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<document ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" approvalRequired="56" approvalRequired_for_textStreams="56" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee &amp; Nützel, Al- Exander" docDate="2020" docId="03B587AB4F2C1565FF5179A3FC35FB42" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@1ef1a562" docStyleName="zootaxa.v2_0.2013.journal_article" docTitle="Naticopsis McCoy 1844" docType="treatment" docVersion="5" lastPageNumber="28" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="28" updateTime="1643439188523" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ketwetsuriya, Chatchalerm</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karapunar, Baran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Charoentitirat, Thasinee</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nützel, Al- Exander</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4766</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>1</mods:start>
<mods:end>47</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.4766.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">ed4d1bf6-284a-4898-952a-58943d5a3026</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">3763907</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803899" ID-GBIF-Taxon="163737130" ID-Zenodo-Dep="3803899" LSID="urn:lsid:plazi:treatment:03B587AB4F2C1565FF5179A3FC35FB42" httpUri="http://treatment.plazi.org/id/03B587AB4F2C1565FF5179A3FC35FB42" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection box="[151,341,477,503]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph blockId="27.[151,341,477,539]" box="[151,341,477,503]" pageId="27" pageNumber="28">
<heading bold="true" box="[151,341,477,503]" fontSize="10" level="2" pageId="27" pageNumber="28" reason="2">
<emphasis bold="true" box="[151,341,477,503]" pageId="27" pageNumber="28">
<taxonomicName authorityName="McCoy" authorityYear="1844" box="[151,276,477,503]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis bold="true" box="[151,276,477,503]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
</taxonomicName>
sp. 2
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,299,513,539]" pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[151,341,477,539]" box="[151,299,513,539]" pageId="27" pageNumber="28">
(
<figureCitation box="[160,240,513,539]" captionStart="FIGURE 20" captionStartId="28.[151,250,1717,1742]" captionTargetBox="[179,1409,309,1693]" captionTargetId="figure@28.[179,1409,309,1693]" captionTargetPageId="28" captionText="FIGURE 20. AC, Naticopsis sp. 1, ESKU-19-LP 47. DF, Naticopsis sp. 2, ESKU-19-LP 46. GI, Naticopsis? sp. 3, ESKU- 19-LP 32. J, Naticopsis? sp. 4, ESKU-19-LP 52. KL, Naticopsis cf. heshanensis Pan &amp; Erwin, 2002, ESKU-19-LP 44. Scale bars represent: 2 mm (AJ); 5 mm (KL)." figureDoi="http://doi.org/10.5281/zenodo.3763947" httpUri="https://zenodo.org/record/3763947/files/figure.png" pageId="27" pageNumber="28">Fig. 20</figureCitation>
DF)
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="materials_examined">
<paragraph blockId="27.[151,1437,584,1223]" box="[151,637,584,611]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,262,584,611]" pageId="27" pageNumber="28">Material.</emphasis>
One specimen: ESKU-19-
<collectionCode box="[561,595,585,611]" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP</collectionCode>
46.
</paragraph>
<paragraph blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
Dimensions (mm): ESKU-19-
<collectionCode box="[539,573,621,647]" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP</collectionCode>
46: height = 9.3; width = 10.3; apical angle = 112
<superScript attach="none" box="[1159,1164,620,634]" fontSize="6" pageId="27" pageNumber="28">º</superScript>
; apertural height = 7.5; apertural width = 7.7.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,345,692,719]" pageId="27" pageNumber="28">Description.</emphasis>
Moderately low-spired shell, naticiform, consisting of three rapidly expanding whorls; whorls smooth, strongly convex, rounded, embracing at periphery; spire low but distinctly elevated; body-whorl inflated, very large and much higher than spire with height about 90% of total height; periphery at about mid-whorl; base convex; suture shallow but impressed; anomphalous; aperture evenly rounded in cross section, lip thin, inner lip arched.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="discussion">
<paragraph blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,315,873,899]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName authorityName="McCoy" authorityYear="1844" box="[322,440,874,899]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[322,440,874,899]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
</taxonomicName>
sp. 2 differs from
<taxonomicName authorityName="McCoy" authorityYear="1844" box="[649,767,874,899]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[649,767,874,899]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
</taxonomicName>
sp.
<quantity box="[813,856,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="28" unit="in" value="1.0">1 in</quantity>
having more flattened upper whorl surface, lacking callus, having a higher whorl expansion rate and a rounded aperture. It resembles
<taxonomicName authority="Worthen in Meek &amp; Worthen (1873)" authorityName="Worthen in Meek &amp; Worthen" authorityYear="1873" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
<emphasis box="[1076,1303,909,935]" italics="true" pageId="27" pageNumber="28">Naticopsis subovata</emphasis>
Worthen
<emphasis box="[1414,1436,910,935]" italics="true" pageId="27" pageNumber="28">in</emphasis>
<bibRefCitation author="Meek &amp; Worthen" box="[151,431,945,971]" firstAuthor="Meek" pageId="27" pageNumber="28" pagination="321 - 619" refId="ref26472" refString="Meek, F. B. &amp; Worthen, A. H. (1873) Descriptions of invertebrates from the Carboniferous System. Illinois Geological Survey, 5 (2), 321 - 619." type="journal article" year="1873">Meek &amp; Worthen (1873)</bibRefCitation>
</taxonomicName>
as illustrated by
<bibRefCitation author="Knight" box="[628,793,945,971]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
Knight (1933
<emphasis box="[779,793,945,971]" italics="true" pageId="27" pageNumber="28">b</emphasis>
</bibRefCitation>
, p. 379, pl. 43, fig. 2aj) in size, shell shape and pleural angle.
<taxonomicName authority="Knight, 1933 b" authorityName="Knight" authorityYear="1933" box="[228,535,981,1007]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
<emphasis box="[228,247,982,1007]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[262,362,981,1007]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
<bibRefCitation author="Knight" box="[371,535,981,1007]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
Knight, 1933
<emphasis box="[521,535,981,1007]" italics="true" pageId="27" pageNumber="28">b</emphasis>
</bibRefCitation>
</taxonomicName>
(p. 377, pl. 43, fig. 3ak) is also similar.
<bibRefCitation author="Knight" box="[1016,1191,981,1007]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
Knight (1933
<emphasis box="[1168,1182,981,1007]" italics="true" pageId="27" pageNumber="28">b</emphasis>
)
</bibRefCitation>
mentioned that these two species,
<taxonomicName authorityName="Worthen in Meek &amp; Worthen" authorityYear="1873" box="[297,432,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
<emphasis box="[297,316,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[331,432,1017,1043]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Knight" authorityYear="1933" box="[488,624,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
<emphasis box="[488,507,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[522,624,1017,1043]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
</taxonomicName>
<emphasis box="[624,628,1017,1043]" italics="true" pageId="27" pageNumber="28">,</emphasis>
are very close to each other but
<taxonomicName authorityName="Worthen in Meek &amp; Worthen" authorityYear="1873" box="[1000,1134,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
<emphasis box="[1000,1019,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[1033,1134,1017,1043]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName authorityName="Knight" authorityYear="1933" box="[151,284,1053,1079]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
<emphasis box="[151,170,1054,1079]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[184,284,1053,1079]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
</taxonomicName>
in having a swelling just above the mid-whorl height, and by the shape of the columellar lip, which is evenly crescent-shaped in
<taxonomicName authorityName="Worthen in Meek &amp; Worthen" authorityYear="1873" box="[451,586,1089,1115]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
<emphasis box="[451,470,1090,1115]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[485,586,1089,1115]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
</taxonomicName>
while it is slightly angled in
<taxonomicName authorityName="Knight" authorityYear="1933" box="[920,1054,1089,1115]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
<emphasis box="[920,939,1090,1115]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[954,1054,1089,1115]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
</taxonomicName>
. Although the parietal area of the present specimen is obscure it seems closer to
<taxonomicName authorityName="Worthen in Meek &amp; Worthen" authorityYear="1873" box="[679,814,1125,1151]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
<emphasis box="[679,698,1126,1151]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[713,814,1125,1151]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
</taxonomicName>
. The present specimen is also similar to the specimens illustrated and described as
<taxonomicName authority="Wanner, 1922" authorityName="Wanner" authorityYear="1922" box="[458,744,1161,1187]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="praealta">
<emphasis box="[458,477,1162,1187]" italics="true" pageId="27" pageNumber="28">N</emphasis>
.
<emphasis box="[489,584,1161,1187]" italics="true" pageId="27" pageNumber="28">praealta</emphasis>
<bibRefCitation author="Wanner" box="[590,744,1161,1187]" firstAuthor="Wanner" pageId="27" pageNumber="28" pagination="1 - 82" refId="ref28413" refString="Wanner, C. (1922) Die Gastropoden und Lamellibranchiaten der Dyas von Timor. Palaontologie von Timor, 11, 1 - 82." type="journal article" year="1922">Wanner, 1922</bibRefCitation>
</taxonomicName>
by
<bibRefCitation author="Batten" box="[783,931,1161,1187]" firstAuthor="Batten" pageId="27" pageNumber="28" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
; p. 14, fig. 16ab) from the Permian of
<collectingRegion box="[1366,1433,1161,1187]" country="Malaysia" name="Perak" pageId="27" pageNumber="28">Perak</collectingRegion>
,
<collectingCountry box="[151,254,1197,1223]" name="Malaysia" pageId="27" pageNumber="28">Malaysia</collectingCountry>
but the latter specimens are more slender and relatively high-spired.
</paragraph>
</subSubSection>
</treatment>
</document>