668 lines
79 KiB
XML
668 lines
79 KiB
XML
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<mods:title id="6DD1AB306FC9322D8173B4FC5D2A4DAC">Two new species of the remarkable scorpion genus Megacormus Karsch, 1881 (Scorpiones: Euscorpiidae)</mods:title>
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<mods:namePart id="CBCBD311413C93E74EAF0EBAF171973E">Teruel, Rolando</mods:namePart>
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<mods:namePart id="FE973308099C0AB2A656756D92D65568">Kovařík, František</mods:namePart>
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<mods:namePart id="E72C1C29D8F96B359B09BA09946CFAC1">Lowe, Graeme</mods:namePart>
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<mods:namePart id="ECD5C69AC3205671C9527F577A80C5A0">Šťáhlavský, František</mods:namePart>
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<mods:title id="90B9EBE573F10CAC6147DFEC420F97A8">Euscorpius</mods:title>
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<mods:part id="10C0CF789495786EE248BB902B76665E">
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<mods:date id="897C8E1BF2CC0A05179D410B5ACA248E">2023</mods:date>
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<heading id="D0FE81B0FFAB3202FC6006D41950DCD3" box="[966,1324,1238,1268]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="4" reason="2">
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<taxonomicName id="4C094D5FFFAB3202FC6006D418A0DCD4" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[966,1244,1238,1267]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">Megacormus orizaba</taxonomicName>
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.
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<emphasis id="B97DEACEFFAB3202FAB506D41959DCD4" bold="true" box="[1299,1317,1239,1267]" pageId="5" pageNumber="4">n</emphasis>
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.
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<paragraph id="8BB636DCFFAB3202FC3506F819C7DD15" blockId="5.[823,1467,1238,1366]" pageId="5" pageNumber="4">
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(Figures 58–59, 61–62, 66, 82,
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<tableCitation id="C68B0367FFAB3202FB4406F8192BDD34" box="[1250,1367,1275,1299]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="4.[113,169,1006,1027]" captionStartId-1="20.[111,167,905,926]" captionText-0="Table 1. Morphometric data for types of Megacormus orizaba sp. n. and Megacormus seductus sp. n. Abbreviations: length (L), width (W), in carapace it corresponds to median width), depth (D)." captionText-1="Table 2. Variation in number of tines on the crown-like structure (cls) of the hemispermatophore of Megacormus species. Data with DNA No. and locality code were obtained from examined hemispermatophores. Other data were obtained from published descriptions or illustrations (1Francke, 1979; 2González-Santillán & Alvarez-Padilla, 2015; 3Sissom, 1994: fig. 8; 4González-Santillán et al., 2017)." pageId="5" pageNumber="4">Tables 1–2</tableCitation>
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)
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<emphasis id="B97DEACEFFAB3202FC62073D1953DD71" bold="true" box="[964,1327,1342,1366]" pageId="5" pageNumber="4">4E89-4654-9375-86633841D26D</emphasis>
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<taxonomicName id="4C094D5FFFAB3202FC96077C1974DDB1" ID-CoL="6R89F" authority=": KovarIk, 2019: 1" authorityName=": KovarIk" authorityPageNumber="1" authorityYear="2019" box="[816,1288,1406,1432]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="granosus">
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<emphasis id="B97DEACEFFAB3202FC96077C1856DDB1" box="[816,1066,1407,1430]" italics="true" pageId="5" pageNumber="4">Megacormus granosus</emphasis>
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: KovařÍk, 2019: 1
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, 11–13, 21–27; figs. 40–41, 51, 90–123 (misidentification).
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west of
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, northeast slope of
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Pico
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[= Citlaltépetl]
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;
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,
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(
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(
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, northeast slope of
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Pico
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[=
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],
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,
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a. s. l.
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, pine forest, under rocks, 2019,
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, No. 1621, figs. 40, 51, 90–91, 94–102, 106–108, 111–114, 115–
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KovařÍk
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, 2019)
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,
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|
||
<specimenCount id="9D0FFD55FFAB3202FC5D05581867DF54" box="[1019,1051,1883,1907]" pageId="5" pageNumber="4" type="female">2♀</specimenCount>
|
||
(paratopotypes, figs. 41, 51, 92–93, 103–105, 109–110, 118–
|
||
<quantity id="4CF19B39FFAB3202FB9B057818C0DFB4" box="[1085,1212,1915,1939]" metricMagnitude="0" metricUnit="m" metricValue="3.0510988" pageId="5" pageNumber="4" unit="in" value="120.122">120, 122 in</quantity>
|
||
<location id="8ED66007FFAB3202FB6E05781963DFB2" LSID="urn:lsid:plazi:treatment:03A087CAFFAB320DFC6006D41D54DEE4:8ED66007FFAB3202FB6E05781963DFB2" box="[1224,1311,1915,1941]" country="Mexico" name="KovarIk" pageId="5" pageNumber="4" stateProvince="Puebla">KovařÍk</location>
|
||
, 2019)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B613C81FFAB3202FAD805781FE7DF94" collectingDate="2022-11-02" collectionCode="FKCP" country="Mexico" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Puebla" typeStatus="holotype">
|
||
<collectionCode id="ED18AE19FFAB3202FAD8057819C3DFB4" box="[1406,1471,1915,1939]" pageId="5" pageNumber="4">FKCP</collectionCode>
|
||
,
|
||
<date id="FFB7101CFFAB3202FC9605981FE7DF94" box="[816,923,1947,1971]" pageId="5" pageNumber="4" value="2022-11-02">
|
||
<collectingDate id="EFF3E9F4FFAB3202FC9605981FE7DF94" box="[816,923,1947,1971]" pageId="5" pageNumber="4" value="2022-11-02">2.XI.2022</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B613C81FFAB3202FC0205981917DF94" box="[932,1387,1947,1971]" collectionCode="R" collectorName="Teruel" country="Mexico" pageId="5" pageNumber="4" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Puebla" typeStatus="holotype">
|
||
<collectionCode id="ED18AE19FFAB3202FC0205981FC4DF95" box="[932,952,1947,1970]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="4">R</collectionCode>
|
||
.
|
||
<collectorName id="26FC530AFFAB3202FC670598187EDF94" box="[961,1026,1947,1971]" pageId="5" pageNumber="4">Teruel</collectorName>
|
||
et al.,
|
||
<specimenCount id="9D0FFD55FFAB3202FBE305981819DF94" box="[1093,1125,1947,1971]" pageId="5" pageNumber="4" type="male">2♂</specimenCount>
|
||
<specimenCount id="9D0FFD55FFAB3202FBC005981803DF94" box="[1126,1151,1947,1971]" pageId="5" pageNumber="4" type="female">8♀</specimenCount>
|
||
7juvs. (paratopotypes)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED18AE19FFAB3202FAD2059819C7DF94" box="[1396,1467,1947,1971]" pageId="5" pageNumber="4">RTOC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136557FFAB3202FC9605DD1884D011" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="8BB636DCFFAB3202FC9605DD1884D011" blockId="5.[816,1474,2014,2102]" pageId="5" pageNumber="4">ETYMOLOGY. The selected epithet is a Latinized noun in apposition, taken directly from the Spanish toponym of the majestic volcano inhabited by this species.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF766654FFA83201FFD705BE1E18DFD7" ID-DOI="http://doi.org/10.5281/zenodo.8222210" ID-Zenodo-Dep="8222210" httpUri="https://zenodo.org/record/8222210/files/figure.png" pageId="6" pageNumber="5" startId="6.[113,190,1981,2003]" targetBox="[170,1395,231,1967]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BB636DCFFA83201FFD705BE1E18DFD7" blockId="6.[113,1471,1981,2032]" pageId="6" pageNumber="5">
|
||
<emphasis id="B97DEACEFFA83201FFD705BE1C94DFF5" bold="true" box="[113,232,1981,2003]" pageId="6" pageNumber="5">Figures 3–6</emphasis>
|
||
:
|
||
<taxonomicName id="4C094D5FFFA83201FF5005BD1DB1DFF4" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[246,461,1982,2003]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA83201FF5005BD1DB1DFF4" box="[246,461,1982,2003]" italics="true" pageId="6" pageNumber="5">Megacormus seductus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA83201FE7205BD1E76DFF4" box="[468,522,1982,2003]" pageId="6" pageNumber="5" rank="species">
|
||
<emphasis id="B97DEACEFFA83201FE7205BD1D97DFF4" bold="true" box="[468,491,1982,2003]" pageId="6" pageNumber="5">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA83201FE5105BD1E78DFF4" bold="true" box="[503,516,1982,2003]" pageId="6" pageNumber="5">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B97DEACEFFA83201FDB705BE1EF5DFF5" bold="true" box="[529,649,1981,2003]" pageId="6" pageNumber="5">Figures 3–4</emphasis>
|
||
. Male holotype in dorsal (3) and ventral (4) views.
|
||
<emphasis id="B97DEACEFFA83201FB2705BE1885DFF5" bold="true" box="[1153,1273,1981,2003]" pageId="6" pageNumber="5">Figures 5–6</emphasis>
|
||
. Female paratype in dorsal (5) and ventral (6) views. Scale bars: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3136557FFA93200FFD702E81E6CDB45" pageId="7" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8BB636DCFFA93200FFD702E81D61DA45" blockId="7.[113,772,235,611]" pageId="7" pageNumber="6">
|
||
DIAGNOSIS. Adult size large for the genus (
|
||
<quantity id="4CF19B39FFA93200FDC002E81E9CD925" box="[614,736,235,259]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="7" pageNumber="6" unit="mm" value="32.0" valueMax="34.0" valueMin="30.0">30–34 mm</quantity>
|
||
in males,
|
||
<quantity id="4CF19B39FFA93200FF6603091D4AD905" box="[192,310,266,290]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="7" pageNumber="6" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
|
||
in females). Coloration entirely blackish, paler and less spotted on chelicerae, pedipalp, legs and pectines. Pedipalp patella with 22–24 external trichobothria (7
|
||
<emphasis id="B97DEACEFFA93200FF2803681CDBD9A4" box="[142,167,363,387]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
|
||
, 2
|
||
<emphasis id="B97DEACEFFA93200FF6E03681C90D9A4" box="[200,236,363,387]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
|
||
, 4–5
|
||
<emphasis id="B97DEACEFFA93200FE8103681D3AD9A5" box="[295,326,363,386]" italics="true" pageId="7" pageNumber="6">em</emphasis>
|
||
, 4
|
||
<emphasis id="B97DEACEFFA93200FEC003681DF8D9A5" box="[358,388,363,386]" italics="true" pageId="7" pageNumber="6">est</emphasis>
|
||
, 4
|
||
<emphasis id="B97DEACEFFA93200FE0303681DC4D9A5" box="[421,440,363,386]" italics="true" pageId="7" pageNumber="6">et</emphasis>
|
||
) and 6–8 ventral trichobothria. Pedipalp fingers of adult male with undulate proximal dentate margins, lobe/notch combination moderate. Pectines with shaft clearly divided into areas and lamellae by conspicuous sulci; tooth count
|
||
<quantity id="4CF19B39FFA93200FE9C03E91DFDDA24" box="[314,385,490,515]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="7" pageNumber="6" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
|
||
males,
|
||
<quantity id="4CF19B39FFA93200FE7F03E91E5CDA24" box="[473,544,490,515]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="7" pageNumber="6" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
|
||
females. Metasomal segment V length/width ratio 2.37–2.40 in males, 2.38–2.40 in females. Telson elongate, narrower than metasomal segment V in both sexes.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FFD700881E6CDB45" blockId="7.[113,772,650,867]" pageId="7" pageNumber="6">
|
||
SEXUAL DIMORPHISM. Female differing from male by:
|
||
<emphasis id="B97DEACEFFA93200FD1D00891EADDA85" bold="true" box="[699,721,650,674]" pageId="7" pageNumber="6">1)</emphasis>
|
||
size larger;
|
||
<emphasis id="B97DEACEFFA93200FF6400A91CA4DAE5" bold="true" box="[194,216,682,706]" pageId="7" pageNumber="6">2)</emphasis>
|
||
coloration darker and more densely infuscate;
|
||
<emphasis id="B97DEACEFFA93200FD4B00A91F7FDAE5" bold="true" box="[749,771,682,706]" pageId="7" pageNumber="6">3)</emphasis>
|
||
pedipalps, legs and metasoma slightly longer and slenderer;
|
||
<emphasis id="B97DEACEFFA93200FD4B00C91F7FDAC5" bold="true" box="[749,771,714,738]" pageId="7" pageNumber="6">4)</emphasis>
|
||
mesosoma much wider and more convex-sided;
|
||
<emphasis id="B97DEACEFFA93200FDDF00E91EF3DB25" bold="true" box="[633,655,746,770]" pageId="7" pageNumber="6">5)</emphasis>
|
||
pedipalps, carapace, tergites and metasoma with granulation sparser and finer;
|
||
<emphasis id="B97DEACEFFA93200FF1001291CB0DB65" bold="true" box="[182,204,810,834]" pageId="7" pageNumber="6">6)</emphasis>
|
||
genital papillae absent;
|
||
<emphasis id="B97DEACEFFA93200FE4301291D87DB65" bold="true" box="[485,507,810,834]" pageId="7" pageNumber="6">7)</emphasis>
|
||
pectines conspicuously smaller, with lower tooth counts (4–5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136557FFA9320EFFD701881956DCC8" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6" type="description">
|
||
<paragraph id="8BB636DCFFA93200FFD701881D6FDBC4" blockId="7.[113,772,906,2019]" pageId="7" pageNumber="6">
|
||
DESCRIPTION (male
|
||
<typeStatus id="54B2887EFFA93200FEE301881DDADB84" box="[325,422,907,931]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
; Figs. 58, 62, 66 and figs. 40, 51, 90–91, 94–102, 107–108, 111–114, 115–
|
||
<quantity id="4CF19B39FFA93200FD8E01A91EE2DBE4" box="[552,670,938,963]" metricMagnitude="0" metricUnit="m" metricValue="2.9748734" pageId="7" pageNumber="6" unit="in" value="117.121">117, 121 in</quantity>
|
||
KovařÍk, 2019;
|
||
<tableCitation id="C68B0367FFA93200FF1201C81D7FDBC4" box="[180,259,971,995]" captionStart="Table 1" captionStartId="4.[113,169,1006,1027]" captionText="Table 1. Morphometric data for types of Megacormus orizaba sp. n. and Megacormus seductus sp. n. Abbreviations: length (L), width (W), in carapace it corresponds to median width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FFD701E91DF1DF64" blockId="7.[113,772,906,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FFD701E91C90DC25" bold="true" box="[113,236,1002,1026]" pageId="7" pageNumber="6">Coloration</emphasis>
|
||
. Base color light brown, paler on chelicerae, pedipalp manus, legs and pectines, pedipalp chelae and telson with a reddish hue. Cheliceral manus finely and irregularly reticulate with dark brown, denser distally; fingers deeply infuscate. Pedipalps very densely and irregularly marbled with blackish brown, but sparser distally (much denser on femur, sparser on manus), with all carinae deeply infuscate and the base of most trichobothria much paler (yellowish); fingers blackish, with distal third much paler progressively (becoming yellowish at the tip). Carapace symmetrically and very densely marbled with dark to blackish brown, anterior and lateral margins largely infuscate; eyes and ocular tubercles black. Tergites symmetrically and very densely marbled with dark to blackish brown, without any conspicuous pale spots that could suggest a stripped pattern. Pectines with shaft heavily and irregularly infuscate, but teeth essentially immaculate. Sternites colored and patterned similarly to tergites. Legs irregularly and very densely marbled with blackish brown, with few pale areas. Metasoma concolorous throughout (i.e., no segments conspicuously darker), irregularly and very densely marbled with blackish brown and with carinae deeply infuscate. Telson vesicle densely and irregularly marbled with dark to blackish brown, with four faint pale stripes corresponding to longitudinal furrows; aculeus orange, with distal third blackish. In general, the dark pattern is so dense that the entire scorpion looks dull black to the unaided eye, especially when still alive.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FFD705491E8CDFC4" blockId="7.[113,772,906,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FFD705491C96DF45" bold="true" box="[113,234,1866,1890]" pageId="7" pageNumber="6">Chelicerae</emphasis>
|
||
. Dentition typical for the genus, teeth short but sharp. Tegument smooth and glossy. Setation very dense ventrally, but essentially lacking dorsally, i.e., reduced to a few macrosetae distally in manus and also along movable finger. Fingers short, robust, evenly curved and subequal in length.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FC9602E91FD3DAA4" blockId="7.[816,1475,234,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FC9602E91FE3D925" bold="true" box="[816,927,234,258]" pageId="7" pageNumber="6">Pedipalps</emphasis>
|
||
. Size and shape standard for the genus. Trichobothrial pattern C, neobothriotaxic majorante: femur with all three trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FBC103281808D964" box="[1127,1140,299,323]" italics="true" pageId="7" pageNumber="6">d</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FB22032818ECD965" box="[1156,1168,299,322]" italics="true" pageId="7" pageNumber="6">e</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FB39032818DAD965" box="[1183,1190,299,322]" italics="true" pageId="7" pageNumber="6">i</emphasis>
|
||
) in positions standard for the genus (at basal-most part of segment); patella with 2 dorsal trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FBB00368185FD9A4" box="[1046,1059,363,387]" italics="true" pageId="7" pageNumber="6">d</emphasis>
|
||
), one internal trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FAC60368191BD9A5" box="[1376,1383,363,386]" italics="true" pageId="7" pageNumber="6">i</emphasis>
|
||
), 21–22 external trichobothria (7/7
|
||
<emphasis id="B97DEACEFFA93200FBF603881815D984" box="[1104,1129,395,419]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
|
||
, 2/2
|
||
<emphasis id="B97DEACEFFA93200FB07038818B9D984" box="[1185,1221,395,419]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
|
||
, 4/5
|
||
<emphasis id="B97DEACEFFA93200FB5D03881966D985" box="[1275,1306,395,418]" italics="true" pageId="7" pageNumber="6">em</emphasis>
|
||
, 4/4
|
||
<emphasis id="B97DEACEFFA93200FAF703881913D985" box="[1361,1391,395,418]" italics="true" pageId="7" pageNumber="6">est</emphasis>
|
||
, 4/4
|
||
<emphasis id="B97DEACEFFA93200FA00038819C5D985" box="[1446,1465,395,418]" italics="true" pageId="7" pageNumber="6">et</emphasis>
|
||
) and 6/7 ventral trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FBDF03A818F5D9E5" box="[1145,1161,427,450]" italics="true" pageId="7" pageNumber="6">V</emphasis>
|
||
); chela manus with 2 dorsal trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FC7703C81F8ED9C4" box="[977,1010,459,483]" italics="true" pageId="7" pageNumber="6">Db</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FBA303C8185CD9C5" box="[1029,1056,459,482]" italics="true" pageId="7" pageNumber="6">Dt</emphasis>
|
||
), 11 external trichobothria (4
|
||
<emphasis id="B97DEACEFFA93200FA2503C819DDD9C4" box="[1411,1441,459,483]" italics="true" pageId="7" pageNumber="6">Eb</emphasis>
|
||
, 1
|
||
<emphasis id="B97DEACEFFA93200FC9603E81F24DA24" box="[816,856,491,515]" italics="true" pageId="7" pageNumber="6">Esb</emphasis>
|
||
, 1
|
||
<emphasis id="B97DEACEFFA93200FCDA03E81FE2DA25" box="[892,926,491,514]" italics="true" pageId="7" pageNumber="6">Est</emphasis>
|
||
, 5
|
||
<emphasis id="B97DEACEFFA93200FC6403E81FA6DA25" box="[962,986,491,514]" italics="true" pageId="7" pageNumber="6">Et</emphasis>
|
||
, with
|
||
<emphasis id="B97DEACEFFA93200FBB903E81843DA27" box="[1055,1087,491,514]" italics="true" pageId="7" pageNumber="6">
|
||
Et
|
||
<subScript id="178D3499FFA93200FB9103F11843DA27" attach="left" box="[1079,1087,498,512]" fontSize="6" pageId="7" pageNumber="6">1</subScript>
|
||
</emphasis>
|
||
displaced to ventral surface), and 4 ventral trichobothria (with
|
||
<emphasis id="B97DEACEFFA93200FBFE0008180CDA07" box="[1112,1136,523,546]" italics="true" pageId="7" pageNumber="6">
|
||
V
|
||
<subScript id="178D3499FFA93200FBCE0011180CDA07" attach="left" box="[1128,1136,530,544]" fontSize="6" pageId="7" pageNumber="6">4</subScript>
|
||
</emphasis>
|
||
displaced to external surface); fixed finger with 4 dorsal trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FB4500281882DA64" box="[1251,1278,555,579]" italics="true" pageId="7" pageNumber="6">db</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FAAB0028194EDA64" box="[1293,1330,555,579]" italics="true" pageId="7" pageNumber="6">dsb</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FAE600281923DA64" box="[1344,1375,555,579]" italics="true" pageId="7" pageNumber="6">dst</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FAC8002819FFDA64" box="[1390,1411,555,579]" italics="true" pageId="7" pageNumber="6">dt</emphasis>
|
||
), two internal trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FB8200481845DA44" box="[1060,1081,587,611]" italics="true" pageId="7" pageNumber="6">ib</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FBE10048182ADA45" box="[1095,1110,587,610]" italics="true" pageId="7" pageNumber="6">it</emphasis>
|
||
) and 4 external trichobothria (
|
||
<emphasis id="B97DEACEFFA93200FA04004819C7DA44" box="[1442,1467,587,611]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FC9600681F28DAA4" box="[816,852,619,643]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FCC700681F03DAA5" box="[865,895,619,642]" italics="true" pageId="7" pageNumber="6">est</emphasis>
|
||
,
|
||
<emphasis id="B97DEACEFFA93200FC2A00681FE3DAA5" box="[908,927,619,642]" italics="true" pageId="7" pageNumber="6">et</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FC960088190BDB44" blockId="7.[816,1475,234,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FC9600881F0BDA85" box="[816,887,651,674]" italics="true" pageId="7" pageNumber="6">Femur</emphasis>
|
||
robust (length/width ratio = 2.34), almost bare, with all surfaces essentially flat to shallowly convex. Dorsointernal, dorsoexternal, ventrointernal and ventroexternal carinae complete (moderate, coarsely granulose). Tegument coriaceous, with abundant fine to medium-sized granules scattered all over, plus abundant medium-sized to coarse, glossy granules irregularly scattered on dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FC960168192ADCC4" blockId="7.[816,1475,234,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FC9601681F01DBA4" box="[816,893,875,899]" italics="true" pageId="7" pageNumber="6">Patella</emphasis>
|
||
robust (length/width ratio = 2.29), almost bare, with dorsal and ventral surfaces essentially flat, external and internal surfaces markedly convex. Dorsointernal, dorsoexternal, external median, ventrointernal and ventroexternal carinae complete (strong to very strong, coarsely granulose to crenulate). Tegument finely, densely and evenly granulose on internal surface, coriaceous on dorsal, external and ventral surfaces, with abundant medium-sized to coarse, glossy granules evenly scattered all over dorsally and abundant fine and rough granules irregularly scattered all over externally and ventrally. Inner dorsal spur large, sharp and straight; inner ventral spur vestigial, reduced to a conical denticle.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA93200FC9606E81FB8DE44" blockId="7.[816,1475,234,2019]" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FC9606E81F0CDD24" box="[816,880,1259,1283]" italics="true" pageId="7" pageNumber="6">Chela</emphasis>
|
||
large and robust, standard for the genus (length/width ratio = 2.87). Manus incrassate-oval in dorsal view, medially wider and narrow-oval in lateral view, sparsely setose and prismatic in cross-section; dorsal internal, dorsal marginal, dorsal secondary, digital, external secondary, ventroexternal and ventrointernal carinae complete (strong, coarsely granulose to crenulate); tegument coriaceous, with a clearly defined reticulate pattern of small, glossy granules scattered all over. Fingers somewhat longer than standard for the genus, shallowly curved and sparsely setose, dentition standard for the genus, basal lobe/notch combination moderate (long and not too deep).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA9320EFC9604691E80D904" blockId="7.[816,1475,234,2019]" lastBlockId="9.[118,777,235,2019]" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
|
||
<emphasis id="B97DEACEFFA93200FC9604691FE2DEA5" bold="true" box="[816,926,1642,1666]" pageId="7" pageNumber="6">Carapace</emphasis>
|
||
. Almost as long as wide (length/width ratio = 1.03), paraboloid in dorsal view. Anterior margin shallowly bilobed, with 2–3 pairs of white macrosetae plus several minor setae; median notch very shallowly U-shaped. Tegument finely and densely granulose, with abundant medium-sized to coarse granules scattered all over. All carinae indistinct or absent, except for the superciliaries, central medians and posterior medians (weakly, finely granulose and not fused). Furrows: anterior median, central median, posterior median and posterior marginal narrow and moderately deep, fused altogether; posterior laterals narrow and moderately deep; other furrows either absent or undefined. Median ocular tubercle moderately raised, with eyes large and separated by less than one ocular diameter; two pairs of large lateral eyes.
|
||
</paragraph>
|
||
<caption id="DF766654FFA6320FFFD205AB1F47DFDF" ID-DOI="http://doi.org/10.5281/zenodo.8222212" ID-Zenodo-Dep="8222212" httpUri="https://zenodo.org/record/8222212/files/figure.png" pageId="8" pageNumber="7" startId="8.[116,193,1960,1982]" targetBox="[132,1474,239,1928]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BB636DCFFA6320FFFD205AB1F47DFDF" blockId="8.[116,1475,1960,2040]" pageId="8" pageNumber="7">
|
||
<emphasis id="B97DEACEFFA6320FFFD205AB1C8BDF9A" bold="true" box="[116,247,1960,1982]" pageId="8" pageNumber="7">Figures 7–16</emphasis>
|
||
:
|
||
<taxonomicName id="4C094D5FFFA6320FFEA305AA1DA7DF9A" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[261,475,1960,1982]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA6320FFEA305AA1DA7DF9A" box="[261,475,1960,1982]" italics="true" pageId="8" pageNumber="7">Megacormus seductus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA6320FFE4705AB1E64DF9A" box="[481,536,1960,1981]" pageId="8" pageNumber="7" rank="species">
|
||
<emphasis id="B97DEACEFFA6320FFE4705AB1D84DF9A" bold="true" box="[481,504,1960,1981]" pageId="8" pageNumber="7">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA6320FFDA305AB1E6EDF9A" bold="true" box="[517,530,1960,1981]" pageId="8" pageNumber="7">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype, pedipalp chela dorsal (7), external (8) and ventral (9) views, patella dorsal (10), external (11) and ventral (12) views, femur and trochanter dorsal (13) and ventral (14) views, movable (15) and fixed (16) fingers dentition. Movable finger (15) is under UV light, other figures are under white light.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636DCFFA7320EFFD003291C99D9A4" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD003291CA5D965" bold="true" box="[118,217,298,322]" pageId="9" pageNumber="8">Sternum</emphasis>
|
||
. Standard for the genus:
|
||
<typeStatus id="54B2887EFFA7320EFE5503281E5DD964" box="[499,545,299,323]" pageId="9" pageNumber="8">type</typeStatus>
|
||
2, large, longer than wide and hexagonal in shape. Tegument densely and finely granulose.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD003891EC4DA04" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD003891D1AD984" bold="true" box="[118,358,394,419]" pageId="9" pageNumber="8">Genital operculum</emphasis>
|
||
. Medium-sized and prominent, remarkably rhomboidal in shape. Halves neither separated nor fused and almost triangular in shape; tegument smooth and glossy. Genital papillae medium-sized, well-protruding, with tips blunt, round. Pre-pectinal plate apparently absent.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD000291E0BDB24" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD000291CAFDA65" bold="true" box="[118,211,554,578]" pageId="9" pageNumber="8">Pectines</emphasis>
|
||
. Size and shape standard for the genus: not reaching coxa-trochanter joint of leg IV, subrectangular and moderately setose, anterior area with 3/2 lamellae, median area with 3/3. Tooth count 6/6, teeth swollen, straight and basally not separated; fulcra entirely absent. Basal plate wider than long; anterior margin essentially straight, posterior margin widely convex; tegument smooth, glossy to coriaceous.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD001081EC6DCC4" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD001081C92DB04" bold="true" box="[118,238,779,803]" pageId="9" pageNumber="8">Mesosoma</emphasis>
|
||
. Tergites almost bare, anterior margin shallowly sinuose, posterior margin straight to very shallowly produced in the median part; median carina very weak and finely subcrenulate to crenulate on I–VI, submedian and lateral carinae strong and finely serratocrenulate on VII; tegument coriaceous, with abundant small to medium-sized, glossy granules scattered all over (denser posterolaterally). Sternites sparsely setose (largely along posterior and lateral margins), posterior margin shallowly convex to widely bilobed; III– VI acarinate, VII with only ventral median carinae perfectly defined, finely granulose to subcrenulate; tegument on III– VI glossy and minutely punctate, on VII coriaceous, with abundant medium-sized, glossy granules scattered all over; spiracles small and elongate-oval; V with smooth patch large, paraboloid in shape, translucent, glossy and not bulky.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD006E81EB7DE84" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD006E81C96DD24" bold="true" box="[118,234,1259,1283]" pageId="9" pageNumber="8">Metasoma</emphasis>
|
||
. Size and shape standard for the genus (length/ width ratio of segments I–V = 0.65, 0.81, 0.95, 1.26 and 2.37), moderately narrower distally and moderately setose; segment I–III wider than long, IV–V longer than wide; all segments wider than deep. Segments I–IV with seven complete to almost complete carinae, V with five: all strong, irregularly and coarsely dentate to serrate, with distalmost denticles not conspicuously enlarged; lateral inframedians entirely absent or undefined from surrounding granulation on all segments; ventral submedians entirely absent; ventral median carina perfectly defined all along I–V; anal arc weakly granulose; laterodistal lobes of V obsolete, blunt-triangular. Intercarinal tegument coriaceous, with abundant minute and fine granules scattered all over. Dorsal furrow complete, moderately narrow and shallow.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD004A81EEEDF44" blockId="9.[118,777,235,2019]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD004A81CC3DEE4" bold="true" box="[118,191,1707,1731]" pageId="9" pageNumber="8">Telson</emphasis>
|
||
. Sparsely setose. Vesicle oval and somewhat depressed; tegument coriaceous, with abundant minute and fine granules scattered all over and four very shallow longitudinal furrows; distal end of vesicle well-defined by a shallow annular ring and weak laterodistal shoulders. Aculeus moderately long, sharp and shallowly curved, with tegument glossy.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFFD005681903D904" blockId="9.[118,777,235,2019]" lastBlockId="9.[821,1479,235,675]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFFD005681CD7DFA4" bold="true" box="[118,171,1899,1923]" pageId="9" pageNumber="8">Legs</emphasis>
|
||
. Somewhat longer and slenderer than standard for the genus, with all carinae weakly granulose to subcrenulate; tegument smooth coriaceous, with dorsal surface of femur sparsely granulose. Prolateral pedal spurs standard-sized to large, thick but sharp. Claws standard-sized (at least one-third the length of its respective telotarsus) and well-curved.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFC930329184DDA84" blockId="9.[821,1479,235,675]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97DEACEFFA7320EFC930329185CD965" bold="true" box="[821,1056,298,322]" pageId="9" pageNumber="8">Hemispermatophore</emphasis>
|
||
. (
|
||
<figureCitation id="13322A59FFA7320EFB9C032818E4D964" box="[1082,1176,299,323]" captionStart="Figures 62–65" captionStartId="18.[109,186,1132,1154]" captionTargetBox="[138,1454,242,1112]" captionTargetId="figure-463@18.[112,1473,221,1121]" captionTargetPageId="18" captionText="Figures 62–65. Hemispermatophores of Megacormus spp. Figure 62. M. orizaba sp. n. 19MD (holotype, No. 1621). Figure 63. M. granosus (No. 2462). Figures 64–65. M. seductus sp. n. 22MA (holotype and paratopotype, Nos. 2460, 2461).Convex views of left hemispermatophores (62, 63) and right hemispermatophores shown in mirror image (64, 65). Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8222224" httpUri="https://zenodo.org/record/8222224/files/figure.png" pageId="9" pageNumber="8">Figs. 62</figureCitation>
|
||
,
|
||
<figureCitation id="13322A59FFA7320EFB0E032818BAD964" box="[1192,1222,299,323]" captionStart="Figures 66–77" captionStartId="19.[113,187,1941,1963]" captionTargetBox="[118,1474,212,1929]" captionTargetId="figure-160@19.[113,1474,210,1931]" captionTargetPageId="19" captionText="Figures 66–77. Hemispermatophore capsules of Megacormus. 66. M. orizaba sp. n. (holotype, No.1621). 67–68. M. gertschi (Nos.1619, 1620).69. M. franckei (paratype, No. 1618).70–71. M. xichu (Nos.1617, 1626).72–73. M. seductus sp. n. (holotype and paratopotype,Nos.2460–2461).74. M. segmentatus (No.1622).75. M. granosus (No.2462).76. M. grubbsi (No.1623).77. M. grubbsi (No.2463).Convex views ofleftcapsules (66, 68–71, 73–75 and 77), and right capsules shown in mirror image (67, 72 and 76).Abbreviations: dpd, distal dentate process; dpp, proximal dentate process; cls, crown-like structure; trd, distal transverse ridge; trp, proximal transverse ridge; tmsd, terminal membrane of the sperm duct. Scale bars: 400 μm." figureDoi="http://doi.org/10.5281/zenodo.8222226" httpUri="https://zenodo.org/record/8222226/files/figure.png" pageId="9" pageNumber="8">66</figureCitation>
|
||
). Lamelliform. Overall proportions of distal lamina, capsule, truncal flexure, trunk and pedicel standard for the genus. Distal lamina relatively narrow, nearly unform in width, with straight anterior margin, lacking a basal constriction. Capsule with moderately broad proximal transverse ridge; proximal dentate process elongate or spatulate, slightly broadened, armed with 7–9 irregular, short, blunt or sharp denticles. Distal dentate process moderately broad, laminate, armed 6–7 irregular, small to large, sharp denticles. Basal carina with crown-like structure bearing 6–8 tines. Terminal membrane of sperm duct with numerous fine spicules.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFC9300C8181CDB48" blockId="9.[821,1479,715,1264]" pageId="9" pageNumber="8">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="4C094D5FFFA7320EFA9D00C81FFADB2E" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFA9D00C81FFADB2E" italics="true" pageId="9" pageNumber="8">Megacormus orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFC2800F21FB7DB2E" box="[910,971,753,777]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFC2800F21FDBDB2E" bold="true" box="[910,935,753,777]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFC1300F21FB8DB2E" bold="true" box="[949,964,753,777]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from all other species of the genus (see the key below). the validity of
|
||
<taxonomicName id="4C094D5FFFA7320EFB8D011B18DCDB08" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[1067,1184,791,815]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFB8D011B18DCDB08" box="[1067,1184,791,815]" italics="true" pageId="9" pageNumber="8">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFB0F011B189ADB08" box="[1193,1254,792,816]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFB0F011B18BEDB17" bold="true" box="[1193,1218,792,816]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFB76011B18A3DB17" bold="true" box="[1232,1247,792,816]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is confirmed also by DNA analysis implemented by Charles University in
|
||
<collectingRegion id="49CDF83EFFA7320EFADA013B19BADB77" box="[1404,1478,824,848]" country="Czech Republic" name="Praha" pageId="9" pageNumber="8">Prague</collectingRegion>
|
||
(manuscript in preparation).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFCC3017B1956DCC8" blockId="9.[821,1479,715,1264]" pageId="9" pageNumber="8">
|
||
<taxonomicName id="4C094D5FFFA7320EFCC3017B1F9FDBA8" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[869,995,887,911]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFCC3017B1F9FDBA8" box="[869,995,887,911]" italics="true" pageId="9" pageNumber="8">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFC55017B1845DBA8" box="[1011,1081,888,912]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFC55017B1870DBB7" bold="true" box="[1011,1036,888,912]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFB85017B184EDBB7" bold="true" box="[1059,1074,888,912]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
inhabits the same locality as
|
||
<taxonomicName id="4C094D5FFFA7320EFA0C017B1FEBDB88" authorityName=": KovarIk" authorityYear="2019" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFA0C017B1FEBDB88" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
, with which it has been confused in the past (see KovařÍk, 2019), but it can be easily distinguished by its larger size (
|
||
<quantity id="4CF19B39FFA7320EFC1701DB1859DBC8" box="[945,1061,984,1008]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="9" pageNumber="8" unit="mm" value="32.0" valueMax="34.0" valueMin="30.0">30–34 mm</quantity>
|
||
in males,
|
||
<quantity id="4CF19B39FFA7320EFB3501D4197ADBC8" box="[1171,1286,983,1007]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="8" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
|
||
in females versus
|
||
<quantity id="4CF19B39FFA7320EFC9301FB1FD6DC28" box="[821,938,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="9" pageNumber="8" unit="mm" value="16.0" valueMax="18.0" valueMin="14.0">14–18 mm</quantity>
|
||
in males,
|
||
<quantity id="4CF19B39FFA7320EFBB901FB18E8DC28" box="[1055,1172,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="9" pageNumber="8" unit="mm" value="20.5" valueMax="23.0" valueMin="18.0">18–23 mm</quantity>
|
||
in females in
|
||
<taxonomicName id="4C094D5FFFA7320EFA9001FB19C2DC28" authorityName=": KovarIk" authorityYear="2019" box="[1334,1470,1016,1039]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFA9001FB19C2DC28" box="[1334,1470,1016,1039]" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
) and mainly by the presence of a pedipalp lobe/notch in adult males of
|
||
<taxonomicName id="4C094D5FFFA7320EFC3C063B186CDC68" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[922,1040,1079,1103]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFC3C063B186CDC68" box="[922,1040,1079,1103]" italics="true" pageId="9" pageNumber="8">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFBBE063B182ADC68" box="[1048,1110,1080,1104]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFBBE063B184DDC77" bold="true" box="[1048,1073,1080,1104]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFBE6063B1833DC77" bold="true" box="[1088,1103,1080,1104]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(Fig. 58) which is absent in adult males of
|
||
<taxonomicName id="4C094D5FFFA7320EFC3B065B1859DC48" authorityName=": KovarIk" authorityYear="2019" box="[925,1061,1112,1135]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFC3B065B1859DC48" box="[925,1061,1112,1135]" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 46). The hemispermatophore of
|
||
<taxonomicName id="4C094D5FFFA7320EFC93067B1FC7DCA8" authorityName=": KovarIk" authorityYear="2019" box="[821,955,1144,1167]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFC93067B1FC7DCA8" box="[821,955,1144,1167]" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
has a broad distal lamina with basal constriction (
|
||
<figureCitation id="13322A59FFA7320EFC9B06941FF1DC88" box="[829,909,1175,1199]" captionStart="Figures 62–65" captionStartId="18.[109,186,1132,1154]" captionTargetBox="[138,1454,242,1112]" captionTargetId="figure-463@18.[112,1473,221,1121]" captionTargetPageId="18" captionText="Figures 62–65. Hemispermatophores of Megacormus spp. Figure 62. M. orizaba sp. n. 19MD (holotype, No. 1621). Figure 63. M. granosus (No. 2462). Figures 64–65. M. seductus sp. n. 22MA (holotype and paratopotype, Nos. 2460, 2461).Convex views of left hemispermatophores (62, 63) and right hemispermatophores shown in mirror image (64, 65). Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8222224" httpUri="https://zenodo.org/record/8222224/files/figure.png" pageId="9" pageNumber="8">Fig. 63</figureCitation>
|
||
,
|
||
<figureCitation id="13322A59FFA7320EFC3C06941FCBDC88" box="[922,951,1175,1199]" captionStart="Figures 66–77" captionStartId="19.[113,187,1941,1963]" captionTargetBox="[118,1474,212,1929]" captionTargetId="figure-160@19.[113,1474,210,1931]" captionTargetPageId="19" captionText="Figures 66–77. Hemispermatophore capsules of Megacormus. 66. M. orizaba sp. n. (holotype, No.1621). 67–68. M. gertschi (Nos.1619, 1620).69. M. franckei (paratype, No. 1618).70–71. M. xichu (Nos.1617, 1626).72–73. M. seductus sp. n. (holotype and paratopotype,Nos.2460–2461).74. M. segmentatus (No.1622).75. M. granosus (No.2462).76. M. grubbsi (No.1623).77. M. grubbsi (No.2463).Convex views ofleftcapsules (66, 68–71, 73–75 and 77), and right capsules shown in mirror image (67, 72 and 76).Abbreviations: dpd, distal dentate process; dpp, proximal dentate process; cls, crown-like structure; trd, distal transverse ridge; trp, proximal transverse ridge; tmsd, terminal membrane of the sperm duct. Scale bars: 400 μm." figureDoi="http://doi.org/10.5281/zenodo.8222226" httpUri="https://zenodo.org/record/8222226/files/figure.png" pageId="9" pageNumber="8">75</figureCitation>
|
||
), similar to that of
|
||
<taxonomicName id="4C094D5FFFA7320EFB2C069B1977DC88" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[1162,1291,1175,1199]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFB2C069B1977DC88" box="[1162,1291,1175,1199]" italics="true" pageId="9" pageNumber="8">M. seductus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFAB5069B192DDC88" box="[1299,1361,1176,1200]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFAB5069B1950DC97" bold="true" box="[1299,1324,1176,1200]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFA9D069B1936DC97" bold="true" box="[1339,1354,1176,1200]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13322A59FFA7320EFAC706941F2EDCE8" captionStart="Figures 62–65" captionStartId="18.[109,186,1132,1154]" captionTargetBox="[138,1454,242,1112]" captionTargetId="figure-463@18.[112,1473,221,1121]" captionTargetPageId="18" captionText="Figures 62–65. Hemispermatophores of Megacormus spp. Figure 62. M. orizaba sp. n. 19MD (holotype, No. 1621). Figure 63. M. granosus (No. 2462). Figures 64–65. M. seductus sp. n. 22MA (holotype and paratopotype, Nos. 2460, 2461).Convex views of left hemispermatophores (62, 63) and right hemispermatophores shown in mirror image (64, 65). Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8222224" httpUri="https://zenodo.org/record/8222224/files/figure.png" pageId="9" pageNumber="8">Figs. 64– 65</figureCitation>
|
||
), contrasting with the narrow distal lamina without basal constriction in
|
||
<taxonomicName id="4C094D5FFFA7320EFC7306DB1835DCC8" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[981,1097,1239,1263]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFC7306DB1F97DCC8" box="[981,1003,1240,1263]" italics="true" pageId="9" pageNumber="8">M</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFC5E06D41835DCC8" box="[1016,1097,1239,1263]" italics="true" pageId="9" pageNumber="8">orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFBF706DB18F1DCC8" box="[1105,1165,1240,1264]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFBF706DB1816DCD7" bold="true" box="[1105,1130,1240,1264]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFBD106DB18FADCD7" bold="true" box="[1143,1158,1240,1264]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13322A59FFA7320EFB3D06D41888DCC8" box="[1179,1268,1239,1263]" captionStart="Figures 62–65" captionStartId="18.[109,186,1132,1154]" captionTargetBox="[138,1454,242,1112]" captionTargetId="figure-463@18.[112,1473,221,1121]" captionTargetPageId="18" captionText="Figures 62–65. Hemispermatophores of Megacormus spp. Figure 62. M. orizaba sp. n. 19MD (holotype, No. 1621). Figure 63. M. granosus (No. 2462). Figures 64–65. M. seductus sp. n. 22MA (holotype and paratopotype, Nos. 2460, 2461).Convex views of left hemispermatophores (62, 63) and right hemispermatophores shown in mirror image (64, 65). Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8222224" httpUri="https://zenodo.org/record/8222224/files/figure.png" pageId="9" pageNumber="8">Figs. 62</figureCitation>
|
||
,
|
||
<figureCitation id="13322A59FFA7320EFB5906D41961DCC8" box="[1279,1309,1239,1263]" captionStart="Figures 66–77" captionStartId="19.[113,187,1941,1963]" captionTargetBox="[118,1474,212,1929]" captionTargetId="figure-160@19.[113,1474,210,1931]" captionTargetPageId="19" captionText="Figures 66–77. Hemispermatophore capsules of Megacormus. 66. M. orizaba sp. n. (holotype, No.1621). 67–68. M. gertschi (Nos.1619, 1620).69. M. franckei (paratype, No. 1618).70–71. M. xichu (Nos.1617, 1626).72–73. M. seductus sp. n. (holotype and paratopotype,Nos.2460–2461).74. M. segmentatus (No.1622).75. M. granosus (No.2462).76. M. grubbsi (No.1623).77. M. grubbsi (No.2463).Convex views ofleftcapsules (66, 68–71, 73–75 and 77), and right capsules shown in mirror image (67, 72 and 76).Abbreviations: dpd, distal dentate process; dpp, proximal dentate process; cls, crown-like structure; trd, distal transverse ridge; trp, proximal transverse ridge; tmsd, terminal membrane of the sperm duct. Scale bars: 400 μm." figureDoi="http://doi.org/10.5281/zenodo.8222226" httpUri="https://zenodo.org/record/8222226/files/figure.png" pageId="9" pageNumber="8">66</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136557FFA7320DFC93071B1D84D9E4" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="distribution">
|
||
<paragraph id="8BB636DCFFA7320EFC93071B194DDD88" blockId="9.[821,1479,1303,1456]" pageId="9" pageNumber="8">
|
||
DISTRIBUTION (
|
||
<figureCitation id="13322A59FFA7320EFC4407141840DD17" box="[994,1084,1303,1328]" captionStart="Figure 82" captionStartId="22.[113,181,1223,1245]" captionTargetBox="[112,1473,247,1206]" captionTargetId="figure-319@22.[112,1473,247,1206]" captionTargetPageId="22" captionText="Figure 82. Distribution of Megacormus specimens examined for this review (see Comparative material examined in Methods, Material & Abbreviations section)." figureDoi="http://doi.org/10.5281/zenodo.8222230" httpUri="https://zenodo.org/record/8222230/files/figure.png" pageId="9" pageNumber="8">Fig. 82</figureCitation>
|
||
).
|
||
<taxonomicName id="4C094D5FFFA7320EFBFC071B18A5DD08" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[1114,1241,1303,1327]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFBFC071B18A5DD08" box="[1114,1241,1303,1327]" italics="true" pageId="9" pageNumber="8">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFB4D071B194EDD08" box="[1259,1330,1304,1328]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFB4D071B1978DD17" bold="true" box="[1259,1284,1304,1328]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFABA071B1957DD17" bold="true" box="[1308,1323,1304,1328]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is currently known only from the type-locality. Nevertheless, a potential additional locality record may have been given misidentified as
|
||
<taxonomicName id="4C094D5FFFA7320EFCF5077B1FB2DDA8" box="[851,974,1399,1423]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gertschi">
|
||
<emphasis id="B97DEACEFFA7320EFCF5077B1FB2DDA8" box="[851,974,1399,1423]" italics="true" pageId="9" pageNumber="8">M. gertschi</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF984B2DFFA7320EFC5E07741928DDB7" author="GONZALEZ-SANTILLAN E." box="[1016,1364,1399,1424]" pageId="9" pageNumber="8" pagination="221 - 237" refId="ref12930" refString="GONZALEZ-SANTILLAN E., J. M. GONZALEZ-RUIZ & L. A. ESCOBEDO-MORALES. 2017. A new species of Megacormus (Scorpiones, Euscorpiidae) from an oak-pine forest in Guanajuato, Mexico with an identification key tothe species in the genus. Zootaxa, 4299 (2): 221 - 237." type="journal article" year="2017">
|
||
González-Santillán
|
||
<emphasis id="B97DEACEFFA7320EFB6A077B197FDDA8" box="[1228,1283,1399,1423]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
, also from
|
||
<collectingRegion id="49CDF83EFFA7320EFC93079B1FB3DD97" box="[821,975,1432,1456]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="9" pageNumber="8">Veracruz State</collectingRegion>
|
||
(see below, in Remarks section).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320EFC9307DB1FB9DF28" blockId="9.[821,1479,1495,2031]" pageId="9" pageNumber="8">
|
||
ECOLOGICAL NOTES. The type-specimens were all collected under rocks of various sizes (usually small to medium), in somewhat humid places inside pine forest (fig.
|
||
<quantity id="4CF19B39FFA7320EFA84041B1918DE08" box="[1314,1380,1559,1584]" metricMagnitude="0" metricUnit="m" metricValue="3.1242" pageId="9" pageNumber="8" unit="in" value="123.0">123 in</quantity>
|
||
KovařÍk, 2019: 27). The scorpions were usually found on the ground, inside short scrapes excavated by themselves, but as an unusual finding, not less than 10 individuals (mostly juveniles and subadults) were found hanging to the underside of the rocks, frequently at the edges. After the rocks were turned, all scorpions played dead and remain motionless (thanatosis or catalepsy).
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA7320DFCC3051B1E40D964" blockId="9.[821,1479,1495,2031]" lastBlockId="10.[113,771,235,451]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
|
||
At the single known locality, this species lives syntopically with
|
||
<taxonomicName id="4C094D5FFFA7320EFCCB053B1F89DF68" authorityName=": KovarIk" authorityYear="2019" box="[877,1013,1848,1871]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFCCB053B1F89DF68" box="[877,1013,1848,1871]" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
, but we observed a very interesting trend. On one hand,
|
||
<taxonomicName id="4C094D5FFFA7320EFC6B055B1828DF48" authorityName=": KovarIk" authorityYear="2019" box="[973,1108,1880,1903]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA7320EFC6B055B1828DF48" box="[973,1108,1880,1903]" italics="true" pageId="9" pageNumber="8">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
is widespread throughout the area and occurs in similar numbers both in oak and pine forests, but most commonly in the less densely vegetated areas of the oak forest. On the other hand,
|
||
<taxonomicName id="4C094D5FFFA7320EFB2B05BB1979DFE8" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[1165,1285,1975,1999]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA7320EFB2B05BB1979DFE8" box="[1165,1285,1975,1999]" italics="true" pageId="9" pageNumber="8">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA7320EFAB605BB192CDFE8" box="[1296,1360,1976,2000]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B97DEACEFFA7320EFAB605BB1955DFF7" bold="true" box="[1296,1321,1976,2000]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA7320EFA9C05BB1935DFF7" bold="true" box="[1338,1353,1976,2000]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is entirely restricted to the pine forest and becomes more and more abundant towards its densest core, while at the edge of the pure pine forest (without any oak trees), both species are found together, even under the same rocks.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA4320DFF0703481D84D9E4" blockId="10.[113,771,235,451]" pageId="10" pageNumber="9">
|
||
This is the first time that two species of
|
||
<taxonomicName id="4C094D5FFFA4320DFDDE03481F7FD945" authorityName="Karsch" authorityYear="1881" box="[632,771,331,354]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEACEFFA4320DFDDE03481F7FD945" box="[632,771,331,354]" italics="true" pageId="10" pageNumber="9">Megacormus</emphasis>
|
||
</taxonomicName>
|
||
are confirmed to occur together in a same locality. No other scorpion species were found there during the two field trips conducted by the authors to this site.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136557FFA4320DFFD703E81D54DEE4" pageId="10" pageNumber="9" type="discussion">
|
||
<paragraph id="8BB636DCFFA4320DFFD703E81F78DAC4" blockId="10.[113,772,491,1731]" pageId="10" pageNumber="9">
|
||
REMARKS. This species was formerly misidentified as
|
||
<taxonomicName id="4C094D5FFFA4320DFD4003E81CADDA05" authorityName=": KovarIk" authorityYear="2019" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA4320DFD4003E81CADDA05" italics="true" pageId="10" pageNumber="9">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
by KovařÍk (2019). During the previous field trips (2018–2019) that yielded the specimens used by him for that paper, both species were actually found, but unfortunately, all small specimens (including the authentic
|
||
<taxonomicName id="4C094D5FFFA4320DFD9000681EC1DAA5" authorityName=": KovarIk" authorityYear="2019" box="[566,701,619,642]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA4320DFD9000681EC1DAA5" box="[566,701,619,642]" italics="true" pageId="10" pageNumber="9">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
) were mistaken for early immatures and not collected. Only during the last field trip (2022), was it noted that there were in fact two distinct species living together and the mistake was discovered.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA4320DFF0700E81D87DBC4" blockId="10.[113,772,491,1731]" pageId="10" pageNumber="9">
|
||
Conversely, it is odd that González-Santillán & ÁlvarezPadilla (2015) redescribed
|
||
<taxonomicName id="4C094D5FFFA4320DFE2901081E68DB05" authorityName=": KovarIk" authorityYear="2019" box="[399,532,779,802]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="granosus">
|
||
<emphasis id="B97DEACEFFA4320DFE2901081E68DB05" box="[399,532,779,802]" italics="true" pageId="10" pageNumber="9">M. granosus</emphasis>
|
||
</taxonomicName>
|
||
from exactly the same locality, but they did not record the existence of the second, much larger species there. González-Santillán & ÁlvarezPadilla (2015: 75, 77) explicitly stated that their specimens were collected exclusively in the oak forest, where according to our data
|
||
<taxonomicName id="4C094D5FFFA4320DFF5C01A81D0FDBE4" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[250,371,939,963]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA4320DFF5C01A81D0FDBE4" box="[250,371,939,963]" italics="true" pageId="10" pageNumber="9">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA4320DFE2601A81DBDDBE5" box="[384,449,939,963]" pageId="10" pageNumber="9" rank="species">
|
||
<emphasis id="B97DEACEFFA4320DFE2601A81DE5DBE4" bold="true" box="[384,409,939,963]" pageId="10" pageNumber="9">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA4320DFE0A01A81DC7DBE4" bold="true" box="[428,443,939,963]" pageId="10" pageNumber="9">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
seems not to occur, thus, it apparently went undetected by them.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA4320DFF0701E81D3CDD44" blockId="10.[113,772,491,1731]" pageId="10" pageNumber="9">
|
||
There is one locality record published as
|
||
<taxonomicName id="4C094D5FFFA4320DFD2501E81F7FDC24" box="[643,771,1003,1027]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gertschi">
|
||
<emphasis id="B97DEACEFFA4320DFD2501E81F7FDC24" box="[643,771,1003,1027]" italics="true" pageId="10" pageNumber="9">M. gertschi</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF984B2DFFA4320DFF3E06081D36DC04" author="SOLEGLAD" box="[152,330,1035,1059]" pageId="10" pageNumber="9" pagination="251 - 303" refId="ref14062" refString="SOLEGLAD, M. E. 1976. A revision of the scorpion subfamily Megacorminae (Scorpionida, Chactidae). Wassmann Journal of Biology, 34 (2): 251 - 303." type="journal article" year="1976">Soleglad (1976)</bibRefCitation>
|
||
and González- Santillán
|
||
<emphasis id="B97DEACEFFA4320DFDC806081ED5DC04" box="[622,681,1035,1059]" italics="true" pageId="10" pageNumber="9">et al.</emphasis>
|
||
(2017), which could well actually belong to its closest-relative
|
||
<taxonomicName id="4C094D5FFFA4320DFD4006281CBEDC44" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA4320DFD4006281CBEDC44" italics="true" pageId="10" pageNumber="9">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA4320DFF6A06481D77DC45" box="[204,267,1099,1123]" pageId="10" pageNumber="9" rank="species">
|
||
<emphasis id="B97DEACEFFA4320DFF6A06481C99DC44" bold="true" box="[204,229,1099,1123]" pageId="10" pageNumber="9">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA4320DFF5306481D78DC44" bold="true" box="[245,260,1099,1123]" pageId="10" pageNumber="9">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: 11.2 km southeast of Las Vigas de RamÍrez, in
|
||
<collectingRegion id="49CDF83EFFA4320DFF2906681D4DDCA4" box="[143,305,1131,1155]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="9">Veracruz State</collectingRegion>
|
||
. This site is located roughly
|
||
<quantity id="4CF19B39FFA4320DFDD206691EC6DCA4" box="[628,698,1130,1155]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="9" unit="km" value="55.0">55 km</quantity>
|
||
to the north of Atotonilco, in the same mountain system (eastern tip of the Trans-Mexican Neovolcanic Belt) and at similar altitude (
|
||
<quantity id="4CF19B39FFA4320DFF7506C81D4FDCC4" box="[211,307,1227,1251]" metricMagnitude="0" metricUnit="m" metricValue="2.42" pageId="10" pageNumber="9" unit="m" value="2.42">2,420 m</quantity>
|
||
); even the reference to a “subadult male” (
|
||
<bibRefCitation id="EF984B2DFFA4320DFFDF06E81D27DD24" author="SOLEGLAD" box="[121,347,1259,1283]" pageId="10" pageNumber="9" pagination="251 - 303" refId="ref14062" refString="SOLEGLAD, M. E. 1976. A revision of the scorpion subfamily Megacorminae (Scorpionida, Chactidae). Wassmann Journal of Biology, 34 (2): 251 - 303." type="journal article" year="1976">Soleglad, 1976: 284</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF984B2DFFA4320DFECC06E81E87DD24" author="GONZALEZ-SANTILLAN E." box="[362,763,1259,1283]" pageId="10" pageNumber="9" pagination="221 - 237" refId="ref12930" refString="GONZALEZ-SANTILLAN E., J. M. GONZALEZ-RUIZ & L. A. ESCOBEDO-MORALES. 2017. A new species of Megacormus (Scorpiones, Euscorpiidae) from an oak-pine forest in Guanajuato, Mexico with an identification key tothe species in the genus. Zootaxa, 4299 (2): 221 - 237." type="journal article" year="2017">
|
||
González-Santillán
|
||
<emphasis id="B97DEACEFFA4320DFDE706E81E06DD24" box="[577,634,1259,1283]" italics="true" pageId="10" pageNumber="9">et al.</emphasis>
|
||
, 2017: 237
|
||
</bibRefCitation>
|
||
) suggests one of the main diagnostic features of
|
||
<taxonomicName id="4C094D5FFFA4320DFDCE07081EA0DD04" authority="Teruel & Kovařík & Lowe & Šťáhlavský, 2023" authorityName="Teruel & Kovařík & Lowe & Šťáhlavský" authorityYear="2023" box="[616,732,1291,1315]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
|
||
<emphasis id="B97DEACEFFA4320DFDCE07081EA0DD04" box="[616,732,1291,1315]" italics="true" pageId="10" pageNumber="9">M. orizaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57B5FFA4320DFD4507081CFADD65" pageId="10" pageNumber="9" rank="species">
|
||
<emphasis id="B97DEACEFFA4320DFD4507081E80DD04" bold="true" box="[739,764,1291,1315]" pageId="10" pageNumber="9">sp</emphasis>
|
||
.
|
||
<emphasis id="B97DEACEFFA4320DFFD707281CFCDD64" bold="true" box="[113,128,1323,1347]" pageId="10" pageNumber="9">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, i.e., the weaker development of the pedipalp finger lobe/ notch combination.
|
||
</paragraph>
|
||
<paragraph id="8BB636DCFFA4320DFF0707681D54DEE4" blockId="10.[113,772,491,1731]" pageId="10" pageNumber="9">
|
||
Incidentally, there is some confusion in the previous literature about this character, i.e., it has been incorrectly assumed that a weak lobe/notch combination indicates a subadult male (see, e.g.,
|
||
<bibRefCitation id="EF984B2DFFA4320DFE2707C81E23DDC5" author="SOLEGLAD" box="[385,607,1482,1507]" pageId="10" pageNumber="9" pagination="251 - 303" refId="ref14062" refString="SOLEGLAD, M. E. 1976. A revision of the scorpion subfamily Megacorminae (Scorpionida, Chactidae). Wassmann Journal of Biology, 34 (2): 251 - 303." type="journal article" year="1976">Soleglad, 1976: 285</bibRefCitation>
|
||
). Nevertheless, by rearing dozens of specimens from birth to adulthood in captivity, we have confirmed that in all
|
||
<taxonomicName id="4C094D5FFFA4320DFDDE04081CC2DE64" pageId="10" pageNumber="9">
|
||
<emphasis id="B97DEACEFFA4320DFDDE04081F7FDE05" box="[632,771,1547,1570]" italics="true" pageId="10" pageNumber="9">Megacormus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
whose male has pedipalp fingers with lobe/notch combination developed, this character state appears only with the terminal ecdysis to adult, i.e., subadult males entirely lack such combination and its fingers resemble those of adult and juvenile females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |