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<document ID-DOI="10.1206/352.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5405335" approvalRequired="503" approvalRequired_for_taxonomicNames="62" approvalRequired_for_textStreams="429" approvalRequired_for_treatments="12" checkinTime="1630345222680" checkinUser="tatiana" docAuthor="Nesbitt, Sterling J." docDate="2011" docId="357D771BFFB1FFBBEDD7F987FEFBF950" docLanguage="en" docName="B352.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (352)" docSource="http://www.bioone.org/doi/abs/10.1206/352.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.3:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="3" docTitle="Arizonasaurus babbitti Welles 1947" docType="treatment" docVersion="6" lastPageNumber="28" masterDocId="C9440F63FFABFFA0EF17FFF9FFBFFF8E" masterDocTitle="The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades" masterLastPageNumber="292" masterPageNumber="1" pageNumber="27" updateTime="1631052925311" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades</mods:title>
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<paragraph blockId="26.[704,1224,1661,1684]" box="[704,1224,1661,1684]" pageId="26" pageNumber="27">
<heading box="[704,1224,1661,1684]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="27" reason="6">
<taxonomicName authority="Welles, 1947" authorityName="Welles" authorityYear="1947" box="[704,1119,1661,1684]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="babbitti">
<emphasis box="[704,963,1661,1684]" italics="true" pageId="26" pageNumber="27">Arizonasaurus babbitti</emphasis>
<bibRefCitation author="Welles, S. P." box="[972,1119,1661,1683]" pageId="26" pageNumber="27" pagination="241 - 294" refId="ref202729" refString="Welles, S. P. 1947. Vertebrates from the Upper Moenkopi Formation of the Northern Arizona. University of California Publications in Geological Science 27: 241 - 294." type="journal article" year="1947">Welles, 1947</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[1136,1216,1661,1683]" captionStart="Fig" captionStartId="20.[152,190,1552,1571]" captionText="Fig. 8. Skull reconstructions of pseudosuchian archosaur terminal taxa: A, Riojasuchus tenuisceps in lateral and B, dorsal views; redrawn from Sereno (1991a); C, Gracilisuchus stipanicicorum in lateral and D, dorsal views; redrawn from Romer (1971b); E, Arizonasaurus babbitti in lateral view; redrawn from Nesbitt (2005); F, Stagonolepis robertsoni in lateral and G, dorsal views; redrawn from Walker (1961); H, Revueltosaurus callenderi in lateral view; based on PEFO 34561. I, Effigia okeeffeae in lateral view; redrawn from Nesbitt and Norell (2006); J, Xilousuchus sapingensis in lateral view; based on IVPP V 6026. Shaded areas indicate incomplete preservation. See appendix for anatomical abbreviations. Scale bars 5 1 cm." pageId="26" pageNumber="27">fig. 8E</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="26.[701,1227,1706,1759]" box="[730,1227,1706,1730]" pageId="26" pageNumber="27">
AGE: Anisian, Middle Triassic (
<bibRefCitation author="Lucas, S. G." box="[1150,1227,1708,1729]" pageId="26" pageNumber="27" pagination="347 - 384" refId="ref193654" refString="Lucas, S. G. 1998 a. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoeocology 143: 347 - 384." type="journal article" year="1998">Lucas,</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph blockId="26.[701,1227,1706,1759]" box="[701,786,1737,1759]" pageId="26" pageNumber="27">1998a).</paragraph>
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
OCCURRENCE: Holbrook Member of the Moenkopi Formation,
<collectingRegion box="[361,458,242,264]" country="United States of America" name="Arizona" pageId="27" pageNumber="28">Arizona</collectingRegion>
; Anton Chico Member of the Moenkopi Formation,
<collectingRegion country="United States of America" name="New Mexico" pageId="27" pageNumber="28">New Mexico</collectingRegion>
(
<bibRefCitation author="Schoch, R." box="[196,422,300,322]" pageId="27" pageNumber="28" refId="ref199646" refString="Schoch, R., et al. 2010. The reptile assemblage from the Moenkopi Formation (Middle Triassic) of New Mexico. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen (DOI: 10.1127 / 0077 - 7749 / 2009 / 0030)." type="book" year="2010">Schoch et al., 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="materials_examined">
<paragraph blockId="27.[93,620,211,1759]" box="[122,540,328,352]" pageId="27" pageNumber="28">
<typeStatus box="[122,250,328,352]" pageId="27" pageNumber="28">HOLOTYPE</typeStatus>
: UCMP 36232, maxilla.
</paragraph>
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
REFERRED MATERIAL: MSM 4590, skull and partial skeleton; see
<bibRefCitation author="Nesbitt, S. J." box="[440,615,390,412]" pageId="27" pageNumber="28" pagination="237" refId="ref195097" refString="Nesbitt, S. J. 2003. Arizonasaurus and its implications for archosaur divergences. Proceedings of the Royal Society of London B Biological Sciences 270 (suppl. 2): S 234 - S 237." type="journal article" year="2003">Nesbitt (2003</bibRefCitation>
,
<bibRefCitation author="Nesbitt, S. J." box="[93,162,419,441]" pageId="27" pageNumber="28" pagination="19 - 47" refId="ref195136" refString="Nesbitt, S. J. 2005 a. The osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti. Historical Biology 17: 19 - 47." type="journal article" year="2005">2005a</bibRefCitation>
) and
<bibRefCitation author="Schoch, R." box="[243,494,419,441]" pageId="27" pageNumber="28" refId="ref199646" refString="Schoch, R., et al. 2010. The reptile assemblage from the Moenkopi Formation (Middle Triassic) of New Mexico. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen (DOI: 10.1127 / 0077 - 7749 / 2009 / 0030)." type="book" year="2010">Schoch et al. (2010)</bibRefCitation>
for other specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="discussion">
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
REMARKS:
<taxonomicName box="[256,419,478,499]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[256,419,478,499]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
represent one of the most completely documented sailbacked archosaurs from the Anisian. Additionally,
<taxonomicName box="[200,363,566,587]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[200,363,566,587]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
is the most common reptile found in the Holbrook and Anton Chico Members of the Moenkopi Formation (
<bibRefCitation author="Nesbitt, S. J." box="[162,344,653,675]" pageId="27" pageNumber="28" pagination="332 - 346" refId="ref195164" refString="Nesbitt, S. J. 2005 b. A new archosaur from the upper Moenkopi Formation (Middle Triassic) of Arizona and its implications for rauisuchian phylogeny and diversification. Neues Jahrbuch fur Geologie und Palaeontologie Monatshefte 2005: 332 - 346." type="journal article" year="2005">Nesbitt, 2005b</bibRefCitation>
;
<bibRefCitation author="Schoch, R." box="[363,608,653,676]" pageId="27" pageNumber="28" refId="ref199646" refString="Schoch, R., et al. 2010. The reptile assemblage from the Moenkopi Formation (Middle Triassic) of New Mexico. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen (DOI: 10.1127 / 0077 - 7749 / 2009 / 0030)." type="book" year="2010">Schoch et al., 2010</bibRefCitation>
). The
<typeStatus box="[158,261,683,705]" pageId="27" pageNumber="28">holotype</typeStatus>
and the referred specimen (MSM 4590) share two characters: a uniquely shaped ascending process of the maxilla that is triangular in cross section and a deep pit at the posterior side of the base of the ascending process of the maxilla. Both these characters are present in
<taxonomicName box="[127,262,886,909]" class="Reptilia" family="Ctenosauriscidae" genus="Xilousuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[127,262,886,909]" italics="true" pageId="27" pageNumber="28">Xilousuchus</emphasis>
</taxonomicName>
suggesting that the two taxa are closely related (see below). The partial lacrimal described by Nesbitt (2005) is actually the prefrontal.
</paragraph>
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
<bibRefCitation author="Nesbitt, S. J." box="[122,295,1005,1027]" pageId="27" pageNumber="28" pagination="237" refId="ref195097" refString="Nesbitt, S. J. 2003. Arizonasaurus and its implications for archosaur divergences. Proceedings of the Royal Society of London B Biological Sciences 270 (suppl. 2): S 234 - S 237." type="journal article" year="2003">Nesbitt (2003)</bibRefCitation>
found
<taxonomicName box="[391,554,1005,1026]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[391,554,1005,1026]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
as a close relative of
<taxonomicName baseAuthorityName="Weinbaum and Hungerbuhler" baseAuthorityYear="2007" box="[285,416,1035,1056]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[285,416,1035,1056]" italics="true" pageId="27" pageNumber="28">Poposaurus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="sensu Nesbitt" baseAuthorityYear="2007" box="[477,619,1033,1056]" class="Reptilia" family="Shuvosauridae" genus="Shuvosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[477,619,1033,1056]" italics="true" pageId="27" pageNumber="28">Shuvosaurus</emphasis>
</taxonomicName>
(
<emphasis bold="true" box="[102,125,1062,1085]" pageId="27" pageNumber="28">5</emphasis>
<taxonomicName authorityName="Long &amp; Murry" authorityYear="1995" box="[136,268,1063,1086]" class="Reptilia" genus="Chatterjeea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[136,268,1063,1086]" italics="true" pageId="27" pageNumber="28">Chatterjeea</emphasis>
</taxonomicName>
) within Suchia. Other analyses found a similar position (
<bibRefCitation author="Nesbitt, S. J. &amp; M. A. Norell" pageId="27" pageNumber="28" pagination="1045 - 1048" refId="ref195294" refString="Nesbitt, S. J., and M. A. Norell. 2006. Extreme convergence in the body plans of an early suchian (Archosauria) and ornithomimid dinosaurs (Theropoda). Proceedings of the Royal Society of London B Biological Sciences 273: 1045 - 1048." type="journal article" year="2006">Nesbitt and Norell, 2006</bibRefCitation>
;
<bibRefCitation author="Irmis, R. B. &amp; W. G. Parker &amp; S. J. Nesbitt &amp; J. Liu" box="[257,477,1122,1144]" pageId="27" pageNumber="28" pagination="3 - 22" refId="ref192454" refString="Irmis, R. B., W. G. Parker, S. J. Nesbitt, and J. Liu. 2007 a. Ornithischian dinosaurs: the Triassic record. Historical Biology 18: 3 - 22." type="journal article" year="2007">Irmis et al., 2007a</bibRefCitation>
;
<bibRefCitation author="Weinbaum, J. C. &amp; A. Hungerbuhler" pageId="27" pageNumber="28" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
; Brusatte et al., 2008).
<bibRefCitation author="Nesbitt, S. J." box="[181,351,1180,1202]" pageId="27" pageNumber="28" pagination="237" refId="ref195097" refString="Nesbitt, S. J. 2003. Arizonasaurus and its implications for archosaur divergences. Proceedings of the Royal Society of London B Biological Sciences 270 (suppl. 2): S 234 - S 237." type="journal article" year="2003">Nesbitt (2003</bibRefCitation>
,
<bibRefCitation author="Nesbitt, S. J." box="[373,442,1180,1202]" pageId="27" pageNumber="28" pagination="19 - 47" refId="ref195136" refString="Nesbitt, S. J. 2005 a. The osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti. Historical Biology 17: 19 - 47." type="journal article" year="2005">2005a</bibRefCitation>
) hypothesized that
<taxonomicName box="[148,311,1210,1231]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[148,311,1210,1231]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
formed a clade with other sail-backed suchians including
<taxonomicName authorityName="Kuhn" authorityYear="1964" class="Reptilia" family="Ctenosauriscidae" genus="Ctenosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="27" pageNumber="28">Ctenosauriscus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Zhang" authorityYear="1975" box="[151,278,1269,1290]" class="Reptilia" family="Lotosauridae" genus="Lotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[151,278,1269,1290]" italics="true" pageId="27" pageNumber="28">Lotosaurus</emphasis>
</taxonomicName>
,
<taxonomicName box="[300,456,1269,1290]" class="Reptilia" family="Ctenosauriscidae" genus="Bromsgroveia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[300,456,1269,1290]" italics="true" pageId="27" pageNumber="28">Bromsgroveia</emphasis>
</taxonomicName>
, and
<taxonomicName class="Reptilia" family="Ctenosauriscidae" genus="Hypselorhachis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="27" pageNumber="28">Hypselorhachis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
<taxonomicName box="[122,285,1328,1349]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[122,285,1328,1349]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
differs from all other archosaurs except
<taxonomicName box="[244,379,1356,1379]" class="Reptilia" family="Ctenosauriscidae" genus="Xilousuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[244,379,1356,1379]" italics="true" pageId="27" pageNumber="28">Xilousuchus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Zhang" authorityYear="1975" box="[395,522,1357,1378]" class="Reptilia" family="Lotosauridae" genus="Lotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[395,522,1357,1378]" italics="true" pageId="27" pageNumber="28">Lotosaurus</emphasis>
</taxonomicName>
,
<taxonomicName class="Reptilia" family="Ctenosauriscidae" genus="Hypselorhachis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="27" pageNumber="28">Hypselorhachis</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Kuhn" authorityYear="1964" box="[262,432,1386,1407]" class="Reptilia" family="Ctenosauriscidae" genus="Ctenosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[262,432,1386,1407]" italics="true" pageId="27" pageNumber="28">Ctenosauriscus</emphasis>
</taxonomicName>
by the presence of a sail created by the elongation of the neural spines of the presacral vertebrae. It differs from
<taxonomicName box="[243,378,1473,1496]" class="Reptilia" family="Ctenosauriscidae" genus="Xilousuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[243,378,1473,1496]" italics="true" pageId="27" pageNumber="28">Xilousuchus</emphasis>
</taxonomicName>
by the absence of a deep pit in the parabasisphenoid ventral to the descending process of the opisthotic.
<taxonomicName box="[93,256,1561,1582]" class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[93,256,1561,1582]" italics="true" pageId="27" pageNumber="28">Arizonasaurus</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Kuhn" authorityYear="1964" box="[417,587,1561,1582]" class="Reptilia" family="Ctenosauriscidae" genus="Ctenosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[417,587,1561,1582]" italics="true" pageId="27" pageNumber="28">Ctenosauriscus</emphasis>
</taxonomicName>
in anteroposteriorly wide neural spines of the midposterior cervical vertebrae. It differs from
<taxonomicName authorityName="Zhang" authorityYear="1975" box="[159,286,1649,1670]" class="Reptilia" family="Lotosauridae" genus="Lotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis box="[159,286,1649,1670]" italics="true" pageId="27" pageNumber="28">Lotosaurus</emphasis>
</taxonomicName>
by the presence of teeth.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="key">
<paragraph blockId="27.[93,620,211,1759]" pageId="27" pageNumber="28">
KEY REFERENCES:
<bibRefCitation author="Welles, S. P." box="[356,510,1679,1701]" pageId="27" pageNumber="28" pagination="241 - 294" refId="ref202729" refString="Welles, S. P. 1947. Vertebrates from the Upper Moenkopi Formation of the Northern Arizona. University of California Publications in Geological Science 27: 241 - 294." type="journal article" year="1947">Welles, 1947</bibRefCitation>
;
<bibRefCitation author="Nesbitt, S. J." pageId="27" pageNumber="28" pagination="237" refId="ref195097" refString="Nesbitt, S. J. 2003. Arizonasaurus and its implications for archosaur divergences. Proceedings of the Royal Society of London B Biological Sciences 270 (suppl. 2): S 234 - S 237." type="journal article" year="2003">Nesbitt, 2003</bibRefCitation>
,
<bibRefCitation author="Nesbitt, S. J." box="[177,249,1708,1730]" pageId="27" pageNumber="28" pagination="19 - 47" refId="ref195136" refString="Nesbitt, S. J. 2005 a. The osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti. Historical Biology 17: 19 - 47." type="journal article" year="2005">2005a</bibRefCitation>
;
<bibRefCitation author="Gower, D. J. &amp; S. J. Nesbitt" box="[275,615,1708,1730]" pageId="27" pageNumber="28" pagination="79 - 87" refId="ref190381" refString="Gower, D. J., and S. J. Nesbitt. 2006. The braincase of Arizonasaurus babbitti - further evidence of the non-monophyly of Rauisuchia. Journal of Vertebrate Paleontology 26: 79 - 87." type="journal article" year="2006">Gower and Nesbitt, 2006</bibRefCitation>
;
<bibRefCitation author="Schoch, R." box="[93,320,1737,1759]" pageId="27" pageNumber="28" refId="ref199646" refString="Schoch, R., et al. 2010. The reptile assemblage from the Moenkopi Formation (Middle Triassic) of New Mexico. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen (DOI: 10.1127 / 0077 - 7749 / 2009 / 0030)." type="book" year="2010">Schoch et al., 2010</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>