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<document id="829E2513C5EBAB1A4F17CE982B440E80" ID="10.11646/zootaxa.4200.1.4" ID-DOI="10.11646/zootaxa.4200.1.4" ID-GBIF-Dataset="07cdb522-b119-4cec-904a-00cdedc9a5b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="177778" ID-ZooBank="BD517CC9-99BD-4254-B55E-9D0445AFA70F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480322655154" checkinUser="plazi" docAuthor="Uyeno, Daisuke" docDate="2016" docId="039987F6FF9DE621FF510E62FC3F15F0" docLanguage="en" docName="zootaxa.4200.1.4.pdf" docOrigin="Zootaxa 4200 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudanthessius imo Uyeno, 2016, n. sp." docType="treatment" docVersion="6" lastPageNumber="128" masterDocId="FFA0FF8EFF92E632FFC60F78FFE51118" masterDocTitle="Copepods (Cyclopoida) associated with top shells (Vestigastropoda: Trochoidea: Tegulidae) from coastal waters in southern Japan, with descriptions of three new species" masterLastPageNumber="130" masterPageNumber="109" pageNumber="124" updateTime="1698394882508" updateUser="plazi">
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<mods:title id="978DE9E4518A140DE77C219A06BD8394">Copepods (Cyclopoida) associated with top shells (Vestigastropoda: Trochoidea: Tegulidae) from coastal waters in southern Japan, with descriptions of three new species</mods:title>
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<mods:name id="93C5973D9A81AEF530085448760F3EDC" type="personal">
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<mods:namePart id="88B990400D797E68375B5AC32508E8E1">Uyeno, Daisuke</mods:namePart>
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<mods:title id="95812BD50E2FA1756FE745E5A92FFC8A">Zootaxa</mods:title>
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<mods:date id="DEAF2E482DD20C831D51C395D4E10291">2016</mods:date>
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<mods:number id="AD61D7CE947AA55F4A8C2C72884D5CE6">4200</mods:number>
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<mods:identifier id="4D37A8F6F3CA1F473135DD1B2C9A9D21" type="DOI">10.11646/zootaxa.4200.1.4</mods:identifier>
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<treatment id="039987F6FF9DE621FF510E62FC3F15F0" ID-DOI="http://doi.org/10.5281/zenodo.6091846" ID-GBIF-Taxon="124864233" ID-Zenodo-Dep="6091846" LSID="urn:lsid:plazi:treatment:039987F6FF9DE621FF510E62FC3F15F0" httpUri="http://treatment.plazi.org/id/039987F6FF9DE621FF510E62FC3F15F0" lastPageId="19" lastPageNumber="128" pageId="15" pageNumber="124">
<subSubSection id="C32A656BFF9DE63DFF510E62FEC2104E" pageId="15" pageNumber="124" type="nomenclature">
<paragraph id="8B8F36E0FF9DE63DFF510E62FE3B102C" blockId="15.[151,478,282,342]" box="[151,478,282,308]" pageId="15" pageNumber="124">
<heading id="D0C7818CFF9DE63DFF510E62FE3B102C" bold="true" box="[151,478,282,308]" fontSize="11" level="1" pageId="15" pageNumber="124" reason="1">
<emphasis id="B944EAF2FF9DE63DFF510E62FE3B102C" bold="true" box="[151,478,282,308]" pageId="15" pageNumber="124">
<taxonomicName id="4C304D63FF9DE63DFF510E62FE71102C" box="[151,404,282,308]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="imo" status="sp. nov.">
<emphasis id="B944EAF2FF9DE63DFF510E62FE71102C" bold="true" box="[151,404,282,308]" italics="true" pageId="15" pageNumber="124">Pseudanthessius imo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775789FF9DE63DFE5D0E62FE3B102C" box="[411,478,282,308]" pageId="15" pageNumber="124" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF510E45FEC2104E" blockId="15.[151,478,282,342]" box="[151,295,317,342]" pageId="15" pageNumber="124">
(
<figureCitation id="130B2A65FF9DE63DFF590E45FEFA104E" box="[159,287,317,342]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="16.[151,250,1898,1920]" captionStartId-1="17.[151,250,1896,1918]" captionStartId-2="18.[151,250,1669,1691]" captionTargetBox-0="[216,1379,201,1867]" captionTargetBox-1="[211,1378,196,1863]" captionTargetBox-2="[205,1371,204,1626]" captionTargetId-0="figure@16.[197,1381,195,1870]" captionTargetId-1="figure@17.[197,1381,193,1868]" captionTargetId-2="figure@18.[196,1382,193,1634]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." captionText-1="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." captionText-2="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri-0="https://zenodo.org/record/177788/files/figure.png" httpUri-1="https://zenodo.org/record/177789/files/figure.png" httpUri-2="https://zenodo.org/record/177790/files/figure.png" pageId="15" pageNumber="124">Figs 1012</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A656BFF9DE63DFF510E07FA781353" pageId="15" pageNumber="124" type="materials_examined">
<paragraph id="8B8F36E0FF9DE63DFF510E07FAAC131D" blockId="15.[151,1437,383,1263]" pageId="15" pageNumber="124">
<materialsCitation id="3B583CBDFF9DE63DFF510E07FE7210F8" ID-GBIF-Occurrence="1412767582" collectingDate="1929-04" collectionCode="NSMT" collectorName="D. Uyeno &amp; S. Yokoyama" country="Iceland" latitude="28.2" location="Saneku" longLatPrecision="1233" longitude="129.2" municipality="Oshima Strait" pageId="15" pageNumber="124" specimenCount="2" typeStatus="holotype">
<emphasis id="B944EAF2FF9DE63DFF510E07FEA01080" bold="true" box="[151,325,383,408]" pageId="15" pageNumber="124">
<typeStatus id="548B8842FF9DE63DFF510E07FF371080" box="[151,210,383,408]" pageId="15" pageNumber="124">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548B8842FF9DE63DFE8A0E07FE5C1080" box="[332,441,383,408]" pageId="15" pageNumber="124" type="holotype">Holotype</typeStatus>
: adult female (
<collectionCode id="ED21AE25FF9DE63DFD9B0EF8FD491080" box="[605,684,384,408]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="124">NSMT</collectionCode>
Cr 24630), ex
<taxonomicName id="4C304D63FF9DE63DFC9A0EF9FB341080" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[860,1233,383,408]" class="Gastropoda" family="Tegulidae" genus="Tectus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="15" pageNumber="124" phylum="Mollusca" rank="species" species="niloticus">
<emphasis id="B944EAF2FF9DE63DFC9A0EF9FBEF1080" box="[860,1034,383,408]" italics="true" pageId="15" pageNumber="124">Tectus niloticus</emphasis>
(Linnaeus, 1767)
</taxonomicName>
(Vestigastropoda: Trochoidea:
<taxonomicName id="4C304D63FF9DE63DFEE30EDCFE7D10A5" box="[293,408,420,445]" class="Gastropoda" family="Tegulidae" kingdom="Animalia" pageId="15" pageNumber="124" phylum="Mollusca" rank="family">Tegulidae</taxonomicName>
), off
<location id="8EEF603BFF9DE63DFE150EDCFDC010A5" LSID="urn:lsid:plazi:treatment:039987F6FF9DE621FF510E62FC3F15F0:8EEF603BFF9DE63DFE150EDCFDC010A5" box="[467,549,420,445]" country="Iceland" latitude="28.2" longLatPrecision="1233" longitude="129.2" municipality="Oshima Strait" name="Saneku" pageId="15" pageNumber="124">Saneku</location>
(
<geoCoordinate id="EE045027FF9DE63DFDF00EDCFD7010A4" box="[566,661,420,444]" direction="north" orientation="latitude" pageId="15" pageNumber="124" precision="925" value="28.2">28°12'N</geoCoordinate>
,
<geoCoordinate id="EE045027FF9DE63DFD640EDCFCE810A4" box="[674,781,420,444]" direction="east" orientation="longitude" pageId="15" pageNumber="124" precision="925" value="129.2">129°12'E</geoCoordinate>
), Kakeroma
<collectingCountry id="F3277670FF9DE63DFC580EDCFC0010A5" box="[926,997,420,445]" name="Iceland" pageId="15" pageNumber="124">Island</collectingCountry>
,
<collectingMunicipality id="6BEBAC9AFF9DE63DFC370EDCFB6A10A5" box="[1009,1167,420,445]" pageId="15" pageNumber="124">Oshima Strait</collectingMunicipality>
,
<collectingDate id="EFCAE9C8FF9DE63DFB5D0EDCFB1810A5" box="[1179,1277,420,445]" pageId="15" pageNumber="124" value="1929-04">
<date id="FF8E1020FF9DE63DFB5D0EDCFB1810A5" box="[1179,1277,420,445]" pageId="15" pageNumber="124" value="1929-04">29 April</date>
</collectingDate>
, 2015, leg.
<collectorName id="26C55336FF9DE63DFA470EDDFF0110F8" pageId="15" pageNumber="124">D. Uyeno</collectorName>
,
<collectorName id="26C55336FF9DE63DFF300EB0FE7210F8" box="[246,407,455,480]" pageId="15" pageNumber="124">S. Yokoyama</collectorName>
</materialsCitation>
. Allotype: adult male (NSMTCr 24631), collection data same as those of holotype. Paratypes: 2 adult females and 2 adult males (NSMTCr 24632), collection data same as those of holotype
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF010D77FA781353" blockId="15.[151,1437,383,1263]" pageId="15" pageNumber="124">
<materialsCitation id="3B583CBDFF9DE63DFF010D77FA7C1353" ID-GBIF-Occurrence="1412767588" collectingDate="3 March" collectionCode="NSMT" collectorName="D. Uyeno &amp; Y. Fujita &amp; T. Naruse" country="China" latitude="26.333334" location="Itchachi" longLatPrecision="1242" longitude="126.71667" municipality="Kume Island" pageId="15" pageNumber="124" specimenCount="2" specimenCount-adult="2">
<emphasis id="B944EAF2FF9DE63DFF010D77FE5F1330" bold="true" box="[199,442,527,552]" pageId="15" pageNumber="124">Additional material.</emphasis>
<specimenCount id="9D36FD69FF9DE63DFE070D68FDEB1330" box="[449,526,527,552]" pageId="15" pageNumber="124" type="adult">1 adult</specimenCount>
female and
<specimenCount id="9D36FD69FF9DE63DFD5F0D68FD031330" box="[665,742,527,552]" pageId="15" pageNumber="124" type="adult">1 adult</specimenCount>
male (
<collectionCode id="ED21AE25FF9DE63DFCF40D68FC641330" box="[818,897,528,552]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="124">NSMT</collectionCode>
Cr 24633), (
<collectionCode id="ED21AE25FF9DE63DFBDF0D68FB8D1330" box="[1049,1128,528,552]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="124">NSMT</collectionCode>
Cr 24633), ex
<taxonomicName id="4C304D63FF9DE63DFADF0D69FA711330" box="[1305,1428,527,552]" class="Gastropoda" family="Tegulidae" genus="Tectus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="15" pageNumber="124" phylum="Mollusca" rank="species" species="niloticus">
<emphasis id="B944EAF2FF9DE63DFADF0D69FA711330" box="[1305,1428,527,552]" italics="true" pageId="15" pageNumber="124">T. niloticus</emphasis>
</taxonomicName>
, off
<location id="8EEF603BFF9DE63DFF7B0D4BFEF21354" LSID="urn:lsid:plazi:treatment:039987F6FF9DE621FF510E62FC3F15F0:8EEF603BFF9DE63DFF7B0D4BFEF21354" box="[189,279,563,588]" country="China" latitude="26.333334" longLatPrecision="1242" longitude="126.71667" municipality="Kume Island" name="Itchachi" pageId="15" pageNumber="124">Itchachi</location>
(
<geoCoordinate id="EE045027FF9DE63DFEE00D4BFE621354" box="[294,391,563,588]" direction="north" orientation="latitude" pageId="15" pageNumber="124" precision="925" value="26.333334">26°20'N</geoCoordinate>
,
<geoCoordinate id="EE045027FF9DE63DFE570D4BFE191354" box="[401,508,563,588]" direction="east" orientation="longitude" pageId="15" pageNumber="124" precision="925" value="126.71667">126°43'E</geoCoordinate>
),
<collectingMunicipality id="6BEBAC9AFF9DE63DFDC90D4CFD441354" box="[527,673,563,588]" pageId="15" pageNumber="124">Kume Island</collectingMunicipality>
, East
<collectingCountry id="F3277670FF9DE63DFD250D4BFCC31354" box="[739,806,563,588]" name="China" pageId="15" pageNumber="124">China</collectingCountry>
Sea,
<collectingDate id="EFCAE9C8FF9DE63DFCA40D4CFC261354" box="[866,963,563,588]" pageId="15" pageNumber="124">
<date id="FF8E1020FF9DE63DFCA40D4CFC261354" box="[866,963,563,588]" pageId="15" pageNumber="124">3 March</date>
</collectingDate>
, 2012, leg.
<collectorName id="26C55336FF9DE63DFB840D4CFB541354" box="[1090,1201,564,588]" pageId="15" pageNumber="124">D. Uyeno</collectorName>
,
<collectorName id="26C55336FF9DE63DFB7A0D4CFAFB1354" box="[1212,1310,563,588]" pageId="15" pageNumber="124">Y. Fujita</collectorName>
,
<collectorName id="26C55336FF9DE63DFAEF0D4CFA7C1353" box="[1321,1433,564,587]" pageId="15" pageNumber="124">T. Naruse</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A656BFF9DE621FF010D2EFEE91220" lastPageId="19" lastPageNumber="128" pageId="15" pageNumber="124" type="description">
<paragraph id="8B8F36E0FF9DE63DFF010D2EFC4F1238" blockId="15.[151,1437,383,1263]" pageId="15" pageNumber="124">
<emphasis id="B944EAF2FF9DE63DFF010D2EFE0F1377" bold="true" box="[199,490,598,623]" pageId="15" pageNumber="124">
Description of
<typeStatus id="548B8842FF9DE63DFEBA0D2EFE001377" box="[380,485,598,623]" pageId="15" pageNumber="124" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female. Body (
<figureCitation id="130B2A65FF9DE63DFD2E0D2EFCDE1377" box="[744,827,598,623]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
A) cyclopiform, 2.94 long, depressed dorso-ventrally with greatest width at cephalosome; external segmentation distinct. Cephalosome distinctly articulated from first pedigerous somite, wider than long, 0.63 × 0.93. First to fourth pedigerous somites and urosomites free. Prosome 1.50 long. Genital double somite wider than long (
<figureCitation id="130B2A65FF9DE63DFD300DB9FCA913C2" box="[758,844,705,730]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
A), 0.47 × 0.32. Abdomen progressively narrower posteriorly, composed of three free somites, 0.25 × 0.26, 0.18 × 0.21, and 0.19 × 0.20, respectively. Caudal ramus (
<figureCitation id="130B2A65FF9DE63DFF590C70FF151238" box="[159,240,776,801]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
A, D) 2.85 times longer than wide, 0.23 × 0.08, with six setae.
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF010C54FB6D15BF" blockId="15.[151,1437,383,1263]" pageId="15" pageNumber="124">
Rostrum (
<figureCitation id="130B2A65FF9DE63DFEF10C53FE6F125C" box="[311,394,811,836]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
B) triangular, bearing distinct apex with round margin. Post-rostral process situated between bases of antennae (
<figureCitation id="130B2A65FF9DE63DFEAC0C37FE59127F" box="[362,444,847,872]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
B). Antennule (
<figureCitation id="130B2A65FF9DE63DFDA10C37FD52127F" box="[615,695,847,872]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
E) 7-segmented; armature formula 4, 14, 6, 3, 4 + 1 aesthetasc, 2 + 1 aesthetasc, 7 + 1 aesthetasc; all setae naked. Antenna (
<figureCitation id="130B2A65FF9DE63DFCEF0C0AFC991293" box="[809,892,882,907]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
F) 4-segmented, composed of coxobasis and 3- segmented endopod; coxobasis large, bearing single naked seta; first endopodal segment bearing simple seta on inner margin; second endopodal segment bearing three inner setae; third endopodal segment 1.44 times longer than wide, 61 × 43 µm, bearing four claws, one of them being elongate and three setae on distal part. Labrum (
<figureCitation id="130B2A65FF9DE63DFAE90CA5FA9B12ED" box="[1327,1406,989,1014]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
G) broad, bearing pair of posterior lobes. Mandible (
<figureCitation id="130B2A65FF9DE63DFD790B78FCEA1501" box="[703,783,1024,1049]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
H) bearing serrated blade with single hyaline tooth at base of convex margin of blade. Maxillule (
<figureCitation id="130B2A65FF9DE63DFD9D0B5CFD491524" box="[603,684,1060,1085]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig. 11</figureCitation>
A) represented by simple lobe armed with single inner and three distal setae. Maxilla (
<figureCitation id="130B2A65FF9DE63DFE4F0B3FFE341578" box="[393,465,1095,1120]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig.11</figureCitation>
B) 2-segmented; basal segment unarmed; terminal segment bearing serrated blade and three setae; inner seta large and spinulose. Maxilliped (
<figureCitation id="130B2A65FF9DE63DFCDE0B13FC8C159B" box="[792,873,1131,1156]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig. 11</figureCitation>
C) 3-segmented, composed of unarmed syncoxa, basis bearing two simple setae, and conical endopodal segment with two simple elements.
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF010BCAFC7E15F6" blockId="15.[151,1437,383,1263]" pageId="15" pageNumber="124">
Legs 1 to 4 (
<figureCitation id="130B2A65FF9DE63DFE9C0BC9FE4C15D2" box="[346,425,1201,1226]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig. 11</figureCitation>
DG) biramous, with 3-segmented rami. Leg 4 endopod unsegmented, almost as long as exopod, and with sinuous margins. Leg armature formula as follows:
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFE880A72FB441438" blockId="15.[159,1252,1290,1445]" box="[334,1185,1290,1312]" pageId="15" pageNumber="124">Coxa Basis Exopod Endopod</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF590A4CFBB214BD" blockId="15.[159,1252,1290,1445]" pageId="15" pageNumber="124">
Leg 1 0-1 1-0 I-0;
<date id="FF8E1020FF9DE63DFC820A4BFC811451" box="[836,868,1331,1353]" pageId="15" pageNumber="124">I-1</date>
; III, I, 4 0-1; 0-1; I, 2, 3 Leg 2 0-1 1-0 I-0;
<date id="FF8E1020FF9DE63DFC820A2AFC811470" box="[836,868,1362,1384]" pageId="15" pageNumber="124">I-1</date>
; III, I, 5 0-1; 0-2; II, I, 3 Leg 3 0-1 1-0 I-0;
<date id="FF8E1020FF9DE63DFC820A08FC81149E" box="[836,868,1392,1414]" pageId="15" pageNumber="124">I-1</date>
; III, I, 5 0-1; 0-2; II, I, 2 Leg 4 0-1 1-0 I-0;
<date id="FF8E1020FF9DE63DFC820AF7FC8114BD" box="[836,868,1423,1445]" pageId="15" pageNumber="124">I-1</date>
; III, I,
<date id="FF8E1020FF9DE63DFC6C0AF7FBB214BD" box="[938,1111,1423,1445]" pageId="15" pageNumber="124">5 II</date>
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF010A94FD1417C2" blockId="15.[151,1436,1516,1967]" pageId="15" pageNumber="124">
Intercoxal sclerites (
<figureCitation id="130B2A65FF9DE63DFE6F0A94FE13171D" box="[425,502,1516,1541]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig. 11</figureCitation>
DG) of legs 1 to 4 unornamented. Leg 1 bearing basis with row of hair-like setules on inner margin. All spines spatulate with serrated margins and terminal flagellum. Both rami of legs 1 to 4 bearing rows of hair-like setules on outer and inner margins, except inner margin of basal segment of endopods on legs 1 to 3. Basal and middle endopodal segments of legs 1 to 3 each bearing row of fine spinules along distal margin and distal pointed projection on outer margin. Terminal endopodal segments of legs 1 and 2 each bearing single pointed projection on outer margin. Leg 5 (
<figureCitation id="130B2A65FF9DE63DFDEF09E5FD9C17AE" box="[553,633,1693,1718]" captionStart="FIGURE 11" captionStartId="17.[151,250,1896,1918]" captionTargetBox="[211,1378,196,1863]" captionTargetId="figure@17.[197,1381,193,1868]" captionTargetPageId="17" captionText="FIGURE 11. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; D G, 100 µm; H, 50 µm." httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="15" pageNumber="124">Fig. 11</figureCitation>
H) represented by simple spine and two naked setae. Leg 6 (
<figureCitation id="130B2A65FF9DE63DFAE809E5FA6517AE" box="[1326,1408,1693,1718]" captionStart="FIGURE 10" captionStartId="16.[151,250,1898,1920]" captionTargetBox="[216,1379,201,1867]" captionTargetId="figure@16.[197,1381,195,1870]" captionTargetPageId="16" captionText="FIGURE 10. Pseudanthessius imo n. sp., adult female, holotype NSMT Cr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm." httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="15" pageNumber="124">Fig. 10</figureCitation>
C) represented by two small elements at genital opening.
</paragraph>
<paragraph id="8B8F36E0FF9DE63DFF01099CFCEE16B6" blockId="15.[151,1436,1516,1967]" pageId="15" pageNumber="124">
<emphasis id="B944EAF2FF9DE63DFF01099CFE3817E5" bold="true" box="[199,477,1764,1789]" pageId="15" pageNumber="124">
Description of
<typeStatus id="548B8842FF9DE63DFEB1099CFE3D17E5" box="[375,472,1764,1789]" pageId="15" pageNumber="124" type="allotype">allotype</typeStatus>
.
</emphasis>
Adult male. Body (
<figureCitation id="130B2A65FF9DE63DFD7B099CFCE917E5" box="[701,780,1764,1789]" captionStart="FIGURE 12" captionStartId="18.[151,250,1669,1691]" captionTargetBox="[205,1371,204,1626]" captionTargetId="figure@18.[196,1382,193,1634]" captionTargetPageId="18" captionText="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="15" pageNumber="124">Fig. 12</figureCitation>
A) cyclopiform, 2.43 long, depressed dorso-ventrally with greatest width at cephalosome; external segmentations distinct. Cephalosome distinctly articulated from first pedigerous somite, wider than long, 0.55 × 0.80. First to fourth pedigerous somite and urosomites free. Prosome 1.14 long. Genital somite (
<figureCitation id="130B2A65FF9DE63DFE0F0837FDFE1670" box="[457,539,1871,1896]" captionStart="FIGURE 12" captionStartId="18.[151,250,1669,1691]" captionTargetBox="[205,1371,204,1626]" captionTargetId="figure@18.[196,1382,193,1634]" captionTargetPageId="18" captionText="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="15" pageNumber="124">Fig. 12</figureCitation>
A, B), 0.23 × 0.35. Abdomen progressively narrower posteriorly, composed of four free somites, 0.23 × 0.28, 0.20 × 0.24, 0.16 × 0.20, and 0.16 × 0.18, respectively. Caudal ramus (
<figureCitation id="130B2A65FF9DE63DFAEA080AFA981693" box="[1324,1405,1906,1931]" captionStart="FIGURE 12" captionStartId="18.[151,250,1669,1691]" captionTargetBox="[205,1371,204,1626]" captionTargetId="figure@18.[196,1382,193,1634]" captionTargetPageId="18" captionText="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="15" pageNumber="124">Fig. 12</figureCitation>
A) 3.77 times longer than wide, 0.22 × 0.06, with six setae.
</paragraph>
<caption id="DF4F6668FF82E622FF510812FB7F16A5" httpUri="https://zenodo.org/record/177788/files/figure.png" pageId="16" pageNumber="125" targetBox="[216,1379,201,1867]" targetPageId="16">
<paragraph id="8B8F36E0FF82E622FF510812FB7F16A5" blockId="16.[151,1436,1898,1981]" pageId="16" pageNumber="125">
<emphasis id="B944EAF2FF82E622FF510812FEC61667" bold="true" box="[151,291,1898,1920]" pageId="16" pageNumber="125">FIGURE 10.</emphasis>
<taxonomicName id="4C304D63FF82E622FEEA0812FDE11698" box="[300,516,1898,1920]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="imo" status="sp. nov.">
<emphasis id="B944EAF2FF82E622FEEA0812FDE11698" box="[300,516,1898,1920]" italics="true" pageId="16" pageNumber="125">Pseudanthessius imo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775789FF82E622FDCB0812FDAF1667" box="[525,586,1898,1919]" pageId="16" pageNumber="125" rank="species">
<emphasis id="B944EAF2FF82E622FDCB0812FDAF1667" bold="true" box="[525,586,1898,1919]" pageId="16" pageNumber="125">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female, holotype NSMTCr 24630. A, habitus, dorsal; B, rostral area; C, genital aperture and leg 6, dorsal; D, left caudal ramus, dorsal; E, right antennule; F, right antenna, anterior; G, labrum, anterior; H, left mandible, anterior. Scale bars: A, 400 µm; B, 100 µm; C, D, F, G, 50 µm; E, 40 µm; H, 20 µm.
</paragraph>
</caption>
<caption id="DF4F6668FF83E623FF510810FBDC16A3" httpUri="https://zenodo.org/record/177789/files/figure.png" pageId="17" pageNumber="126" targetBox="[211,1378,196,1863]" targetPageId="17">
<paragraph id="8B8F36E0FF83E623FF510810FBDC16A3" blockId="17.[151,1436,1896,1979]" pageId="17" pageNumber="126">
<emphasis id="B944EAF2FF83E623FF510810FEC71665" bold="true" box="[151,290,1896,1918]" pageId="17" pageNumber="126">FIGURE 11.</emphasis>
<taxonomicName id="4C304D63FF83E623FEEB0810FDE01666" box="[301,517,1896,1918]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="126" phylum="Arthropoda" rank="species" species="imo" status="sp. nov.">
<emphasis id="B944EAF2FF83E623FEEB0810FDE01666" box="[301,517,1896,1918]" italics="true" pageId="17" pageNumber="126">Pseudanthessius imo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775789FF83E623FDC90810FDA81665" box="[527,589,1896,1917]" pageId="17" pageNumber="126" rank="species">
<emphasis id="B944EAF2FF83E623FDC90810FDA81665" bold="true" box="[527,589,1896,1917]" pageId="17" pageNumber="126">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female, holotype NSMTCr 24630. A, right maxillule, anterior; B, right maxilla, posterior; C, right maxilliped, posterior; D, left leg 1, anterior; E, left leg 2, anterior; F, left leg 3, anterior; G, left leg 4, anterior; H, right leg 5, dorsal. Scale bars: A, 20 µm; B, C, 40 µm; DG, 100 µm; H, 50 µm.
</paragraph>
</caption>
<caption id="DF4F6668FF80E620FF5109FDFB5017A2" httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="18" pageNumber="127" targetBox="[205,1371,204,1626]" targetPageId="18">
<paragraph id="8B8F36E0FF80E620FF5109FDFB5017A2" blockId="18.[151,1436,1669,1722]" pageId="18" pageNumber="127">
<emphasis id="B944EAF2FF80E620FF5109FDFEC51782" bold="true" box="[151,288,1669,1691]" pageId="18" pageNumber="127">FIGURE 12.</emphasis>
<taxonomicName id="4C304D63FF80E620FEE109FDFE191783" box="[295,508,1669,1691]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="18" pageNumber="127" phylum="Arthropoda" rank="species" species="imo" status="sp. nov.">
<emphasis id="B944EAF2FF80E620FEE109FDFE191783" box="[295,508,1669,1691]" italics="true" pageId="18" pageNumber="127">Pseudanthessius imo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775789FF80E620FDC509FDFDD81782" box="[515,573,1669,1690]" pageId="18" pageNumber="127" rank="species">
<emphasis id="B944EAF2FF80E620FDC509FDFDD81782" bold="true" box="[515,573,1669,1690]" pageId="18" pageNumber="127">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male, allotype NSMTCr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm.
</paragraph>
</caption>
<paragraph id="8B8F36E0FF80E620FF01099DFDFB1696" blockId="18.[151,1436,1765,2007]" pageId="18" pageNumber="127">
Antennule, antenna, mandible, maxillule, and maxilla as in female. Antenna bearing third endopodal segment 1.48 times longer than wide, 57 × 38 µm. Maxilliped (
<figureCitation id="130B2A65FF80E620FCD20872FC82163A" box="[788,871,1802,1827]" captionStart="FIGURE 12" captionStartId="18.[151,250,1669,1691]" captionTargetBox="[205,1371,204,1626]" captionTargetId="figure@18.[196,1382,193,1634]" captionTargetPageId="18" captionText="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="18" pageNumber="127">Fig. 12</figureCitation>
D) 4-segmented; basal segment unarmed; second segment bearing two setae and row of fine spinules; third segment small without armature; terminal claw curved bearing two basal setae. Armature formula of legs 1 to 5 as in female. Leg 6 (
<figureCitation id="130B2A65FF80E620FBE3082AFB9C1672" box="[1061,1145,1874,1899]" captionStart="FIGURE 12" captionStartId="18.[151,250,1669,1691]" captionTargetBox="[205,1371,204,1626]" captionTargetId="figure@18.[196,1382,193,1634]" captionTargetPageId="18" captionText="FIGURE 12. Pseudanthessius imo n. sp., adult male, allotype NSMT Cr 2463. A, habitus, dorsal; B, genital somite, ventral; C, right leg 6, ventral; D, left maxilliped, inner. Scale bars: A, 400 µm; B, 100 µm; C, 20 µm; D, 40 µm." httpUri="https://zenodo.org/record/177790/files/figure.png" pageId="18" pageNumber="127">Fig. 12</figureCitation>
B, C) represented by two simple setae on genital operculum.
</paragraph>
<paragraph id="8B8F36E0FF80E621FF0108E2FC6A10B5" blockId="18.[151,1436,1765,2007]" lastBlockId="19.[151,1437,151,1256]" lastPageId="19" lastPageNumber="128" pageId="18" pageNumber="127">
<emphasis id="B944EAF2FF80E620FF0108E2FEA816AB" bold="true" box="[199,333,1946,1971]" pageId="18" pageNumber="127">Variability.</emphasis>
Morphology of
<typeStatus id="548B8842FF80E620FDD108E3FD6616AB" box="[535,643,1947,1971]" pageId="18" pageNumber="127" type="paratype">paratypes</typeStatus>
and other female specimens as in
<typeStatus id="548B8842FF80E620FBE908E2FB7016AB" box="[1071,1173,1946,1971]" pageId="18" pageNumber="127" type="holotype">holotype</typeStatus>
. Measurements of the
<typeStatus id="548B8842FF80E620FF5108C7FEE716CF" box="[151,258,1983,2007]" pageId="18" pageNumber="127" type="paratype">paratypes</typeStatus>
and other specimens (
<emphasis id="B944EAF2FF80E620FDD408C7FDC516CE" box="[530,544,1983,2006]" italics="true" pageId="18" pageNumber="127">n</emphasis>
= 3) as follows: body length 2.082.51 (2.31 ± 0.22); cephalosome length 0.580.65 (0.63 ± 0.04); cephalosome width 0.820.98 (0.90 ± 0.08); prosome length 1.051.33 (1.23 ± 0.15); genital double somite length 0.300.36 (0.33 ± 0.03); genital double somite width 0.290.37 (0.34 ± 0.04); first urosomite length 0.150.21 (0.19 ± 0.04); first urosomite width 0.270.30 (0.29 ± 0.02); second urosomite length 0.120.17 (0.14 ± 0.03); second urosomite width 0.220.28 (0.25 ± 0.03); anal somite length 0.170.19 (0.18 ± 0.01); anal somite width 0.200.23 (0.22 ± 0.01); caudal ramus length 0.210.26 (0.24 ± 0.02); caudal ramus width 0.07; length of third endopodal segment of antenna 5774 (65 ± 8) µm; width of third endopodal segment of antenna 3861 (46 ± 13) µm. Caudal ramus 3.003.56 (3.35 ± 0.31) times longer than wide. Third endopodal segment of antenna 1.211.65 (1.46 ± 0.23) times longer than wide.
</paragraph>
<paragraph id="8B8F36E0FF81E621FF010ECFFEE91220" blockId="19.[151,1437,151,1256]" pageId="19" pageNumber="128">
Morphology of
<typeStatus id="548B8842FF81E621FE440EC0FE0810C8" box="[386,493,440,464]" pageId="19" pageNumber="128" type="paratype">paratypes</typeStatus>
and other male specimens as in
<typeStatus id="548B8842FF81E621FCBD0ECFFC3C10C8" box="[891,985,439,464]" pageId="19" pageNumber="128" type="allotype">allotype</typeStatus>
. Measurements of male
<typeStatus id="548B8842FF81E621FB3A0EC0FA8D10C8" box="[1276,1384,440,464]" pageId="19" pageNumber="128" type="paratype">paratypes</typeStatus>
and additional specimens (
<emphasis id="B944EAF2FF81E621FE550EA5FE4410EC" box="[403,417,477,500]" italics="true" pageId="19" pageNumber="128">n</emphasis>
= 3) as follows: body length 2.082.30 (2.15 ± 0.13); cephalosome length 0.510.63 (0.58 ± 0.06); cephalosome width 0.680.85 (0.75 ± 0.09); prosome length 0.961.14 (1.03 ± 0.10); genital somite length 0.180.26 (0.22 ± 0.04); genital somite width 0.310.43 (0.37 ± 0.06); first urosomite length 0.17; first urosomite width 0.250.31 (0.28 ± 0.03); second urosomite length 0.180.20 (0.19 ± 0.01); second urosomite width 0.230.28 (0.25 ± 0.02); third urosomite length 0.130.16 (0.14 ± 0.02); third urosomite width 0.180.23 (0.20 ± 0.03); anal somite length 0.120.15 (0.14 ± 0.01); anal somite width 0.170.19 (0.18 ± 0.01); caudal ramus length 0.180.24 (0.21 ± 0.03); caudal ramus width 0.060.07 (0.07 ± 0.00); length of third endopodal segment of antenna 4971 (61 ± 11) µm; width of third endopodal segment of antenna 3642 (38 ± 3) µm. Caudal ramus 2.743.69 (3.24 ± 0.48) times longer than wide. Third endopodal segment of antenna 1.331.73 (1.59 ± 0.22) times longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C32A656BFF81E621FF010C3CFDD71565" pageId="19" pageNumber="128" type="discussion">
<paragraph id="8B8F36E0FF81E621FF010C3CFD271540" blockId="19.[151,1437,151,1256]" pageId="19" pageNumber="128">
<emphasis id="B944EAF2FF81E621FF010C3CFEDE1245" bold="true" box="[199,315,836,861]" pageId="19" pageNumber="128">Remarks.</emphasis>
<taxonomicName id="4C304D63FF81E621FE830C3CFDD61244" box="[325,563,836,861]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="19" pageNumber="128" phylum="Arthropoda" rank="species" species="imo" status="sp. nov.">
<emphasis id="B944EAF2FF81E621FE830C3CFDD61244" box="[325,563,836,861]" italics="true" pageId="19" pageNumber="128">Pseudanthessius imo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775789FF81E621FDFB0C3CFD651245" box="[573,640,836,861]" pageId="19" pageNumber="128" rank="species">
<emphasis id="B944EAF2FF81E621FDFB0C3CFD651245" bold="true" box="[573,640,836,861]" pageId="19" pageNumber="128">n. sp.</emphasis>
</taxonomicNameLabel>
has four spines on the third exopodal segment of leg 4, a character shared with
<taxonomicName id="4C304D63FF81E621FED80C11FCB21298" authority="Shin &amp; Kim, 2004 (Shin &amp; Kim, 2004)" authorityName="Shin &amp; Kim, 2004 (Shin &amp; Kim" authorityYear="2004" box="[286,855,871,896]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="19" pageNumber="128" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B944EAF2FF81E621FED80C11FE7F1298" box="[286,410,873,896]" italics="true" pageId="19" pageNumber="128">P. spinosus</emphasis>
<bibRefCitation id="EFA14B11FF81E621FE660C1FFD881298" author="Shin" box="[416,621,871,896]" pageId="19" pageNumber="128" refString="Shin, S. &amp; Kim, I. - H. (2004) Pseudanthessius spinosus, a new species of Copepoda (Poecilostomatoida, Pseudanthessiidae) associated with the echinoid Clypeaster japonicus from Korea. Korean Journal of Biological Sciences, 8, 13 - 18. http: // dx. doi. org / 10.1080 / 12265071.2004.9647728" type="journal article" year="2004">Shin &amp; Kim, 2004</bibRefCitation>
(
<bibRefCitation id="EFA14B11FF81E621FDBA0C1FFCB51298" author="Shin" box="[636,848,871,896]" pageId="19" pageNumber="128" refString="Shin, S. &amp; Kim, I. - H. (2004) Pseudanthessius spinosus, a new species of Copepoda (Poecilostomatoida, Pseudanthessiidae) associated with the echinoid Clypeaster japonicus from Korea. Korean Journal of Biological Sciences, 8, 13 - 18. http: // dx. doi. org / 10.1080 / 12265071.2004.9647728" type="journal article" year="2004">Shin &amp; Kim, 2004</bibRefCitation>
)
</taxonomicName>
. The new species differs from
<taxonomicName id="4C304D63FF81E621FB6A0C11FAC31298" box="[1196,1318,873,896]" class="Maxillopoda" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Poecilostomatoida" pageId="19" pageNumber="128" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B944EAF2FF81E621FB6A0C11FAC31298" box="[1196,1318,873,896]" italics="true" pageId="19" pageNumber="128">P. spinosus</emphasis>
</taxonomicName>
by having the following characters: caudal ramus 2.5 to 4 times longer than wide (vs. less than 2 times); presence of postrostral process between bases of antenna (vs. process absent); third endopodal segment of antenna less than 2 times longer than wide, with four claws and three setae (vs. about 3 times longer than wide, bearing two claws, one clawlike seta, and three setae); bearing mandible with single hyaline tooth on convex margin (vs. with four small spinules); leg 4 bearing unsegmented endopod with sinuous both lateral margins (vs. with straight margins and two conical distal processes) (see
<bibRefCitation id="EFA14B11FF81E621FE240B47FD531540" author="Shin" box="[482,694,1087,1112]" pageId="19" pageNumber="128" refString="Shin, S. &amp; Kim, I. - H. (2004) Pseudanthessius spinosus, a new species of Copepoda (Poecilostomatoida, Pseudanthessiidae) associated with the echinoid Clypeaster japonicus from Korea. Korean Journal of Biological Sciences, 8, 13 - 18. http: // dx. doi. org / 10.1080 / 12265071.2004.9647728" type="journal article" year="2004">Shin &amp; Kim, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36E0FF81E621FF010B1CFDD71565" blockId="19.[151,1437,151,1256]" box="[199,562,1124,1149]" pageId="19" pageNumber="128">
<emphasis id="B944EAF2FF81E621FF010B1CFE6E1565" bold="true" box="[199,395,1124,1149]" pageId="19" pageNumber="128">Attachment site.</emphasis>
Mantle cavity.
</paragraph>
</subSubSection>
<subSubSection id="C32A656BFF81E621FF010BFFFC3F15F0" pageId="19" pageNumber="128" type="etymology">
<paragraph id="8B8F36E0FF81E621FF010BFFFC5D15DD" blockId="19.[151,1437,151,1256]" pageId="19" pageNumber="128">
<emphasis id="B944EAF2FF81E621FF010BFFFEA315B8" bold="true" box="[199,326,1159,1184]" pageId="19" pageNumber="128">Etymology</emphasis>
. The specific name means “sweet potato” in Japanese. It alludes to shape of the irregularly constricted endopod of leg 4. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F36E0FF81E621FF010BB7FC3F15F0" blockId="19.[151,1437,151,1256]" box="[199,986,1231,1256]" pageId="19" pageNumber="128">
<emphasis id="B944EAF2FF81E621FF010BB7FD0115F0" bold="true" box="[199,740,1231,1256]" pageId="19" pageNumber="128">Newly established Japanese name for species.</emphasis>
<emphasis id="B944EAF2FF81E621FD2D0BB7FC3615F0" box="[747,979,1231,1256]" italics="true" pageId="19" pageNumber="128">Imo-ashi-kenmijinko</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>