528 lines
62 KiB
XML
528 lines
62 KiB
XML
<document id="F7761DD0C0BA535B6D26245522249514" ID-DOI="10.11646/zootaxa.5352.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="8426638" ID-ZooBank="74DCF84E-60ED-49EA-B5E2-A794A60E4D06" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1696942088598" checkinUser="plazi" docAuthor="Matson, Tanner A." docDate="2023" docId="039987ADFFC5E02ED398FC75FE75FB5B" docLanguage="en" docName="zootaxa.5352.4.7.pdf" docOrigin="Zootaxa 5352 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achagua magna Matson 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="571" masterDocId="FFA0FFD5FFC7E028D30FFFA7FF89FFA1" masterDocTitle="The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae)" masterLastPageNumber="576" masterPageNumber="565" pageNumber="567" updateTime="1697115518778" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="557781DFE03554964B7D2483A0DA62F0">The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae)</mods:title>
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<mods:name id="A5E24BE3472A4711CEDD2B289DC4DA0C" type="personal">
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<mods:namePart id="A5C8E67E6A2C6139D819DFD67220E750">Matson, Tanner A.</mods:namePart>
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<mods:title id="FBA91EC58235F70AF72758B0DFF90A32">Zootaxa</mods:title>
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<mods:part id="6568A40979D2AAA94E23CA1E5DA718D8">
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<mods:date id="3F3CA4150489125D54343EB810087541">2023</mods:date>
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<mods:number id="37A9B9BDCD8B96BA4CD59B0EF5315018">2023-10-05</mods:number>
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<mods:number id="E91550AB79387E24E0C69004399B3A44">5352</mods:number>
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<mods:start id="EC8CC9080E44B174F63BCC2866CF0535">565</mods:start>
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<mods:url id="59B4D128939AECE8ECE6C2BC2408E5FC">http://dx.doi.org/10.11646/zootaxa.5352.4.7</mods:url>
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<mods:classification id="0F24AD21A4C610573CA87908F00C3636">journal article</mods:classification>
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<mods:identifier id="FF8F89DACA9D4934E583AA95B427D3A3" type="DOI">10.11646/zootaxa.5352.4.7</mods:identifier>
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<mods:identifier id="2AB81B704592453883092C496966806D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039987ADFFC5E02ED398FC75FE75FB5B" LSID="urn:lsid:plazi:treatment:039987ADFFC5E02ED398FC75FE75FB5B" httpUri="http://treatment.plazi.org/id/039987ADFFC5E02ED398FC75FE75FB5B" lastPageId="6" lastPageNumber="571" pageId="2" pageNumber="567">
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<subSubSection id="C32A6530FFC5E02AD398FC75FD9FFC4C" box="[151,534,978,1005]" pageId="2" pageNumber="567" type="nomenclature">
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<paragraph id="8B8F36BBFFC5E02AD398FC75FD9FFC4C" blockId="2.[151,770,978,1077]" box="[151,534,978,1005]" pageId="2" pageNumber="567">
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<heading id="D0C781D7FFC5E02AD398FC75FD9FFC4C" bold="true" box="[151,534,978,1005]" fontSize="11" level="1" pageId="2" pageNumber="567" reason="1">
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<emphasis id="B944EAA9FFC5E02AD398FC75FD9FFC4C" bold="true" box="[151,534,978,1005]" pageId="2" pageNumber="567">
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<taxonomicName id="4C304D38FFC5E02AD398FC75FE4FFC4C" authority="Matson" authorityName="Matson" authorityYear="2023" box="[151,454,978,1005]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna" status="n. sp.">
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<emphasis id="B944EAA9FFC5E02AD398FC75FED6FC4C" bold="true" box="[151,351,978,1005]" italics="true" pageId="2" pageNumber="567">Achagua magna</emphasis>
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Matson
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27757D2FFC5E02AD2DCFC74FD9FFC4C" box="[467,534,979,1005]" pageId="2" pageNumber="567" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A6530FFC5E02AD398FC50FC8BFB94" pageId="2" pageNumber="567" type="description">
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<paragraph id="8B8F36BBFFC5E02AD398FC50FEEFFBB0" blockId="2.[151,770,978,1077]" box="[151,358,1015,1041]" pageId="2" pageNumber="567">
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(
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<figureCitation id="130B2A3EFFC5E02AD390FC50FF66FBB0" box="[159,239,1015,1041]" captionStart="FIGURES 1–4" captionStartId="3.[152,270,1870,1894]" captionTargetBox="[156,1432,179,1844]" captionTargetId="figure-16@3.[156,1432,179,1844]" captionTargetPageId="3" captionText="FIGURES 1–4. Achagua adult habitus. (a) dorsal view, (b) ventral view. (1) A. obsoleta Rindge, AMNH_IZC 00353009, paratype, (Colombia: Cundinamarca), AMNH; (2) A. magna n. sp., AMNH_IZC 00353021, holotype, (Peru: Cusco), AMNH; (3) A. cooperae n. sp., USNMENT01771270, holotype, (Costa Rica: Alajuela), USNM; (4) A. velata n. sp., holotype, (French Guiana: Régina), MNHN. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.8426640" httpUri="https://zenodo.org/record/8426640/files/figure.png" pageId="2" pageNumber="567">Figs. 2</figureCitation>
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,
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<figureCitation id="130B2A3EFFC5E02AD3F4FC50FE82FBB0" box="[251,267,1015,1041]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="2" pageNumber="567">6</figureCitation>
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,
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<figureCitation id="130B2A3EFFC5E02AD218FC50FEBDFBB0" box="[279,308,1015,1041]" captionStart="FIGURE 11" captionStartId="9.[152,255,1039,1063]" captionTargetBox="[151,1436,181,1015]" captionTargetId="figure-16@9.[151,1436,181,1015]" captionTargetPageId="9" captionText="FIGURE 11. Geographic distribution of Achagua: A. obsoleta Rindge (star), A. magna n. sp. (triangle), A. cooperae n. sp. (circle), and A. velata n. sp. (square). Single dots may represent>1 individual. Scale bar = 600 km." figureDoi="http://doi.org/10.5281/zenodo.8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="2" pageNumber="567">11</figureCitation>
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,
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<figureCitation id="130B2A3EFFC5E02AD24FFC50FED4FBB0" box="[320,349,1015,1041]" captionStart="FIGURE 12" captionStartId="9.[152,255,1750,1774]" captionTargetBox="[172,1413,1136,1720]" captionTargetId="figure-60@9.[162,1426,1124,1726]" captionTargetPageId="9" captionText="FIGURE 12. COI neighbor-joining tree generated from all publicly available Achagua sequences. Taxon names followed by BOLD Process ID and country of origin." figureDoi="http://doi.org/10.5281/zenodo.8426648" httpUri="https://zenodo.org/record/8426648/files/figure.png" pageId="2" pageNumber="567">12</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8F36BBFFC5E02AD398FBBCFC8BFB94" blockId="2.[151,770,978,1077]" box="[151,770,1050,1077]" pageId="2" pageNumber="567">
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LSID:
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<uuid id="FF960C6EFFC5E02AD3EBFBBDFC8BFB94" box="[228,770,1050,1077]" pageId="2" pageNumber="567">EBF2D227-50CC-420C-83F7-2939D2F9C3AD</uuid>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32A6530FFC5E02AD398FBC5FE29F9F0" pageId="2" pageNumber="567" type="diagnosis">
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<paragraph id="8B8F36BBFFC5E02AD398FBC5FB6CFAF4" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="567">
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<emphasis id="B944EAA9FFC5E02AD398FBC5FE9BFBDD" bold="true" box="[151,274,1122,1148]" pageId="2" pageNumber="567">Diagnosis.</emphasis>
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<taxonomicName id="4C304D38FFC5E02AD217FBC4FE47FBDD" authorityName="Matson" authorityYear="2023" box="[280,462,1123,1149]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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<emphasis id="B944EAA9FFC5E02AD217FBC4FE47FBDD" box="[280,462,1123,1149]" italics="true" pageId="2" pageNumber="567">Achagua magna</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName id="4C304D38FFC5E02AD195FBC4FC91FBDC" box="[666,792,1123,1149]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC5E02AD195FBC4FC91FBDC" box="[666,792,1123,1149]" italics="true" pageId="2" pageNumber="567">A. obsoleta</emphasis>
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</taxonomicName>
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. While
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<taxonomicName id="4C304D38FFC5E02AD060FBC4FC53FBDD" authorityName="Matson" authorityYear="2023" box="[879,986,1123,1149]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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<emphasis id="B944EAA9FFC5E02AD060FBC4FC53FBDD" box="[879,986,1123,1149]" italics="true" pageId="2" pageNumber="567">A. magna</emphasis>
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</taxonomicName>
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inhabits the eastern slopes of the central Andes in
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<collectingCountry id="F327762BFFC5E02AD208FB20FEEFFB00" box="[263,358,1159,1185]" name="Ecuador" pageId="2" pageNumber="567">Ecuador</collectingCountry>
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,
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<collectingCountry id="F327762BFFC5E02AD27BFB20FE23FB00" box="[372,426,1159,1185]" name="Peru" pageId="2" pageNumber="567">Peru</collectingCountry>
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, and
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<collectingCountry id="F327762BFFC5E02AD2E5FB20FDB6FB00" box="[490,575,1159,1185]" name="Bolivia" pageId="2" pageNumber="567">Bolivia</collectingCountry>
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,
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<taxonomicName id="4C304D38FFC5E02AD142FB20FD44FB00" box="[589,717,1159,1185]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC5E02AD142FB20FD44FB00" box="[589,717,1159,1185]" italics="true" pageId="2" pageNumber="567">A. obsoleta</emphasis>
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</taxonomicName>
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is thus far only known from the northern Andes in
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<collectingCountry id="F327762BFFC5E02AD628FB20FA10FB00" box="[1319,1433,1159,1185]" name="Colombia" pageId="2" pageNumber="567">Colombia</collectingCountry>
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.
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<taxonomicName id="4C304D38FFC5E02AD398FB0CFED9FB65" authorityName="Matson" authorityYear="2023" box="[151,336,1195,1221]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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<emphasis id="B944EAA9FFC5E02AD398FB0CFED9FB65" box="[151,336,1195,1221]" italics="true" pageId="2" pageNumber="567">Achagua magna</emphasis>
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</taxonomicName>
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(forewing,
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<quantity id="4CC89B5EFFC5E02AD2D7FB0CFDDDFB64" box="[472,596,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="2" pageNumber="567" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21–22 mm</quantity>
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) is noticeably larger than
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<taxonomicName id="4C304D38FFC5E02AD08EFB0CFB88FB64" box="[897,1025,1195,1221]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC5E02AD08EFB0CFB88FB64" box="[897,1025,1195,1221]" italics="true" pageId="2" pageNumber="567">A. obsoleta</emphasis>
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</taxonomicName>
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(forewing,
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||
<quantity id="4CC89B5EFFC5E02AD786FB0CFA81FB64" box="[1161,1288,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="2" pageNumber="567" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
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), and though subtle, the pearly-white ground color of the forewing upperside appears to be marked more strongly with intermittent black scaling over the veins; especially in the antemedial and postmedial areas.
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<taxonomicName id="4C304D38FFC5E02AD734FB54FB78FAAD" authorityName="Matson" authorityYear="2023" box="[1083,1265,1267,1293]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
|
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<emphasis id="B944EAA9FFC5E02AD734FB54FB78FAAD" box="[1083,1265,1267,1293]" italics="true" pageId="2" pageNumber="567">Achagua magna</emphasis>
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</taxonomicName>
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also appears to bear a more strongly marked, brown to black, terminal area of the hindwing. In males, the lateral margin of the large, subtriangular, anellar processes appears to be less deeply concave in
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<taxonomicName id="4C304D38FFC5E02AD095FA9CFB8CFAF5" authorityName="Matson" authorityYear="2023" box="[922,1029,1339,1365]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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||
<emphasis id="B944EAA9FFC5E02AD095FA9CFB8CFAF5" box="[922,1029,1339,1365]" italics="true" pageId="2" pageNumber="567">A. magna</emphasis>
|
||
</taxonomicName>
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than in
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<taxonomicName id="4C304D38FFC5E02AD76FFA9CFB56FAF4" box="[1120,1247,1339,1365]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC5E02AD76FFA9CFB56FAF4" box="[1120,1247,1339,1365]" italics="true" pageId="2" pageNumber="567">A. obsoleta</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B8F36BBFFC5E02AD3C8FAF8FE29F9F0" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="567">
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||
<taxonomicName id="4C304D38FFC5E02AD3C8FAF8FE09FAD9" authorityName="Matson" authorityYear="2023" box="[199,384,1375,1401]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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||
<emphasis id="B944EAA9FFC5E02AD3C8FAF8FE09FAD9" box="[199,384,1375,1401]" italics="true" pageId="2" pageNumber="567">Achagua magna</emphasis>
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</taxonomicName>
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can be distinguished from both
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<taxonomicName id="4C304D38FFC5E02AD1F9FAF8FCF6FAD9" authorityName="Matson" authorityYear="2023" box="[758,895,1375,1401]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="cooperae">
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||
<emphasis id="B944EAA9FFC5E02AD1F9FAF8FCF6FAD9" box="[758,895,1375,1401]" italics="true" pageId="2" pageNumber="567">A. cooperae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C304D38FFC5E02AD0B5FAF8FBA9FAD8" authorityName="Matson" authorityYear="2023" box="[954,1056,1375,1401]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="velata">
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<emphasis id="B944EAA9FFC5E02AD0B5FAF8FBA9FAD8" box="[954,1056,1375,1401]" italics="true" pageId="2" pageNumber="567">A. velata</emphasis>
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</taxonomicName>
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by the absence of a conspicuous antemedial black spot along the forewing inner margin. Additionally, the male genitalia of
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<taxonomicName id="4C304D38FFC5E02AD794FA24FA8EFA3D" authorityName="Matson" authorityYear="2023" box="[1179,1287,1411,1437]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
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<emphasis id="B944EAA9FFC5E02AD794FA24FA8EFA3D" box="[1179,1287,1411,1437]" italics="true" pageId="2" pageNumber="567">A. magna</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="130B2A3EFFC5E02AD617FA24FAD5FA3C" box="[1304,1372,1411,1437]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="2" pageNumber="567">Fig. 6</figureCitation>
|
||
) bear a spatulate dorsal process of the uncus, a costa with a postmedial digitate protuberance, and a denticulate patch of cornuti on the vesica. In
|
||
<taxonomicName id="4C304D38FFC5E02AD2A9FA6CFDA5FA45" authorityName="Matson" authorityYear="2023" box="[422,556,1483,1509]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="cooperae">
|
||
<emphasis id="B944EAA9FFC5E02AD2A9FA6CFDA5FA45" box="[422,556,1483,1509]" italics="true" pageId="2" pageNumber="567">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130B2A3EFFC5E02AD135FA6CFDF4FA44" box="[570,637,1483,1509]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="2" pageNumber="567">Fig. 7</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C304D38FFC5E02AD1B6FA6CFC92FA44" authorityName="Matson" authorityYear="2023" box="[697,795,1483,1509]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="velata">
|
||
<emphasis id="B944EAA9FFC5E02AD1B6FA6CFC92FA44" box="[697,795,1483,1509]" italics="true" pageId="2" pageNumber="567">A. velata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130B2A3EFFC5E02AD025FA6CFCE5FA44" box="[810,876,1483,1509]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="2" pageNumber="567">Fig. 8</figureCitation>
|
||
), the dorsal process of the uncus is not spatulate or obviously swollen, the costa lacks a postmedial digitate protuberance, and the vesica lacks cornuti. There are other, more subtle differences that can be observed in
|
||
<figureCitation id="130B2A3EFFC5E02AD1BEF9B4FCB1F98C" box="[689,824,1555,1581]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="2" pageNumber="567">Figures 6–8</figureCitation>
|
||
, which further serve to differentiate
|
||
<taxonomicName id="4C304D38FFC5E02AD7DCF9B4FAB6F98D" authorityName="Matson" authorityYear="2023" box="[1235,1343,1555,1581]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC5E02AD7DCF9B4FAB6F98D" box="[1235,1343,1555,1581]" italics="true" pageId="2" pageNumber="567">A. magna</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C304D38FFC5E02AD68BF9B4FF76F9F1" authorityName="Matson" authorityYear="2023" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="cooperae">
|
||
<emphasis id="B944EAA9FFC5E02AD68BF9B4FF76F9F1" italics="true" pageId="2" pageNumber="567">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C304D38FFC5E02AD239F990FE10F9F0" authorityName="Matson" authorityYear="2023" box="[310,409,1591,1617]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="velata">
|
||
<emphasis id="B944EAA9FFC5E02AD239F990FE10F9F0" box="[310,409,1591,1617]" italics="true" pageId="2" pageNumber="567">A. velata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC5E02DD3C8F9FDFE4AF871" lastPageId="5" lastPageNumber="570" pageId="2" pageNumber="567" type="description">
|
||
<paragraph id="8B8F36BBFFC5E02AD3C8F9FDFCD3F9D4" blockId="2.[151,1437,1122,2013]" box="[199,858,1626,1653]" pageId="2" pageNumber="567">
|
||
<emphasis id="B944EAA9FFC5E02AD3C8F9FDFE32F9D5" bold="true" box="[199,443,1626,1652]" pageId="2" pageNumber="567">Description. MALE.</emphasis>
|
||
Forewing length,
|
||
<quantity id="4CC89B5EFFC5E02AD187F9FCFC88F9D4" box="[648,769,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="2" pageNumber="567" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21–22 mm</quantity>
|
||
(n = 8).
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC5E02AD3C8F9D8FACAF8A4" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="567">
|
||
<emphasis id="B944EAA9FFC5E02AD3C8F9D8FE87F938" bold="true" box="[199,270,1663,1689]" pageId="2" pageNumber="567">Head.</emphasis>
|
||
Antenna mostly bipectinate, but with gradually diminishing rami that are absent in distal quarter of antenna; scales above fuscous, rami ochreous. Vertex mostly white with central gray spot; frons thinly scaled, white. Labial palpus decumbent, 1.5x diameter of eye, and with mixture of white and gray scales on outer surface and white on inner surface. Chaetosemata in transverse row; cephalic collar mostly white with few gray scales.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC5E02AD3C8F8A8FB59F8EC" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="567">
|
||
<emphasis id="B944EAA9FFC5E02AD3C8F8A8FEAEF888" bold="true" box="[199,295,1807,1833]" pageId="2" pageNumber="567">Thorax.</emphasis>
|
||
Patagium, tegula, and mesothorax admixture of gray and white scales. Legs mostly white and mottled with gray; epiphysis well-developed; hind tibia with large hair pencil; tibial spur formula 0–2–4.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC5E02AD3C8F8F1FC11F87C" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="567">
|
||
<emphasis id="B944EAA9FFC5E02AD3C8F8F1FEB6F8D1" bold="true" box="[199,319,1878,1904]" pageId="2" pageNumber="567">Forewing.</emphasis>
|
||
Pearly-white; widely scattered with inconspicuous light gray scales and with intermittent black scaling over parts of veins. Basal, costal, and terminal area of outer margin broadly maculated with brown scales; basal area, apical area, and tornus, more uniformly brown. Underside more uniform pearly-white and without brown basal scaling. Apical area with more diffuse brown scaling than upperside.
|
||
</paragraph>
|
||
<caption id="DF4F6633FFC4E02BD397F8E9FDDFF873" ID-DOI="http://doi.org/10.5281/zenodo.8426640" ID-Zenodo-Dep="8426640" httpUri="https://zenodo.org/record/8426640/files/figure.png" pageId="3" pageNumber="568" startId="3.[152,270,1870,1894]" targetBox="[156,1432,179,1844]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B8F36BBFFC4E02BD397F8E9FDDFF873" blockId="3.[151,1437,1870,2002]" pageId="3" pageNumber="568">
|
||
<emphasis id="B944EAA9FFC4E02BD397F8E9FECDF8C7" bold="true" box="[152,324,1870,1894]" pageId="3" pageNumber="568">FIGURES 1–4.</emphasis>
|
||
<taxonomicName id="4C304D38FFC4E02BD242F8E9FE2EF8C7" authorityName="Rindge" authorityYear="1983" box="[333,423,1870,1894]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC4E02BD242F8E9FE2EF8C7" box="[333,423,1870,1894]" italics="true" pageId="3" pageNumber="568">Achagua</emphasis>
|
||
</taxonomicName>
|
||
adult habitus. (a) dorsal view, (b) ventral view.
|
||
<emphasis id="B944EAA9FFC4E02BD0A8F8E9FC4CF8C7" bold="true" box="[935,965,1870,1894]" pageId="3" pageNumber="568">(1)</emphasis>
|
||
<taxonomicName id="4C304D38FFC4E02BD0C1F8E8FB7BF8C7" authority="Rindge, AMNH" authorityName="Rindge, AMNH" box="[974,1266,1870,1894]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC4E02BD0C1F8E8FBCDF8C7" box="[974,1092,1870,1894]" italics="true" pageId="3" pageNumber="568">A. obsoleta</emphasis>
|
||
Rindge, AMNH
|
||
</taxonomicName>
|
||
_IZC 00353009, paratype, (Colombia: Cundinamarca), AMNH;
|
||
<emphasis id="B944EAA9FFC4E02BD175F8D5FD1EF82B" bold="true" box="[634,663,1906,1930]" pageId="3" pageNumber="568">(2)</emphasis>
|
||
<taxonomicName id="4C304D38FFC4E02BD191F8D4FC89F82B" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[670,768,1907,1930]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="magna" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC4E02BD191F8D4FC89F82B" box="[670,768,1907,1930]" italics="true" pageId="3" pageNumber="568">A. magna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC4E02BD008F8D5FCC8F82B" bold="true" box="[775,833,1906,1930]" pageId="3" pageNumber="568">
|
||
<taxonomicNameLabel id="A27757D2FFC4E02BD008F8D5FCC8F82B" box="[775,833,1906,1930]" pageId="3" pageNumber="568" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, AMNH_IZC 00353021, holotype, (Peru: Cusco), AMNH;
|
||
<emphasis id="B944EAA9FFC4E02BD398F831FF3CF80F" bold="true" box="[151,181,1942,1966]" pageId="3" pageNumber="568">(3)</emphasis>
|
||
<taxonomicName id="4C304D38FFC4E02BD3B4F830FEBFF80F" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[187,310,1943,1966]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="cooperae" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC4E02BD3B4F830FEBFF80F" box="[187,310,1943,1966]" italics="true" pageId="3" pageNumber="568">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC4E02BD232F831FEFEF80F" bold="true" box="[317,375,1942,1966]" pageId="3" pageNumber="568">
|
||
<taxonomicNameLabel id="A27757D2FFC4E02BD232F831FEFEF80F" box="[317,375,1942,1966]" pageId="3" pageNumber="568" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, USNMENT01771270, holotype, (Costa Rica: Alajuela), USNM;
|
||
<emphasis id="B944EAA9FFC4E02BD713F831FBB3F80F" bold="true" box="[1052,1082,1942,1966]" pageId="3" pageNumber="568">(4)</emphasis>
|
||
<taxonomicName id="4C304D38FFC4E02BD74FF830FB12F80F" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[1088,1179,1942,1966]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="velata" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC4E02BD74FF830FB12F80F" box="[1088,1179,1942,1966]" italics="true" pageId="3" pageNumber="568">A. velata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC4E02BD7AEF831FB52F80F" bold="true" box="[1185,1243,1942,1966]" pageId="3" pageNumber="568">
|
||
<taxonomicNameLabel id="A27757D2FFC4E02BD7AEF831FB52F80F" box="[1185,1243,1942,1966]" pageId="3" pageNumber="568" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, (French Guiana: Régina), MNHN. Scale bar = 2 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6633FFC3E02CD397F8EAFE9DF871" ID-DOI="http://doi.org/10.5281/zenodo.8426642" ID-Zenodo-Dep="8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="4" pageNumber="569" startId="4.[152,270,1869,1893]" targetBox="[158,1432,190,1832]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8F36BBFFC3E02CD397F8EAFE9DF871" blockId="4.[151,1436,1869,2001]" pageId="4" pageNumber="569">
|
||
<emphasis id="B944EAA9FFC3E02CD397F8EAFEC9F8C4" bold="true" box="[152,320,1869,1893]" pageId="4" pageNumber="569">FIGURES 5–8.</emphasis>
|
||
<taxonomicName id="4C304D38FFC3E02CD249F8EAFE29F8C4" authorityName="Rindge" authorityYear="1983" box="[326,416,1869,1893]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC3E02CD249F8EAFE29F8C4" box="[326,416,1869,1893]" italics="true" pageId="4" pageNumber="569">Achagua</emphasis>
|
||
</taxonomicName>
|
||
male genitalia. (a) genital capsule, (b) phallus.
|
||
<emphasis id="B944EAA9FFC3E02CD070F8EAFC15F8C4" bold="true" box="[895,924,1869,1893]" pageId="4" pageNumber="569">(5)</emphasis>
|
||
<taxonomicName id="4C304D38FFC3E02CD0ADF8EAFBEFF8C4" authority="Rindge" authorityName="Rindge" box="[930,1126,1869,1893]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC3E02CD0ADF8EAFB9DF8C4" box="[930,1044,1869,1893]" italics="true" pageId="4" pageNumber="569">A. obsoleta</emphasis>
|
||
Rindge
|
||
</taxonomicName>
|
||
, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH;
|
||
<emphasis id="B944EAA9FFC3E02CD2C8F8D6FE6CF828" bold="true" box="[455,485,1905,1929]" pageId="4" pageNumber="569">(6)</emphasis>
|
||
<taxonomicName id="4C304D38FFC3E02CD2E4F8D6FDC4F828" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[491,589,1905,1929]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="magna" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC3E02CD2E4F8D6FDC4F828" box="[491,589,1905,1929]" italics="true" pageId="4" pageNumber="569">A. magna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC3E02CD15CF8D6FD04F828" bold="true" box="[595,653,1905,1929]" pageId="4" pageNumber="569">
|
||
<taxonomicNameLabel id="A27757D2FFC3E02CD15CF8D6FD04F828" box="[595,653,1905,1929]" pageId="4" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH;
|
||
<emphasis id="B944EAA9FFC3E02CD7F2F8D6FA92F828" bold="true" box="[1277,1307,1905,1929]" pageId="4" pageNumber="569">(7)</emphasis>
|
||
<taxonomicName id="4C304D38FFC3E02CD62DF8D6FA15F828" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[1314,1436,1905,1929]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="cooperae" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC3E02CD62DF8D6FA15F828" box="[1314,1436,1905,1929]" italics="true" pageId="4" pageNumber="569">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC3E02CD398F832FF46F80C" bold="true" box="[151,207,1941,1965]" pageId="4" pageNumber="569">
|
||
<taxonomicNameLabel id="A27757D2FFC3E02CD398F832FF46F80C" box="[151,207,1941,1965]" pageId="4" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, TAM-2023-251, AMNH_IZC 00353028, paratype, AMNH;
|
||
<emphasis id="B944EAA9FFC3E02CD036F832FCDFF80C" bold="true" box="[825,854,1941,1965]" pageId="4" pageNumber="569">(8)</emphasis>
|
||
<taxonomicName id="4C304D38FFC3E02CD054F832FC3DF80C" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[859,948,1941,1965]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="velata" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC3E02CD054F832FC3DF80C" box="[859,948,1941,1965]" italics="true" pageId="4" pageNumber="569">A. velata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D2FFC3E02CD0B5F832FC78F80D" box="[954,1009,1941,1965]" pageId="4" pageNumber="569" rank="species">
|
||
<emphasis id="B944EAA9FFC3E02CD0B5F832FC62F80C" bold="true" box="[954,1003,1941,1965]" pageId="4" pageNumber="569">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, TAM-2023-289, holotype, MNHN. Scale bar = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6633FFC2E02DD397FA3BFF4FFA5D" ID-DOI="http://doi.org/10.5281/zenodo.8426644" ID-Zenodo-Dep="8426644" httpUri="https://zenodo.org/record/8426644/files/figure.png" pageId="5" pageNumber="570" startId="5.[152,270,1436,1460]" targetBox="[256,1331,198,1417]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8F36BBFFC2E02DD397FA3BFF4FFA5D" blockId="5.[151,1437,1436,1532]" pageId="5" pageNumber="570">
|
||
<emphasis id="B944EAA9FFC2E02DD397FA3BFEC7FA15" bold="true" box="[152,334,1436,1460]" pageId="5" pageNumber="570">FIGURES 9–10.</emphasis>
|
||
<taxonomicName id="4C304D38FFC2E02DD25BFA3BFE27FA15" authorityName="Rindge" authorityYear="1983" box="[340,430,1436,1460]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="570" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC2E02DD25BFA3BFE27FA15" box="[340,430,1436,1460]" italics="true" pageId="5" pageNumber="570">Achagua</emphasis>
|
||
</taxonomicName>
|
||
female genitalia.
|
||
<emphasis id="B944EAA9FFC2E02DD16BFA3BFD0BFA15" bold="true" box="[612,642,1436,1460]" pageId="5" pageNumber="570">(9)</emphasis>
|
||
<taxonomicName id="4C304D38FFC2E02DD187FA3AFCF4FA15" authority="Rindge, No." authorityName="Rindge, No." box="[648,893,1436,1460]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="570" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC2E02DD187FA3AFD72FA15" box="[648,763,1436,1460]" italics="true" pageId="5" pageNumber="570">A. obsoleta</emphasis>
|
||
Rindge, No.
|
||
</taxonomicName>
|
||
20288, NHMUK010293435, NHMUK; photo credit: Louise Berridge (NHMUK);
|
||
<emphasis id="B944EAA9FFC2E02DD2CDFA67FE64FA79" bold="true" box="[450,493,1472,1496]" pageId="5" pageNumber="570">(10)</emphasis>
|
||
<taxonomicName id="4C304D38FFC2E02DD2FCFA66FDE6FA79" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[499,623,1473,1496]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="570" phylum="Arthropoda" rank="species" species="cooperae" status="n. sp.">
|
||
<emphasis id="B944EAA9FFC2E02DD2FCFA66FDE6FA79" box="[499,623,1473,1496]" italics="true" pageId="5" pageNumber="570">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAA9FFC2E02DD179FA67FD39FA79" bold="true" box="[630,688,1472,1496]" pageId="5" pageNumber="570">
|
||
<taxonomicNameLabel id="A27757D2FFC2E02DD179FA67FD39FA79" box="[630,688,1472,1496]" pageId="5" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, TAM-2023-276, USNMENT01771269, paratype, USNM. Scale bar = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F36BBFFC2E02DD3C8F98DFBACF9C9" blockId="5.[151,1436,1578,2000]" pageId="5" pageNumber="570">
|
||
<emphasis id="B944EAA9FFC2E02DD3C8F98DFECCF9E5" bold="true" box="[199,325,1578,1604]" pageId="5" pageNumber="570">Hindwing.</emphasis>
|
||
Pearly-white except for brown and black terminal band. Underside patterned similar to upperside, but darkened area around outer margin more diffuse and tornal area much whiter.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC2E02DD3C8F9D5FCA2F974" blockId="5.[151,1436,1578,2000]" pageId="5" pageNumber="570">
|
||
<emphasis id="B944EAA9FFC2E02DD3C8F9D5FECAF92D" bold="true" box="[199,323,1650,1676]" pageId="5" pageNumber="570">Abdomen.</emphasis>
|
||
First segment white with pair of black elongate dorsal spots, second segment black with narrowly triangular white median patch, remainder of abdomen with mixture of white, gray, black, and brown scales; white below. Third sternite of male abdomen with comb of setae.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC2E02DD3C8F979FD33F80D" blockId="5.[151,1436,1578,2000]" pageId="5" pageNumber="570">
|
||
<emphasis id="B944EAA9FFC2E02DD3C8F979FEBCF959" bold="true" box="[199,309,1758,1784]" pageId="5" pageNumber="570">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3EFFC2E02DD24AF979FE03F959" box="[325,394,1758,1784]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="5" pageNumber="570">Fig. 6</figureCitation>
|
||
). Uncus abruptly widened at base; dorsal process thin through middle and apically spatulate; ventral process thumb-like. Base of gnathos subquadrangular, with upcurved, heavily sclerotized, pointed apical projection; apex of projection lightly dentate. Valve large and elongate with heavily sclerotized costa bearing postmedial digitate protuberance. Anellar processes large and subtriangular with elongate, slender posterior projection; lateral margin only slightly concave. Juxta with elongate, medial cylindrical process. Vesica with large denticulate basal patch and weakly sclerotized medial patch.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC2E02DD3C8F811FE4AF871" blockId="5.[151,1436,1578,2000]" box="[199,451,1974,2000]" pageId="5" pageNumber="570">
|
||
<emphasis id="B944EAA9FFC2E02DD3C8F811FECFF871" bold="true" box="[199,326,1974,2000]" pageId="5" pageNumber="570">FEMALE.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC1E02ED398FF30FCFEFD95" pageId="6" pageNumber="571" type="materials_examined">
|
||
<paragraph id="8B8F36BBFFC1E02ED398FF30FED9FF10" blockId="6.[151,336,151,177]" box="[151,336,151,177]" pageId="6" pageNumber="571">
|
||
<heading id="D0C781D7FFC1E02ED398FF30FED9FF10" bold="true" box="[151,336,151,177]" fontSize="11" level="1" pageId="6" pageNumber="571" reason="1">
|
||
<emphasis id="B944EAA9FFC1E02ED398FF30FED9FF10" bold="true" box="[151,336,151,177]" pageId="6" pageNumber="571">
|
||
<typeStatus id="548B8819FFC1E02ED398FF30FF5DFF10" box="[151,212,151,177]" pageId="6" pageNumber="571">Type</typeStatus>
|
||
Material.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B583CE6FFC1E02ED398FF78FAA7FEBC" box="[151,1326,223,288]" collectingDate="1962-08" collectionCode="AMNH" collectorName="L. E. Pena" country="Peru" elevation="732" location="Quincemil" pageId="6" pageNumber="571" specimenCode="IZC 00353021" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
|
||
<paragraph id="8B8F36BBFFC1E02ED398FF78FE8DFF58" blockId="6.[151,1326,223,288]" box="[151,260,223,249]" pageId="6" pageNumber="571">
|
||
<heading id="D0C781D7FFC1E02ED398FF78FE8DFF58" bold="true" box="[151,260,223,249]" fontSize="11" level="1" pageId="6" pageNumber="571" reason="1">
|
||
<emphasis id="B944EAA9FFC1E02ED398FF78FE8DFF58" bold="true" box="[151,260,223,249]" pageId="6" pageNumber="571">
|
||
<typeStatus id="548B8819FFC1E02ED398FF78FE8DFF58" box="[151,260,223,249]" pageId="6" pageNumber="571">Holotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC1E02ED398FEA3FAA7FEBC" blockId="6.[151,1326,223,288]" box="[151,1326,259,288]" pageId="6" pageNumber="571">
|
||
<collectingCountry id="F327762BFFC1E02ED398FEA3FF56FEBF" box="[151,223,260,286]" name="Peru" pageId="6" pageNumber="571">PERU</collectingCountry>
|
||
• ♁;
|
||
<collectingRegion id="49F4F859FFC1E02ED215FEA4FEEAFEBC" box="[282,355,259,285]" country="Peru" name="Cusco" pageId="6" pageNumber="571">Cusco</collectingRegion>
|
||
,
|
||
<location id="8EEF6060FFC1E02ED261FEA4FE61FEBC" LSID="urn:lsid:plazi:treatment:039987ADFFC5E02ED398FC75FE75FB5B:8EEF6060FFC1E02ED261FEA4FE61FEBC" box="[366,488,259,285]" country="Peru" name="Quincemil" pageId="6" pageNumber="571" stateProvince="Cusco">Quincemil</location>
|
||
; elev.
|
||
<quantity id="4CC89B5EFFC1E02ED121FEA4FD08FEBC" box="[558,641,259,285]" metricMagnitude="2" metricUnit="m" metricValue="7.3152" pageId="6" pageNumber="571" unit="ft" value="2400.0">
|
||
<elevation id="001DD188FFC1E02ED121FEA4FD08FEBC" box="[558,641,259,285]" metricMagnitude="2" metricUnit="m" metricValue="7.3152" pageId="6" pageNumber="571" unit="ft" value="2400.0">2400 ft</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FF8E107BFFC1E02ED184FEA4FC8CFEBC" box="[651,773,259,285]" pageId="6" pageNumber="571" value="1962-08">
|
||
<collectingDate id="EFCAE993FFC1E02ED184FEA4FC8CFEBC" box="[651,773,259,285]" pageId="6" pageNumber="571" value="1962-08">Aug. 1962</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26C5536DFFC1E02ED01FFEA3FCF7FEBF" box="[784,894,260,286]" pageId="6" pageNumber="571">L.E. Pena</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED21AE7EFFC1E02ED0B3FEA4FB9BFEBC" box="[956,1042,259,285]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
_
|
||
<specimenCode id="DB969EC0FFC1E02ED72FFEA4FB4EFEBF" box="[1056,1223,259,286]" pageId="6" pageNumber="571">IZC 00353021</specimenCode>
|
||
;
|
||
<collectionCode id="ED21AE7EFFC1E02ED7DFFEA4FAA0FEBC" box="[1232,1321,259,285]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8B8F36BBFFC1E02ED398FEE5FEC2FEFF" blockId="6.[151,1437,321,565]" box="[151,331,321,350]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED398FEE5FEA7FEFA" bold="true" box="[151,302,321,348]" pageId="6" pageNumber="571">
|
||
<typeStatus id="548B8819FFC1E02ED398FEE5FE86FEFD" box="[151,271,322,348]" pageId="6" pageNumber="571">Paratypes</typeStatus>
|
||
(8
|
||
</emphasis>
|
||
♁)
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC1E02ED398FEC1FCFEFD95" blockId="6.[151,1437,321,565]" pageId="6" pageNumber="571">
|
||
<collectingCountry id="F327762BFFC1E02ED398FEC1FF56FE21" box="[151,223,358,384]" name="Peru" pageId="6" pageNumber="571">PERU</collectingCountry>
|
||
• 5♁; same collection data as
|
||
<typeStatus id="548B8819FFC1E02ED128FEC1FD04FE21" box="[551,653,358,384]" pageId="6" pageNumber="571">holotype</typeStatus>
|
||
; Genitalia: FHR no. 12173, TAM-2023-255, TAM-2023-268;
|
||
<collectionCode id="ED21AE7EFFC1E02ED636FEC1FA07FE21" box="[1337,1422,358,384]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
_ IZC 00353022–
|
||
<collectionCode id="ED21AE7EFFC1E02ED246FE2DFE16FE04" box="[329,415,394,421]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
_IZC 00353025, USNMENT01771249;
|
||
<collectionCode id="ED21AE7EFFC1E02ED050FE2DFC3EFE05" box="[863,951,394,420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
, USNM • ♁;
|
||
<collectingRegion id="49F4F859FFC1E02ED745FE2DFB1DFE05" box="[1098,1172,394,420]" country="Peru" name="Cusco" pageId="6" pageNumber="571">Cuzco</collectingRegion>
|
||
, Cosnipata; (-12.9018°, -71.4117°); elev.
|
||
<quantity id="4CC89B5EFFC1E02ED258FE09FE29FE69" box="[343,416,430,456]" metricMagnitude="2" metricUnit="m" metricValue="7.24" pageId="6" pageNumber="571" unit="m" value="724.0">724 m</quantity>
|
||
;
|
||
<date id="FF8E107BFFC1E02ED2A4FE09FDCCFE69" box="[427,581,430,456]" pageId="6" pageNumber="571" value="2016-08-28">28 Aug. 2016</date>
|
||
; Gunnar Brehm leg.; BOLD Process ID: PEMOA404-16; JPM •
|
||
<collectingCountry id="F327762BFFC1E02ED61AFE09FA15FE69" box="[1301,1436,430,456]" name="Ecuador" pageId="6" pageNumber="571">ECUADOR</collectingCountry>
|
||
• ♁;
|
||
<collectingRegion id="49F4F859FFC1E02ED3C2FE75FE2AFE4D" box="[205,419,466,492]" country="Ecuador" name="Zamora-Chinchipe" pageId="6" pageNumber="571">Zamora Chinchipe</collectingRegion>
|
||
; (-3.97388°, -79.084°); elev.
|
||
<quantity id="4CC89B5EFFC1E02ED1E4FE75FCCDFE4D" box="[747,836,466,493]" metricMagnitude="3" metricUnit="m" metricValue="1.925" pageId="6" pageNumber="571" unit="m" value="1925.0">1925 m</quantity>
|
||
;
|
||
<date id="FF8E107BFFC1E02ED05FFE75FC6EFE4D" box="[848,999,466,492]" pageId="6" pageNumber="571" value="2002-10-29">29 Oct. 2002</date>
|
||
; Nadine Hilt & Claudia Ramenda leg.; BOLD Process ID: NGEOE359-12; JPM •
|
||
<collectingCountry id="F327762BFFC1E02ED18EFE51FD7AFDB1" box="[641,755,502,528]" name="Bolivia" pageId="6" pageNumber="571">BOLIVIA</collectingCountry>
|
||
• ♁; N.P. Carrasco; elev.
|
||
<quantity id="4CC89B5EFFC1E02ED719FE51FBD3FDB1" box="[1046,1114,502,528]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="571" unit="m" value="900.0">900m</quantity>
|
||
; (
|
||
<geoCoordinate id="EE04507CFFC1E02ED760FE51FB7DFDB1" box="[1135,1268,502,528]" degrees="17" direction="south" minutes="06" orientation="latitude" pageId="6" pageNumber="571" precision="15" seconds="44" value="-17.112223">17°06’44”S</geoCoordinate>
|
||
<geoCoordinate id="EE04507CFFC1E02ED7F3FE51FA06FDB0" box="[1276,1423,502,529]" degrees="65" direction="west" minutes="33" orientation="longitude" pageId="6" pageNumber="571" precision="15" seconds="55" value="-65.56528">65°33’55”W</geoCoordinate>
|
||
);
|
||
<date id="FF8E107BFFC1E02ED398FDBDFEA4FD95" box="[151,301,538,565]" pageId="6" pageNumber="571" value="2010-10-05">05 Oct. 2010</date>
|
||
; Aare Lindt leg.; Genetic Voucher: TTBO002; TUZ.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC1E02ED3C8FDFFFBF1FD36" pageId="6" pageNumber="571" type="distribution">
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FDFFFBF1FD36" blockId="6.[151,1437,600,1275]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FDFFFED0FDD3" bold="true" box="[199,345,600,626]" pageId="6" pageNumber="571">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3EFFC1E02ED266FDFFFE30FDD2" box="[361,441,600,627]" captionStart="FIGURE 11" captionStartId="9.[152,255,1039,1063]" captionTargetBox="[151,1436,181,1015]" captionTargetId="figure-16@9.[151,1436,181,1015]" captionTargetPageId="9" captionText="FIGURE 11. Geographic distribution of Achagua: A. obsoleta Rindge (star), A. magna n. sp. (triangle), A. cooperae n. sp. (circle), and A. velata n. sp. (square). Single dots may represent>1 individual. Scale bar = 600 km." figureDoi="http://doi.org/10.5281/zenodo.8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="6" pageNumber="571">Fig. 11</figureCitation>
|
||
).
|
||
<taxonomicName id="4C304D38FFC1E02ED2DEFDFFFD01FDD3" authorityName="Matson" authorityYear="2023" box="[465,648,600,626]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC1E02ED2DEFDFFFD01FDD3" box="[465,648,600,626]" italics="true" pageId="6" pageNumber="571">Achagua magna</emphasis>
|
||
</taxonomicName>
|
||
inhabits the eastern slopes of the central Andes from the Eastern Cordillera real montane forests of
|
||
<collectingCountry id="F327762BFFC1E02ED2E1FDDBFDC2FD37" box="[494,587,636,662]" name="Ecuador" pageId="6" pageNumber="571">Ecuador</collectingCountry>
|
||
south through the Peruvian and Bolivian Yungas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC1E02ED3C8FD07FB38FD1B" box="[199,1201,672,699]" pageId="6" pageNumber="571" type="biology_ecology">
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FD07FB38FD1B" blockId="6.[151,1437,600,1275]" box="[199,1201,672,699]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FD07FEAEFD1B" bold="true" box="[199,295,672,698]" pageId="6" pageNumber="571">Biology.</emphasis>
|
||
Life history information is wanting. Adult records are from August and October.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC1E02ED3C8FD62FDA6FCA2" pageId="6" pageNumber="571" type="etymology">
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FD62FDA6FCA2" blockId="6.[151,1437,600,1275]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FD62FEC7FD7E" bold="true" box="[199,334,709,735]" pageId="6" pageNumber="571">Etymology.</emphasis>
|
||
The specific epithet
|
||
<emphasis id="B944EAA9FFC1E02ED14CFD62FD06FD7F" box="[579,655,709,734]" italics="true" pageId="6" pageNumber="571">magna</emphasis>
|
||
is from the Latin “magnus,” meaning “large,” as this species is the largest known member of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6530FFC1E02ED3C8FCAAFE75FB5B" pageId="6" pageNumber="571" type="discussion">
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FCAAFD74FC33" blockId="6.[151,1437,600,1275]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FCAAFD9AFC87" bold="true" box="[199,531,780,807]" pageId="6" pageNumber="571">Molecular characterization.</emphasis>
|
||
This species is represented in BOLD by the BIN: BOLD:ABW8871 (n =3,
|
||
<collectingCountry id="F327762BFFC1E02ED66DFCAAFA11FC86" box="[1378,1432,781,807]" name="Peru" pageId="6" pageNumber="571">Peru</collectingCountry>
|
||
,
|
||
<collectingCountry id="F327762BFFC1E02ED398FC97FF7FFCEB" box="[151,246,816,842]" name="Ecuador" pageId="6" pageNumber="571">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F327762BFFC1E02ED20FFC97FEDEFCEB" box="[256,343,816,842]" name="Bolivia" pageId="6" pageNumber="571">Bolivia</collectingCountry>
|
||
). Specimens from
|
||
<collectingCountry id="F327762BFFC1E02ED12AFC97FDFEFCEB" box="[549,631,816,842]" name="Bolivia" pageId="6" pageNumber="571">Bolivia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F327762BFFC1E02ED1A4FC96FD57FCEA" box="[683,734,817,843]" name="Peru" pageId="6" pageNumber="571">Peru</collectingCountry>
|
||
differ from the only sequenced Ecuadorian individual by about 1.2%. The distance to the nearest neighbor,
|
||
<taxonomicName id="4C304D38FFC1E02ED189FCF3FCD0FCCF" authorityName="Matson" authorityYear="2023" box="[646,857,852,878]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="cooperae">
|
||
<emphasis id="B944EAA9FFC1E02ED189FCF3FCD0FCCF" box="[646,857,852,878]" italics="true" pageId="6" pageNumber="571">Achagua cooperae</emphasis>
|
||
</taxonomicName>
|
||
is about 5.9%, however, the presumable sister species of
|
||
<taxonomicName id="4C304D38FFC1E02ED3E7FCDEFEDAFC33" authorityName="Matson" authorityYear="2023" box="[232,339,889,915]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC1E02ED3E7FCDEFF70FC32" box="[232,249,889,915]" italics="true" pageId="6" pageNumber="571">A</emphasis>
|
||
.
|
||
<emphasis id="B944EAA9FFC1E02ED208FCDEFEDAFC33" box="[263,339,889,914]" italics="true" pageId="6" pageNumber="571">magna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C304D38FFC1E02ED26EFCDEFE56FC33" box="[353,479,888,915]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC1E02ED26EFCDEFE56FC33" box="[353,479,888,915]" italics="true" pageId="6" pageNumber="571">A. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
, has not been sequenced.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FC3AFCCDFB83" blockId="6.[151,1437,600,1275]" pageId="6" pageNumber="571">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FC3AFEB2FC16" bold="true" box="[199,315,925,951]" pageId="6" pageNumber="571">Remarks.</emphasis>
|
||
Multi-locus molecular data for this species were used in the phylogenetic studies of
|
||
<bibRefCitation id="EFA14B4AFFC1E02ED7FFFC3BFEA9FC7A" author="Murillo-Ramos, Ld. C. & Brehm, G. & Sihvonen, P. & Hausmann, A. & Holm, S. & Ghanavi, H. R. & Ounap, E. & Truuverk, A. & Staude, H. & Friedrich, E. & Tammaru, T. & Wahlberg, N." pageId="6" pageNumber="571" pagination="7386" refId="ref6742" refString="Murillo-Ramos, Ld. C., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Ghanavi, H. R., Ounap, E., Truuverk, A., Staude, H., Friedrich, E., Tammaru, T. & Wahlberg, N. (2019) A comprehensive molecular phylogeny of Geometridae (Lepidoptera) with a focus on enigmatic small subfamilies. PeerJ, 7, e 7386. https: // doi. org / 10.7717 / peerj. 7386" type="journal article" year="2019">
|
||
Murillo-Ramos
|
||
<emphasis id="B944EAA9FFC1E02ED398FC66FF41FC7B" box="[151,200,960,986]" italics="true" pageId="6" pageNumber="571">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFA14B4AFFC1E02ED25AFC67FDBDFC7A" author="Brehm, G. & Murillo-Ramos, L. & Sihvonen, P. & Hausmann, A. & Schmidt, B. C. & Ounap, E. & Moser A. & Mortter, R. & Bolt, D. & Bodner, F. & Lindt, A. & Parra, L. E. & Wahlberg, N." box="[341,564,960,987]" pageId="6" pageNumber="571" pagination="457 - 486" refId="ref6406" refString="Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B. C., Ounap, E., Moser A., Mortter, R., Bolt, D., Bodner, F., Lindt, A., Parra, L. E. & Wahlberg, N. (2019) New World geometrid moths (Lepidoptera: Geometridae): Molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Systematics and Phylogeny, 77 (3), 457 - 486. https: // doi. org / 10.26049 / ASP 77 - 3 - 2019 - 5" type="journal article" year="2019">
|
||
Brehm
|
||
<emphasis id="B944EAA9FFC1E02ED2A7FC66FE50FC7B" box="[424,473,960,986]" italics="true" pageId="6" pageNumber="571">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
. However, the sequenced individual of
|
||
<taxonomicName id="4C304D38FFC1E02ED0E4FC67FBC6FC7B" authorityName="Rindge" authorityYear="1983" box="[1003,1103,960,986]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC1E02ED0E4FC67FBC6FC7B" box="[1003,1103,960,986]" italics="true" pageId="6" pageNumber="571">Achagua</emphasis>
|
||
</taxonomicName>
|
||
was misidentified as
|
||
<taxonomicName id="4C304D38FFC1E02ED630FC66FF48FC5F" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC1E02ED630FC66FF48FC5F" italics="true" pageId="6" pageNumber="571">A. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
in these studies. This is unsurprising considering the similarity between
|
||
<taxonomicName id="4C304D38FFC1E02ED0E1FC42FBD0FC5F" authorityName="Matson" authorityYear="2023" box="[1006,1113,997,1023]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC1E02ED0E1FC42FBD0FC5F" box="[1006,1113,997,1023]" italics="true" pageId="6" pageNumber="571">A. magna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C304D38FFC1E02ED781FC42FA85FC5F" box="[1166,1292,996,1023]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC1E02ED781FC42FA85FC5F" box="[1166,1292,996,1023]" italics="true" pageId="6" pageNumber="571">A. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
, and the fact that
|
||
<taxonomicName id="4C304D38FFC1E02ED3C7FBAEFECFFB83" box="[200,326,1032,1059]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC1E02ED3C7FBAEFECFFB83" box="[200,326,1032,1059]" italics="true" pageId="6" pageNumber="571">A. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
was the only described
|
||
<taxonomicName id="4C304D38FFC1E02ED15BFBAFFD31FB83" authorityName="Rindge" authorityYear="1983" box="[596,696,1032,1058]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC1E02ED15BFBAFFD31FB83" box="[596,696,1032,1058]" italics="true" pageId="6" pageNumber="571">Achagua</emphasis>
|
||
</taxonomicName>
|
||
at that time.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC1E02ED3C8FB8BFE75FB5B" blockId="6.[151,1437,600,1275]" pageId="6" pageNumber="571">
|
||
<taxonomicName id="4C304D38FFC1E02ED3C8FB8BFEF7FBE7" authorityName="Matson" authorityYear="2023" box="[199,382,1068,1094]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC1E02ED3C8FB8BFEF7FBE7" box="[199,382,1068,1094]" italics="true" pageId="6" pageNumber="571">Achagua magna</emphasis>
|
||
</taxonomicName>
|
||
is the name given to the undescribed species mentioned in
|
||
<bibRefCitation id="EFA14B4AFFC1E02ED710FB8BFB4FFBE6" author="Rindge, F. H." box="[1055,1222,1068,1095]" pageId="6" pageNumber="571" pagination="147 - 262" refId="ref7036" refString="Rindge, F. H. (1983) A generic revision of the New World Nacophorini (Lepidoptera: Geometridae). Bulletin of the American Museum of Natural History, 175, 147 - 262." type="journal article" year="1983">Rindge (1983)</bibRefCitation>
|
||
. While Rindge initially mentioned his series of
|
||
<taxonomicName id="4C304D38FFC1E02ED2D2FBF6FDCEFBCB" authorityName="Matson" authorityYear="2023" box="[477,583,1105,1131]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC1E02ED2D2FBF6FDCEFBCB" box="[477,583,1105,1131]" italics="true" pageId="6" pageNumber="571">A. magna</emphasis>
|
||
</taxonomicName>
|
||
as being from
|
||
<collectingCountry id="F327762BFFC1E02ED1E4FBF7FCC1FBCB" box="[747,840,1104,1130]" name="Ecuador" pageId="6" pageNumber="571">Ecuador</collectingCountry>
|
||
in the distribution section of his generic treatment, he later referred to it as being from
|
||
<collectingCountry id="F327762BFFC1E02ED10DFBD2FDBCFB2E" box="[514,565,1141,1167]" name="Peru" pageId="6" pageNumber="571">Peru</collectingCountry>
|
||
in the diagnosis and remarks sections of his species’ treatment of
|
||
<taxonomicName id="4C304D38FFC1E02ED618FBD2FA1CFB2F" box="[1303,1429,1140,1167]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B944EAA9FFC1E02ED618FBD2FA1CFB2F" box="[1303,1429,1140,1167]" italics="true" pageId="6" pageNumber="571">A. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
. After examining specimens that Rindge likely studied at the
|
||
<collectionCode id="ED21AE7EFFC1E02ED04EFB3EFC10FB12" box="[833,921,1177,1203]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="571">AMNH</collectionCode>
|
||
, where he curated
|
||
<taxonomicName id="4C304D38FFC1E02ED762FB3FFB7EFB13" authorityName="Linnaeus" authorityYear="1758" box="[1133,1271,1176,1202]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
|
||
, there were no specimens from
|
||
<collectingCountry id="F327762BFFC1E02ED25DFB1BFE38FB77" box="[338,433,1212,1238]" name="Ecuador" pageId="6" pageNumber="571">Ecuador</collectingCountry>
|
||
, only from
|
||
<collectingCountry id="F327762BFFC1E02ED13BFB1AFDE2FB76" box="[564,619,1213,1239]" name="Peru" pageId="6" pageNumber="571">Peru</collectingCountry>
|
||
. However, during this study, material from
|
||
<collectingCountry id="F327762BFFC1E02ED758FB1BFB3DFB77" box="[1111,1204,1212,1238]" name="Ecuador" pageId="6" pageNumber="571">Ecuador</collectingCountry>
|
||
was later confirmed from other museum collections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |