292 lines
36 KiB
XML
292 lines
36 KiB
XML
<document id="12CD6573F45372C80D480987AE7ADD2B" ID-DOI="10.11646/zootaxa.5352.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8407172" ID-ZooBank="AB121961-ECCA-4CE2-9190-14704DAE55F2" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1696428524420" checkinUser="plazi" docAuthor="Pinheiro, Mileudiane O. & Silveira, Orlando T." docDate="2023" docId="CD6C04370B0CFFEDE1B9FA887FCEF9E9" docLanguage="en" docName="zootaxa.5352.2.3.pdf" docOrigin="Zootaxa 5352 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mischocyttarus heliconius Richards 1941" docType="treatment" docVersion="3" lastPageNumber="207" masterDocId="31557C4F0B0FFFE9E12EFF957C20FF9C" masterDocTitle="Taxonomic notes on the group of Mischocyttarus heliconius Richards 1941, with description of four new species (Hymenoptera: Vespidae, Polistinae)" masterLastPageNumber="220" masterPageNumber="203" pageNumber="206" updateTime="1697029632406" updateUser="carolina" zenodo-license-document="CLOSED">
|
||
<mods:mods id="8A2989ECF6E980AD7DF0B0C1F97CAF34" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="07E40C788F0EC9881A6500B20790A330">
|
||
<mods:title id="B8F280B72F65F9C0ED0C6D06E8501B2F">Taxonomic notes on the group of Mischocyttarus heliconius Richards 1941, with description of four new species (Hymenoptera: Vespidae, Polistinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="884F338C642374D97D5776309C485308" type="personal">
|
||
<mods:role id="B0BAD2F41E22D7CF7BBA4EF61E872378">
|
||
<mods:roleTerm id="01ED97CFFF4CD96FF54B2FE8DEA8E66E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6889700351CD800A08FDFD54503156EA">Pinheiro, Mileudiane O.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="274835DC649E7C11ABA016A4199C6323" type="personal">
|
||
<mods:role id="B73C178F01411761C305603DBA6E4BE9">
|
||
<mods:roleTerm id="742DF0348FC89165C3FEDB51352C186F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="115379B0478EF04B0A1C6933671059FB">Silveira, Orlando T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="5200ED537C2147F15357AA203CCE771F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A901DF30AF58374F25074195E0D53A73" type="host">
|
||
<mods:titleInfo id="30BEDE65928FE56F62E12C9254C82D0B">
|
||
<mods:title id="E568AAEE8A8117550BD64079A533590F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="29B79F9F9A3F1055E4065B5D0BA36D24">
|
||
<mods:date id="660B4570D330CEB8CE2F48E8E0B4DDAD">2023</mods:date>
|
||
<mods:detail id="00F6E33D8930DEB4BA43E13F34B48CFB" type="pubDate">
|
||
<mods:number id="0775438F93226E1C5A08717E42EEB7F4">2023-10-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="9A68D826D19783463FFD0FB1B8C060D6" type="volume">
|
||
<mods:number id="9E14D94C0E5C3A08A7D51671F70A360C">5352</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="37ADFD300913B1D3EB4CA67C09A5FF48" type="issue">
|
||
<mods:number id="5C45E9CB2E24E846912CB6F80DDEEF52">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="139D920E06B313CCF7B7EF352629D544" unit="page">
|
||
<mods:start id="189B4569E9593206D6DD3F1AE69D0C16">203</mods:start>
|
||
<mods:end id="308A56D7D4D8663239E5ED9D5ACD33C1">220</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="E94EE702A71273C01B7C9EDF631E3359">
|
||
<mods:url id="6E2E8B3E13DF19BF8245A5E22CBCBFD8">http://dx.doi.org/10.11646/zootaxa.5352.2.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="C9EE4153D8A253E5D764CFE3F4B0A791">journal article</mods:classification>
|
||
<mods:identifier id="3C4E5E67AAAE8E5133A4B9CA1BD16E6C" type="DOI">10.11646/zootaxa.5352.2.3</mods:identifier>
|
||
<mods:identifier id="4038ED7F47E1A9498B25AA51CE885B72" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="7F15405FE204429BF2BBDDB928A7F44E" type="Zenodo-Dep">8407172</mods:identifier>
|
||
<mods:identifier id="35304BC2D6D01A59B56D2A3FCABF07B7" type="ZooBank">AB121961-ECCA-4CE2-9190-14704DAE55F2</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CD6C04370B0CFFEDE1B9FA887FCEF9E9" ID-DOI="http://doi.org/10.5281/zenodo.8407192" ID-Zenodo-Dep="8407192" LSID="urn:lsid:plazi:treatment:CD6C04370B0CFFEDE1B9FA887FCEF9E9" httpUri="http://treatment.plazi.org/id/CD6C04370B0CFFEDE1B9FA887FCEF9E9" lastPageId="4" lastPageNumber="207" pageId="3" pageNumber="206">
|
||
<subSubSection id="0DDFE6AA0B0CFFEAE1B9FA887EB9FAAB" box="[151,665,1309,1336]" pageId="3" pageNumber="206" type="nomenclature">
|
||
<paragraph id="457AB5210B0CFFEAE1B9FA887EB9FAAB" blockId="3.[151,665,1309,1336]" box="[151,665,1309,1336]" pageId="3" pageNumber="206">
|
||
<heading id="1E32024D0B0CFFEAE1B9FA887EB9FAAB" bold="true" box="[151,665,1309,1336]" fontSize="11" level="1" pageId="3" pageNumber="206" reason="1">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FA887EB9FAAB" ID-CoL="43MX9" authority="Richards, 1941" authorityName="Richards" authorityYear="1941" box="[151,665,1309,1336]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA887EB9FAAB" bold="true" box="[151,665,1309,1336]" pageId="3" pageNumber="206">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA887DF6FAA4" bold="true" box="[151,470,1309,1336]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
<bibRefCitation id="2154C8D00B0CFFEAE0F3FA887EB9FAAB" author="Richards, O. W." box="[477,665,1309,1335]" pageId="3" pageNumber="206" pagination="124 - 130" refId="ref13643" refString="Richards, O. W. (1941) The classification of the genus Mischocyttarus Saussure (Hymenopt., Vespidae). Proceedings of the Royal Entomological Society of London (B), 10, 124 - 130. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00705. x" type="journal article" year="1941">Richards, 1941</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DDFE6AA0B0CFFEAE1B9FAF77DACF945" pageId="3" pageNumber="206" type="reference_group">
|
||
<paragraph id="457AB5210B0CFFEAE1B9FAF77DCCFA09" blockId="3.[151,1437,1378,1644]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9FAF77DCCFA09" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9FAF77E7EFAE4" author="Richards, O. W." box="[151,606,1378,1400]" page="125" pageId="3" pageNumber="206" year="1941">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FAF77E7EFAE4" ID-CoL="43MX9" authority="Richards, 1941: 125" authorityName="Richards" authorityPageNumber="125" authorityYear="1941" box="[151,606,1378,1400]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9FAF77DB4FAEB" box="[151,404,1378,1399]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
<materialsCitation id="F5ADBF7C0B0CFFEAE0B4FAF77E7EFAE4" box="[410,606,1378,1400]" collectingDate="1929-09-20" collectionCode="R, NHM" country="Guyana" location="Moraballi Creek" pageId="3" pageNumber="206" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<bibRefCitation id="2154C8D00B0CFFEAE0B4FAF77E7EFAE4" author="Richards, O. W." box="[410,606,1378,1400]" pageId="3" pageNumber="206" pagination="124 - 130" refId="ref13643" refString="Richards, O. W. (1941) The classification of the genus Mischocyttarus Saussure (Hymenopt., Vespidae). Proceedings of the Royal Entomological Society of London (B), 10, 124 - 130. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00705. x" type="journal article" year="1941">Richards, 1941: 125</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 128.
|
||
<typeStatus id="9A7E0B830B0CFFEAE3B6FAF77ED5FAE4" box="[664,757,1378,1400]" pageId="3" pageNumber="206">Holotype</typeStatus>
|
||
:
|
||
<specimenCount id="53C37EA80B0CFFEAE3D0FAF67F30FAE4" box="[766,784,1379,1400]" pageId="3" pageNumber="206" type="female">♀</specimenCount>
|
||
,
|
||
<collectingCountry id="3DD2F5B10B0CFFEAE232FAF77F49FAEB" box="[796,873,1378,1399]" name="Guyana" pageId="3" pageNumber="206">Guyana</collectingCountry>
|
||
, Essequibo R.,
|
||
<location id="401AE3FA0B0CFFEAE2D0FAF778BEFAE4" LSID="urn:lsid:plazi:treatment:CD6C04370B0CFFEDE1B9FA887FCEF9E9:401AE3FA0B0CFFEAE2D0FAF778BEFAE4" box="[1022,1182,1378,1400]" country="Guyana" name="Moraballi Creek" pageId="3" pageNumber="206">Moraballi Creek</location>
|
||
,
|
||
<date id="317B93E10B0CFFEAE586FAF7792FFAE4" box="[1192,1295,1378,1400]" pageId="3" pageNumber="206" value="1929-09-20">
|
||
<collectingDate id="213F6A090B0CFFEAE586FAF7792FFAE4" box="[1192,1295,1378,1400]" pageId="3" pageNumber="206" value="1929-09-20">20.ix.1929</collectingDate>
|
||
</date>
|
||
(Oxford Univ. Exped.) (NHM), examined.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9FA097E95FA28" blockId="3.[151,1437,1378,1644]" box="[151,693,1436,1460]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9FA097E95FA28" box="[151,693,1436,1460]" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9FA097E5DFA28" author="Richards, O. W." box="[151,637,1436,1460]" page="311" pageId="3" pageNumber="206" year="1945">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FA097E5DFA28" ID-CoL="43MX9" authority=": Richards, 1945: 311" authorityName="Richards" authorityPageNumber="311" authorityYear="1945" box="[151,637,1436,1460]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA097D82FA28" box="[151,418,1436,1460]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
:
|
||
<bibRefCitation id="2154C8D00B0CFFEAE081FA097E5DFA28" author="Richards, O. W." box="[431,637,1436,1460]" pageId="3" pageNumber="206" pagination="295 - 462" refId="ref13703" refString="Richards, O. W. (1945) A revision of the genus Mischocyttarus de Saussure (Hymen., Vespidae). Transactions of the Royal Entomological Society London, 95, 295 - 462. https: // doi. org / 10.1111 / j. 1365 - 2311.1945. tb 00264. x" type="journal article" year="1945">Richards, 1945: 311</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 338.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9FA2E7F68FA4F" blockId="3.[151,1437,1378,1644]" box="[151,840,1467,1491]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9FA2E7F68FA4F" box="[151,840,1467,1491]" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9FA2E7F30FA4F" author="Richards, O. W." box="[151,784,1467,1491]" page="311" pageId="3" pageNumber="206" year="1945">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FA2E7F30FA4F" ID-CoL="43MX9" authority="var. heliconius Richards, 1945: 311" authorityName="var. heliconius Richards" authorityPageNumber="311" authorityYear="1945" box="[151,784,1467,1491]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="variety" species="heliconius" variety="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA2E7D82FA4F" box="[151,418,1467,1491]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
var.
|
||
<emphasis id="77B169330B0CFFEAE0FAFA2E7E1CFA4F" box="[468,572,1467,1491]" italics="true" pageId="3" pageNumber="206">heliconius</emphasis>
|
||
<bibRefCitation id="2154C8D00B0CFFEAE36CFA2E7F30FA4F" author="Richards, O. W." box="[578,784,1467,1491]" pageId="3" pageNumber="206" pagination="295 - 462" refId="ref13703" refString="Richards, O. W. (1945) A revision of the genus Mischocyttarus de Saussure (Hymen., Vespidae). Transactions of the Royal Entomological Society London, 95, 295 - 462. https: // doi. org / 10.1111 / j. 1365 - 2311.1945. tb 00264. x" type="journal article" year="1945">Richards, 1945: 311</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 339.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9FA4F7FD0FA6E" blockId="3.[151,1437,1378,1644]" box="[151,1008,1497,1522]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9FA4F7FD0FA6E" box="[151,1008,1497,1522]" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9FA4F7ED5FA6E" author="Richards, O. W." box="[151,757,1497,1522]" page="311" pageId="3" pageNumber="206" year="1945">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FA4F7ED5FA6E" authority="Richards, 1945: 311" authorityName="Richards" authorityPageNumber="311" authorityYear="1945" box="[151,757,1497,1522]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="variety" species="heliconius" variety="iperuae">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA4F7D82FA6E" box="[151,418,1498,1522]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
var.
|
||
<emphasis id="77B169330B0CFFEAE0FAFA4F7E01FA6D" box="[468,545,1498,1521]" italics="true" pageId="3" pageNumber="206">iperuae</emphasis>
|
||
<bibRefCitation id="2154C8D00B0CFFEAE306FA4F7ED5FA6E" author="Richards, O. W." box="[552,757,1497,1522]" pageId="3" pageNumber="206" pagination="295 - 462" refId="ref13703" refString="Richards, O. W. (1945) A revision of the genus Mischocyttarus de Saussure (Hymen., Vespidae). Transactions of the Royal Entomological Society London, 95, 295 - 462. https: // doi. org / 10.1111 / j. 1365 - 2311.1945. tb 00264. x" type="journal article" year="1945">Richards, 1945: 311</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 340 (misidentification).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9FA6D7E77F98C" blockId="3.[151,1437,1378,1644]" box="[151,599,1528,1552]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9FA6D7E77F98C" box="[151,599,1528,1552]" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9FA6D7E77F98C" author="Zikan, J. F." box="[151,599,1528,1552]" page="185" pageId="3" pageNumber="206" year="1949">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9FA6D7E77F98C" ID-CoL="43MX9" authority=": Zikan, 1949: 185" authorityName=": Zikan" authorityPageNumber="185" authorityYear="1949" box="[151,599,1528,1552]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9FA6D7D82F98C" box="[151,418,1528,1552]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
:
|
||
<bibRefCitation id="2154C8D00B0CFFEAE081FA6D7E77F98C" author="Zikan, J. F." box="[431,599,1528,1552]" pageId="3" pageNumber="206" pagination="1 - 125" refId="ref14221" refString="Zikan, J. F. (1949) O genero Mischocyttarus Saussure (Hymenoptera, Vespidae), com a descricao de 82 especies novas. Boletim do Parque Nacional do Itatiaia, 1, 1 - 125." type="journal article" year="1949">Zikán, 1949:185</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9F9827F46F9B3" blockId="3.[151,1437,1378,1644]" box="[151,870,1559,1583]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9F9827F46F9B3" box="[151,870,1559,1583]" pageId="3" pageNumber="206">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9F9827D82F9B3" ID-CoL="43MX9" authorityName=": Zikan" authorityYear="1949" box="[151,418,1559,1583]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9F9827D82F9B3" box="[151,418,1559,1583]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
</taxonomicName>
|
||
morph
|
||
<treatmentCitation id="C46493300B0CFFEAE0DFF9827F0FF9B3" author="Richards, O. W." box="[497,815,1559,1583]" page="282" pageId="3" pageNumber="206" year="1978">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE0DFF9827F0FF9B3" ID-CoL="43MX9" authority=": Richards 1978: 282" authorityName=": Richards" authorityPageNumber="282" authorityYear="1978" box="[497,815,1559,1583]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE0DFF9827E79F9B3" box="[497,601,1559,1583]" italics="true" pageId="3" pageNumber="206">heliconius</emphasis>
|
||
:
|
||
<bibRefCitation id="2154C8D00B0CFFEAE348F9827F0FF9B3" author="Richards, O. W." box="[614,815,1559,1583]" pageId="3" pageNumber="206" refId="ref13760" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards 1978: 282</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 284.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9F9A3783BF9D2" blockId="3.[151,1437,1378,1644]" box="[151,1051,1590,1614]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9F9A3783BF9D2" box="[151,1051,1590,1614]" pageId="3" pageNumber="206">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9F9A37D82F9D2" ID-CoL="43MX9" authorityName=": Richards" authorityYear="1978" box="[151,418,1590,1614]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9F9A37D82F9D2" box="[151,418,1590,1614]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
</taxonomicName>
|
||
morph.
|
||
<treatmentCitation id="C46493300B0CFFEAE0D9F9A37F01F9D2" author="Richards, O. W." box="[503,801,1590,1614]" page="282" pageId="3" pageNumber="206" year="1978">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE0D9F9A37F01F9D2" authority=": Richards, 1978: 282" authorityName=": Richards" authorityPageNumber="282" authorityYear="1978" box="[503,801,1590,1614]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="iperuae">
|
||
<emphasis id="77B169330B0CFFEAE0D9F9A37E64F9D1" box="[503,580,1590,1613]" italics="true" pageId="3" pageNumber="206">iperuae</emphasis>
|
||
:
|
||
<bibRefCitation id="2154C8D00B0CFFEAE37CF9A37F01F9D2" author="Richards, O. W." box="[594,801,1590,1614]" pageId="3" pageNumber="206" refId="ref13760" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978: 282</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 284 (misidentification).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9F9C17F8BF9F0" blockId="3.[151,1437,1378,1644]" box="[151,939,1620,1644]" pageId="3" pageNumber="206">
|
||
<treatmentCitationGroup id="65D5920F0B0CFFEAE1B9F9C17F8BF9F0" box="[151,939,1620,1644]" pageId="3" pageNumber="206">
|
||
<treatmentCitation id="C46493300B0CFFEAE1B9F9C17E53F9F0" author="Silveira, O. T." box="[151,627,1620,1644]" page="363" pageId="3" pageNumber="206" year="1998">
|
||
<taxonomicName id="82C5CEA20B0CFFEAE1B9F9C17E53F9F0" ID-CoL="43MX9" authority=": Silveira, 1998: 363" authorityName=": Silveira" authorityPageNumber="363" authorityYear="1998" box="[151,627,1620,1644]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0CFFEAE1B9F9C17D82F9F0" box="[151,418,1620,1644]" italics="true" pageId="3" pageNumber="206">Mischocyttarus heliconius</emphasis>
|
||
:
|
||
<bibRefCitation id="2154C8D00B0CFFEAE081F9C17E53F9F0" author="Silveira, O. T." box="[431,627,1620,1644]" pageId="3" pageNumber="206" pagination="359 - 367" refId="ref13792" refString="Silveira, O. T. (1998) Mischocyttarus (Mischocyttarus) aripuanaensis. A new social wasp from western-central Brazil, and redescription of Mischocyttarus lindigi Richards (Hym., Vespidae, Polistinae). Papeis Avulsos de Zoologia, Sao Paulo, 40, 359 - 367." type="journal article" year="1998">Silveira, 1998: 363</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 364; 2008: 541; 2010: 55, 56.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEAE1B9F90F7DACF945" blockId="3.[151,1437,1690,2041]" pageId="3" pageNumber="206">
|
||
The redescription below is based on examination of the
|
||
<typeStatus id="9A7E0B830B0CFFEAE20EF90F7FA2F928" box="[800,898,1690,1716]" pageId="3" pageNumber="206">holotype</typeStatus>
|
||
and previous published work (
|
||
<bibRefCitation id="2154C8D00B0CFFEAE5C6F90F79B8F929" author="Richards, O. W." box="[1256,1432,1690,1717]" pageId="3" pageNumber="206" pagination="295 - 462" refId="ref13703" refString="Richards, O. W. (1945) A revision of the genus Mischocyttarus de Saussure (Hymen., Vespidae). Transactions of the Royal Entomological Society London, 95, 295 - 462. https: // doi. org / 10.1111 / j. 1365 - 2311.1945. tb 00264. x" type="journal article" year="1945">Richards, 1945</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2154C8D00B0CFFEAE1B9F92A7CF2F945" author="Richards, O. W." box="[151,210,1727,1753]" pageId="3" pageNumber="206" refId="ref13760" refString="Richards, O. W. (1978) The social wasps of the Americas, excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">1978</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2154C8D00B0CFFEAE1F0F92B7DA0F945" author="Silveira, O. T." box="[222,384,1726,1753]" pageId="3" pageNumber="206" pagination="510 - 549" refId="ref13898" refString="Silveira, O. T. (2008) Phylogeny of wasps of the genus Mischocyttarus de Saussure (Hymenoptera, Vespidae, Polistinae). Revista Brasileira de Entomologia, 52, 510 - 549. https: // doi. org / 10.1590 / S 0085 - 56262008000400004" type="journal article" year="2008">Silveira, 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DDFE6AA0B0CFFEDE1E9F9777D08FBC5" lastPageId="4" lastPageNumber="207" pageId="3" pageNumber="206" type="diagnosis">
|
||
<paragraph id="457AB5210B0CFFEAE1E9F9777903F8F4" blockId="3.[151,1437,1690,2041]" pageId="3" pageNumber="206">
|
||
<emphasis id="77B169330B0CFFEAE1E9F9777D1AF960" bold="true" box="[199,314,1762,1788]" pageId="3" pageNumber="206">Diagnosis</emphasis>
|
||
. Ground color brown with yellow marks, metasoma distally black; clypeus narrow, with disk pale yellow, and with dense silvery bristles; posterior ocelli separated by distance less than or equal to one ocellar diameter; occipital carina hardly developed; pronotal carina angularly raised at center; metasomal tergum 1 and 2 (basally) light brown, distal third of 2 to 6, black; inner hind tarsal claw with tip rounded and expanded.
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0CFFEDE1E9F8E67EF2FEF9" blockId="3.[151,1437,1690,2041]" lastBlockId="4.[151,1437,151,1653]" lastPageId="4" lastPageNumber="207" pageId="3" pageNumber="206">FEMALE DESCRIPTION. Length of fore wing: 12.0–13.0 mm; head in frontal view slightly higher than wide (FHH/INTOW: 1.10); clypeus higher than wide (HCLP/WCLP: ca. 1.16), apex narrowly rounded; upper free portion of lateral margin relatively long and straight in its course to tentorial pit; this closer to the eye than to the antennal socket; malar space narrow; occiput with dorsal carina low, hardly developed; posterior ocelli separated by a distance less than or equal to one ocellar diameter; POL/OOL: 0.30; anterior margin of pronotum medially with lamella wide and reflexed, without a secondary margin immediately behind; pronotum with lateral fovea; pronotal carina continuous from side to side, not lobed on the humeral region, angularly elevated at center; mesoscutum as long as wide (LMS/WMS: 1.0), lateral margin adjacent to tegula with well-developed lamella; hind leg inner tarsal claw with tip rounded and expanded; first metasomal segment ca. 2.7 times wider at the apex than at the base. Fore wing moderately elongated (LDIS/HMP: ca. 2.50).</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FEFA7F70FE4D" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
<emphasis id="77B169330B0BFFEDE1E9FEFA7D11FE15" box="[199,305,367,393]" italics="true" pageId="4" pageNumber="207">Sculpture</emphasis>
|
||
. Integument surface granular; clypeus with small and sparse punctures, apical region reticulate; mandible with surface reticulate, shiny and with sparse large punctures; frons sculpture similar to clypeus; mesopleuron finely granulated with scattered small punctures.
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FE4E7E5FFDFD" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
<emphasis id="77B169330B0BFFEDE1E9FE4E7D08FE68" box="[199,296,475,500]" italics="true" pageId="4" pageNumber="207">Vestiture</emphasis>
|
||
. Body with whitish pubescence; mandible with rather larger and sparse bristles; surface of clypeus with short and dense silvery hairs; distal margin of the clypeus with a row of larger bristles; pronotum and mesopleuron with short and dense hairs, and some larger bristles; mesoscutum with short and dense hairs; metasoma with numerous short, obliquely oriented bristles.
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FDFE7979FC55" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
<emphasis id="77B169330B0BFFEDE1E9FDFE7D28FD19" box="[199,264,619,645]" italics="true" pageId="4" pageNumber="207">Color</emphasis>
|
||
. Brown with pale yellow marks; antenna light brown except dorsally dark brown, and the scape ventrally yellow; disk of clypeus pale yellow; mandible with same color as clypeus except for brown edges; band adjacent to the inner margin of the eye (from the upper free margin of clypeus up to the vertex), two small spots on the upper inner portion of the interantennary space adjacent to the antennal sockets, evanescent band adjacent to the outer margin of the eye, area associated to the pronotal carina, two narrow stripes on the mesoscutum and shorter ones adjoining tegulae, axillae, anterior band on scutellum, anterior band on metanotum, wide lungshape paired spots on propodeum, weak evanescent mark on the dorsal mesepisternal plate, mark on posterior region of the mesopleuron, mark on the apical portion of the tibiae, pale yellow; first metasomal segment (with narrow and inconspicuous yellow distal band) and anterior portion of the second, light brown dorsally and ventrally; posterior portion of second metasomal segment and segments 3–6, black dorsal and ventrally. Wings hyaline, with light brown venation.
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FC467DBBFC71" blockId="4.[151,1437,151,1653]" box="[199,411,979,1005]" pageId="4" pageNumber="207">MALE. Unknown.</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FC627D08FBC5" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
NEST. A small comb, with symmetrical development; color brown, made of irregular wood pieces; pedicel elongated, centric, and with varnished aspect (
|
||
<bibRefCitation id="2154C8D00B0BFFEDE385FB8E7F7CFBA9" author="Richards, O. W." box="[683,860,1050,1077]" pageId="4" pageNumber="207" pagination="295 - 462" refId="ref13703" refString="Richards, O. W. (1945) A revision of the genus Mischocyttarus de Saussure (Hymen., Vespidae). Transactions of the Royal Entomological Society London, 95, 295 - 462. https: // doi. org / 10.1111 / j. 1365 - 2311.1945. tb 00264. x" type="journal article" year="1945">Richards, 1945</bibRefCitation>
|
||
; Richards, unpublished manuscript; and examined photograph).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DDFE6AA0B0BFFEDE1E9FBF77E8DFA91" pageId="4" pageNumber="207" type="distribution">
|
||
<paragraph id="457AB5210B0BFFEDE1E9FBF77D20FB3D" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
<emphasis id="77B169330B0BFFEDE1E9FBF77D79FBE0" bold="true" box="[199,345,1122,1148]" pageId="4" pageNumber="207">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="3DD2F5B10B0BFFEDE047FBF67DC4FBE1" box="[361,484,1123,1149]" name="Guyana" pageId="4" pageNumber="207">GUYANA</collectingCountry>
|
||
: Essequibo R., Moraballi Creek; Potaro, R., trail from Tukeit to Kaieteur Savannah (
|
||
<figureCitation id="DDFEA9A40B0BFFEDE1B1FB127CD4FB3D" box="[159,244,1159,1185]" captionStart="FIGURE 27" captionStartId="16.[152,255,1308,1332]" captionTargetBox="[195,1393,181,1284]" captionTargetId="figure-19@16.[195,1393,181,1284]" captionTargetPageId="16" captionText="FIGURE 27. Partial map of South America with known distributions of the species of the Mischocyttarus heliconius species group." figureDoi="http://doi.org/10.5281/zenodo.8407184" httpUri="https://zenodo.org/record/8407184/files/figure.png" pageId="4" pageNumber="207">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FB3E7E8DFA91" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
Photographs of the
|
||
<typeStatus id="9A7E0B830B0BFFEDE08EFB3E7E22FB59" box="[416,514,1195,1221]" pageId="4" pageNumber="207">holotype</typeStatus>
|
||
can be examined through the following internet link (site of the NHM; accessed in
|
||
<date id="317B93E10B0BFFEDE1B9FB5A7DA1FB75" box="[151,385,1231,1257]" pageId="4" pageNumber="207" value="2022-09-27">September 27, 2022</date>
|
||
): https://data.nhm.ac.uk/dataset/56e711e6-c847-4f99-915a-6894bb5c5dea/resource/05ff2255- c38a-40c9-b657-4ccb55ab2feb/record/9133906
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DDFE6AA0B0BFFEDE1E9FA827FCEF9E9" pageId="4" pageNumber="207" type="discussion">
|
||
<paragraph id="457AB5210B0BFFEDE1E9FA827F2DFAE5" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
<emphasis id="77B169330B0BFFEDE1E9FA827D14FAAD" bold="true" box="[199,308,1303,1329]" pageId="4" pageNumber="207">Remarks</emphasis>
|
||
. The
|
||
<typeStatus id="9A7E0B830B0BFFEDE05CFA827DF4FAAD" box="[370,468,1303,1329]" pageId="4" pageNumber="207">holotype</typeStatus>
|
||
was previously examined by O. T. S., and recently by photographs. The nest associated with the
|
||
<typeStatus id="9A7E0B830B0BFFEDE1D7FAAE7D7BFAC9" box="[249,347,1339,1365]" pageId="4" pageNumber="207">holotype</typeStatus>
|
||
described by Richards in 1945 was collected on the abaxial surface of a leaf of
|
||
<taxonomicName id="82C5CEA20B0BFFEDE5FBFAAE794BFAC9" box="[1237,1387,1339,1365]" class="Insecta" family="Nymphalidae" genus="Heliconia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="77B169330B0BFFEDE5FBFAAE7963FAC9" box="[1237,1347,1339,1365]" italics="true" pageId="4" pageNumber="207">Heliconia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
The species is hitherto only known from the type specimens.
|
||
</paragraph>
|
||
<paragraph id="457AB5210B0BFFEDE1E9FA167FCEF9E9" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="207">
|
||
Richards’ descriptions (1941, 1945, 1978) emphasized coloration rather than structural characteristics. He also mentioned that the metasoma is testaceous, but it is in fact mostly black. Richards’ works recognized two varieties (or morphs) of
|
||
<taxonomicName id="82C5CEA20B0BFFEDE067FA5E7DC4FA79" authorityName=": Silveira" authorityYear="1998" box="[329,484,1483,1509]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0BFFEDE067FA5E7DC4FA79" box="[329,484,1483,1509]" italics="true" pageId="4" pageNumber="207">M. heliconius</emphasis>
|
||
</taxonomicName>
|
||
(“
|
||
<taxonomicName id="82C5CEA20B0BFFEDE32FFA5E7E89FA79" authorityName="var. heliconius Richards" authorityYear="1945" box="[513,681,1483,1509]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="variety" species="heliconius" variety="heliconius">
|
||
var.
|
||
<emphasis id="77B169330B0BFFEDE318FA5E7E89FA79" box="[566,681,1483,1509]" italics="true" pageId="4" pageNumber="207">heliconius</emphasis>
|
||
</taxonomicName>
|
||
” and “
|
||
<taxonomicName id="82C5CEA20B0BFFEDE3D2FA5E7FA6FA78" authorityName="Richards" authorityYear="1945" box="[764,902,1483,1509]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="variety" species="heliconius" variety="iperuae">
|
||
var.
|
||
<emphasis id="77B169330B0BFFEDE21EFA5E7FA6FA78" box="[816,902,1483,1508]" italics="true" pageId="4" pageNumber="207">iperuae</emphasis>
|
||
</taxonomicName>
|
||
”), but in the 1978 book he already said about
|
||
<taxonomicName id="82C5CEA20B0BFFEDE1B9FA7A7CCDF994" authorityName=": Richards" authorityYear="1978" box="[151,237,1519,1544]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="iperuae">
|
||
<emphasis id="77B169330B0BFFEDE1B9FA7A7CCDF994" box="[151,237,1519,1544]" italics="true" pageId="4" pageNumber="207">iperuae</emphasis>
|
||
</taxonomicName>
|
||
that “
|
||
<emphasis id="77B169330B0BFFEDE018FA7A7F0BF994" box="[310,811,1519,1545]" italics="true" pageId="4" pageNumber="207">This form might really be a distinct species</emphasis>
|
||
…”.
|
||
<bibRefCitation id="2154C8D00B0BFFEDE240FA7A7839F995" author="Silveira, O. T." box="[878,1049,1519,1545]" pageId="4" pageNumber="207" pagination="55 - 59" refId="ref13948" refString="Silveira, O. T. (2010) On Richards's concept of Mischocyttarus undulatus (Ducke), with the description of a new species of the group of M. iheringi Zikan (Hymenoptera, Vespidae). Studies on Neotropical Fauna and Environment, 45, 55 - 59. https: // doi. org / 10.1080 / 01650520903500666" type="journal article" year="2010">Silveira (2010)</bibRefCitation>
|
||
confirmed the conditions of both
|
||
<taxonomicName id="82C5CEA20B0BFFEDE1B9F9867D2AF9B1" authorityName=": Silveira" authorityYear="1998" box="[151,266,1555,1581]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0BFFEDE1B9F9867D2AF9B1" box="[151,266,1555,1581]" italics="true" pageId="4" pageNumber="207">heliconius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82C5CEA20B0BFFEDE06AF9867DBAF9B0" authorityName=": Richards" authorityYear="1978" box="[324,410,1555,1580]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="iperuae">
|
||
<emphasis id="77B169330B0BFFEDE06AF9867DBAF9B0" box="[324,410,1555,1580]" italics="true" pageId="4" pageNumber="207">iperuae</emphasis>
|
||
</taxonomicName>
|
||
as distinct specific taxa.
|
||
<taxonomicName id="82C5CEA20B0BFFEDE392F9867F76F9B1" authorityName=": Silveira" authorityYear="1998" box="[700,854,1555,1581]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="heliconius">
|
||
<emphasis id="77B169330B0BFFEDE392F9867F76F9B1" box="[700,854,1555,1581]" italics="true" pageId="4" pageNumber="207">M. heliconius</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="82C5CEA20B0BFFEDE2DEF986784DF9B0" authorityName=": Richards" authorityYear="1978" box="[1008,1133,1555,1580]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="iperuae">
|
||
<emphasis id="77B169330B0BFFEDE2DEF986784DF9B0" box="[1008,1133,1555,1580]" italics="true" pageId="4" pageNumber="207">M. iperuae</emphasis>
|
||
</taxonomicName>
|
||
by the clypeus higher and narrower, covered by dense silvery bristles, the posterior ocelli closer to each other (separated by less than one ocellar diameter), and the inner hind tarsal claw being expanded at the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |