306 lines
31 KiB
XML
306 lines
31 KiB
XML
<document ID-DOI="10.1206/352.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5405335" approvalRequired="503" approvalRequired_for_taxonomicNames="62" approvalRequired_for_textStreams="429" approvalRequired_for_treatments="12" checkinTime="1630345222680" checkinUser="tatiana" docAuthor="Nesbitt, Sterling J." docDate="2011" docId="357D771BFFB0FFBCEDFBFF2CFD90FA4A" docLanguage="en" docName="B352.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (352)" docSource="http://www.bioone.org/doi/abs/10.1206/352.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.3:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="3" docTitle="Poposaurus gracilis Mehl 1915" docType="treatment" docVersion="5" lastPageNumber="29" masterDocId="C9440F63FFABFFA0EF17FFF9FFBFFF8E" masterDocTitle="The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades" masterLastPageNumber="292" masterPageNumber="1" pageNumber="28" updateTime="1631052925311" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nesbitt, Sterling J.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2011-04-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2011</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>352</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>292</mods:end>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/abs/10.1206/352.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/352.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5405335</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:357D771BFFB0FFBCEDFBFF2CFD90FA4A" httpUri="http://treatment.plazi.org/id/357D771BFFB0FFBCEDFBFF2CFD90FA4A" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<subSubSection pageId="27" pageNumber="28" type="nomenclature">
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<paragraph blockId="27.[748,1112,212,235]" box="[748,1112,212,235]" pageId="27" pageNumber="28">
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<heading box="[748,1112,212,235]" centered="true" fontSize="9" level="2" pageId="27" pageNumber="28" reason="2">
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<taxonomicName authority="Mehl, 1915" authorityName="Mehl" authorityYear="1915" box="[748,1112,212,235]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[748,970,212,235]" italics="true" pageId="27" pageNumber="28">Poposaurus gracilis</emphasis>
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<bibRefCitation author="Mehl, M. G." box="[980,1112,213,235]" pageId="27" pageNumber="28" pagination="516 - 522" refId="ref194517" refString="Mehl, M. G. 1915. Poposaurus gracilis, a new reptile from the Triassic of Wyoming. Journal of Geology 23: 516 - 522." type="journal article" year="1915">Mehl, 1915</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph blockId="27.[667,1194,257,1759]" box="[696,1194,257,281]" pageId="27" pageNumber="28">AGE: Carnian–early Norian, Late Triassic.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="description">
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<paragraph blockId="27.[667,1194,257,1759]" pageId="27" pageNumber="28">
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<materialsCitation collectorName="Popo Agie Formation" location="Wyoming" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Wyoming">
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OCCURRENCE:
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<collectorName box="[893,1188,289,311]" pageId="27" pageNumber="28">Popo Agie Formation</collectorName>
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,
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<collectingRegion box="[667,785,319,341]" country="United States of America" name="Wyoming" pageId="27" pageNumber="28">Wyoming</collectingRegion>
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</materialsCitation>
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;
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<materialsCitation collectorName="Blue Mesa Member" location="Chinle Formation" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Arizona">
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<collectorName box="[797,1030,319,341]" pageId="27" pageNumber="28">Blue Mesa Member</collectorName>
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of the
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<location LSID="urn:lsid:plazi:treatment:357D771BFFB0FFBCEDFBFF2CFD90FA4A:B80B90D6FFB0FFBBEB4BFEC6FCA2FEFC" name="Chinle Formation" pageId="27" pageNumber="28" stateProvince="Arizona">Chinle Formation</location>
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,
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<collectingRegion box="[813,911,348,370]" country="United States of America" name="Arizona" pageId="27" pageNumber="28">Arizona</collectingRegion>
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</materialsCitation>
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;
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<materialsCitation collectorName="Mesa Redondo Member" location="Chinle Formation" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Arizona">
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<collectorName pageId="27" pageNumber="28">Mesa Redondo Member</collectorName>
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of the
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<location LSID="urn:lsid:plazi:treatment:357D771BFFB0FFBCEDFBFF2CFD90FA4A:B80B90D6FFB0FFBBEC55FE83FB98FE1E" box="[834,1063,378,400]" name="Chinle Formation" pageId="27" pageNumber="28" stateProvince="Arizona">Chinle Formation</location>
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,
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<collectingRegion box="[1091,1189,378,400]" country="United States of America" name="Arizona" pageId="27" pageNumber="28">Arizona</collectingRegion>
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</materialsCitation>
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;
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<materialsCitation collectorName="Tecovas Formation" location="Dockum Group" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Texas">
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<collectorName box="[667,900,407,429]" pageId="27" pageNumber="28">Tecovas Formation</collectorName>
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of the
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<location LSID="urn:lsid:plazi:treatment:357D771BFFB0FFBCEDFBFF2CFD90FA4A:B80B90D6FFB0FFBBECF5FE6EFB1BFE22" box="[994,1188,407,429]" name="Dockum Group" pageId="27" pageNumber="28" stateProvince="Texas">Dockum Group</location>
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,
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<collectingRegion box="[667,738,437,458]" country="United States of America" name="Texas" pageId="27" pageNumber="28">Texas</collectingRegion>
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</materialsCitation>
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;
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<materialsCitation collectorName="Monitor Butte Member" location="Chinle Formation" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Utah">
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<collectorName box="[751,1030,437,459]" pageId="27" pageNumber="28">Monitor Butte Member</collectorName>
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of the
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<location LSID="urn:lsid:plazi:treatment:357D771BFFB0FFBCEDFBFF2CFD90FA4A:B80B90D6FFB0FFBBEB4AFE4CFCA2FE66" name="Chinle Formation" pageId="27" pageNumber="28" stateProvince="Utah">Chinle Formation</location>
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, southern
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<collectingRegion box="[921,985,466,488]" country="United States of America" name="Utah" pageId="27" pageNumber="28">Utah</collectingRegion>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="materials_examined">
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<paragraph blockId="27.[667,1194,257,1759]" pageId="27" pageNumber="28">
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<typeStatus box="[696,824,494,518]" pageId="27" pageNumber="28">HOLOTYPE</typeStatus>
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: FMNH 357, two dorsal vertebrae, one caudal vertebra, a left ilium, the proximal portion of a left femur, a right femur, distal portion of the ischia.
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</paragraph>
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<paragraph blockId="27.[667,1194,257,1759]" pageId="27" pageNumber="28">
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REFERRED MATERIAL: TTU-P 10419, vertebrae, pelvic elements;
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<accessionNumber box="[955,1115,645,667]" httpUri="http://www.ncbi.nlm.nih.gov/protein/TMM43683" pageId="27" pageNumber="28">TMM 43683</accessionNumber>
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-1, vertebrae and nearly complete pelvis; various UCMP elements from A269 (see
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<bibRefCitation author="Long, R. A. & P. A. Murry" pageId="27" pageNumber="28" pagination="1 - 254" refId="ref193609" refString="Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
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);
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<accessionNumber box="[843,991,733,755]" httpUri="http://www.ncbi.nlm.nih.gov/protein/YPM57100" pageId="27" pageNumber="28">YPM 57100</accessionNumber>
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, nearly complete skeleton lacking the skull.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
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<paragraph blockId="27.[667,1194,257,1759]" pageId="27" pageNumber="28">
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REMARKS:
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<taxonomicName authorityName="Mehl" authorityYear="1915" box="[830,1052,792,815]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[830,1052,792,815]" italics="true" pageId="27" pageNumber="28">Poposaurus gracilis</emphasis>
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</taxonomicName>
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was named from a fragmentary specimen consisting of pelvic elements, the femora, and a few vertebrae (
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<bibRefCitation author="Mehl, M. G." box="[799,943,881,903]" pageId="27" pageNumber="28" pagination="516 - 522" refId="ref194517" refString="Mehl, M. G. 1915. Poposaurus gracilis, a new reptile from the Triassic of Wyoming. Journal of Geology 23: 516 - 522." type="journal article" year="1915">Mehl, 1915</bibRefCitation>
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). The differences in morphology from other Triassic archosaurs led various authors to identify
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<taxonomicName authorityName="Mehl" authorityYear="1915" box="[1040,1158,939,962]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[1040,1158,939,962]" italics="true" pageId="27" pageNumber="28">P. gracilis</emphasis>
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</taxonomicName>
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as an ornithischian (
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<bibRefCitation author="Nopsca, F." box="[874,1038,970,992]" pageId="27" pageNumber="28" refId="ref195896" refString="Nopsca, F. v. 1921. Zur systematischen Stellung von Poposaurus (Mehl). Zentralblatt fur Mineralogie, Geologie und Palaontologie 1921: 348 pp." type="book" year="1921">Nopsca, 1921</bibRefCitation>
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), a stegosaur (
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<bibRefCitation author="Huene, F." box="[675,832,999,1021]" pageId="27" pageNumber="28" pagination="53 - 58" refId="ref191527" refString="Huene, F. v. 1950. Die Entstehung der Ornithischia schon fruh in der Trias. Neues Jahrbuch fur Geologie und Palaontologie Monatshefte 1950: 53 - 58." type="journal article" year="1950">Huene, 1950</bibRefCitation>
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), a theropod (
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<bibRefCitation author="Colbert, E. H." box="[1012,1180,999,1021]" pageId="27" pageNumber="28" pagination="59 - 78" refId="ref187654" refString="Colbert, E. H. 1961. The Triassic reptile Poposaurus. Fieldiana 14: 59 - 78." type="journal article" year="1961">Colbert, 1961</bibRefCitation>
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), and a pseudosuchian (
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<bibRefCitation author="Walker, A. D." box="[946,1113,1029,1051]" pageId="27" pageNumber="28" pagination="470 - 476" refId="ref202326" refString="Walker, A. D. 1969. The reptile fauna of the '' Lower Keuper' ' Sandstone. Geological Magazine 106: 470 - 476." type="journal article" year="1969">Walker, 1969</bibRefCitation>
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). New specimens of
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<taxonomicName authorityName="Mehl" authorityYear="1915" box="[839,963,1057,1080]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[839,963,1057,1080]" italics="true" pageId="27" pageNumber="28">P. gracilis</emphasis>
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</taxonomicName>
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and other closely related taxa confirmed the pseudosuchian affinity of the taxon (
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<bibRefCitation author="Galton, P. M." box="[948,1112,1117,1139]" pageId="27" pageNumber="28" pagination="234 - 245" refId="ref189223" refString="Galton, P. M. 1977. On Staurikosaurus pricei, an early saurischian dinosaur from the Triassic of Brazil, with notes on the Herrerasauridae and Poposauridae. Palaontologische Zeitschrift 51: 234 - 245." type="journal article" year="1977">Galton, 1977</bibRefCitation>
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;
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<bibRefCitation author="Long, R. A. & P. A. Murry" pageId="27" pageNumber="28" pagination="1 - 254" refId="ref193609" refString="Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
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||
;
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<bibRefCitation author="Nesbitt, S. J. & M. A. Norell" box="[888,1189,1146,1168]" pageId="27" pageNumber="28" pagination="1045 - 1048" refId="ref195294" refString="Nesbitt, S. J., and M. A. Norell. 2006. Extreme convergence in the body plans of an early suchian (Archosauria) and ornithomimid dinosaurs (Theropoda). Proceedings of the Royal Society of London B Biological Sciences 273: 1045 - 1048." type="journal article" year="2006">Nesbitt and Norell, 2006</bibRefCitation>
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;
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||
<bibRefCitation author="Weinbaum, J. C. & A. Hungerbuhler" box="[667,1110,1176,1198]" pageId="27" pageNumber="28" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
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). With the exception of a nearly complete skeleton lacking the skull (
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<bibRefCitation author="Joyce, W. & J. Gauthier" pageId="27" pageNumber="28" pagination="83" refId="ref192716" refString="Joyce, W., and J. Gauthier. 2006. A nearly complete skeleton of Poposaurus gracilis from the Late Triassic of Utah. Journal of Vertebrate Paleontology 26: 83 A." type="journal article" year="2006">Joyce and Gauthier, 2006</bibRefCitation>
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), nearly all specimens of
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<taxonomicName authorityName="Mehl" authorityYear="1915" box="[1068,1193,1264,1287]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[1068,1193,1264,1287]" italics="true" pageId="27" pageNumber="28">P. gracilis</emphasis>
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</taxonomicName>
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consist of pelvic material, a few vertebrae, and partial limbs (
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<bibRefCitation author="Weinbaum, J. C. & A. Hungerbuhler" pageId="27" pageNumber="28" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
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||
).
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</paragraph>
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<paragraph blockId="27.[667,1194,257,1759]" lastBlockId="28.[128,654,212,1477]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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The element that was identified as the pubes in the
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<typeStatus box="[858,960,1413,1435]" pageId="27" pageNumber="28">holotype</typeStatus>
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(FMNH 357) is actually the ischium; therefore, the pubis is not represented in the
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<typeStatus box="[928,1030,1472,1494]" pageId="27" pageNumber="28">holotype</typeStatus>
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material. The element that was once identified at the pubis bears a large distal expansion (
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<emphasis bold="true" box="[1022,1045,1529,1552]" pageId="27" pageNumber="28">5</emphasis>
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pubic boot), and this expansion has greatly influenced the interpretation of its relationships in older (
|
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<bibRefCitation author="Colbert, E. H." box="[675,837,1619,1641]" pageId="27" pageNumber="28" pagination="59 - 78" refId="ref187654" refString="Colbert, E. H. 1961. The Triassic reptile Poposaurus. Fieldiana 14: 59 - 78." type="journal article" year="1961">Colbert, 1961</bibRefCitation>
|
||
) and more recent (
|
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<bibRefCitation author="Weinbaum, J. C. & A. Hungerbuhler" pageId="27" pageNumber="28" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
|
||
) studies. Indeed, the ischium bears a greatly enlarged distal expansion. Ironically, new specimens confirm that a large distal expansion (
|
||
<emphasis bold="true" box="[1005,1028,1736,1759]" pageId="27" pageNumber="28">5</emphasis>
|
||
pubic boot) is present in
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" box="[256,375,212,235]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
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||
<emphasis box="[256,375,212,235]" italics="true" pageId="28" pageNumber="29">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<accessionNumber box="[395,556,213,235]" httpUri="http://www.ncbi.nlm.nih.gov/protein/TMM43683" pageId="28" pageNumber="29">TMM 43683</accessionNumber>
|
||
-1;
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/YPM57100" pageId="28" pageNumber="29">YPM 57100</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="28.[128,654,212,1477]" pageId="28" pageNumber="29">
|
||
<bibRefCitation author="Dawley, R. M. & J. M. Zawiskie & J. W. Cosgriff" box="[157,412,272,295]" pageId="28" pageNumber="29" pagination="1428 - 1431" refId="ref188280" refString="Dawley, R. M., J. M. Zawiskie, and J. W. Cosgriff. 1979. A rauisuchid thecodont from the Upper Triassic Popo Agie Formation of Wyoming. Journal of Paleontology 53: 1428 - 1431." type="journal article" year="1979">Dawley et al. (1979)</bibRefCitation>
|
||
described
|
||
<taxonomicName class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
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||
, another ‘‘rauisuchian’’ from the same formation (Popo Agie Formation) as the
|
||
<typeStatus box="[128,230,361,383]" pageId="28" pageNumber="29">holotype</typeStatus>
|
||
of
|
||
<emphasis box="[276,403,360,383]" italics="true" pageId="28" pageNumber="29">
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" box="[276,399,360,383]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
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||
.
|
||
</emphasis>
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Later,
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<bibRefCitation author="Zawiskie, J. M. & R. M. Dawley" pageId="28" pageNumber="29" refId="ref203710" refString="Zawiskie, J. M., and R. M. Dawley. 2003. On the skull and holotype of Heptasuchus clarki (Rauisuchia, Poposauridae) from the Upper Triassic Popo Agie Formation, Natrona Co. Wyoming. Southwest Palreontological Symposium 2003 Guide to Presentations, no page numbers assigned." type="proceedings paper" year="2003">Zawiskie and Dawley (2003)</bibRefCitation>
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||
hypothesized that the skull of
|
||
<taxonomicName box="[128,273,419,442]" class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[128,273,419,442]" italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the body of
|
||
<emphasis italics="true" pageId="28" pageNumber="29">
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Although only a few elements (e.g., pubis, ulna) are directly comparable between the unique specimen of
|
||
<taxonomicName box="[418,563,508,531]" class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[418,563,508,531]" italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis italics="true" pageId="28" pageNumber="29">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, there are important differences between the pubes. Both taxa have a distal expansion of the pubis; however, the distal expansion in
|
||
<taxonomicName box="[309,454,626,649]" class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[309,454,626,649]" italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
|
||
is robust and rounded like that of
|
||
<taxonomicName box="[396,568,655,678]" class="Reptilia" genus="Batrachotomus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[396,568,655,678]" italics="true" pageId="28" pageNumber="29">Batrachotomus</emphasis>
|
||
</taxonomicName>
|
||
rather than the mediolaterally compressed distal expansion of
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" box="[304,429,715,738]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[304,429,715,738]" italics="true" pageId="28" pageNumber="29">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<accessionNumber box="[455,624,715,737]" httpUri="http://www.ncbi.nlm.nih.gov/protein/TMM43683" pageId="28" pageNumber="29">TMM 43683</accessionNumber>
|
||
-1). Furthermore, the preserved portions of the skull of
|
||
<taxonomicName box="[242,387,773,796]" class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[242,387,773,796]" italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
|
||
(maxilla, premaxilla, braincase) are much like that of
|
||
<taxonomicName class="Reptilia" genus="Batrachotomus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="28" pageNumber="29">Batrachotomus</emphasis>
|
||
</taxonomicName>
|
||
and not much like those of the putative close relatives of
|
||
<taxonomicName baseAuthorityName="Weinbaum and Hungerbuhler" baseAuthorityYear="2007" box="[339,470,863,884]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[339,470,863,884]" italics="true" pageId="28" pageNumber="29">Poposaurus</emphasis>
|
||
</taxonomicName>
|
||
such as
|
||
<taxonomicName class="Reptilia" family="Ctenosauriscidae" genus="Arizonasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="28" pageNumber="29">Arizonasaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="(Nesbitt, 2007)" baseAuthorityName="Nesbitt" baseAuthorityYear="2007" box="[303,582,892,914]" class="Reptilia" family="Shuvosauridae" genus="Effigia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[303,381,892,913]" italics="true" pageId="28" pageNumber="29">Effigia</emphasis>
|
||
(Nesbitt, 2007)
|
||
</taxonomicName>
|
||
. Furthermore, it is reasonable to assume that more than two paracrocodylomorph taxa exist in a single assemblage as demonstrated by the cooccurrence of
|
||
<taxonomicName baseAuthorityName="Chatterjee" baseAuthorityYear="1985" box="[451,590,1010,1033]" class="Reptilia" family="Rauisuchidae" genus="Postosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[451,590,1010,1033]" italics="true" pageId="28" pageNumber="29">Postosuchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Weinbaum and Hungerbuhler" baseAuthorityYear="2007" box="[128,259,1040,1061]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[128,259,1040,1061]" italics="true" pageId="28" pageNumber="29">Poposaurus</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName authority="Quarry (Long and Murry, 1995)" authorityName="Quarry (Long and Murry" authorityYear="1995" class="Reptilia" family="Kannemeyeriidae" genus="Placerias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[363,468,1039,1062]" italics="true" pageId="28" pageNumber="29">Placerias</emphasis>
|
||
Quarry (
|
||
<bibRefCitation author="Long, R. A. & P. A. Murry" pageId="28" pageNumber="29" pagination="1 - 254" refId="ref193609" refString="Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Chatterjee" baseAuthorityYear="1985" box="[449,588,1069,1092]" class="Reptilia" family="Rauisuchidae" genus="Postosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[449,588,1069,1092]" italics="true" pageId="28" pageNumber="29">Postosuchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="sensu Nesbitt" baseAuthorityYear="2007" box="[128,270,1098,1121]" class="Reptilia" family="Shuvosauridae" genus="Shuvosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[128,270,1098,1121]" italics="true" pageId="28" pageNumber="29">Shuvosaurus</emphasis>
|
||
</taxonomicName>
|
||
in the Post (
|
||
<emphasis bold="true" box="[440,463,1098,1121]" pageId="28" pageNumber="29">5</emphasis>
|
||
Miller) Quarry (
|
||
<bibRefCitation author="Long, R. A. & P. A. Murry" box="[136,445,1128,1150]" pageId="28" pageNumber="29" pagination="1 - 254" refId="ref193609" refString="Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
|
||
). Therefore, the hypothesis that
|
||
<taxonomicName box="[327,472,1157,1180]" class="Reptilia" family="Rauisuchidae" genus="Heptasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis box="[327,472,1157,1180]" italics="true" pageId="28" pageNumber="29">Heptasuchus</emphasis>
|
||
</taxonomicName>
|
||
represents the skull of
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" box="[225,342,1187,1210]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[225,342,1187,1210]" italics="true" pageId="28" pageNumber="29">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
is rejected here.
|
||
</paragraph>
|
||
<paragraph blockId="28.[128,654,212,1477]" pageId="28" pageNumber="29">
|
||
<taxonomicName authorityName="Mehl" authorityYear="1915" box="[157,381,1217,1240]" class="Reptilia" family="Poposauridae" genus="Poposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rauisuchiformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[157,381,1217,1240]" italics="true" pageId="28" pageNumber="29">Poposaurus gracilis</emphasis>
|
||
</taxonomicName>
|
||
possesses two autapomorphies: a thick lateral ridge posterior to the acetabulum and a pit on the proximal part of the ischium for reception of the convex ischial peduncle of the ilium (
|
||
<bibRefCitation author="Weinbaum, J. C. & A. Hungerbuhler" pageId="28" pageNumber="29" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="28" pageNumber="29" type="key">
|
||
<paragraph blockId="28.[128,654,212,1477]" pageId="28" pageNumber="29">
|
||
KEY REFERENCES:
|
||
<bibRefCitation author="Mehl, M. G." box="[396,538,1395,1418]" pageId="28" pageNumber="29" pagination="516 - 522" refId="ref194517" refString="Mehl, M. G. 1915. Poposaurus gracilis, a new reptile from the Triassic of Wyoming. Journal of Geology 23: 516 - 522." type="journal article" year="1915">Mehl, 1915</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Colbert, E. H." pageId="28" pageNumber="29" pagination="59 - 78" refId="ref187654" refString="Colbert, E. H. 1961. The Triassic reptile Poposaurus. Fieldiana 14: 59 - 78." type="journal article" year="1961">Colbert, 1961</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Galton, P. M." box="[199,358,1425,1447]" pageId="28" pageNumber="29" pagination="234 - 245" refId="ref189223" refString="Galton, P. M. 1977. On Staurikosaurus pricei, an early saurischian dinosaur from the Triassic of Brazil, with notes on the Herrerasauridae and Poposauridae. Palaontologische Zeitschrift 51: 234 - 245." type="journal article" year="1977">Galton, 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Long, R. A. & P. A. Murry" box="[371,650,1425,1447]" pageId="28" pageNumber="29" pagination="1 - 254" refId="ref193609" refString="Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Weinbaum, J. C. & A. Hungerbuhler" box="[128,556,1455,1477]" pageId="28" pageNumber="29" pagination="131 - 145" refId="ref202626" refString="Weinbaum, J. C., and A. Hungerbuhler. 2007. A revision of Poposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U. S. A. Palaontologische Zeitschrift 81 / 2: 131 - 145." type="journal article" year="2007">Weinbaum and Hungerbühler, 2007</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |