504 lines
53 KiB
XML
504 lines
53 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5521714" ID-GBIF-Dataset="ae9f6659-8606-4ba6-8967-e48abd5b147f" ID-GBIF-Taxon="159871754" ID-Zenodo-Dep="5521714" checkinTime="1559292594248" checkinUser="jeremy" docAuthor="Pérez Gutiérrez, L. A" docDate="2012" docId="B25387F3FFCDFFCAFED68648FE18683A" docLanguage="en" docName="Pérez-Gutiérrez - 2012 - iArchilestes chocoanusi, a new damselfly from Colombia (Odonata Lestidae).pdf" docOrigin="Odonatologica 41 (4)" docStyle="DocumentStyle{}" docTitle="Archilestes chocoanus Gutiérrez, 2012, SP. NOV." docType="treatment" docVersion="12" lastPageId="5" lastPageNumber="351" masterDocId="4E6AFF8BFFCCFFCFFFB18436FFB3690F" masterDocTitle="Archilestes chocoanus, a new damselfly from Colombia (Odonata: Lestidae)" masterLastPageNumber="352" masterPageNumber="347" pageId="1" pageNumber="348" updateTime="1632325217131" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Archilestes chocoanus, a new damselfly from Colombia (Odonata: Lestidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pérez Gutiérrez, L. A</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>41</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>347</mods:start>
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<mods:end>352</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3236059</mods:identifier>
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<mods:identifier type="GBIF-Dataset">ae9f6659-8606-4ba6-8967-e48abd5b147f</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3236059</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5521714" ID-GBIF-Taxon="159871754" ID-Zenodo-Dep="5521714" LSID="urn:lsid:plazi:treatment:B25387F3FFCDFFCAFED68648FE18683A" httpUri="http://treatment.plazi.org/id/B25387F3FFCDFFCAFED68648FE18683A" lastPageId="5" lastPageNumber="351" pageId="1" pageNumber="348">
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<subSubSection box="[359,825,638,662]" pageId="1" pageNumber="348" type="nomenclature">
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<paragraph blockId="1.[359,825,638,692]" box="[359,825,638,662]" pageId="1" pageNumber="348">
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<heading allCaps="true" box="[359,825,638,662]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="348" reason="1">
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<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[359,709,638,661]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="chocoanus" status="spec. nov.">
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<emphasis box="[359,709,638,661]" italics="true" pageId="1" pageNumber="348">ARHILESTES CHOCOANUS</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[722,825,638,662]" pageId="1" pageNumber="348" rank="species">SP. NOV.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[542,641,673,692]" pageId="1" pageNumber="348" type="description">
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||
<paragraph blockId="1.[359,825,638,692]" box="[542,641,673,692]" pageId="1" pageNumber="348">
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||
<figureCitation box="[542,641,673,692]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[514,547,812,831]" captionStartId-1="3.[155,187,1002,1021]" captionStartId-2="4.[514,547,753,772]" captionTargetBox-1="[155,647,219,961]" captionTargetId-0="figure@2.[514,1006,219,780]" captionTargetId-1="figure@3.[155,647,219,961]" captionTargetId-2="figure@4.[514,1006,219,723]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." captionText-1="Fig. 2. Archilestes chocoanus sp.n., male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view." captionText-2="Fig. 3. Fig. 3. Archilestes chocoanus sp. n. [Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view." httpUri-0="https://zenodo.org/record/3236061/files/figure.png" httpUri-1="https://zenodo.org/record/3236063/files/figure.png" httpUri-2="https://zenodo.org/record/3339507/files/figure.png" pageId="1" pageNumber="348">Figures 1-3</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="1" pageNumber="348" type="materials_examined">
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||
<paragraph blockId="1.[155,1029,730,964]" pageId="1" pageNumber="348">
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Material.
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<materialsCitation ID-GBIF-Occurrence="2433380407" collectingDate="2005-08-10" collectionCode="UARC" collectorName="Pérez, L. A." country="Colombia" elevation="129" latitude="5.316944" location="Salero" longitude="-76.61444" pageId="1" pageNumber="348" specimenCount="1" specimenCount-male="1" stateProvince="Choco Department" typeStatus="Holotype">
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−
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<typeStatus box="[315,396,732,751]" pageId="1" pageNumber="348">
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<emphasis bold="true" box="[315,396,732,751]" pageId="1" pageNumber="348">Holotype</emphasis>
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</typeStatus>
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<specimenCount box="[400,418,731,750]" pageId="1" pageNumber="348">♂</specimenCount>
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,
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<collectingCountry box="[427,550,732,751]" name="Colombia" pageId="1" pageNumber="348" value="Colombia">COLOMBIA</collectingCountry>
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,
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<collectingRegion box="[556,730,731,751]" pageId="1" pageNumber="348">Chocó Department</collectingRegion>
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, Salero (05°19¢
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<geoCoordinate box="[862,907,730,750]" degrees="01" direction="north" minutes="2" orientation="latitude" pageId="1" pageNumber="348" precision="925" value="1.0333333">
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01²
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<collectionCode box="[890,907,730,750]" pageId="1" pageNumber="348">N</collectionCode>
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</geoCoordinate>
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, 76°36¢
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<geoCoordinate box="[973,1022,730,750]" degrees="52" direction="west" minutes="2" orientation="longitude" pageId="1" pageNumber="348" precision="925" value="-52.033333">
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52²
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<collectionCode box="[1001,1022,730,750]" pageId="1" pageNumber="348">W</collectionCode>
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</geoCoordinate>
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) alt.
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<quantity box="[187,241,758,778]" metricMagnitude="2" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="348" unit="m" value="129.0">129 m</quantity>
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,
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<date box="[249,367,758,777]" pageId="1" pageNumber="348" value="2005-08-10">10-VIII-2005</date>
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.L.A. Pérez leg.(
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<collectionCode box="[515,581,758,777]" collectionName="UARC" pageId="1" pageNumber="348">UARC</collectionCode>
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). −
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="2433380409" box="[619,923,758,778]" collectingDate="2005-08-10" collectionCode="MIZA" collectorName="Pérez, L. A." country="Colombia" elevation="129 m" latitude="5.316944" location="Salero" longitude="-76.61444" pageId="1" pageNumber="348" specimenCount="1" specimenCount-male="1" stateProvince="Choco Department" typeStatus="Paratype">
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<typeStatus box="[619,698,758,777]" pageId="1" pageNumber="348">
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<emphasis bold="true" box="[619,698,758,777]" pageId="1" pageNumber="348">Paratype</emphasis>
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</typeStatus>
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<specimenCount box="[702,720,758,777]" pageId="1" pageNumber="348">♂</specimenCount>
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, same data (
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<collectionCode box="[826,889,758,777]" collectionName="Venezuela, Maracay, Museuo del Instituto de Zoologia Agricola" pageId="1" pageNumber="348">MIZA</collectionCode>
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).−
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="2433380408" collectingDate="2005-08-10" collectionCode="UARC" collectorName="Pérez, L. A." country="Colombia" elevation="129 m" latitude="5.316944" location="Salero" longitude="-76.61444" pageId="1" pageNumber="348" specimenCount="1" specimenCount-female="1" stateProvince="Choco Department" typeStatus="Allotype">
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<typeStatus box="[927,1001,758,777]" pageId="1" pageNumber="348">
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<emphasis bold="true" box="[927,1001,758,777]" pageId="1" pageNumber="348">Allotype</emphasis>
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</typeStatus>
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<specimenCount box="[1005,1023,758,777]" pageId="1" pageNumber="348">♀</specimenCount>
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, same data (
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<collectionCode box="[252,318,785,804]" collectionName="UARC" pageId="1" pageNumber="348">UARC</collectionCode>
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)
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</materialsCitation>
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. − Other species examined:
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<taxonomicName authority="Tamazunchale" authorityName="Gloyd" authorityYear="1944" box="[569,650,785,804]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="regalis">
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<emphasis box="[569,650,785,804]" italics="true" pageId="1" pageNumber="348">A. regalis</emphasis>
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</taxonomicName>
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<specimenCount box="[653,671,784,803]" pageId="1" pageNumber="348">♂</specimenCount>
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, Tamazunchale, San Luís Potosí, Mexico
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<date box="[155,272,811,830]" pageId="1" pageNumber="348" value="1958-08-30">
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30-
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<date box="[183,272,811,830]" pageId="1" pageNumber="348">VIII-1958</date>
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</date>
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, Beatty leg. & Beatty det., 1959. JR- 08291. −
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<taxonomicName authorityName="McLachlan" authorityYear="1895" box="[688,813,811,831]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="californicus">
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<emphasis box="[688,813,811,831]" italics="true" pageId="1" pageNumber="348">A. californicus</emphasis>
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</taxonomicName>
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<specimenCount box="[819,837,811,830]" pageId="1" pageNumber="348">♂</specimenCount>
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,
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<specimenCount box="[847,865,811,830]" pageId="1" pageNumber="348">♀</specimenCount>
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(in tandem) Calif: Stanislaus co., Del Puerto Cyn. at N. Fork of Del Puerto Creek about
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<quantity box="[768,823,838,857]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="1" pageNumber="348" unit="mi" value="14.0">14 mi.</quantity>
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W. of Patterson, 21/
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<date pageId="1" pageNumber="348" value="1978-09-23">
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<date bridgedPair="-" day="23" month="09" monthName="IX" pageId="1" pageNumber="348" year="1923">
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23-
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<date box="[155,230,865,884]" pageId="1" pageNumber="348">
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IX-
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<date box="[186,230,865,884]" pageId="1" pageNumber="348" value="1978" year="1978">1978</date>
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</date>
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</date>
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</date>
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, R.W. Garrison leg. & det.,
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<date box="[472,517,865,884]" pageId="1" pageNumber="348">1978</date>
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. −
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<taxonomicName authority="Choroni, La Rinconada" authorityName="Choroni, La Rinconada" box="[550,672,865,884]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="tuberalatus">
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<emphasis box="[550,672,865,884]" italics="true" pageId="1" pageNumber="348">A. tuberalatus</emphasis>
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</taxonomicName>
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<specimenCount box="[676,694,864,883]" pageId="1" pageNumber="348">♂</specimenCount>
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, Choroní, La Rinconada,
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<quantity box="[926,981,865,884]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="348" unit="m" value="350.0">350 m</quantity>
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, AR,
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<date box="[155,266,892,911]" pageId="1" pageNumber="348" value="2003-12-18">18-XII-2003</date>
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, J. De Marmels leg. & det. −
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<taxonomicName authority="Rancho Grande" authorityName="Rambur" authorityYear="1842" box="[538,628,891,911]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis box="[538,628,891,911]" italics="true" pageId="1" pageNumber="348">A. grandis</emphasis>
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</taxonomicName>
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<specimenCount box="[634,652,891,910]" pageId="1" pageNumber="348">♂</specimenCount>
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, Rancho Grande, Aragua, Venezuela,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="348" unit="m" value="500.0">500 m</quantity>
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,
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<date box="[184,287,918,937]" pageId="1" pageNumber="348" value="1951-09-16">16-IX-1951</date>
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, Y. Fernandez leg., J. Racenis det., 1954. −
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<taxonomicName authorityName="De Marmels" authorityYear="1982" box="[682,797,918,937]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="guayaraca">
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<emphasis box="[682,797,918,937]" italics="true" pageId="1" pageNumber="348">A. guayaraca</emphasis>
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</taxonomicName>
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<specimenCount box="[804,822,918,937]" pageId="1" pageNumber="348">♂</specimenCount>
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(
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<typeStatus box="[833,913,918,937]" pageId="1" pageNumber="348">holotype</typeStatus>
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) Guayaraca, Auyantepuí, Bolivar, Venezuela,
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<date box="[442,543,945,964]" pageId="1" pageNumber="348" value="1956-04-25">25-IV-1956</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="348" type="etymology">
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<paragraph blockId="1.[155,1029,1003,1563]" pageId="1" pageNumber="348">Etymology. − The species name is dedicated to the amazing but endangered exuberant Chocoan Pluvial Forest.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="350" pageId="1" pageNumber="348" type="description">
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<paragraph blockId="1.[155,1029,1003,1563]" pageId="1" pageNumber="348">
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MALE (
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<typeStatus box="[258,361,1059,1083]" pageId="1" pageNumber="348">holotype</typeStatus>
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). − H e a d (
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<figureCitation box="[522,601,1059,1083]" captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="1" pageNumber="348">Fig. 1a</figureCitation>
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). − Labium yellow, base of mandible pale green, tip black; labrum pale green, margin rimmed with a thin black line; anteclypeus pale green, postclypeus entirely shiny black. Gena ochre; frons, vertex and top of head dark metallic green, antenna with scape and pedicel black, flagellum brown. A small, pale spot externally at each lateral ocellus. Rear of head yellow, black near maxillar insertion.
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</paragraph>
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<paragraph blockId="1.[155,1029,1003,1563]" lastBlockId="2.[132,1006,221,1558]" lastPageId="2" lastPageNumber="349" pageId="1" pageNumber="348">
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Thorax. − Anterior lobe of prothorax rounded, black (
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||
<figureCitation box="[803,881,1251,1275]" captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="1" pageNumber="348">Fig. 1b</figureCitation>
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), middle lobe metallic green with longitudinal mid line and lateral region yellow, posterior lobe dark metallic green with small central yellow spot; proepisternum and proepimeron yellow with pruinescense (
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<figureCitation box="[490,563,1347,1371]" captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="1" pageNumber="348">Fig. 1c</figureCitation>
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); procoxa largely yellow with pruinescence. Pterothorax largely yellow laterally with broad green metallic stripes bordered black. Mesepisternum in dorsal view largely metallic green, middorsal carina yellow, in lateral view green metallic with thin yellow stripe along humeral suture (
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<figureCitation box="[164,246,1475,1499]" captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="1" pageNumber="348">Fig. 1d</figureCitation>
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||
). Mesokatepisternum largely black with metallic green luster, inferior portion yellow. Mesepimeron entirely metallic green with a narrow, shiny black stripe along interpleural suture. Mesocoxae yellow with external black stripe (
|
||
<figureCitation captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" lastPageId="2" lastPageNumber="349" pageId="1" pageNumber="348">Fig. 1d</figureCitation>
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). Metepisternum largely yel- low with inferior margin along pleural suture with shiny black stripe extending to superior mar- gin of metepimeron. Distal half of paracoxal suture with shiny black stripe. Metepimeron and metasternum largely yellow with thin, black longitudinal line in middle, and with a lateral black spot. Metakatepimeron yellow and pruinose. Meso- and metascutum yellow, meso- and metaescutellum yellow.
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</paragraph>
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<paragraph blockId="2.[132,1006,221,1558]" pageId="2" pageNumber="349">
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Legs (
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<figureCitation box="[220,299,669,694]" captionStart="Fig" captionStartId="3.[155,187,1002,1021]" captionTargetBox="[155,647,219,961]" captionTargetId="figure@3.[155,647,219,961]" captionTargetPageId="3" captionText="Fig. 2. Archilestes chocoanus sp.n., male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view." httpUri="https://zenodo.org/record/3236063/files/figure.png" pageId="2" pageNumber="349">Fig. 2a</figureCitation>
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). − Mainly dark brown to black. Trochanter with ventral surface yellow, and posterior side black; femur in proximal quarter with pale inner side, meso and metafemur with 8-10 spines of approximately the same size on its outer surface. Wings (
|
||
<figureCitation box="[244,331,925,949]" captionStart="Fig" captionStartId="3.[155,187,1002,1021]" captionTargetBox="[155,647,219,961]" captionTargetId="figure@3.[155,647,219,961]" captionTargetPageId="3" captionText="Fig. 2. Archilestes chocoanus sp.n., male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view." httpUri="https://zenodo.org/record/3236063/files/figure.png" pageId="2" pageNumber="349">Fig. 2b</figureCitation>
|
||
). − Hyaline, veins black, pterostigma dark reddish brown, proximal side of quadrangle
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="349" unit="mm" value="0.87">0.87 mm</quantity>
|
||
(same in
|
||
<typeStatus box="[276,365,989,1013]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
and
|
||
<typeStatus box="[416,517,990,1014]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
); posterior side
|
||
<quantity box="[685,779,990,1014]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="2" pageNumber="349" unit="mm" value="1.87">1.87 mm</quantity>
|
||
(
|
||
<quantity box="[792,864,989,1013]" metricMagnitude="-2" metricUnit="m" metricValue="4.1148" pageId="2" pageNumber="349" unit="in" value="1.62">1.62 in</quantity>
|
||
<typeStatus box="[870,959,989,1013]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
and
|
||
<quantity box="[132,204,1021,1045]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" pageId="2" pageNumber="349" unit="in" value="1.75">1.75 in</quantity>
|
||
<typeStatus box="[210,308,1022,1046]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
) and anterior side
|
||
<quantity box="[509,604,1021,1045]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="349" unit="mm" value="0.75">0.75 mm</quantity>
|
||
(in
|
||
<typeStatus box="[644,741,1022,1046]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
and
|
||
<typeStatus box="[793,882,1021,1045]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
0.62). Petiolation reaching level of arculus. First antenodal space
|
||
<quantity box="[728,818,1054,1078]" metricMagnitude="-3" metricUnit="m" metricValue="4.87" pageId="2" pageNumber="349" unit="mm" value="4.87">4.87mm</quantity>
|
||
(
|
||
<quantity box="[829,903,1053,1077]" metricMagnitude="-1" metricUnit="m" metricValue="1.2065" pageId="2" pageNumber="349" unit="in" value="4.75">4.75 in</quantity>
|
||
<typeStatus box="[908,1005,1054,1078]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
and same in
|
||
<typeStatus box="[270,363,1085,1109]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
), third antenodal space
|
||
<quantity box="[625,720,1085,1109]" metricMagnitude="-3" metricUnit="m" metricValue="5.87" pageId="2" pageNumber="349" unit="mm" value="5.87">5.87 mm</quantity>
|
||
(
|
||
<quantity box="[734,809,1085,1109]" metricMagnitude="-1" metricUnit="m" metricValue="1.42748" pageId="2" pageNumber="349" unit="in" value="5.62">5.62 in</quantity>
|
||
<typeStatus box="[816,905,1085,1109]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
and
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.36398" pageId="2" pageNumber="349" unit="in" value="5.37">5.37 in</quantity>
|
||
<typeStatus box="[158,260,1118,1142]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
). Broadening ratio 1.05 (
|
||
<quantity box="[526,585,1117,1142]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="2" pageNumber="349" unit="in" value="1.1">1.1 in</quantity>
|
||
<typeStatus box="[590,689,1118,1142]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
,
|
||
<quantity box="[698,769,1117,1141]" metricMagnitude="-2" metricUnit="m" metricValue="2.667" pageId="2" pageNumber="349" unit="in" value="1.05">1.05 in</quantity>
|
||
<typeStatus box="[775,868,1117,1141]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
). Radial side of pterostigma
|
||
<quantity box="[301,383,1149,1173]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="349" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(same in
|
||
<typeStatus box="[488,577,1149,1173]" pageId="2" pageNumber="349">allotype</typeStatus>
|
||
and
|
||
<typeStatus box="[630,732,1150,1174]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
). The elongation ratio of the first post-quadrangular cell, defined as the relative lengths of the anterior to the distal vein of that cell (sensu
|
||
<bibRefCitation author="DONNELLY, T. W." box="[482,699,1214,1238]" journalOrPublisher="Fla Ent." pageId="2" pageNumber="349" pagination="412 - 417" part="64" refId="ref2708" refString="DONNELLY, T. W., 1981. A new species of Archilestes from Mexico and Central America, with further notes on the status of Cyptolestes Williamson (Odonata: Lestidae). Fla Ent. 64 (3): 412 - 417." title="A new species of Archilestes from Mexico and Central America, with further notes on the status of Cyptolestes Williamson (Odonata: Lestidae)" type="journal article" year="1981">
|
||
DONNELLY,
|
||
<date box="[644,699,1214,1238]" pageId="2" pageNumber="349">1981</date>
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="GARRISON, R. W." box="[708,921,1214,1238]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="2" pageNumber="349" pagination="1 - 12" part="702" refId="ref2755" refString="GARRISON, R. W., 1982. Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae). Occ. Pap. Mus. Zool. Univ. Mich. 702: 1 - 12." title="Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae)" type="journal article" year="1982">
|
||
GARRISON,
|
||
<date box="[867,921,1214,1238]" pageId="2" pageNumber="349" value="1982">1982</date>
|
||
</bibRefCitation>
|
||
) ranges from 2.0 to 2.33 (1.79-2.0 in
|
||
<typeStatus box="[451,553,1246,1270]" pageId="2" pageNumber="349">paratype</typeStatus>
|
||
). − Fw: Pt surmounting almost 4 crossveins; two complete Ax, 21 Px. RP
|
||
<subScript attach="left" box="[506,514,1293,1307]" fontSize="6" pageId="2" pageNumber="349">2</subScript>
|
||
arising from RP
|
||
<subScript attach="left" box="[693,701,1293,1307]" fontSize="6" pageId="2" pageNumber="349">1</subScript>
|
||
between Px 3 and 4. − Hw: Pt surmounting 4 crossveins, two complete Ax; 17 Px. RP
|
||
<subScript attach="left" box="[756,764,1325,1339]" fontSize="6" pageId="2" pageNumber="349">2</subScript>
|
||
arising in RP
|
||
<quantity box="[911,945,1309,1339]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="349" unit="in" value="1.0">
|
||
<subScript attach="left" box="[911,919,1325,1339]" fontSize="6" pageId="2" pageNumber="349">1</subScript>
|
||
in
|
||
</quantity>
|
||
3 Px.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="2" pageNumber="349" targetBox="[515,1003,220,777]" targetPageId="2">
|
||
<paragraph blockId="2.[132,1006,221,1558]" pageId="2" pageNumber="349">
|
||
Fig. 1.
|
||
<taxonomicName authority="Pérez-Gutiérrez, 2012" authorityName="Pérez Gutiérrez" authorityYear="2012" box="[581,771,812,831]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="chocoanus" status="sp. n.">
|
||
<emphasis box="[581,771,812,831]" italics="true" pageId="2" pageNumber="349">Archilestes chocoanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[778,828,812,831]" pageId="2" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="2.[132,1006,221,1558]" pageId="2" pageNumber="349">
|
||
Abdomen. − Abdominal segments largely green, lateral sides of S1 and S2 partially yellow with dark spots (
|
||
<figureCitation box="[523,602,1373,1397]" captionStart="Fig" captionStartId="2.[514,547,812,831]" captionTargetBox="[515,1003,220,777]" captionTargetId="figure@2.[514,1006,219,780]" captionTargetPageId="2" captionText="Fig. 1. Archilestes chocoanus sp. n., male holotype: (a) head, dorsal view; − (b) pterothorax, dorsal view; − (c) same, ventral view; − (d) pterothorax and abdominal segments 1 and 2 showing color pattern, right lateral view." httpUri="https://zenodo.org/record/3236061/files/figure.png" pageId="2" pageNumber="349">Fig. 1d</figureCitation>
|
||
), lateral side of S3 with a yellow oval spot close to posterior distal transverse carina, terga with a thin yellow line along the ventral margin, widest in S8; ventral pruinecense on S8-10, S10 black. Posterior hamule brown, lobulated and concave (
|
||
<figureCitation box="[628,706,1469,1494]" captionStart="Fig" captionStartId="3.[155,187,1002,1021]" captionTargetBox="[155,647,219,961]" captionTargetId="figure@3.[155,647,219,961]" captionTargetPageId="3" captionText="Fig. 2. Archilestes chocoanus sp.n., male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view." httpUri="https://zenodo.org/record/3236063/files/figure.png" pageId="2" pageNumber="349">Fig. 2c</figureCitation>
|
||
). Apical segment of genital ligula oval. (
|
||
<figureCitation box="[268,332,1501,1525]" captionStart="Fig" captionStartId="3.[155,187,1002,1021]" captionTargetBox="[155,647,219,961]" captionTargetId="figure@3.[155,647,219,961]" captionTargetPageId="3" captionText="Fig. 2. Archilestes chocoanus sp.n., male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view." httpUri="https://zenodo.org/record/3236063/files/figure.png" pageId="2" pageNumber="349">Fig. 2</figureCitation>
|
||
d-f).
|
||
</paragraph>
|
||
<paragraph blockId="2.[132,1006,221,1558]" lastPageId="3" lastPageNumber="350" pageId="2" pageNumber="349">
|
||
Cercus entirely black, slightly directed upwards (
|
||
<figureCitation box="[764,853,1533,1558]" captionStart="Fig" captionStartId="4.[514,547,753,772]" captionTargetId="figure@4.[514,1006,219,723]" captionTargetPageId="4" captionText="Fig. 3. Fig. 3. Archilestes chocoanus sp. n. [Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view." httpUri="https://zenodo.org/record/3339507/files/figure.png" pageId="2" pageNumber="349">Fig. 3a</figureCitation>
|
||
) and pointed with acute basal tooth mesally in the first third of its length (
|
||
<figureCitation captionStart="Fig" captionStartId="4.[514,547,753,772]" captionTargetId="figure@4.[514,1006,219,723]" captionTargetPageId="4" captionText="Fig. 3. Fig. 3. Archilestes chocoanus sp. n. [Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view." httpUri="https://zenodo.org/record/3339507/files/figure.png" pageId="3" pageNumber="350">Fig. 3b</figureCitation>
|
||
), middle region of cercus con- cave, followed by a post-median expansion armed with 7-8 strong spines on internal margin and dorsally with a prominent tu- bercle excavated, paraproct ves- tigial (
|
||
<figureCitation box="[746,821,477,502]" captionStart="Fig" captionStartId="4.[514,547,753,772]" captionTargetId="figure@4.[514,1006,219,723]" captionTargetPageId="4" captionText="Fig. 3. Fig. 3. Archilestes chocoanus sp. n. [Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view." httpUri="https://zenodo.org/record/3339507/files/figure.png" pageId="3" pageNumber="350">Fig. 3c</figureCitation>
|
||
), in lateral view ex- tending scarcely beyond at base of cercus, approximately 24 % of cercus length; tips of cerci in natural position crossed.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="350">
|
||
Measurements (in mm). − Fw 38.3, Hw 36.7, abdomen (excluding ap- pendages) 51.5, cercus 2.1 (61% longer than
|
||
<collectionCode box="[722,734,716,735]" pageId="3" pageNumber="350">S</collectionCode>
|
||
10), paraproct 0.5.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="350">
|
||
FEMALE
|
||
<emphasis bold="true" box="[806,892,744,768]" pageId="3" pageNumber="350">
|
||
<typeStatus box="[806,892,744,768]" pageId="3" pageNumber="350">allotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[905,983,744,768]" captionStart="Fig" captionStartId="4.[514,547,753,772]" captionTargetId="figure@4.[514,1006,219,723]" captionTargetPageId="4" captionText="Fig. 3. Fig. 3. Archilestes chocoanus sp. n. [Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view." httpUri="https://zenodo.org/record/3339507/files/figure.png" pageId="3" pageNumber="350">Fig. 3d</figureCitation>
|
||
). − Appearance and size as in male, ovipositor with a distintive col- or pattern, lateral valves armed with 12-14 small spines on ven- tral margin, all approximate- ly the same size.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="350">
|
||
Measurements (in mm). − Ab- domen without appendages 42.0, cercus 1.06, paraprocts 0.87.Elongation ratio of the first post-quadrangular cell 2.0-2.1. VARIATION. − Male
|
||
<typeStatus box="[932,1028,1075,1099]" pageId="3" pageNumber="350">paratype</typeStatus>
|
||
as
|
||
<typeStatus box="[705,804,1107,1131]" pageId="3" pageNumber="350">holotype</typeStatus>
|
||
, but RP2 vein arising in 3 Px, in 4 Px and 3 Px in left and right Hw respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption httpUri="https://zenodo.org/record/3236063/files/figure.png" pageId="3" pageNumber="350" targetBox="[155,647,219,961]" targetPageId="3">
|
||
<paragraph pageId="3" pageNumber="350">
|
||
Fig. 2.
|
||
<taxonomicName authority="Pérez-Gutiérrez, 2012" authorityName="Pérez Gutiérrez" authorityYear="2012" box="[213,395,1002,1021]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus" status="sp.n.">
|
||
<emphasis box="[213,395,1002,1021]" italics="true" pageId="3" pageNumber="350">Archilestes chocoanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[398,443,1003,1022]" pageId="3" pageNumber="350" rank="species">sp.n.</taxonomicNameLabel>
|
||
, male holotype:(a) pos- terior right leg; − (b) left wings, ventral view; − (c) left posterior hamule, right lateral view; − (d) genital ligula, right lateral view;− (e) same, distal portion, ventral view; − (f) same, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="4" lastPageNumber="351" pageId="3" pageNumber="350" type="diagnosis">
|
||
<paragraph lastBlockId="4.[132,1006,221,1558]" lastPageId="4" lastPageNumber="351" pageId="3" pageNumber="350">
|
||
DIAGNOSIS. − Morphologically the new species is closely related to
|
||
<emphasis italics="true" pageId="3" pageNumber="350">A. guayara- ca</emphasis>
|
||
<bibRefCitation author="DE MARMELS, J." box="[187,445,1235,1259]" journalOrPublisher="Boln Ent. venez. (N. S.)" pageId="3" pageNumber="350" pagination="111 - 113" part="2" refId="ref2667" refString="DE MARMELS, J., 1982. Archilestes guayaraca, nuevo Zygoptera de Venezuela (Odonata: Lestidae). Boln Ent. venez. (N. S.) 2 (14): 111 - 113." title="Archilestes guayaraca, nuevo Zygoptera de Venezuela (Odonata: Lestidae)" type="journal article" year="1982">
|
||
DE
|
||
<bibRefCitation author="DE MARMELS, J." box="[234,445,1235,1259]" journalOrPublisher="Boln Ent. venez. (N. S.)" pageId="3" pageNumber="350" pagination="111 - 113" part="2" refId="ref2667" refString="DE MARMELS, J., 1982. Archilestes guayaraca, nuevo Zygoptera de Venezuela (Odonata: Lestidae). Boln Ent. venez. (N. S.) 2 (14): 111 - 113." title="Archilestes guayaraca, nuevo Zygoptera de Venezuela (Odonata: Lestidae)" type="journal article" year="1982">
|
||
MARMELS,
|
||
<date box="[390,445,1235,1259]" pageId="3" pageNumber="350">1982</date>
|
||
</bibRefCitation>
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName authority="DONNELLY, 1981" authorityName="Donnelly" authorityYear="1981" box="[457,816,1235,1259]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="latialatus">
|
||
<emphasis box="[457,591,1235,1259]" italics="true" pageId="3" pageNumber="350">A. latialatus</emphasis>
|
||
<bibRefCitation author="DONNELLY, T. W." box="[599,816,1235,1259]" journalOrPublisher="Fla Ent." pageId="3" pageNumber="350" pagination="412 - 417" part="64" refId="ref2708" refString="DONNELLY, T. W., 1981. A new species of Archilestes from Mexico and Central America, with further notes on the status of Cyptolestes Williamson (Odonata: Lestidae). Fla Ent. 64 (3): 412 - 417." title="A new species of Archilestes from Mexico and Central America, with further notes on the status of Cyptolestes Williamson (Odonata: Lestidae)" type="journal article" year="1981">
|
||
DONNELLY,
|
||
<date box="[764,816,1235,1259]" pageId="3" pageNumber="350">1981</date>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[874,1028,1235,1259]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="tuberalatus">
|
||
<emphasis box="[874,1028,1235,1259]" italics="true" pageId="3" pageNumber="350">A. tuberalatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="WILLIAMSON, E. B." box="[164,410,1267,1291]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="3" pageNumber="350" pagination="1 - 9" part="96" refId="ref3015" refString="WILLIAMSON, E. B., 1921. Two new neotropical genera of Lestinae (Odonata). Occ. Pap. Mus. Zool. Univ. Mich. 96: 1 - 9." title="Two new neotropical genera of Lestinae (Odonata)" type="journal article" year="1921">
|
||
WILLIAMSON,
|
||
<date box="[358,410,1267,1291]" pageId="3" pageNumber="350">1921</date>
|
||
</bibRefCitation>
|
||
). With this latter it shares the presence of an robust basal tooth (specimens examined from Choroní, Aragua), although in
|
||
<taxonomicName box="[841,989,1299,1323]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="tuberalatus">
|
||
<emphasis box="[841,989,1299,1323]" italics="true" pageId="3" pageNumber="350">A. tuberalatus</emphasis>
|
||
</taxonomicName>
|
||
the basal tooth can be vestigial, visible just as a slightly rounded projection. GARRI- SON (
|
||
<date box="[223,277,1363,1387]" pageId="3" pageNumber="350">1982</date>
|
||
) has suggested that the tooth can show variations in size and that these variations are geographical [i. e.
|
||
<taxonomicName authorityName="Donnelly" authorityYear="1981" box="[500,631,1395,1419]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="latialatus">
|
||
<emphasis box="[500,631,1395,1419]" italics="true" pageId="3" pageNumber="350">A. latialatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="GARRISON, R. W." box="[644,896,1395,1419]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="3" pageNumber="350" pagination="1 - 12" part="702" refId="ref2755" refString="GARRISON, R. W., 1982. Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae). Occ. Pap. Mus. Zool. Univ. Mich. 702: 1 - 12." title="Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae)" type="journal article" year="1982">
|
||
GARRISON,
|
||
<date box="[803,855,1395,1419]" pageId="3" pageNumber="350">1982</date>
|
||
: 10
|
||
</bibRefCitation>
|
||
)]. The para- proct is, in
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[274,416,1427,1451]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[274,416,1427,1451]" italics="true" pageId="3" pageNumber="350">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
, underdeveloped, as in
|
||
<taxonomicName authorityName="De Marmels" authorityYear="1982" box="[673,816,1427,1450]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="guayaraca">
|
||
<emphasis box="[673,816,1427,1450]" italics="true" pageId="3" pageNumber="350">A. guayaraca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[828,980,1427,1451]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="tuberalatus">
|
||
<emphasis box="[828,980,1427,1451]" italics="true" pageId="3" pageNumber="350">A. tuberalatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Donnelly" authorityYear="1981" box="[155,286,1459,1483]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="latialatus">
|
||
<emphasis box="[155,286,1459,1483]" italics="true" pageId="3" pageNumber="350">A. latialatus</emphasis>
|
||
</taxonomicName>
|
||
. The character which most closely relates
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[750,892,1459,1483]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[750,892,1459,1483]" italics="true" pageId="3" pageNumber="350">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<emphasis italics="true" pageId="3" pageNumber="350">A. lati- alatus</emphasis>
|
||
is, however, the acute and curved cercus (
|
||
<bibRefCitation author="GARRISON, R. W." box="[660,875,1491,1515]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="3" pageNumber="350" pagination="1 - 12" part="702" refId="ref2755" refString="GARRISON, R. W., 1982. Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae). Occ. Pap. Mus. Zool. Univ. Mich. 702: 1 - 12." title="Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae)" type="journal article" year="1982">
|
||
GARRISON,
|
||
<date box="[819,875,1491,1515]" pageId="3" pageNumber="350" value="1982">1982</date>
|
||
</bibRefCitation>
|
||
: fig. 22), being notoriously longest in
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[391,530,1523,1547]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[391,530,1523,1547]" italics="true" pageId="3" pageNumber="350">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
. Marginal cell after quadrangle is in
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" lastPageId="4" lastPageNumber="351" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis authorityName="Pérez-Gutiérrez" authorityYear="2012" class="Insecta" family="Lestidae" genus="Archilestes" italics="true" kingdom="Animalia" lastPageId="4" lastPageNumber="351" order="Odonata" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="chocoanus">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
longer and narrower than in all other species described to date, but most similar to
|
||
<emphasis italics="true" pageId="4" pageNumber="351">A. gran- dis</emphasis>
|
||
(
|
||
<bibRefCitation author="RAMBUR, J." box="[175,365,318,342]" journalOrPublisher="Roret, Paris" pageId="4" pageNumber="351" refId="ref2935" refString="RAMBUR, J., 1842. Histoire naturelle des insectes. nevropteres. Roret, Paris." title="Histoire naturelle des insectes. nevropteres" type="book" year="1842">
|
||
RAMBUR,
|
||
<date box="[312,365,318,342]" pageId="4" pageNumber="351">1842</date>
|
||
</bibRefCitation>
|
||
) and
|
||
<emphasis italics="true" pageId="4" pageNumber="351">A. re- galis</emphasis>
|
||
(
|
||
<bibRefCitation author="GLOYD, L. K." box="[196,365,350,374]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="4" pageNumber="351" pagination="1 - 4" part="483" refId="ref2852" refString="GLOYD, L. K., 1944. A new species of Archilestes from Mexico (Odonata: Lestinae). Occ. Pap. Mus. Zool. Univ. Mich. 483: 1 - 4." title="A new species of Archilestes from Mexico (Odonata: Lestinae)" type="journal article" year="1944">
|
||
<collectionCode box="[196,294,350,374]" pageId="4" pageNumber="351">GLOYD</collectionCode>
|
||
,
|
||
<date box="[308,365,350,374]" pageId="4" pageNumber="351">1944</date>
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[132,1006,221,1558]" pageId="4" pageNumber="351">
|
||
In color pattern,
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[341,484,381,405]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[341,484,381,405]" italics="true" pageId="4" pageNumber="351">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
comes close to
|
||
<taxonomicName authorityName="Garrison" authorityYear="1982" box="[297,406,413,437]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="neblina">
|
||
<emphasis box="[297,406,413,437]" italics="true" pageId="4" pageNumber="351">A. neblina</emphasis>
|
||
</taxonomicName>
|
||
GAR- RISON,
|
||
<date box="[242,300,446,470]" pageId="4" pageNumber="351">1982</date>
|
||
, nonetheless it is important to note that on mesepisternum
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[251,396,509,533]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[251,396,509,533]" italics="true" pageId="4" pageNumber="351">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
has the green stripe complete, whereas
|
||
<taxonomicName authorityName="Garrison" authorityYear="1982" box="[132,243,573,597]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="neblina">
|
||
<emphasis box="[132,243,573,597]" italics="true" pageId="4" pageNumber="351">A. neblina</emphasis>
|
||
</taxonomicName>
|
||
has a thin stripe bordered by the posterior pleural suture. Also in
|
||
<taxonomicName authorityName="Garrison" authorityYear="1982" box="[269,376,637,661]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="neblina">
|
||
<emphasis box="[269,376,637,661]" italics="true" pageId="4" pageNumber="351">A. neblina</emphasis>
|
||
</taxonomicName>
|
||
the metasternum is largely black while in
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[132,276,701,725]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[132,276,701,725]" italics="true" pageId="4" pageNumber="351">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
it is mostly yellow, resembling
|
||
<taxonomicName authorityName="De Marmels" authorityYear="1982" box="[266,417,734,757]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="guayaraca">
|
||
<emphasis box="[266,417,734,757]" italics="true" pageId="4" pageNumber="351">A. guayaraca</emphasis>
|
||
</taxonomicName>
|
||
. Pos- terior hamule bear a short and rounded tip, as in
|
||
<emphasis box="[332,484,798,821]" italics="true" pageId="4" pageNumber="351">
|
||
<taxonomicName authorityName="De Marmels" authorityYear="1982" box="[332,480,798,821]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="guayaraca">A. guayaraca</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The genital ligula of
|
||
<emphasis italics="true" pageId="4" pageNumber="351">A. chocoa- nus</emphasis>
|
||
(ligula ovate with ven- tral arm not touching tip) be- longs to group 3 as proposed by
|
||
<bibRefCitation author="GARRISON, R. W." box="[132,357,958,982]" journalOrPublisher="Occ. Pap. Mus. Zool. Univ. Mich." pageId="4" pageNumber="351" pagination="1 - 12" part="702" refId="ref2755" refString="GARRISON, R. W., 1982. Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae). Occ. Pap. Mus. Zool. Univ. Mich. 702: 1 - 12." title="Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae)" type="journal article" year="1982">GARRISON (1982)</bibRefCitation>
|
||
, although the post-mortem variation of this structure is suspected to be particularly problematic when defining intrageneric groups. In the two
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[181,325,1021,1045]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[181,325,1021,1045]" italics="true" pageId="4" pageNumber="351">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
males examined, the ventral arm is in contact with the tip of the ligula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption httpUri="https://zenodo.org/record/3339507/files/figure.png" pageId="4" pageNumber="351" targetBox="[519,1002,220,722]" targetPageId="4">
|
||
<paragraph blockId="4.[132,1006,221,1558]" pageId="4" pageNumber="351">
|
||
Fig. 3. Fig. 3.
|
||
<taxonomicName authority="Pérez-Gutiérrez, 2012" authorityName="Pérez Gutiérrez" authorityYear="2012" box="[636,823,753,772]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus" status="sp. n.">
|
||
<emphasis box="[636,823,753,772]" italics="true" pageId="4" pageNumber="351">Archilestes chocoanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[827,874,753,772]" pageId="4" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
|
||
[Figs a-c: male holotype, Fig. d: female allotype]: (a) distal portion of abdomen, lateral view; − (b) right circus, dorsal view (right), ventral view (left); − (c) paraprocts and right circus, ventral view; − (d) distal portion of abdomen, left lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="5" lastPageNumber="352" pageId="4" pageNumber="351" type="discussion">
|
||
<paragraph blockId="4.[132,1006,221,1558]" pageId="4" pageNumber="351">
|
||
FINAL REMARKS. − The specimens of this colorful species were collected in a secondary forest. Days before our visit to the locality, according to our guide, there was a bulldozer opening way through the dense forest. This destructive practice is conducted to determine locations of new platinum prospecting camps and for introducing machinery. We ingressed through one of these roads, and there we had the first sighting of
|
||
<taxonomicName authorityName="Pérez Gutiérrez" authorityYear="2012" box="[336,475,1245,1269]" class="Insecta" family="Lestidae" genus="Archilestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="chocoanus">
|
||
<emphasis box="[336,475,1245,1269]" italics="true" pageId="4" pageNumber="351">A. chocoanus</emphasis>
|
||
</taxonomicName>
|
||
, perched on leaves, ca
|
||
<quantity box="[712,752,1246,1270]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="351" unit="m" value="2.0">2 m</quantity>
|
||
above ground, of about
|
||
<quantity box="[132,185,1278,1302]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="351" unit="m" value="10.0">10 m</quantity>
|
||
high
|
||
<taxonomicName box="[243,368,1278,1302]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="351" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="K.S.Kunth" authorityYear="1822" box="[420,573,1277,1301]" class="Magnoliopsida" family="Bombacaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="351" phylum="Tracheophyta" rank="family">Bombacaceae</taxonomicName>
|
||
trees. The violent and powerful flight of this species is noteworthy, and detection of two males was possible thanks to the audible impact produced when they landed on leaves of woody vegetation. Their evasive behavior, flying to the trees tops when disturbed, is also uncommon in the genus.
|
||
</paragraph>
|
||
<paragraph blockId="4.[132,1006,221,1558]" lastBlockId="5.[155,1028,221,309]" lastPageId="5" lastPageNumber="352" pageId="4" pageNumber="351">
|
||
In
|
||
<date box="[186,288,1437,1462]" pageId="4" pageNumber="351" value="2007-07">July 2007</date>
|
||
and
|
||
<date box="[343,449,1437,1462]" pageId="4" pageNumber="351" value="2010-07">July 2010</date>
|
||
, two more expeditions were carried out to the area in order to find the larva, but without success. Adults were not seen either. Unfortunately, the progressive loss of forest, due to mining activities and timbering practiced by the “colonos”, makes the rediscovery of the species at the
|
||
<typeStatus box="[957,1005,1534,1558]" pageId="4" pageNumber="351">type</typeStatus>
|
||
locality most unlikely. Future efforts must be made to determine the extent of its distribution range. Provisionally it is considered here as an endemic element of biogeographic Chocó.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |