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<document id="7B2C60A2A0424D40CEE58ADB22AA9EB6" ID-DOI="10.15298/rusentj.31.2.05" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179662" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596660692" checkinUser="felipe" docAuthor="Kabak, I. I. &amp; Schmidt, J." docDate="2022" docId="C16DC544FA26FF98FF26FCB98332CE09" docLanguage="en" docName="RussEntomolJ.31.2.124-127.pdf" docOrigin="Russian Entomological Journal 31 (2)" docSource="http://dx.doi.org/10.15298/rusentj.31.2.05" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Cymindis dolpoensis Kabak et Schmidt 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="127" masterDocId="3D54BD3CFA27FF9BFFE5FFB48227CC05" masterDocTitle="A new species of the genus Cymindis Latreille, 1805 from Nepal (Coleoptera: Carabidae: Lebiini)" masterLastPageNumber="127" masterPageNumber="124" pageNumber="125" updateTime="1722637701445" updateUser="ExternalLinkService">
<mods:mods id="66C9DD486C595C036296593281B301F1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C406687D4852E1790E6CA27AFA48025">
<mods:title id="FD99C76661AB809F30B9330E0F103A87">A new species of the genus Cymindis Latreille, 1805 from Nepal (Coleoptera: Carabidae: Lebiini)</mods:title>
</mods:titleInfo>
<mods:name id="C469A53E7605901BB9AAED262FF8A2FD" type="personal">
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</mods:role>
<mods:namePart id="2823C3D113C8387A5FD52A168F39ED59">Kabak, I. I.</mods:namePart>
</mods:name>
<mods:name id="83A22DDDB27DAA187E84E99D19DD2990" type="personal">
<mods:role id="7FC322C5EF6470E0B9EC3DB49B42FE26">
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</mods:role>
<mods:namePart id="572963A23A0FFB9AE8D0FCF5E779AB5A">Schmidt, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F98742FB77317BD16E442BA7A4A7DFD3">text</mods:typeOfResource>
<mods:relatedItem id="4F582695B4A6BE98DA9567CAC9D9B0D6" type="host">
<mods:titleInfo id="9156E90DBC88A7B5B7333838946A9F7F">
<mods:title id="1646DC901FB902113F377312AC135457">Russian Entomological Journal</mods:title>
</mods:titleInfo>
<mods:part id="2E294498063B7397564128226B935485">
<mods:date id="AC40D2C075C80C05B96E0CA194672186">2022</mods:date>
<mods:detail id="EBA69B25DBD55242E1777981AACD24A4" type="pubDate">
<mods:number id="028F58046A5FE8D939106470DDA41061">2022-06-30</mods:number>
</mods:detail>
<mods:detail id="888C5F6EA1237FACBF0BBB348BD19393" type="volume">
<mods:number id="CC5B614E2F20CE05E7EE300C2F4C52F3">31</mods:number>
</mods:detail>
<mods:detail id="9619DF20328DBD8B96E3D547B3F91F44" type="issue">
<mods:number id="9AE25201BFA0C3D35241DD3306DF4767">2</mods:number>
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<mods:start id="9DDCCD39353C8337B38DE8A563444B65">124</mods:start>
<mods:end id="88866AE0CCFD1DD9679A63E0E5586A43">127</mods:end>
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<mods:location id="A13AB511E3BFB4BC910ECE2D63586DCE">
<mods:url id="89DF5D1D663FBDE59C8E07B3F7B4ECA1">http://dx.doi.org/10.15298/rusentj.31.2.05</mods:url>
</mods:location>
<mods:classification id="2118BCD53FFEFD790AB246D7E8A73BC7">journal article</mods:classification>
<mods:identifier id="A30464C9097A2ED850141C39BD28CB38" type="DOI">10.15298/rusentj.31.2.05</mods:identifier>
<mods:identifier id="8CFF332ED148872C8B83CA658F3D2612" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="DBC20D6444B4D64E8CC57FD248797127" type="Zenodo-Dep">13179662</mods:identifier>
<mods:identifier id="135FE93BAEC3F05AC2A988EA8BAB3BC6" type="ZooBank">2E6B1F66-3A95-46DF-AEBF-D49AC94DA438</mods:identifier>
</mods:mods>
<treatment id="C16DC544FA26FF98FF26FCB98332CE09" LSID="urn:lsid:plazi:treatment:C16DC544FA26FF98FF26FCB98332CE09" httpUri="http://treatment.plazi.org/id/C16DC544FA26FF98FF26FCB98332CE09" lastPageId="3" lastPageNumber="127" pageId="1" pageNumber="125">
<subSubSection id="01DE27D9FA26FF9AFF26FCB98127CF20" box="[195,768,781,805]" pageId="1" pageNumber="125" type="nomenclature">
<paragraph id="497B7452FA26FF9AFF26FCB98127CF20" blockId="1.[189,775,744,2053]" box="[195,768,781,805]" pageId="1" pageNumber="125">
<heading id="1233C33EFA26FF9AFF26FCB98127CF20" box="[195,768,781,805]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="125" reason="4">
<taxonomicName id="8EC40FD1FA26FF9AFF26FCB98098CF20" authority="Kabak et Schmidt" authorityName="Kabak et Schmidt" authorityYear="2022" box="[195,703,781,805]" class="Insecta" family="Carabidae" genus="Cymindis" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="125" phylum="Arthropoda" rank="species" species="dolpoensis" status="sp. nov.">
<emphasis id="7BB0A840FA26FF9AFF26FCB9830FCF20" box="[195,296,781,805]" italics="true" pageId="1" pageNumber="125">Cymindis</emphasis>
(s. str.)
<emphasis id="7BB0A840FA26FF9AFE67FCB983D3CF20" box="[386,500,781,805]" italics="true" pageId="1" pageNumber="125">dolpoensis</emphasis>
Kabak et Schmidt
</taxonomicName>
,
<emphasis id="7BB0A840FA26FF9AFD2FFCB98127CF20" bold="true" box="[714,768,781,805]" pageId="1" pageNumber="125">
<taxonomicNameLabel id="6083153BFA26FF9AFD2FFCB98127CF20" box="[714,768,781,805]" pageId="1" pageNumber="125" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA26FF9AFE4AFC9F8033CF46" box="[431,532,811,835]" pageId="1" pageNumber="125" type="description">
<paragraph id="497B7452FA26FF9AFE4AFC9F8033CF46" blockId="1.[189,775,744,2053]" box="[431,532,811,835]" pageId="1" pageNumber="125">
<figureCitation id="D1FF68D7FA26FF9AFE4AFC9F8037CF46" box="[431,528,811,835]" captionStart="Figs 1–2" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[191,1394,331,1971]" captionTargetId="figure-57@2.[188,1398,330,1984]" captionTargetPageId="2" captionText="Figs 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — habitus of ♂, paratype; 2 — aedeagal median lobe of the holotype, lateral view. Рис. 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — обЩий вид, ♂, паратип; 2 — медиальнаЯ долЯ Эдеагуса голотипа, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179666" httpUri="https://zenodo.org/record/13179666/files/figure.png" pageId="1" pageNumber="125">Figs 1–2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA26FF9AFF07FCE4803ECFDE" pageId="1" pageNumber="125" type="materials_examined">
<paragraph id="497B7452FA26FF9AFF07FCE4803ECFDE" blockId="1.[189,775,744,2053]" pageId="1" pageNumber="125">
<materialsCitation id="F9AC7E0FFA26FF9AFF07FCE48032CFDE" collectingDate="1970-06" collectingDateMax="2007-10-08" collectingDateMin="1970-06" collectionCode="V, SMNS, ZIN" collectorName="S. Tamang" country="Nepal" elevation="3700" latitude="29.208334" location="lake Phoksumdo" longLatPrecision="20" longitude="82.95833" municipality="Phoksundo" pageId="1" pageNumber="125" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
MATERIAL.
<typeStatus id="967FCAF0FA26FF9AFEB8FCE4838ACF66" box="[349,429,848,867]" pageId="1" pageNumber="125" type="holotype">Holotype</typeStatus>
<specimenCount id="5FC2BFDBFA26FF9AFE51FCF983EFCF66" box="[436,456,845,867]" count="1" pageId="1" pageNumber="125" type="male">♂</specimenCount>
, “
<collectingCountry id="31D334C2FA26FF9AFE3EFCE48035CF66" box="[475,530,848,867]" name="Nepal" pageId="1" pageNumber="125">Nepal</collectingCountry>
, Dolpo, Ringmo, Phoksumdo-See,
<collectionCode id="2FD5EC97FA26FF9AFEEEFCDC833BCF7E" box="[267,284,872,891]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="125" type="Museum">V</collectionCode>
/
<date id="3D7A5292FA26FF9AFEC6FCDC834BCF7E" box="[291,364,872,891]" pageId="1" pageNumber="125" value="1970-06">
<collectingDate id="2D3EAB7AFA26FF9AFEC6FCDC834BCF7E" box="[291,364,872,891]" pageId="1" pageNumber="125" value="1970-06">VI.1970</collectingDate>
</date>
,
<quantity id="8E3CD9B7FA26FF9AFE9EFCDC83DACF7E" box="[379,509,872,891]" metricMagnitude="3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="1" pageNumber="125" unit="m" value="3700.0" valueMax="3800.0" valueMin="3600.0">
<elevation id="C2E99361FA26FF9AFE9EFCDC83DACF7E" box="[379,509,872,891]" metricMagnitude="3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="1" pageNumber="125" unit="m" value="3700.0" valueMax="3800.0" valueMin="3600.0">3600–3800 m</elevation>
</quantity>
, Nepal-Expeditionen Jochen Martens” [
<location id="4C1B2289FA26FF9AFEF2FC348380CF96" LSID="urn:lsid:plazi:treatment:C16DC544FA26FF98FF26FCB98332CE09:4C1B2289FA26FF9AFEF2FC348380CF96" box="[279,423,896,915]" country="Nepal" latitude="29.208334" longLatPrecision="20" longitude="82.95833" municipality="Phoksundo" name="lake Phoksumdo" pageId="1" pageNumber="125">lake Phoksumdo</location>
or
<collectingMunicipality id="A91FEE28FA26FF9AFE26FC348001CF96" box="[451,550,896,915]" pageId="1" pageNumber="125">Phoksundo</collectingMunicipality>
: ca.
<quantity id="8E3CD9B7FA26FF9AFDAEFC3480A9CF96" box="[587,654,896,915]" metricMagnitude="3" metricUnit="m" metricValue="3.6119999999999997" pageId="1" pageNumber="125" unit="m" value="3612.0">
<elevation id="C2E99361FA26FF9AFDAEFC3480A9CF96" box="[587,654,896,915]" metricMagnitude="3" metricUnit="m" metricValue="3.6119999999999997" pageId="1" pageNumber="125" unit="m" value="3612.0">3612 m</elevation>
</quantity>
,
<geoCoordinate id="2CF01295FA26FF9AFD73FC348124CF96" box="[662,771,896,915]" degrees="29" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="125" precision="15" seconds="30" value="29.208334">N29°12´30´´</geoCoordinate>
,
<geoCoordinate id="2CF01295FA26FF9AFF58FC238300CFAF" box="[189,295,919,938]" degrees="82" direction="east" minutes="57" orientation="longitude" pageId="1" pageNumber="125" precision="15" seconds="30" value="82.95833">E82°57´30´´</geoCoordinate>
] (
<collectionCode id="2FD5EC97FA26FF9AFED2FC2C8351CFAE" box="[311,374,920,939]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="125">SMNS</collectionCode>
).
<typeStatus id="967FCAF0FA26FF9AFE60FC2C83FBCFAE" box="[389,476,920,939]" pageId="1" pageNumber="125" type="paratype">Paratypes</typeStatus>
: 2(2)
<specimenCount id="5FC2BFDBFA26FF9AFDE8FC218015CFAE" box="[525,562,917,939]" count="1" pageId="1" pageNumber="125" type="male">♂♂</specimenCount>
, collected with holotype (cJS,
<collectionCode id="2FD5EC97FA26FF9AFF0AFC048330CFC6" box="[239,279,944,963]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="125" type="Museum">ZIN</collectionCode>
);
<specimenCount id="5FC2BFDBFA26FF9AFECDFC048375CFC6" box="[296,338,941,963]" count="2" pageId="1" pageNumber="125" type="female">2 ♀♀</specimenCount>
, “
<collectingCountry id="31D334C2FA26FF9AFE82FC0483B9CFC6" box="[359,414,944,963]" name="Nepal" pageId="1" pageNumber="125">Nepal</collectingCountry>
, Dolpo,
<date id="3D7A5292FA26FF9AFE09FC048019CFC6" box="[492,574,944,963]" pageId="1" pageNumber="125" value="2007-10-08">
<collectingDate id="2D3EAB7AFA26FF9AFE09FC048019CFC6" box="[492,574,944,963]" pageId="1" pageNumber="125" value="2007-10-08">8.X.2007</collectingDate>
</date>
, Ringmo, Phoksumdo,
<quantity id="8E3CD9B7FA26FF9AFF58FC738326CFDE" box="[189,257,967,987]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="125" unit="m" value="3500.0">
<elevation id="C2E99361FA26FF9AFF58FC738326CFDE" box="[189,257,967,987]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="125" unit="m" value="3500.0">3500 m</elevation>
</quantity>
, leg.
<collectorName id="E4311184FA26FF9AFED4FC7C83B6CFDE" box="[305,401,968,987]" pageId="1" pageNumber="125">S. Tamang</collectorName>
” (cJS,
<collectionCode id="2FD5EC97FA26FF9AFE34FC7C8028CFDE" box="[465,527,968,987]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="125">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA26FF9AFF07FC5481E7CB22" pageId="1" pageNumber="125" type="description">
<paragraph id="497B7452FA26FF9AFF07FC54834CC8CE" blockId="1.[189,775,744,2053]" pageId="1" pageNumber="125">
DESCRIPTION. Small-sized species, body length 6.8– 7.3 (7.1) mm, hind wings reduced, habitus rather narrow, subparallel–subconvex, appendages averaged (
<figureCitation id="D1FF68D7FA26FF9AFD62FBA180E5C82F" box="[647,706,1045,1067]" captionStart="Figs 1–2" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[191,1394,331,1971]" captionTargetId="figure-57@2.[188,1398,330,1984]" captionTargetPageId="2" captionText="Figs 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — habitus of ♂, paratype; 2 — aedeagal median lobe of the holotype, lateral view. Рис. 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — обЩий вид, ♂, паратип; 2 — медиальнаЯ долЯ Эдеагуса голотипа, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179666" httpUri="https://zenodo.org/record/13179666/files/figure.png" pageId="1" pageNumber="125">Fig. 1</figureCitation>
). Dorsum reddish-brown with somewhat paler clypeus, mouthparts, margins and basal part of pronotum, suture, lateral margins and base of elytra. Legs and antennae monochromatic brownish-yellow. Ventral side of body rufous medially, brown laterally. Lower surface sparsely punctate, pubescence feebly developed.
</paragraph>
<paragraph id="497B7452FA26FF9AFF07FB7B83E4CAE5" blockId="1.[189,775,744,2053]" pageId="1" pageNumber="125">
Head medium-sized, PW/HW = 1.22–1.26 (1.24); eyes long, moderately protruding, glabrous, YL/3AL = 1.29– 1.33 (1.3) in males,
<quantity id="8E3CD9B7FA26FF9AFE62FAB18022C91E" box="[391,517,1285,1307]" metricMagnitude="-2" metricUnit="m" metricValue="3.0860999999999996" metricValueMax="3.1242" metricValueMin="3.048" pageId="1" pageNumber="125" unit="in" value="1.2149999999999999" valueMax="1.23" valueMin="1.2">1.20–1.23 in</quantity>
females; tempora shorter than eyes, slightly convex, with several hairs. Anterolateral margins of forehead moderately curved, slightly reflexed. Upper-side of head convex, frontal foveae moderately deep, supraorbital furrow impressed, medium in length. Frons and vertex glabrous, sparsely and gently punctate, transversely wrinkled medially. Two pairs of long supraorbital setae. Antennae comparatively short, in males longer, EL/AL = 1.11–1.13 (1.12)
<emphasis id="7BB0A840FA26FF9AFD86FA6F8050C9F5" box="[611,631,1499,1520]" italics="true" pageId="1" pageNumber="125">vs</emphasis>
.
<quantity id="8E3CD9B7FA26FF9AFD63FA6E8121C9F5" box="[646,774,1498,1520]" metricMagnitude="-2" metricUnit="m" metricValue="3.0099" metricValueMax="3.0225999999999997" metricValueMin="2.9972" pageId="1" pageNumber="125" unit="in" value="1.185" valueMax="1.19" valueMin="1.18">1.18–1.19 in</quantity>
females. Scape average in length, subcylindric, with several small hairs and a long preapical seta. Genae with a few hairs. Labial tooth shorter than lateral lobes, rounded at apex, bordered near mid-length, with a pair of small setae near base; submentum quadrisetose. Ultimate segment of labial palpi in male stout, not dilated toward apex, with a few small hairs. Penultimate labial palpomere with long setae on anterior margin. Apical maxillary palpomere fusiform, sparsely pubescent.
</paragraph>
<paragraph id="497B7452FA26FF9AFF07F951869CCE44" blockId="1.[189,775,744,2053]" lastBlockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">Pronotum moderately wide, PW/PL = 1.20–1.28 (1.23), broadest in anterior third. Sides rounded in anterior half, subrectilinear or bisinuate in basal half, incision before laterobasal angles long, not deep, the latter obtusangular, rounded or blunt at apex, feebly protruded laterally. Anterior margin subrectilinear medially, anterior angles widely rounded, moderately produced anteriorly, their margins ciliate. Basal margin of pronotum moderately wide, PW/PB = 1.36–1.42 (1.40), subconvex and slightly lobed medially, obliquely curved latarally, its border interrupted near basal foveae. Lateral sides of pronotum moderately explanate, margins distinctly reflexed.Disc convex medially, median line deeply impressed, slightly shortened anteriorly and posteriorly. Apical transverse impression moderately deep. Basal foveae small and sharp, separated from lateral flattening. Basal transverse impression deep, vaguely outlined, interrupted medially. Punctures rather rough and dense along anterior, posterior and lateral margins, becoming fainter, sparser and irregular on disc. Surface of pronotum glabrous, distinctly transversely wrinkled. Two pairs of lateral setae, one in anterior half of pronotum, and one in laterobasal angles.</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FDF281B7C803" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">Elytra moderately long, oval, convex, flattened on disc, broadest near posterior third, EL/EW = 1.40–1.41 (1.40), EL/ PL = 2.73–2.85 (2.80), EW/PW = 1.59–1.67 (1.62). Lateral margins subrectilinear medially, shoulders rounded, protrud- ed anteriorly, ciliate. Apices oblique, straight in middle part, glabrous or very faintly ciliate near sutural angle; both exterior and sutural apical angles of each elytron broadly round- ed. Marginal gutter moderate in width, narrowed near humeri, lateral margins reflexed throughout. Basal border complete, moderately sinuate. Elytral striae deep, very faintly punctate. Both parascutellary striole and parascutellary setiferous pores present. Intervals evenly convex, irregularly punctate, punctures arranged mostly in 1 row; interval 3 with 3–4 small discal setiferous pores, attached to stria 3. Elytral surface glabrous. Umbilicate series consisting of 12–14 pores. One apical pore in stria 7 on the level either of stria 3 or of interval 3.</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FBBF8610C839" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">Scutellum glabrous and smooth. Sides of prothorax glabrous. Metacoxae bisetose.</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FBF58187C889" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">Microsculpture (males and females) on dorsum indistinct, only on scutellum traces of isodiametric meshes hardly perceptible.</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FB258189C8F2" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
Visible abdominal sternites with a single pair of paramedian setae, anal ventrite bisetose in males, variable in the two available females: in
<specimenCount id="5FC2BFDBFA26FF9AFBEFFB7286B4C8D9" box="[1034,1171,1222,1244]" count="1" pageId="1" pageNumber="125" type="generic">one specimen</specimenCount>
bisetose, in the other quadrisetose.
</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FB4F8665C929" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">Dorsal surface of tarsi with long and sparse hairs; inner margin of claws denticulate.</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FA8581C1C993" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
Median lobe of aedeagus rather thin, slightly curved, apical lamella long and wide, with apical hook on dorsal margin; copulatory piece wide, its anterior incision moderately deep (
<figureCitation id="D1FF68D7FA26FF9AFC7BFA3581FDC993" box="[926,986,1409,1431]" captionStart="Figs 1–2" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[191,1394,331,1971]" captionTargetId="figure-57@2.[188,1398,330,1984]" captionTargetPageId="2" captionText="Figs 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — habitus of ♂, paratype; 2 — aedeagal median lobe of the holotype, lateral view. Рис. 1–2. Cymindis (s. str.) dolpoensis sp.n.: 1 — обЩий вид, ♂, паратип; 2 — медиальнаЯ долЯ Эдеагуса голотипа, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179666" httpUri="https://zenodo.org/record/13179666/files/figure.png" pageId="1" pageNumber="125">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="497B7452FA26FF9AFCB7FA2F81E7CB22" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
COMPARATIVE NOTES. Among all
<taxonomicName id="8EC40FD1FA26FF9AFB2AFA2F8751C9B4" box="[1231,1398,1435,1457]" pageId="1" pageNumber="125">
<emphasis id="7BB0A840FA26FF9AFB2AFA2F870DC9B5" box="[1231,1322,1435,1456]" italics="true" pageId="1" pageNumber="125">Cymindis</emphasis>
species
</taxonomicName>
known from Himalaya, the new species seems to be most closely related to
<taxonomicName id="8EC40FD1FA26FF9AFC30FA658733C9E3" authority="Andrewes, 1935" authorityName="Andrewes" authorityYear="1935" box="[981,1300,1488,1511]" class="Insecta" family="Carabidae" genus="Cymindis" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="125" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="7BB0A840FA26FF9AFC30FA6581C2C9E3" box="[981,997,1489,1510]" italics="true" pageId="1" pageNumber="125">C</emphasis>
. (s. str.)
<emphasis id="7BB0A840FA26FF9AFBDCFA658655C9E3" box="[1081,1138,1489,1510]" italics="true" pageId="1" pageNumber="125">nitens</emphasis>
<bibRefCitation id="2D5509A3FA26FF9AFB92FA658733C9E3" author="Andrewes H. E." box="[1143,1300,1488,1511]" pageId="1" pageNumber="125" refId="ref2222" refString="Andrewes H. E. 1935. Cymindis (Lebiinae, Carab., Col.) // Visser C., Visser-Hooft J. (eds.). Wissenschaftliche Ergebnisse der niederlandischen Expeditionen in den Karakorum und die angrenzenden Gebiete, 1922, 1925 und 1929 / 30. Zoologie. Leipzig: F. A. Brockhaus. S. 279." type="journal volume" year="1935">Andrewes, 1935</bibRefCitation>
</taxonomicName>
described from Sringar in Kashmir [
<bibRefCitation id="2D5509A3FA26FF9AFBDCFA5F86C4CA04" author="Andrewes H. E." box="[1081,1251,1515,1537]" pageId="1" pageNumber="125" refId="ref2222" refString="Andrewes H. E. 1935. Cymindis (Lebiinae, Carab., Col.) // Visser C., Visser-Hooft J. (eds.). Wissenschaftliche Ergebnisse der niederlandischen Expeditionen in den Karakorum und die angrenzenden Gebiete, 1922, 1925 und 1929 / 30. Zoologie. Leipzig: F. A. Brockhaus. S. 279." type="journal volume" year="1935">Andrewes, 1935</bibRefCitation>
]. Both species share the similar size and proportions of body and gestalt of male genitalia: median lobe of aedegus rather narrow, slightly curved, its apical lamella wide, hooked apically. The new species easily differs from
<taxonomicName id="8EC40FD1FA26FF9AFBCEF9E28658CA69" authorityName="Andrewes" authorityYear="1935" box="[1067,1151,1622,1644]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="125" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="7BB0A840FA26FF9AFBCEF9E2861CCA6E" box="[1067,1083,1622,1643]" italics="true" pageId="1" pageNumber="125">C</emphasis>
.
<emphasis id="7BB0A840FA26FF9AFBA3F9E38658CA69" box="[1094,1151,1623,1644]" italics="true" pageId="1" pageNumber="125">nitens</emphasis>
</taxonomicName>
by lacking of hind wings, almost monochromous body coloration without lightened spots on elytra, not dilated apically ultimate labial palpomere in male, less protruded humeri, reduced pubescence of dorsum. Additionally, the medial lobe of aedeagus in
<taxonomicName id="8EC40FD1FA26FF9AFACAF9758153CAF4" authorityName="Kabak &amp; Schmidt" authorityYear="2022" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="125" phylum="Arthropoda" rank="species" species="dolpoensis" status="sp.n.">
<emphasis id="7BB0A840FA26FF9AFACAF9758718CAD3" box="[1327,1343,1729,1750]" italics="true" pageId="1" pageNumber="125">C</emphasis>
.
<emphasis id="7BB0A840FA26FF9AFAAAF9758153CAF4" italics="true" pageId="1" pageNumber="125">dolpoensis</emphasis>
</taxonomicName>
<emphasis id="7BB0A840FA26FF9AFC98F96F8189CAF5" bold="true" box="[893,942,1755,1776]" pageId="1" pageNumber="125">
<taxonomicNameLabel id="6083153BFA26FF9AFC98F96F8189CAF5" box="[893,942,1755,1776]" pageId="1" pageNumber="125" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is thinner, less curved, its apical lamella distinctly longer, copulatory piece wider, its anterior margin less deeply incised.
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA26FF9AFCB7F89F8777CBC2" pageId="1" pageNumber="125" type="distribution">
<paragraph id="497B7452FA26FF9AFCB7F89F818DCB94" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
<materialsCitation id="F9AC7E0FFA26FF9AFCB7F89F8181CB94" country="Nepal" county="Himalaya" location="Phoksumdo Lake" municipality="Dolpo District" pageId="1" pageNumber="125" specimenCount="1" stateProvince="Karnali">
DISTRIBUTION. The new species is only known from the
<typeStatus id="967FCAF0FA26FF9AFCB1F8F28159CB5E" box="[852,894,1862,1883]" pageId="1" pageNumber="125">type</typeStatus>
locality: mountains slopes surrounding the
<location id="4C1B2289FA26FF9AFAD8F8F28182CB72" LSID="urn:lsid:plazi:treatment:C16DC544FA26FF98FF26FCB98332CE09:4C1B2289FA26FF9AFAD8F8F28182CB72" country="Nepal" county="Himalaya" municipality="Dolpo District" name="Phoksumdo Lake" pageId="1" pageNumber="125" stateProvince="Karnali">Phoksumdo Lake</location>
,
<collectingMunicipality id="A91FEE28FA26FF9AFC4BF8D5861CCB72" box="[942,1083,1889,1911]" pageId="1" pageNumber="125">Dolpo District</collectingMunicipality>
,
<collectingRegion id="8B00BAB0FA26FF9AFBA1F8D586CDCB72" box="[1092,1258,1889,1911]" country="Nepal" name="Karnali" pageId="1" pageNumber="125">Karnali Province</collectingRegion>
,
<collectingCountry id="31D334C2FA26FF9AFB16F8D58708CB72" box="[1267,1327,1889,1911]" name="Nepal" pageId="1" pageNumber="125">Nepal</collectingCountry>
, Great- er
<collectingCounty id="A01A0CDEFA26FF9AFCA0F8CF8181CB94" box="[837,934,1915,1937]" pageId="1" pageNumber="125">Himalaya</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="497B7452FA26FF9AFCB7F8228777CBC2" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
BIONOMICS. Poorly noted.
<taxonomicName id="8EC40FD1FA26FF9AFB96F822871ACBAE" authorityName="Kabak &amp; Schmidt" authorityYear="2022" box="[1139,1341,1942,1963]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="125" phylum="Arthropoda" rank="species" species="dolpoensis" status="sp. nov.">
<emphasis id="7BB0A840FA26FF9AFB96F822871ACBAE" box="[1139,1341,1942,1963]" italics="true" pageId="1" pageNumber="125">Cymindis dolpoensis</emphasis>
</taxonomicName>
<emphasis id="7BB0A840FA26FF9AFAA3F8228751CBAE" bold="true" box="[1350,1398,1942,1963]" pageId="1" pageNumber="125">
<taxonomicNameLabel id="6083153BFA26FF9AFAA3F8228751CBAE" box="[1350,1398,1942,1963]" pageId="1" pageNumber="125" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
was found at elevations between 3500 and
<quantity id="8E3CD9B7FA26FF9AFB37F805873BCBC3" box="[1234,1308,1969,1990]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="125" unit="m" value="3800.0">3800 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA26FF98FCB7F87F8082CDD2" lastPageId="3" lastPageNumber="127" pageId="1" pageNumber="125" type="etymology">
<paragraph id="497B7452FA26FF9AFCB7F87F8743CBF9" blockId="1.[812,1399,315,2044]" pageId="1" pageNumber="125">
<materialsCitation id="F9AC7E0FFA26FF9AFCB7F87F8778CBF9" country="Nepal" location="Dolpo region" municipality="Himalaya" pageId="1" pageNumber="125" specimenCount="1">
ETYMOLOGY. The new species is named after its
<typeStatus id="967FCAF0FA26FF9AFAA8F8788750CBE4" box="[1357,1399,1996,2017]" pageId="1" pageNumber="125">type</typeStatus>
locality, the
<location id="4C1B2289FA26FF9AFC43F8528600CBF9" LSID="urn:lsid:plazi:treatment:C16DC544FA26FF98FF26FCB98332CE09:4C1B2289FA26FF9AFC43F8528600CBF9" box="[934,1063,2022,2044]" country="Nepal" municipality="Himalaya" name="Dolpo region" pageId="1" pageNumber="125">Dolpo region</location>
in the western
<collectingCountry id="31D334C2FA26FF9AFB5BF85286DFCBF9" box="[1214,1272,2022,2044]" name="Nepal" pageId="1" pageNumber="125">Nepal</collectingCountry>
<collectingMunicipality id="A91FEE28FA26FF9AFB1BF8528778CBF9" box="[1278,1375,2022,2044]" pageId="1" pageNumber="125">Himalaya</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<caption id="1DBB24DAFA25FF99FF07F8558761C415" ID-DOI="http://doi.org/10.5281/zenodo.13179666" ID-Zenodo-Dep="13179666" httpUri="https://zenodo.org/record/13179666/files/figure.png" pageId="2" pageNumber="126" startId="2.[226,263,2017,2036]" targetBox="[191,1394,331,1971]" targetPageId="2" targetType="figure">
<paragraph id="497B7452FA25FF99FF07F8558761C415" blockId="2.[226,1378,2015,2064]" pageId="2" pageNumber="126">
Figs 1–2.
<taxonomicName id="8EC40FD1FA25FF99FEDFF8558014CBF1" authorityName="Kabak &amp; Schmidt" authorityYear="2022" box="[314,563,2017,2036]" class="Insecta" family="Carabidae" genus="Cymindis" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="dolpoensis" status="sp. nov.">
<emphasis id="7BB0A840FA25FF99FEDFF85583ACCBF1" box="[314,395,2017,2036]" italics="true" pageId="2" pageNumber="126">Cymindis</emphasis>
(s. str.)
<emphasis id="7BB0A840FA25FF99FE32F8558014CBF1" box="[471,563,2017,2036]" italics="true" pageId="2" pageNumber="126">dolpoensis</emphasis>
</taxonomicName>
<emphasis id="7BB0A840FA25FF99FDDCF8558043CBF1" bold="true" box="[569,612,2017,2036]" pageId="2" pageNumber="126">
<taxonomicNameLabel id="6083153BFA25FF99FDDCF8558043CBF1" box="[569,612,2017,2036]" pageId="2" pageNumber="126" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1 — habitus of ♂, paratype; 2 — aedeagal median lobe of the holotype, lateral view. Рис. 1–2.
<taxonomicName id="8EC40FD1FA25FF99FED9F8498012C415" authorityName="Kabak &amp; Schmidt" authorityYear="2022" box="[316,565,2045,2064]" class="Insecta" family="Carabidae" genus="Cymindis" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="dolpoensis" status="sp. nov.">
<emphasis id="7BB0A840FA25FF99FED9F84983AAC415" box="[316,397,2045,2064]" italics="true" pageId="2" pageNumber="126">Cymindis</emphasis>
(s. str.)
<emphasis id="7BB0A840FA25FF99FE3CF8498012C415" box="[473,565,2045,2064]" italics="true" pageId="2" pageNumber="126">dolpoensis</emphasis>
</taxonomicName>
<emphasis id="7BB0A840FA25FF99FDDEF8498041C415" bold="true" box="[571,614,2045,2064]" pageId="2" pageNumber="126">
<taxonomicNameLabel id="6083153BFA25FF99FDDEF8498041C415" box="[571,614,2045,2064]" pageId="2" pageNumber="126" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 1 — обЩий вид, ♂, паратип; 2 — медиальнаЯ долЯ Эдеагуса голотипа, сбоку.
</paragraph>
</caption>
<paragraph id="497B7452FA24FF98FF07FE8F8082CDD2" blockId="3.[189,775,315,524]" pageId="3" pageNumber="127">
<emphasis id="7BB0A840FA24FF98FF07FE8F8382CD54" bold="true" box="[226,421,315,337]" pageId="3" pageNumber="127">Acknowledgments.</emphasis>
<materialsCitation id="F9AC7E0FFA24FF98FE4AFE888086CDD2" collectionCode="SMNS" country="Russia" location="Genrik Davidian" municipality="St-Petersburg" pageId="3" pageNumber="127" specimenCount="1">
We are very grateful to Wolfgang Schawaller (
<collectionCode id="2FD5EC97FA24FF98FED1FEE2835DCD6E" box="[308,378,342,363]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="127">SMNS</collectionCode>
), for the loan of carabid material from the Himalaya for study, Ben Brugge (Zoölogisch Museum Amsterdam, the Netherlands) who loaned us the
<typeStatus id="967FCAF0FA24FF98FD9FFE388083CDA4" box="[634,676,396,417]" pageId="3" pageNumber="127">type</typeStatus>
specimen of
<taxonomicName id="8EC40FD1FA24FF98FF3CFE128351CDB9" authorityName="Andrewes" authorityYear="1935" box="[217,374,422,444]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="7BB0A840FA24FF98FF3CFE128351CDB9" box="[217,374,422,444]" italics="true" pageId="3" pageNumber="127">Cymindis nitens</emphasis>
</taxonomicName>
, and to
<location id="4C1B2289FA24FF98FE2CFE12804ACDB9" LSID="urn:lsid:plazi:treatment:C16DC544FA26FF98FF26FCB98332CE09:4C1B2289FA24FF98FE2CFE12804ACDB9" box="[457,621,422,444]" country="Russia" municipality="St-Petersburg" name="Genrik Davidian" pageId="3" pageNumber="127">Genrik Davidian</location>
(
<collectingMunicipality id="A91FEE28FA24FF98FD9EFE128124CDB9" box="[635,771,422,444]" pageId="3" pageNumber="127">St-Petersburg</collectingMunicipality>
,
<collectingCountry id="31D334C2FA24FF98FF58FE7582D8CDD2" box="[189,255,449,471]" name="Russia" pageId="3" pageNumber="127">Russia</collectingCountry>
) for his help in preparing the photographs
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01DE27D9FA24FF98FF07FE6F8332CE09" pageId="3" pageNumber="127" type="description">
<paragraph id="497B7452FA24FF98FF07FE6F8332CE09" blockId="3.[189,775,315,524]" pageId="3" pageNumber="127">
<emphasis id="7BB0A840FA24FF98FF07FE6F8390CDF4" bold="true" box="[226,439,475,497]" pageId="3" pageNumber="127">Competing interests.</emphasis>
The authors declare no competing interests.
</paragraph>
</subSubSection>
</treatment>
</document>
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