGURE 2" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1437,193,1221]" captionTargetId="figure@5.[151,1437,193,1221]" captionTargetPageId="5" captionText="FIGURE 2. Asiagomphus acco alive in hand. [ac] male, [ef] female. (a) Ba Be NP, 21-VII-2013; (b) Quang Nam Prov., 21- VI-2016; (c) Tam Dao NP, 18-V-2014; (d) Yen Bai Prov., 20-VI-2015; (e) Xuan Son NP, 22-V-2015; (f) Lang Son Prov., 2-V- 2015." httpUri="https://zenodo.org/record/1441615/files/figure.png" pageId="2" pageNumber="303">2ac</figureCitation>
). The combination of largely black abdomen and black metepisternum is enough to identify the male, but in addition the sub-basal ventral tooth to the cerci in combination with the distinct slender branches of the epiproct in lateral view, which are very straight both dorsally and ventrally (
<figureCitation id="135A2A4860678C03FC8ED08D1A84FE52" box="[894,978,439,464]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="2" pageNumber="303">Fig. 6h</figureCitation>
), can be used for in-hand identification. The female has the same thorax pattern as the male, but has largely yellow S12, with only a black dorso-lateral spot on S1 and some black centrally on the dorsum of S2 at the posterior margin. S36 all have paired posterior dorsal spots, somewhat increasing in size from S3 to S6. S78 and S10 are black, or rarely S8 with a small dorsal yellow spot, and S9 has a relatively small postero-dorsal spot. The female too is quite consistently patterned (
<figureCitation id="135A2A4860678C03FA94D37D19F3FD06" captionStart="FIGURE 1" captionStartId="3.[151,250,1992,2014]" captionTargetBox="[204,1391,193,1976]" captionTargetId="figure@3.[204,1391,193,1976]" captionTargetPageId="3" captionText="FIGURE 1. Asiagomphus acco copula, Xuan Son NP, 30-V-2015." httpUri="https://zenodo.org/record/1441613/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
,
<figureCitation id="135A2A4860678C03FF43D35619BCFD07" box="[179,234,620,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1437,193,1221]" captionTargetId="figure@5.[151,1437,193,1221]" captionTargetPageId="5" captionText="FIGURE 2. Asiagomphus acco alive in hand. [ac] male, [ef] female. (a) Ba Be NP, 21-VII-2013; (b) Quang Nam Prov., 21- VI-2016; (c) Tam Dao NP, 18-V-2014; (d) Yen Bai Prov., 20-VI-2015; (e) Xuan Son NP, 22-V-2015; (f) Lang Son Prov., 2-V- 2015." httpUri="https://zenodo.org/record/1441615/files/figure.png" pageId="2" pageNumber="303">2df</figureCitation>
).
</paragraph>
<paragraph id="8BDE36CD60678C03FF37D3B51D19FCBA" blockId="2.[151,1437,151,968]" pageId="2" pageNumber="303">
<bibRefCitation id="EFF04B3C60678C03FF37D3B51822FD2A" box="[199,372,655,680]" pageId="2" pageNumber="303" refId="ref17031" refString="Asahina, S. (1996) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 3. Aeschnidae [sic], Corduliidae and Libellulidae. Bulletin of the National Science Museum, Tokyo, Series A, 22, 69 - 80." type="journal article">Asahina (1996)</bibRefCitation>
originally described
<taxonomicName id="4C614D4E60678C03FD90D3B51A64FD2A" box="[608,818,655,680]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60678C03FD90D3B51A64FD2A" box="[608,818,655,680]" italics="true" pageId="2" pageNumber="303">Asiagomphus acco</emphasis>
</taxonomicName>
as a subspecies of
<emphasis id="B915EADF60678C03FBF7D3AA1DE3FD2A" box="[1031,1205,655,680]" italics="true" pageId="2" pageNumber="303">
<taxonomicName id="4C614D4E60678C03FBF7D3AA1DE7FD2A" baseAuthorityName="Williamson" baseAuthorityYear="1907" box="[1031,1201,655,680]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="xanthenatus">A. xanthenatus</taxonomicName>
.
</emphasis>
However, this was a strange taxonomic solution, as that species has clearly different cerci without a ventral sub-basal tooth and different epiproct (compare
<figureCitation id="135A2A4860678C03FE9ED3ED189AFD72" box="[366,460,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="2" pageNumber="303">Figs. 5h</figureCitation>
,
<figureCitation id="135A2A4860678C03FE2AD3ED18A0FD72" box="[474,502,727,752]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="2" pageNumber="303">6h</figureCitation>
with
<figureCitation id="135A2A4860678C03FDC9D3ED1BC0FD72" box="[569,662,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="2" pageNumber="303">Figs. 5e</figureCitation>
,
<figureCitation id="135A2A4860678C03FD53D3ED1B97FD72" box="[675,705,727,752]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="2" pageNumber="303">6e</figureCitation>
). It is also completely differently patterned (compare
<figureCitation id="135A2A4860678C03FAD9D3ED1C21FD72" box="[1321,1399,727,752]" captionStart="FIGURE 1" captionStartId="3.[151,250,1992,2014]" captionTargetBox="[204,1391,193,1976]" captionTargetId="figure@3.[204,1391,193,1976]" captionTargetPageId="3" captionText="FIGURE 1. Asiagomphus acco copula, Xuan Son NP, 30-V-2015." httpUri="https://zenodo.org/record/1441613/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
,
<figureCitation id="135A2A4860678C03FA77D3E21CC1FD72" box="[1415,1431,728,752]" captionStart="FIGURE 2" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1437,193,1221]" captionTargetId="figure@5.[151,1437,193,1221]" captionTargetPageId="5" captionText="FIGURE 2. Asiagomphus acco alive in hand. [ac] male, [ef] female. (a) Ba Be NP, 21-VII-2013; (b) Quang Nam Prov., 21- VI-2016; (c) Tam Dao NP, 18-V-2014; (d) Yen Bai Prov., 20-VI-2015; (e) Xuan Son NP, 22-V-2015; (f) Lang Son Prov., 2-V- 2015." httpUri="https://zenodo.org/record/1441615/files/figure.png" pageId="2" pageNumber="303">2</figureCitation>
,
<figureCitation id="135A2A4860678C03FF67D3C6198AFC97" box="[151,220,764,789]" captionStart="FIGURE 13" captionStartId="22.[151,250,1766,1788]" captionTargetBox="[151,1437,481,1745]" captionTargetId="figure@22.[151,1437,481,1745]" captionTargetPageId="22" captionText="FIGURE 13. Asiagomphus spp. males in nature. (a) A. acco, Xuan Son NP, 8-VI-2014; (b) A. acco, Pia Oac NR, 8-VII-2014; (c) A. kosterini, Lam Dong Prov., 17-IV-2016; (d) A. pacificus, Cao Bang Prov., 19-V-2015; (e) A. reinhardti, Gia Lai Prov., 20- VI-2016; (f) A. reinhardti, Gia Lai Prov., 13-V-2017." httpUri="https://zenodo.org/record/1441637/files/figure.png" pageId="2" pageNumber="303">13ab</figureCitation>
with
<figureCitation id="135A2A4860678C03FEEBD3C6182FFC97" box="[283,377,764,789]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [ad, g] male, [ef, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="2" pageNumber="303">Fig. 10d</figureCitation>
) and has different hamuli (compare
<figureCitation id="135A2A4860678C03FCE0D3C61AD3FC97" box="[784,901,764,789]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (ab) Asiagomphus acco, Tam Dao NP, 18-V-2014; (cd) A. kosterini sp. nov. holotype; (ef) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="2" pageNumber="303">Figs. 3ab</figureCitation>
with
<figureCitation id="135A2A4860678C03FC35D3C61D6FFC97" box="[965,1081,764,789]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (ab) Asiagomphus acco, Tam Dao NP, 18-V-2014; (cd) A. kosterini sp. nov. holotype; (ef) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="2" pageNumber="303">Figs. 3ef</figureCitation>
). I strongly support the view by
<bibRefCitation id="EFF04B3C60678C03FF67D21A181FFCBA" box="[151,329,799,824]" pageId="2" pageNumber="303" refId="ref17122" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Phan, Q. T. (2011) Anisoptera of Cuc Phuong National Park, North Vietnam. International Dragonfly Fund - Report, Newsletter of the International Dragonfly Fund, 33, 1 - 18." type="journal article">
Do
<emphasis id="B915EADF60678C03FF30D21B19AEFCBA" box="[192,248,799,824]" italics="true" pageId="2" pageNumber="303">et al.</emphasis>
(2011)
</bibRefCitation>
that
<taxonomicName id="4C614D4E60678C03FE71D21A1883FCBA" box="[385,469,800,824]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60678C03FE71D21A1883FCBA" box="[385,469,800,824]" italics="true" pageId="2" pageNumber="303">A. acco</emphasis>
</taxonomicName>
should be considered a separate species in its own right.
</paragraph>
<paragraph id="8BDE36CD60678C03FF37D27E1D51FC4A" blockId="2.[151,1437,151,968]" pageId="2" pageNumber="303">
<emphasis id="B915EADF60678C03FF37D27E18EBFCDF" bold="true" box="[199,445,836,861]" pageId="2" pageNumber="303">Habitat and ecology.</emphasis>
<taxonomicName id="4C614D4E60678C03FE34D27E1BCEFCDE" box="[452,664,836,861]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60678C03FE34D27E1BCEFCDE" box="[452,664,836,861]" italics="true" pageId="2" pageNumber="303">Asiagomphus acco</emphasis>
</taxonomicName>
is a spring species, locally common, appearing in the second half of April and flying to the end of July. Sampling bias has been towards northern
<collectingCountry id="F376765D60678C03FC1ED25D1D06FC02" box="[1006,1104,871,896]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
, but the species was observed as far south as
<collectingRegion id="49A5F82F60678C03FEB5D2B61B6AFC27" box="[325,572,908,933]" country="Vietnam" name="Quang Nam" pageId="2" pageNumber="303">
Quang
<collectingRegion id="49A5F82F60678C03FE69D2B71B6AFC27" box="[409,572,908,933]" inferredBySuffix="true" pageId="2" pageNumber="303">Nam Province</collectingRegion>
</collectingRegion>
in central
<collectingCountry id="F376765D60678C03FD48D2B61A4CFC27" box="[696,794,908,933]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
. It inhabits clear, shallow streams, generally under deep forest cover, and is very approachable, unlike many other species in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>