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<mods:title id="2CE223169A0991BC64164636AC5D55DB">Systematic Revision Of The Andean Scorpion Genus Orobothriurus Maury, 1976 (Bothriuridae), With Discussion Of The Altitude Record For Scorpions</mods:title>
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<taxonomicName id="47984FD41F4DFF921A9FFCC8FE0D4129" ID-CoL="6SVW2" authority="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini, 2011" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[156,475,769,792]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana" status="sp. nov.">
<emphasis id="B2ECE8451F4DFF921A9FFCC8FE0D4129" bold="true" box="[156,475,769,792]" italics="true" pageId="70" pageNumber="70">Orobothriurus quewerukana</emphasis>
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,
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<figureCitation id="18A328D21F4DFF921A65FCE8FF214107" box="[102,247,800,823]" captionStart="Fig" captionStartId="27.[150,187,1252,1271]" captionTargetBox="[150,1206,212,1221]" captionTargetId="figure-196@27.[150,1207,212,1222]" captionTargetPageId="27" captionText="Fig.15. Orobothriurus Maury, 1976, carapace, dorsal aspect.A. Orobothriurus paessleri (Kraepelin, 1911), ♂ (CDA 186). B. Orobothriurus alticola (Pocock, 1899), ♂ (AMNH). C. Orobothriurus quewerukana, n. sp., paratype ♂ (AMNH). D. Orobothriurus tamarugal, n. sp., holotype ♂ (MACN-Ar). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611161" httpUri="https://zenodo.org/record/4611161/files/figure.png" pageId="70" pageNumber="70">Figures 15C</figureCitation>
,
<figureCitation id="18A328D21F4DFF921B06FCE8FEE14107" box="[261,311,801,822]" captionStart="Fig" captionStartId="31.[150,188,1526,1545]" captionTargetBox="[150,1206,211,1496]" captionTargetId="figure-1@31.[150,1207,210,1496]" captionTargetPageId="31" captionText="Fig. 19. Orobothriurus Maury, 1976, sternite VII and metasomal segments I and II, ventral aspect. A. Orobothriurus curvidigitus (Kraepelin, 1911), ♀ (MHNC). B. Orobothriurus paessleri (Kraepelin, 1911), ♀ (MHNC). C. Orobothriurus quewerukana, n. sp., paratype ♂ (MHNC). D. Orobothriurus huascaran, n. sp., paratype ♀ (MACN-Ar). E. Orobothriurus wawita Acosta and Ochoa, 2000, paratype ♀ (MHNC). F. Orobothriurus tamarugal, n. sp., holotype ♂ (MACN-Ar). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611174" httpUri="https://zenodo.org/record/4611174/files/figure.png" pageId="70" pageNumber="70">19C</figureCitation>
,
<figureCitation id="18A328D21F4DFF921B47FCE8FEAF4107" box="[324,377,801,822]" captionStart="Fig" captionStartId="32.[116,155,1617,1636]" captionTargetBox="[140,1145,212,1586]" captionTargetId="figure-1@32.[140,1146,212,1587]" captionTargetPageId="32" captionText="Fig. 20. Orobothriurus Maury, 1976, metasomal segment V, ventral aspect. A. Orobothriurus alticola (Pocock, 1899), ♀ (MACN-Ar). B. Orobothriurus compagnuccii, n. sp., paratype ♂ (MACN-Ar). C. Orobothriurus ramirezi, n. sp., paratype ♂ (MACN-Ar). D. Orobothriurus quewerukana, n. sp., paratype ♂ (MHNC). E. Orobothriurus tamarugal, n. sp., holotype ♂ (MACN-Ar). F. O. ramirezi, n. sp., paratype ♀ (MACN-Ar). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611178" httpUri="https://zenodo.org/record/4611178/files/figure.png" pageId="70" pageNumber="70">20D</figureCitation>
,
<figureCitation id="18A328D21F4DFF921B84FCE8FE614107" box="[391,439,801,822]" captionStart="Fig" captionStartId="36.[116,155,1588,1607]" captionTargetBox="[131,1155,211,1557]" captionTargetId="figure-1@36.[131,1156,210,1558]" captionTargetPageId="36" captionText="Fig. 23. Orobothriurus Maury, 1976, telson, lateral aspect. A. Orobothriurus alticola (Pocock, 1899), ♂ (MACN-Ar). B. Orobothriurus calchaqui, n. sp., paratype ♂ (MACN-Ar). C. Orobothriurus compagnuccii, n. sp., paratype ♂ (MACN-Ar). D. Orobothriurus huascaran, n. sp., paratype ♂ (MACN-Ar). E. Orobothriurus grismadoi Ojanguren Afillastro et al., 2009, paratype ♂ (MACN-Ar). F. Orobothriurus quewerukana, n. sp., paratype ♂ (MHNC). G. Orobothriurus ramirezi, n. sp., paratype ♂ (MACN-Ar). H. Orobothriurus tamarugal, n. sp., holotype ♂ (MACN-Ar). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611190" httpUri="https://zenodo.org/record/4611190/files/figure.png" pageId="70" pageNumber="70">23F</figureCitation>
,
<figureCitation id="18A328D21F4DFF921BC6FCE8FE2F4107" box="[453,505,801,822]" captionStart="Fig" captionStartId="37.[150,188,1617,1636]" captionTargetBox="[207,1148,212,1586]" captionTargetId="figure-1@37.[207,1148,212,1587]" captionTargetPageId="37" captionText="Fig. 24. Orobothriurus Maury, 1976, telson, lateral aspect. A. Orobothriurus calchaqui, n. sp., paratype ♀ (MACN-Ar). B. Orobothriurus compagnuccii, n. sp., paratype ♀ (MACN-Ar). C. Orobothriurus huascaran, n. sp., paratype ♀ (MACN-Ar). D. Orobothriurus quewerukana, n. sp., paratype ♀ (MHNC). E. Orobothriurus ramirezi, n. sp., paratype ♀ (MACN-Ar). F. Orobothriurus tamarugal, n. sp., paratype ♀ (MZUC). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611192" httpUri="https://zenodo.org/record/4611192/files/figure.png" pageId="70" pageNumber="70">24D</figureCitation>
,
<figureCitation id="18A328D21F4DFF921804FCE8FD884107" box="[519,606,801,822]" captionStart="Fig" captionStartId="56.[116,155,1166,1185]" captionTargetBox="[127,1159,211,1135]" captionTargetId="figure-216@56.[127,1160,210,1136]" captionTargetPageId="56" captionText="Fig. 40. Orobothriurus Maury, 1976, sinistral hemispermatophores. AC. Orobothriurus huascaran, n. sp., paratype ♂ (MACN-Ar). D, E. Orobothriurus quewerukana, n. sp., paratype ♂ (MACN-Ar). A, D. Ental aspect. B, E. Ectal aspect. C. Dorsal aspect. Scale bar 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611244" httpUri="https://zenodo.org/record/4611244/files/figure.png" pageId="70" pageNumber="70">40D, E</figureCitation>
,
<figureCitation id="18A328D21F4DFF921AFFFCF7FE924162" box="[252,324,830,851]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="57.[150,188,1676,1695]" captionStartId-1="60.[689,728,988,1007]" captionStartId-2="61.[150,188,1670,1689]" captionTargetBox-0="[178,1177,211,1645]" captionTargetBox-1="[670,1190,211,957]" captionTargetBox-2="[214,1142,212,1640]" captionTargetId-0="figure-1@57.[178,1177,210,1646]" captionTargetId-1="figure-643@60.[670,1190,210,958]" captionTargetId-2="figure-1@61.[214,1142,212,1640]" captionTargetPageId-0="57" captionTargetPageId-1="60" captionTargetPageId-2="61" captionText-0="Fig. 41. Orobothriurus quewerukana, n. sp., habitus. A, B. Holotype ♂ (AMNH). C, D. Paratype ♀ (MHNC). A, C. Dorsal aspect. B, D. Ventral aspect. Scale bars 5 10 mm." captionText-1="Fig. 42. Orobothriurus quewerukana, n. sp., paratype ♂ (MHNC), dextral pedipalp femur (A) and patella (BD). A, B. Dorsal aspect. C. External aspect. D. Ventral aspect. Scale bar 5 1 mm." captionText-2="Fig. 43. Orobothriurus quewerukana, n. sp., dextral pedipalp chela. A, B. Paratype ♀ (MHNC). CE. Paratype ♂ (MHNC). A. Dorsal aspect. B, C. External aspect. D. Ventral aspect. E. Ventrointernal aspect. Scale bar 5 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4611248" figureDoi-1="http://doi.org/10.5281/zenodo.4611250" figureDoi-2="http://doi.org/10.5281/zenodo.4611254" httpUri-0="https://zenodo.org/record/4611248/files/figure.png" httpUri-1="https://zenodo.org/record/4611250/files/figure.png" httpUri-2="https://zenodo.org/record/4611254/files/figure.png" pageId="70" pageNumber="70">4143</figureCitation>
,
<figureCitation id="18A328D21F4DFF921B52FCF7FEB94165" box="[337,367,830,852]" captionStart="Fig" captionStartId="81.[150,189,937,956]" captionTargetBox="[147,1207,212,907]" captionTargetId="figure-373@81.[147,1207,212,907]" captionTargetPageId="81" captionText="Fig. 53. Orobothriurus Maury, 1976, locality records in southern Peru and northern Chile. Orobothriurus atiquipa Ochoa and Acosta, 2002, square; Orobothriurus curvidigitus (Kraepelin, 1911), circles; Orobothriurus paessleri (Kraepelin, 1911), crosses; Orobothriurus quewerukana, n. sp., triangles; Orobothriurus tamarugal, n. sp., diamonds." figureDoi="http://doi.org/10.5281/zenodo.4611286" httpUri="https://zenodo.org/record/4611286/files/figure.png" pageId="70" pageNumber="70">53</figureCitation>
; table 3
</paragraph>
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<treatmentCitation id="013912461F4DFF921A5EFCA2FDD2414F" author="Maury, E. A." box="[93,516,875,895]" page="21" pageId="70" pageNumber="70" year="1976">
<taxonomicName id="47984FD41F4DFF921A5EFCA2FDD2414F" authority=": Maury, 1976: 21" authorityName="Maury" authorityPageNumber="21" authorityYear="1976" box="[93,516,875,895]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="dumayi">
<emphasis id="B2ECE8451F4DFF921A5EFCA2FE97414E" box="[93,321,875,895]" italics="true" pageId="70" pageNumber="70">Orobothriurus dumayi</emphasis>
:
<bibRefCitation id="E40949A61F4DFF921B52FCA5FDD2414F" author="Maury, E. A." box="[337,516,875,895]" pageId="70" pageNumber="70" pagination="14 - 25" refId="ref44084" refString="Maury, E. A. 1976. Escorpiones y escorpionismo en el Peru V. Orobothriurus, un nuevo genero de escorpiones altoandinos (Bothriuridae). Revista Peruana de Entomologia 18 (1): 14 - 25." type="journal article" year="1976">Maury, 1976: 21</bibRefCitation>
</taxonomicName>
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, figs. 40 44;
<treatmentCitation id="013912461F4DFF921AA0FC4FFE2641A8" author="Acosta, L. E. &amp; J. A. Ochoa" box="[163,496,902,921]" page="136" pageId="70" pageNumber="70" year="2000">
<bibRefCitation id="E40949A61F4DFF921AA0FC4FFE2641A8" author="Acosta, L. E. &amp; J. A. Ochoa" box="[163,496,902,921]" pageId="70" pageNumber="70" pagination="135 - 144" refId="ref41945" refString="Acosta, L. E., and J. A. Ochoa. 2000. Nueva especie de Orobothriurus, Maury, del Peru. (Scorpiones, Bothriuridae). Revue Arachnologique 13 (10): 135 - 144." type="journal article" year="2000">Acosta and Ochoa, 2000: 136</bibRefCitation>
</treatmentCitation>
, 143. [misidentification].
</treatmentCitationGroup>
</paragraph>
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<materialsCitation id="30F03E0A1F4DFF921A79FC02FE9746FF" ID-GBIF-Occurrence="3053988311" collectingDate="1967-08-08" collectingDateMax="2005-01-16" collectingDateMin="1967-08-08" collectionCode="AMNH" country="Chile" county="Parinacota Province" elevation="3732" latitude="-69.0" location="Iquique Province" municipality="Putre" pageId="70" pageNumber="70" specimenCount="4" typeStatus="holotype">
TYPE MATERIAL:
<emphasis id="B2ECE8451F4DFF921B56FC04FF794631" bold="true" pageId="70" pageNumber="70">
<collectingCountry id="F88F74C71F4DFF921B56FC04FE6741D2" box="[341,433,973,995]" name="Chile" pageId="70" pageNumber="70">CHILE</collectingCountry>
:
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(Tarapaca´):
</emphasis>
<emphasis id="B2ECE8451F4DFF921AC0FC22FE614631" box="[195,439,1003,1024]" italics="true" pageId="70" pageNumber="70">
<collectingCounty id="69464CDB1F4DFF921AC0FC22FE614631" box="[195,439,1003,1024]" pageId="70" pageNumber="70">Parinacota Province</collectingCounty>
</emphasis>
:
<typeStatus id="5F238AF51F4DFF921BD1FC23FD964631" box="[466,576,1002,1024]" pageId="70" pageNumber="70" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="E689AC921F4DFF921A66FBC1FF10462F" box="[101,198,1032,1054]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
),
<collectingMunicipality id="6043AE2D1F4DFF921AEAFBC1FEFD462C" box="[233,299,1032,1053]" pageId="70" pageNumber="70">Putre</collectingMunicipality>
,
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W, 18
<emphasis id="B2ECE8451F4DFF921BF7FBC1FE29462C" bold="true" box="[500,511,1032,1053]" pageId="70" pageNumber="70">°</emphasis>
13926.80S 69
<emphasis id="B2ECE8451F4DFF921A7BFBECFF55460B" bold="true" box="[120,131,1061,1082]" pageId="70" pageNumber="70">°</emphasis>
32947.10W,
<quantity id="476099B21F4DFF921B08FBECFEB5460A" box="[267,355,1061,1083]" metricMagnitude="3" metricUnit="m" metricValue="3.732" pageId="70" pageNumber="70" unit="m" value="3732.0">
<elevation id="0BB5D3641F4DFF921B08FBECFEB5460A" box="[267,355,1061,1083]" metricMagnitude="3" metricUnit="m" metricValue="3.732" pageId="70" pageNumber="70" unit="m" value="3732.0">3732 m</elevation>
</quantity>
,
<date id="F42612971F4DFF921B73FBECFE0D460A" box="[368,475,1061,1083]" pageId="70" pageNumber="70" value="2005-01-16">
<collectingDate id="E462EB7F1F4DFF921B73FBECFE0D460A" box="[368,475,1061,1083]" pageId="70" pageNumber="70" value="2005-01-16">16.i.2005</collectingDate>
</date>
,
<collectorName id="2D6D51811F4DFF921BE4FBECFF294669" pageId="70" pageNumber="70">A. Ojanguren Affilastro</collectorName>
,
<collectorName id="2D6D51811F4DFF921B0FFB8AFE774669" box="[268,417,1090,1112]" pageId="70" pageNumber="70">C.I. Mattoni</collectorName>
and
<collectorName id="2D6D51811F4DFF921BDCFB8AFDB14669" box="[479,615,1090,1112]" pageId="70" pageNumber="70">J.A. Ochoa</collectorName>
, UV and under stones.
<typeStatus id="5F238AF51F4DFF921B81FBA9FE2E4644" box="[386,504,1120,1141]" pageId="70" pageNumber="70" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B2ECE8451F4DFF921808FBA9FE8046A2" bold="true" pageId="70" pageNumber="70">
<collectingCountry id="F88F74C71F4DFF921808FBA9FDB04647" box="[523,614,1120,1142]" name="Chile" pageId="70" pageNumber="70">CHILE</collectingCountry>
:
<collectorName id="2D6D51811F4DFF921A5EFBB5FF1246A3" box="[93,196,1148,1171]" pageId="70" pageNumber="70">Región I</collectorName>
(Tarapaca´):
</emphasis>
<emphasis id="B2ECE8451F4DFF921B61FBB7FDF046A2" box="[354,550,1150,1171]" italics="true" pageId="70" pageNumber="70">
<location id="8547628C1F4DFF921B61FBB7FDF046A2" LSID="urn:lsid:plazi:treatment:083185411F4DFF9E1A9FFCC8FB8047F4:8547628C1F4DFF921B61FBB7FDF046A2" box="[354,550,1150,1171]" country="Chile" county="Parinacota Province" latitude="-69.0" municipality="Putre" name="Iquique Province" pageId="70" pageNumber="70">Iquique Province</location>
</emphasis>
:
<location id="8547628C1F4DFF92183AFBB7FECD4680" LSID="urn:lsid:plazi:treatment:083185411F4DFF9E1A9FFCC8FB8047F4:8547628C1F4DFF92183AFBB7FECD4680" country="Chile" county="Parinacota Province" latitude="-69.0" municipality="Putre" name="Termas de Mamina" pageId="70" pageNumber="70">
Termas
<collectorName id="2D6D51811F4DFF921A90FB52FECD4680" box="[147,283,1179,1201]" pageId="70" pageNumber="70">de Mamiña</collectorName>
</location>
,
<date id="F42612971F4DFF921B24FB52FE764680" box="[295,416,1179,1201]" pageId="70" pageNumber="70" value="1967-08-08">
<collectingDate id="E462EB7F1F4DFF921B24FB52FE764680" box="[295,416,1179,1201]" pageId="70" pageNumber="70" value="1967-08-08">8.viii.1967</collectingDate>
</date>
,
<collectorName id="2D6D51811F4DFF921BA8FB52FDC94680" box="[427,543,1179,1201]" pageId="70" pageNumber="70">J.C. Ortiz</collectorName>
,
<specimenCount id="969EFFDE1F4DFF921828FB52FDBA4680" box="[555,620,1179,1201]" pageId="70" pageNumber="70" type="generic">4 juv.</specimenCount>
(MACN-Ar 6846)
</materialsCitation>
;
<materialsCitation id="30F03E0A1F4DFF921B5AFB71FE1F447A" ID-GBIF-Occurrence="3053988363" collectingDate="2000-02-26" collectingDateMax="2005-01-19" collectingDateMin="2000-02-26" collectionCode="LBRE, MHNC, MZUC, AMNH" country="Peru" county="Mamina" elevation="2822" latitude="-69.0" location="Tacna Province" municipality="Parinacota Province" pageId="70" pageNumber="70" specimenCount="55" specimenCount-female="14" stateProvince="Tacna" typeStatus="holotype">
<collectingCounty id="69464CDB1F4DFF921B5AFB71FE6946FF" box="[345,447,1208,1230]" pageId="70" pageNumber="70">Mamiña</collectingCounty>
, 20
<emphasis id="B2ECE8451F4DFF921BF7FB71FE2946FC" bold="true" box="[500,511,1208,1229]" pageId="70" pageNumber="70">°</emphasis>
04933.60S 69
<emphasis id="B2ECE8451F4DFF921A7BFB1CFF5546DB" bold="true" box="[120,131,1237,1258]" pageId="70" pageNumber="70">°</emphasis>
12905.40W,
<quantity id="476099B21F4DFF921B08FB1CFEB546DA" box="[267,355,1237,1259]" metricMagnitude="3" metricUnit="m" metricValue="2.822" pageId="70" pageNumber="70" unit="m" value="2822.0">
<elevation id="0BB5D3641F4DFF921B08FB1CFEB546DA" box="[267,355,1237,1259]" metricMagnitude="3" metricUnit="m" metricValue="2.822" pageId="70" pageNumber="70" unit="m" value="2822.0">2822 m</elevation>
</quantity>
,
<date id="F42612971F4DFF921B73FB1CFE0C46DA" box="[368,474,1237,1259]" pageId="70" pageNumber="70" value="2005-01-19">
<collectingDate id="E462EB7F1F4DFF921B73FB1CFE0C46DA" box="[368,474,1237,1259]" pageId="70" pageNumber="70" value="2005-01-19">19.i.2005</collectingDate>
</date>
,
<collectorName id="2D6D51811F4DFF921BE4FB1CFF294738" pageId="70" pageNumber="70">A. Ojanguren Affilastro</collectorName>
,
<collectorName id="2D6D51811F4DFF921B0FFB3AFE774738" box="[268,417,1267,1289]" pageId="70" pageNumber="70">C.I. Mattoni</collectorName>
and
<collectorName id="2D6D51811F4DFF921BDDFB3AFDB14738" box="[478,615,1267,1289]" pageId="70" pageNumber="70">J.A. Ochoa</collectorName>
, UV and under stones, small wet valley with old terrace cultivation, 2 ♂,
<specimenCount id="969EFFDE1F4DFF921BCDFAE4FE284775" box="[462,510,1322,1348]" pageId="70" pageNumber="70" type="female">2 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921816FAE4FDBA4772" box="[533,620,1325,1347]" pageId="70" pageNumber="70" type="generic">14 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF921A66FA82FF634750" box="[101,181,1355,1377]" pageId="70" pageNumber="70">LBRE</collectionCode>
), 2 ♂,
<specimenCount id="969EFFDE1F4DFF921B00FA82FEFD4750" box="[259,299,1351,1377]" pageId="70" pageNumber="70" type="female">2 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921B39FA82FEAC4750" box="[314,378,1355,1377]" pageId="70" pageNumber="70" type="generic">8 juv.</specimenCount>
(MACN-Ar), 2 ♂,
<specimenCount id="969EFFDE1F4DFF92185EFA82FFA7474E" pageId="70" pageNumber="70" type="female">1 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921A85FAA1FED8474E" box="[134,270,1381,1407]" pageId="70" pageNumber="70" type="female">1 subad. ♀</specimenCount>
(
<collectionCode id="E689AC921F4DFF921B20FAA1FE55474F" box="[291,387,1384,1406]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="70" pageNumber="70">MHNC</collectionCode>
), 1 ♂,
<specimenCount id="969EFFDE1F4DFF921BE1FAA1FDC7474E" box="[482,529,1381,1407]" pageId="70" pageNumber="70" type="female">1 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921826FAA1FDBA474F" box="[549,620,1384,1406]" pageId="70" pageNumber="70" type="generic">6 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF921A66FA4CFF1447AA" box="[101,194,1413,1435]" country="Chile" name="Museo de Zoologia, Universidad de Concepcion" pageId="70" pageNumber="70">MZUC</collectionCode>
).
<emphasis id="B2ECE8451F4DFF921AD4FA4FFE1747AA" box="[215,449,1414,1435]" italics="true" pageId="70" pageNumber="70">
<collectingMunicipality id="6043AE2D1F4DFF921AD4FA4FFE1747AA" box="[215,449,1414,1435]" pageId="70" pageNumber="70">Parinacota Province</collectingMunicipality>
</emphasis>
: same data as holotype, 2 ♂,
<specimenCount id="969EFFDE1F4DFF921B03FA6AFEF2478B" box="[256,292,1440,1466]" pageId="70" pageNumber="70" type="female">2♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921B2CFA6AFEBD4788" box="[303,363,1443,1465]" pageId="70" pageNumber="70" type="generic">6juv.</specimenCount>
(MACN-Ar),2 ♂,
<specimenCount id="969EFFDE1F4DFF92183CFA6AFDB5478B" box="[575,611,1440,1466]" pageId="70" pageNumber="70" type="female">1♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921A5EFA09FF7647E7" box="[93,160,1472,1494]" pageId="70" pageNumber="70" type="generic">3 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF921AB2FA09FED747E7" box="[177,257,1472,1494]" pageId="70" pageNumber="70">LBRE</collectionCode>
), 1 ♂,
<specimenCount id="969EFFDE1F4DFF921B54FA09FE5447E6" box="[343,386,1469,1495]" pageId="70" pageNumber="70" type="female">1 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921B90FA09FE0047E7" box="[403,470,1472,1494]" pageId="70" pageNumber="70" type="generic">3 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF921BE4FA09FD9147E7" box="[487,583,1472,1494]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="70" pageNumber="70">MHNC</collectionCode>
),
<specimenCount id="969EFFDE1F4DFF92185EFA09FF7C47C2" pageId="70" pageNumber="70" type="generic">1 subad.</specimenCount>
♂ (
<collectionCode id="E689AC921F4DFF921AE0FA14FE9347C2" box="[227,325,1501,1523]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
).
<emphasis id="B2ECE8451F4DFF921B5DFA14FF4A4421" bold="true" pageId="70" pageNumber="70">
<collectingCountry id="F88F74C71F4DFF921B5DFA14FE7847C3" box="[350,430,1501,1522]" name="Peru" pageId="70" pageNumber="70">PERU</collectingCountry>
:
<collectingRegion id="425CFAB51F4DFF921BBCFA14FDD047C3" box="[447,518,1501,1522]" country="Peru" name="Tacna" pageId="70" pageNumber="70">Tacna</collectingRegion>
Department:
</emphasis>
<emphasis id="B2ECE8451F4DFF921AB0FA32FEA54421" box="[179,371,1531,1552]" italics="true" pageId="70" pageNumber="70">
<location id="8547628C1F4DFF921AB0FA32FEA54421" LSID="urn:lsid:plazi:treatment:083185411F4DFF9E1A9FFCC8FB8047F4:8547628C1F4DFF921AB0FA32FEA54421" box="[179,371,1531,1552]" country="Peru" county="Mamina" latitude="-69.0" municipality="Parinacota Province" name="Tacna Province" pageId="70" pageNumber="70" stateProvince="Tacna">Tacna Province</location>
</emphasis>
: Palca, 17
<emphasis id="B2ECE8451F4DFF92180AFA32FDC24421" bold="true" box="[521,532,1531,1552]" pageId="70" pageNumber="70">°</emphasis>
469450S 69
<emphasis id="B2ECE8451F4DFF921A7BF9D1FF55441C" bold="true" box="[120,131,1560,1581]" pageId="70" pageNumber="70">°</emphasis>
579040W,
<quantity id="476099B21F4DFF921AF0F9D1FE9E441C" box="[243,328,1560,1581]" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" pageId="70" pageNumber="70" unit="m" value="3230.0">
<elevation id="0BB5D3641F4DFF921AF0F9D1FE9E441C" box="[243,328,1560,1581]" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" pageId="70" pageNumber="70" unit="m" value="3230.0">3230 m</elevation>
</quantity>
,
<date id="F42612971F4DFF921B50F9D1FE13441F" box="[339,453,1560,1582]" pageId="70" pageNumber="70" value="2000-02-26">
<collectingDate id="E462EB7F1F4DFF921B50F9D1FE13441F" box="[339,453,1560,1582]" pageId="70" pageNumber="70" value="2000-02-26">26.ii.2000</collectingDate>
</date>
,
<collectorName id="2D6D51811F4DFF921BCDF9D1FD82441F" box="[462,596,1560,1582]" pageId="70" pageNumber="70">J.A. Ochoa</collectorName>
,
<specimenCount id="969EFFDE1F4DFF92185EF9D1FFA4447D" pageId="70" pageNumber="70" type="female">1 ♀</specimenCount>
(
<collectionCode id="E689AC921F4DFF921A7CF9FCFF37447A" box="[127,225,1589,1611]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
), 1 ♂,
<specimenCount id="969EFFDE1F4DFF921B29F9FCFE84447D" box="[298,338,1586,1612]" pageId="70" pageNumber="70" type="female">2 ♀</specimenCount>
(
<collectionCode id="E689AC921F4DFF921B63F9FCFE17447A" box="[352,449,1589,1611]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="70" pageNumber="70">MHNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="802734571F4DFF921A79F99BFDBD44EE" blockId="70.[93,620,971,1759]" pageId="70" pageNumber="70">
ADDITIONAL MATERIAL:
<emphasis id="B2ECE8451F4DFF921BCEF99AFEFB44B6" bold="true" pageId="70" pageNumber="70">
<collectingCountry id="F88F74C71F4DFF921BCEF99AFDFE4458" box="[461,552,1619,1641]" name="Chile" pageId="70" pageNumber="70">CHILE</collectingCountry>
: Región I (Tarapaca´):
</emphasis>
<emphasis id="B2ECE8451F4DFF921B37F9BBFE2444B6" box="[308,498,1650,1671]" italics="true" pageId="70" pageNumber="70">Iquique Province</emphasis>
: Mamiña, 20
<emphasis id="B2ECE8451F4DFF921A7BF947FF554492" bold="true" box="[120,131,1678,1699]" pageId="70" pageNumber="70">°</emphasis>
04933.60
<geoCoordinate id="E5AC52901F4DFF921AE3F946FECC4492" box="[224,282,1678,1700]" degrees="69" direction="south" orientation="latitude" pageId="70" pageNumber="70" precision="55555" value="-69.0">
S 69
<emphasis id="B2ECE8451F4DFF921B0CF947FECC4492" bold="true" box="[271,282,1678,1699]" pageId="70" pageNumber="70">°</emphasis>
</geoCoordinate>
12905.40W,
<quantity id="476099B21F4DFF921B9DF946FE244495" box="[414,498,1679,1700]" metricMagnitude="3" metricUnit="m" metricValue="2.822" pageId="70" pageNumber="70" unit="m" value="2822.0">2822 m</quantity>
,
<date id="F42612971F4DFF921BFFF947FDB04495" box="[508,614,1678,1700]" pageId="70" pageNumber="70" value="2005-01-19">19.i.2005</date>
, A. Ojanguren Affilastro, C.I. Mattoni, and J.A. Ochoa, UV and under stones, small wet
</paragraph>
<paragraph id="802734571F4DFF921898FF1CFCC743E4" blockId="70.[667,1194,207,1759]" pageId="70" pageNumber="70">
valleywith oldterrace cultivation,
<specimenCount id="969EFFDE1F4DFF921E1AFF1CFBA142DA" box="[1049,1143,213,235]" pageId="70" pageNumber="70" type="generic">1 subad.</specimenCount>
♂,
<specimenCount id="969EFFDE1F4DFF921E98FF1CFD294338" pageId="70" pageNumber="70" type="female">1 subad. ♀</specimenCount>
,
<specimenCount id="969EFFDE1F4DFF921909FF3BFC914339" box="[778,839,242,264]" pageId="70" pageNumber="70" type="generic">2 juv.</specimenCount>
♂,
<specimenCount id="969EFFDE1F4DFF92196FFF3BFC174338" box="[876,961,239,265]" pageId="70" pageNumber="70" type="female">2 juv. ♀</specimenCount>
(
<collectionCode id="E689AC921F4DFF9219CFFF3BFBFC4336" box="[972,1066,242,263]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
[LP 4307]),
<specimenCount id="969EFFDE1F4DFF921898FEC6FD0B4314" box="[667,733,271,293]" pageId="70" pageNumber="70" type="generic">5 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF9218EEFEC6FC984314" box="[749,846,271,293]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
),
<specimenCount id="969EFFDE1F4DFF921967FEC6FC704314" box="[868,934,271,293]" pageId="70" pageNumber="70" type="generic">5 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF9219B4FEC6FBCE4314" box="[951,1048,271,293]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="70" pageNumber="70">MHNC</collectionCode>
).
<emphasis id="B2ECE8451F4DFF921E2FFED9FD284373" italics="true" pageId="70" pageNumber="70">Parinacota Province</emphasis>
: Putre,
<quantity id="476099B21F4DFF921979FEE5FC104372" box="[890,966,300,323]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="70" pageNumber="70" unit="km" value="6.0">6 km</quantity>
W, 18
<emphasis id="B2ECE8451F4DFF921E31FEE5FBEB4370" bold="true" box="[1074,1085,300,321]" pageId="70" pageNumber="70">°</emphasis>
13926.80
<geoCoordinate id="E5AC52901F4DFF921E99FEE4FD17436E" degrees="69" direction="south" orientation="latitude" pageId="70" pageNumber="70" precision="55555" value="-69.0">
S 69
<emphasis id="B2ECE8451F4DFF9218B5FE83FD17436E" bold="true" box="[694,705,330,351]" pageId="70" pageNumber="70">°</emphasis>
</geoCoordinate>
32947.10W,
<quantity id="476099B21F4DFF921940FE83FC43436E" box="[835,917,330,351]" metricMagnitude="3" metricUnit="m" metricValue="3.732" pageId="70" pageNumber="70" unit="m" value="3732.0">3732m</quantity>
,
<date id="F42612971F4DFF92199EFE83FBD14351" box="[925,1031,330,352]" pageId="70" pageNumber="70" value="2005-01-16">16.i.2005</date>
,A. Ojanguren Affilastro, C.I. Mattoni and J.A. Ochoa, UV and under stones,
<specimenCount id="969EFFDE1F4DFF921964FE4CFC1043AB" box="[871,966,388,410]" pageId="70" pageNumber="70" type="generic">1 subad.</specimenCount>
♂,
<specimenCount id="969EFFDE1F4DFF9219EEFE4CFBFD43AB" box="[1005,1067,388,410]" pageId="70" pageNumber="70" type="generic">4 juv.</specimenCount>
♂,
<specimenCount id="969EFFDE1F4DFF921E51FE4CFB7F43AA" box="[1106,1193,385,411]" pageId="70" pageNumber="70" type="female">1 juv. ♀</specimenCount>
(
<collectionCode id="E689AC921F4DFF9218A0FE6BFCD74386" box="[675,769,418,439]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
[LP 4306]),
<specimenCount id="969EFFDE1F4DFF92199CFE6BFC304389" box="[927,998,418,440]" pageId="70" pageNumber="70" type="generic">4 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF9219F9FE6BFB9C4389" box="[1018,1098,418,440]" pageId="70" pageNumber="70">LBRE</collectionCode>
),
<specimenCount id="969EFFDE1F4DFF921E60FE6BFB7F4389" box="[1123,1193,418,440]" pageId="70" pageNumber="70" type="generic">3 juv.</specimenCount>
(
<collectionCode id="E689AC921F4DFF9218A0FE76FCD243E4" box="[675,772,447,469]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="70" pageNumber="70">MHNC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F4DFF9218BBFE12FBC840B7" pageId="70" pageNumber="70" type="etymology">
<paragraph id="802734571F4DFF9218BBFE12FBC840B7" blockId="70.[667,1194,207,1759]" pageId="70" pageNumber="70">
ETYMOLOGY: The specific name is a noun in apposition composed of two Quechua words,
<emphasis id="B2ECE8451F4DFF9218FAFDD1FCE8401C" box="[761,830,536,557]" italics="true" pageId="70" pageNumber="70">quewe</emphasis>
, meaning curved, and
<emphasis id="B2ECE8451F4DFF921E8BFDD1FD04407A" italics="true" pageId="70" pageNumber="70">rukana</emphasis>
, meaning finger. It refers the strong curvature of the movable finger of the pedipalp chela in the adult male.
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F4DFF9318BBFD45FD884057" lastPageId="71" lastPageNumber="71" pageId="70" pageNumber="70" type="diagnosis">
<paragraph id="802734571F4DFF9318BBFD45FD884057" blockId="70.[667,1194,207,1759]" lastBlockId="71.[128,654,212,1759]" lastPageId="71" lastPageNumber="71" pageId="70" pageNumber="70">
DIAGNOSIS:
<taxonomicName id="47984FD41F4DFF921958FD44FB7F4095" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[859,1193,653,676]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF921958FD44FB7F4095" box="[859,1193,653,676]" italics="true" pageId="70" pageNumber="70">Orobothriurus quewerukana</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="47984FD41F4DFF92199AFD65FB9040F0" authorityName=": Lourenco and Dastych" authorityYear="2001" box="[921,1094,682,705]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F4DFF92199AFD65FB9040F0" box="[921,1094,682,705]" italics="true" pageId="70" pageNumber="70">O. curvidigitus</emphasis>
</taxonomicName>
and
<taxonomicName id="47984FD41F4DFF921E8FFD65FCD940EE" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4DFF921E8FFD65FCD940EE" italics="true" pageId="70" pageNumber="70">O. tamarugal</emphasis>
</taxonomicName>
(fig. 5), with which it shares the following characters: anterior margin of carapace with weak median projection (epistome; figs. 14C, 15C, D); pedipalp chela shape similar, with movable finger strongly curved in male (figs. 26C, 43C, 51A); pedipalp femur and patella more elongated in male than female; sternite VII with VSM carinae absent or obsolete (fig. 19A, C); metasomal segments IIV with DL and ML carinae complete, granular, and VL carinae obsolete, smooth.
<taxonomicName id="47984FD41F4DFF92198FFBC3FD04460F" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF92198FFBC3FD04460F" italics="true" pageId="70" pageNumber="70">Orobothriurus quewerukana</emphasis>
</taxonomicName>
may be separated from
<taxonomicName id="47984FD41F4DFF921E08FBE1FB7F460F" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1035,1193,1063,1086]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4DFF921E08FBE1FB7F460F" box="[1035,1193,1063,1086]" italics="true" pageId="70" pageNumber="70">O. tamarugal</emphasis>
</taxonomicName>
by the shape of the lamina of the hemispermatophore (figs. 40E, 41D). The pedicel of the apex is narrower, and the frontal crest more developed and elongated in
<taxonomicName id="47984FD41F4DFF921E28FB54FD3646E1" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF921E28FB54FD3646E1" italics="true" pageId="70" pageNumber="70">O. quewerukana</emphasis>
</taxonomicName>
; the ventral border of the distal portion of the apex is slightly curved in
<taxonomicName id="47984FD41F4DFF921E2BFB11FD36473A" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF921E2BFB11FD36473A" italics="true" pageId="70" pageNumber="70">O. quewerukana</emphasis>
</taxonomicName>
, but straight in
<taxonomicName id="47984FD41F4DFF9219A2FB3CFBEC473A" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[929,1082,1268,1291]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4DFF9219A2FB3CFBEC473A" box="[929,1082,1268,1291]" italics="true" pageId="70" pageNumber="70">O. tamarugal</emphasis>
</taxonomicName>
; and one or two small folds, observed in the distal crest of 80
<emphasis id="B2ECE8451F4DFF9218DBFAE6FD264775" box="[728,752,1327,1348]" italics="true" pageId="70" pageNumber="70">%</emphasis>
of the specimens of
<taxonomicName id="47984FD41F4DFF9219E4FAF9FB744777" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[999,1186,1327,1350]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF9219E4FAF9FB744777" box="[999,1186,1327,1350]" italics="true" pageId="70" pageNumber="70">O. quewerukana</emphasis>
</taxonomicName>
, are absent in the distal crest of
<taxonomicName id="47984FD41F4DFF921E0AFA84FB774752" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1033,1185,1356,1379]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4DFF921E0AFA84FB774752" box="[1033,1185,1356,1379]" italics="true" pageId="70" pageNumber="70">O. tamarugal</emphasis>
</taxonomicName>
. The hemispermatophore of
<taxonomicName id="47984FD41F4DFF9219EFFAA3FB7F47B1" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1004,1193,1385,1408]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF9219EFFAA3FB7F47B1" box="[1004,1193,1385,1408]" italics="true" pageId="70" pageNumber="70">O. quewerukana</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="47984FD41F4DFF921983FA4EFBF047AC" authorityName=": Lourenco and Dastych" authorityYear="2001" box="[896,1062,1414,1437]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F4DFF921983FA4EFBF047AC" box="[896,1062,1414,1437]" italics="true" pageId="70" pageNumber="70">O. curvidigitus</emphasis>
</taxonomicName>
, except for the slightly shorter frontal crest and the absence of folds in the distal crest (figs. 28B, 40E). The two species may be distinguished by the shape and granulation of the male telson vesicle, which is slightly deeper, with a length/height ratio of 3.594.00 (mean 5 3.76), and granulation on the entire ventral surface (fig. 23F), in
<taxonomicName id="47984FD41F4DFF92199DF9BBFB8C44B9" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[926,1114,1649,1672]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4DFF92199DF9BBFB8C44B9" box="[926,1114,1649,1672]" italics="true" pageId="70" pageNumber="70">O. quewerukana</emphasis>
</taxonomicName>
, compared with
<taxonomicName id="47984FD41F4DFF921935F946FC3E4494" authorityName=": Lourenco and Dastych" authorityYear="2001" box="[822,1000,1678,1701]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F4DFF921935F946FC3E4494" box="[822,1000,1678,1701]" italics="true" pageId="70" pageNumber="70">O. curvidigitus</emphasis>
</taxonomicName>
, in which the length/height ratio is 3.724.23 (mean 5 3.96) and granulation is present only on the anterior third of the ventral surface. The male pedipalp chela carinae are finely and densely granular in
<taxonomicName id="47984FD41F4CFF931B05FEC6FE684314" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[262,446,270,293]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4CFF931B05FEC6FE684314" box="[262,446,270,293]" italics="true" pageId="71" pageNumber="71">O. quewerukana</emphasis>
</taxonomicName>
and
<taxonomicName id="47984FD41F4CFF931BF4FEC6FD5B4314" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[503,653,270,293]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4CFF931BF4FEC6FD5B4314" box="[503,653,270,293]" italics="true" pageId="71" pageNumber="71">O. tamarugal</emphasis>
</taxonomicName>
(figs. 43D, E, 51B, C), but finely and sparsely granular to smooth in
<taxonomicName id="47984FD41F4CFF931B93FE83FDEF4351" authorityName=": Lourenco and Dastych" authorityYear="2001" box="[400,569,329,352]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F4CFF931B93FE83FDEF4351" box="[400,569,329,352]" italics="true" pageId="71" pageNumber="71">O. curvidigitus</emphasis>
</taxonomicName>
. These species may also be separated by the pigmentation pattern.
<taxonomicName id="47984FD41F4CFF931B51FE4AFD5843AB" authorityName=": Lourenco and Dastych" authorityYear="2001" box="[338,654,387,410]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F4CFF931B51FE4AFD5843AB" box="[338,654,387,410]" italics="true" pageId="71" pageNumber="71">Orobothriurus curvidigitus</emphasis>
</taxonomicName>
exhibits well-developed pigmentation on the carapace, tergites, metasoma, and pedipalps (figs. 11B, 12C, 13E), compared to
<taxonomicName id="47984FD41F4CFF931872FE15FEC3403E" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F4CFF931872FE15FEC3403E" italics="true" pageId="71" pageNumber="71">O. quewerukana</emphasis>
</taxonomicName>
in which the pigmentation, especially that of the carapace and tergites, is faint or barely discernible (fig. 41) and
<taxonomicName id="47984FD41F4CFF931872FDFAFF224057" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F4CFF931872FDFAFF224057" italics="true" pageId="71" pageNumber="71">O. tamarugal</emphasis>
</taxonomicName>
, in which it is absent (fig. 48).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F4CFF9E1A9FFDA5FCE141FF" lastPageId="74" lastPageNumber="74" pageId="71" pageNumber="71" type="description">
<paragraph id="802734571F4CFF931A9FFDA5FE25408C" blockId="71.[128,654,212,1759]" pageId="71" pageNumber="71">
DESCRIPTION: Based on
<typeStatus id="5F238AF51F4CFF931BC8FDA7FDE440B5" box="[459,562,622,644]" pageId="71" pageNumber="71" type="holotype">holotype</typeStatus>
♂ and
<typeStatus id="5F238AF51F4CFF931A83FD45FF234090" box="[128,245,652,673]" pageId="71" pageNumber="71" type="paratype">paratypes</typeStatus>
. Measurements of
<typeStatus id="5F238AF51F4CFF931BD0FD42FDEC4090" box="[467,570,651,673]" pageId="71" pageNumber="71" type="holotype">holotype</typeStatus>
♂ and
<typeStatus id="5F238AF51F4CFF931A83FD60FF31408F" box="[128,231,681,702]" pageId="71" pageNumber="71" type="paratype">paratype</typeStatus>
<specimenCount id="969EFFDE1F4CFF931AECFD6CFED2408E" box="[239,260,677,703]" pageId="71" pageNumber="71" type="female"></specimenCount>
recorded in table 3.
</paragraph>
<paragraph id="802734571F4CFF931A9EFD0CFE834127" blockId="71.[128,654,212,1759]" pageId="71" pageNumber="71">
<emphasis id="B2ECE8451F4CFF931A9EFD0CFEFD40ED" box="[157,299,709,732]" italics="true" pageId="71" pageNumber="71">Total length</emphasis>
: ♂, 38.0947.00 mm (
<geoCoordinate id="E5AC52901F4CFF93183EFD0EFD5F40EA" box="[573,649,709,732]" degrees="5" direction="north" minutes="8" orientation="latitude" pageId="71" pageNumber="71" precision="925" value="5.133333">
<emphasis id="B2ECE8451F4CFF93183EFD0EFD9D40ED" box="[573,587,711,732]" italics="true" pageId="71" pageNumber="71">n</emphasis>
5 8
</geoCoordinate>
, mean 5
<quantity id="476099B21F4CFF931AE6FD2AFE8D40C8" box="[229,347,739,761]" metricMagnitude="-2" metricUnit="m" metricValue="4.159" pageId="71" pageNumber="71" unit="mm" value="41.59">41.59 mm</quantity>
);
<specimenCount id="969EFFDE1F4CFF931B6CFD29FE5240CB" box="[367,388,736,762]" pageId="71" pageNumber="71" type="female"></specimenCount>
,
<quantity id="476099B21F4CFF931B90FD2AFD8740C8" box="[403,593,739,761]" metricMagnitude="-2" metricUnit="m" metricValue="4.435" metricValueMax="5.15" metricValueMin="3.72" pageId="71" pageNumber="71" unit="mm" value="44.35" valueMax="51.5" valueMin="37.2">37.2051.50 mm</quantity>
(
<geoCoordinate id="E5AC52901F4CFF931862FD2DFF464124" degrees="5" direction="north" minutes="8" orientation="latitude" pageId="71" pageNumber="71" precision="925" value="5.133333">
<emphasis id="B2ECE8451F4CFF931862FD2DFDB940C8" box="[609,623,740,761]" italics="true" pageId="71" pageNumber="71">n</emphasis>
5 8
</geoCoordinate>
, mean 5 43.35).
</paragraph>
<paragraph id="802734571F4CFF931A9EFCD4FC2544EE" blockId="71.[128,654,212,1759]" lastBlockId="71.[701,1228,1145,1759]" pageId="71" pageNumber="71">
<emphasis id="B2ECE8451F4CFF931A9EFCD4FF084105" box="[157,222,797,820]" italics="true" pageId="71" pageNumber="71">Color</emphasis>
: General color yellowish with light brown spots on carapace, tergites, and pedipalps; metasomal segments, ventral pigmentation slightly darker, especially in specimens from Palca (fig. 41). Carapace faintly spotted, especially laterally and posteriorly; anterior margin with two spots submedially (fig. 41A, C); posterolateral surfaces faintly pigmented; median ocular tubercle and lateral ocelli dark brown to black; anterior third of anteromedian longitudinal sulcus pigmented along borders; posteromedian longitudinal sulcus faintly pigmented or unpigmented; posterolateral sulci unpigmented. Chelicerae unpigmented. Pedipalp coxa, trochanter and femur usually unpigmented or faintly pigmented along DI and DE carinae (some juveniles); patella external and, usually, internal surfaces faintly spotted; chela unpigmented. Legs, only ventral margin of femur, and dorsal and ventral margins of patella pigmented (unpigmented in some adults). Tergites IVI each with two faint spots sublaterally, extending along anterior twothirds, becoming broader near anterior margins, delimiting broad, unpigmented median stripe (fig. 41A, C); VII unpigmented (adults) or with two faint spots sublaterally (juveniles). Sternum, genital opercula, pectines and sternites IIIVII unpigmented (fig. 41B, D). Metasomal segments IIII, dorsal surfaces each with two faint subtriangular spots medially, separated by narrow unpigmented line (juveniles) or unpigmented (adults, fig. 41); lateral surfaces unpigmented, except near VL carinae; ventral surface with narrow VM stripe on III (absent on I and II), not contiguous with lateral pigmentation, and two VL stripes on IIII (occasionally faint or absent on I) becoming broader in posterior third; specimens from Mamiña usually less pigmented: VL stripe absent on segments I and II; VM stripe often absent on III. Segment IV, dorsal surface unpigmented; lateral and ventral surfaces as for segment III. Segment V, dorsal surface unpigmented or faintly pigmented posteriorly; lateral surfaces pigmented in distal third; ventral surface with three stripes, two VL and one VM, becoming slightly broader in posterior half but not joining with lateral pigmentation. Telson vesicle unpigmented or with lateral surfaces faintly pigmented; aculeus sclerotized, dark reddish brown.
</paragraph>
<caption id="D4E764DF1F4CFF9318D7FC15FB4A460E" ID-DOI="http://doi.org/10.5281/zenodo.4611274" ID-Zenodo-Dep="4611274" httpUri="https://zenodo.org/record/4611274/files/figure.png" pageId="71" pageNumber="71" startId="71.[724,762,988,1007]" targetBox="[705,1225,211,957]" targetPageId="71">
<paragraph id="802734571F4CFF9318D7FC15FB4A460E" blockId="71.[701,1228,987,1087]" pageId="71" pageNumber="71">
Fig. 49.
<taxonomicName id="47984FD41F4CFF93193EFC12FBEB41DE" authority="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini, 2011" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[829,1085,987,1007]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="tamarugal" status="sp. nov.">
<emphasis id="B2ECE8451F4CFF93193EFC12FBEB41DE" box="[829,1085,987,1007]" italics="true" pageId="71" pageNumber="71">Orobothriurus tamarugal</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A9DF553E1F4CFF931E4FFC15FB5E41DE" box="[1100,1160,988,1007]" pageId="71" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
, holotype ♂ (MACN-Ar), dextral pedipalp femur (
<emphasis id="B2ECE8451F4CFF931EB1FC3FFB124638" bold="true" box="[1202,1220,1014,1033]" pageId="71" pageNumber="71">A</emphasis>
) and patella (
<emphasis id="B2ECE8451F4CFF93193DFBD8FCBB4615" bold="true" box="[830,877,1041,1060]" pageId="71" pageNumber="71">BD</emphasis>
).
<emphasis id="B2ECE8451F4CFF931982FBD8FC604615" bold="true" box="[897,950,1041,1060]" pageId="71" pageNumber="71">A, B.</emphasis>
Dorsal aspect.
<emphasis id="B2ECE8451F4CFF931E56FBD8FBBB4615" bold="true" box="[1109,1133,1041,1060]" pageId="71" pageNumber="71">C.</emphasis>
External aspect.
<emphasis id="B2ECE8451F4CFF93190FFBE5FCF0460E" bold="true" box="[780,806,1068,1087]" pageId="71" pageNumber="71">D.</emphasis>
Ventral aspect. Scale bar 5 1 mm.
</paragraph>
</caption>
<caption id="D4E764DF1F43FF9C1A77FB8DFCD94643" ID-DOI="http://doi.org/10.5281/zenodo.4611276" ID-Zenodo-Dep="4611276" httpUri="https://zenodo.org/record/4611276/files/figure.png" pageId="72" pageNumber="72" startId="72.[116,155,1092,1111]" targetBox="[206,1080,211,1061]" targetPageId="72">
<paragraph id="802734571F43FF9C1A77FB8DFCD94643" blockId="72.[93,1193,1089,1138]" pageId="72" pageNumber="72">
Fig. 50.
<taxonomicName id="47984FD41F43FF9C1AE3FB8AFE324666" authority="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini, 2011" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[224,484,1091,1111]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="72" pageNumber="72" phylum="Arthropoda" rank="species" species="tamarugal" status="sp. nov.">
<emphasis id="B2ECE8451F43FF9C1AE3FB8AFE324666" box="[224,484,1091,1111]" italics="true" pageId="72" pageNumber="72">Orobothriurus tamarugal</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A9DF553E1F43FF9C1BF5FB8DFDE04666" box="[502,566,1092,1111]" pageId="72" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♀ (MZUC), dextral pedipalp chela.
<emphasis id="B2ECE8451F43FF9C1E3DFB8DFB804666" bold="true" box="[1086,1110,1092,1111]" pageId="72" pageNumber="72">A.</emphasis>
Dorsal aspect.
<emphasis id="B2ECE8451F43FF9C1AAFFB96FF154643" bold="true" box="[172,195,1119,1138]" pageId="72" pageNumber="72">B.</emphasis>
External aspect.
<emphasis id="B2ECE8451F43FF9C1B78FB97FE454640" bold="true" box="[379,403,1118,1137]" pageId="72" pageNumber="72">C.</emphasis>
Internal aspect. Scale bar 5 1 mm.
</paragraph>
</caption>
<paragraph id="802734571F43FF9C1A79FB52FF3046FE" blockId="72.[93,620,1179,1760]" pageId="72" pageNumber="72">
<emphasis id="B2ECE8451F43FF9C1A79FB52FF244683" box="[122,242,1179,1202]" italics="true" pageId="72" pageNumber="72">Chelicerae</emphasis>
: Movable finger with two subdistal teeth.
</paragraph>
<paragraph id="802734571F43FF9C1A79FB11FF75447A" blockId="72.[93,620,1179,1760]" pageId="72" pageNumber="72">
<emphasis id="B2ECE8451F43FF9C1A79FB11FF3046DC" box="[122,230,1240,1261]" italics="true" pageId="72" pageNumber="72">Carapace</emphasis>
: Surfaces finely granular, more coarsely so along anterior margin, anteromedian longitudinal and median ocular sulci (♂) or smooth in anterior third, finely granular elsewhere (
<specimenCount id="969EFFDE1F43FF9C1B45FA80FE8D4752" box="[326,347,1353,1379]" pageId="72" pageNumber="72" type="female"></specimenCount>
). Anterior margin with weak median projection (epistome; fig. 14C). Anteromedian longitudinal sulcus complete, moderately developed; median ocular and posteromedian longitudinal sulci well developed; posterolateral sulci obsolete. Median ocular tubercle raised, situated anteromedially; median ocelli two ocular diameters apart.
</paragraph>
<paragraph id="802734571F43FF9D1A79F99DFBE647C2" blockId="72.[93,620,1179,1760]" lastBlockId="73.[701,1228,1472,1759]" lastPageId="73" lastPageNumber="73" pageId="72" pageNumber="72">
<emphasis id="B2ECE8451F43FF9C1A79F99DFF3E445A" box="[122,232,1620,1643]" italics="true" pageId="72" pageNumber="72">Pedipalps</emphasis>
: Femur, DI, DE, and VI carinae complete (fig. 42A), DI carina more strongly developed than DE and VI carinae; VM carina vestigial, finely granular proximally; EM carina complete, granular (♂) or obsolete, finely granular (
<specimenCount id="969EFFDE1F43FF9C1911FB50FCF14682" box="[786,807,1177,1203]" pageId="72" pageNumber="72" type="female"></specimenCount>
, juveniles); dorsal surface finely granular; internal surface sparsely granular medially. Patella, DI, VI, and VE carinae complete, granular (fig. 42BD); DE carina complete, less developed than DI carina (♂) or obsolete (
<specimenCount id="969EFFDE1F43FF9C192EFAE2FC944774" box="[813,834,1323,1349]" pageId="72" pageNumber="72" type="female"></specimenCount>
); EM carina vestigial, reduced to few small granules medially (♂) or absent (
<specimenCount id="969EFFDE1F43FF9C18A7FAAFFD6E47B1" box="[676,696,1382,1408]" pageId="72" pageNumber="72" type="female"></specimenCount>
); DPP carina comprising prominent granule and additional small granules proximally; VPP carina vestigial, comprising two or three small granules proximally; internal surface with prominent granule adjacent to trichobothrium
<emphasis id="B2ECE8451F43FF9C18F2FA35FD2E4420" box="[753,760,1532,1553]" italics="true" pageId="72" pageNumber="72">i</emphasis>
near DI carina. Chela manus slightly rounded, fingers relatively elongated (fig. 43); length/width ratio: ♂, 3.424.19, (
<geoCoordinate id="E5AC52901F43FF9C1E1CF9FEFB89447D" box="[1055,1119,1589,1612]" degrees="5" direction="north" minutes="9" orientation="latitude" pageId="72" pageNumber="72" precision="925" value="5.15">
<emphasis id="B2ECE8451F43FF9C1E1CF9FEFBFB447D" box="[1055,1069,1591,1612]" italics="true" pageId="72" pageNumber="72">n</emphasis>
5 9
</geoCoordinate>
, mean 5 3.71),
<specimenCount id="969EFFDE1F43FF9C1906F998FCCF445A" box="[773,793,1617,1643]" pageId="72" pageNumber="72" type="female"></specimenCount>
, 3.544.22 (
<geoCoordinate id="E5AC52901F43FF9C19A9F99DFC27445B" box="[938,1009,1619,1642]" degrees="5" direction="north" minutes="6" orientation="latitude" pageId="72" pageNumber="72" precision="925" value="5.1">
<emphasis id="B2ECE8451F43FF9C19A9F99DFC6E4458" box="[938,952,1620,1641]" italics="true" pageId="72" pageNumber="72">n</emphasis>
5 6
</geoCoordinate>
, mean 5 3.87); length/height ratio: ♂, 2.953.32, (
<geoCoordinate id="E5AC52901F43FF9C1E4CF9BBFB7244B7" box="[1103,1188,1648,1671]" degrees="5" direction="north" minutes="9" orientation="latitude" pageId="72" pageNumber="72" precision="925" value="5.15">
<emphasis id="B2ECE8451F43FF9C1E4CF9BBFB8B44B6" box="[1103,1117,1650,1671]" italics="true" pageId="72" pageNumber="72">n</emphasis>
5 9
</geoCoordinate>
, mean 5 3.07),
<specimenCount id="969EFFDE1F43FF9C194DF942FCB44494" box="[846,866,1675,1701]" pageId="72" pageNumber="72" type="female"></specimenCount>
, 2.873.34 (
<geoCoordinate id="E5AC52901F43FF9C19F7F946FBED4495" box="[1012,1083,1677,1700]" degrees="5" direction="north" minutes="6" orientation="latitude" pageId="72" pageNumber="72" precision="925" value="5.1">
<emphasis id="B2ECE8451F43FF9C19F7F946FBD44495" box="[1012,1026,1679,1700]" italics="true" pageId="72" pageNumber="72">n</emphasis>
5 6
</geoCoordinate>
, mean 5 3.16); most carinae obsolete, finely granular (less so in
<specimenCount id="969EFFDE1F43FF9C1917F90FFCFE44D1" box="[788,808,1734,1760]" pageId="72" pageNumber="72" type="female"></specimenCount>
) or absent; VM carina restricted to proximal two-thirds of manus, VI carina to proximal third; DS, DMA, and DI carinae complete, finely and densely granular, especially near base of fixed finger (♂) or smooth (
<specimenCount id="969EFFDE1F42FF9D1A8BF9FAFF4A447C" box="[136,156,1587,1613]" pageId="73" pageNumber="73" type="female"></specimenCount>
); D carina smooth; internal surface with acuminate apophysis (♂) or low bulge (
<specimenCount id="969EFFDE1F42FF9D183AF999FD9B445B" box="[569,589,1616,1642]" pageId="73" pageNumber="73" type="female"></specimenCount>
) near articulation of movable finger (fig. 43A, D); movable finger (♂) strongly curved, creating small gap with fixed finger when fingers closed (fig. 43C); fingers, dentate margins each with median denticle row and 45 pairs of internal and external accessory denticles.
</paragraph>
<caption id="D4E764DF1F42FF9D1A95FA85FCBA474B" ID-DOI="http://doi.org/10.5281/zenodo.4611280" ID-Zenodo-Dep="4611280" httpUri="https://zenodo.org/record/4611280/files/figure.png" pageId="73" pageNumber="73" startId="73.[150,188,1356,1375]" targetBox="[191,1164,211,1325]" targetPageId="73">
<paragraph id="802734571F42FF9D1A95FA85FCBA474B" blockId="73.[128,1228,1351,1402]" pageId="73" pageNumber="73">
Fig. 51.
<taxonomicName id="47984FD41F42FF9D1AFDFA82FE2B476E" authority="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini, 2011" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[254,509,1355,1375]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="73" pageNumber="73" phylum="Arthropoda" rank="species" species="tamarugal" status="sp. nov.">
<emphasis id="B2ECE8451F42FF9D1AFDFA82FE2B476E" box="[254,509,1355,1375]" italics="true" pageId="73" pageNumber="73">Orobothriurus tamarugal</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A9DF553E1F42FF9D1809FA85FD92476E" box="[522,580,1356,1375]" pageId="73" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
, holotype ♂ (MACN-Ar), dextral pedipalp chela.
<emphasis id="B2ECE8451F42FF9D1E57FA85FBBA476E" bold="true" box="[1108,1132,1356,1375]" pageId="73" pageNumber="73">A.</emphasis>
External aspect.
<emphasis id="B2ECE8451F42FF9D1ACCFAAFFF334748" bold="true" box="[207,229,1382,1401]" pageId="73" pageNumber="73">B.</emphasis>
Ventral aspect.
<emphasis id="B2ECE8451F42FF9D1B91FAAFFE7D4748" bold="true" box="[402,427,1382,1401]" pageId="73" pageNumber="73">C.</emphasis>
Ventrointernal aspect. Scale bar 5 1 mm.
</paragraph>
</caption>
<paragraph id="802734571F42FF9D18D8FA32FC2F44B7" blockId="73.[701,1228,1472,1759]" pageId="73" pageNumber="73">
<emphasis id="B2ECE8451F42FF9D18D8FA32FCA14423" box="[731,887,1531,1554]" italics="true" pageId="73" pageNumber="73">Trichobothria</emphasis>
: Femur with 3 trichobothria, patella with 19, chela with 27 (figs. 42, 43). Chela trichobothrium
<emphasis id="B2ECE8451F42FF9D19C9F9FEFC33447D" box="[970,997,1591,1612]" italics="true" pageId="73" pageNumber="73">Et</emphasis>
<subScript id="1C1C36121F42FF9D19E5F989FC26447F" attach="left" box="[998,1008,1600,1614]" fontSize="6" pageId="73" pageNumber="73">3</subScript>
situated proximal to
<emphasis id="B2ECE8451F42FF9D18DDF99DFCD24458" box="[734,772,1620,1641]" italics="true" pageId="73" pageNumber="73">Est</emphasis>
(in same axis as
<emphasis id="B2ECE8451F42FF9D19D3F99DFC204458" box="[976,1014,1620,1641]" italics="true" pageId="73" pageNumber="73">Est</emphasis>
in one specimen);
<emphasis id="B2ECE8451F42FF9D18BEF9B9FD3E44B6" box="[701,744,1648,1671]" italics="true" pageId="73" pageNumber="73">Esb</emphasis>
situated dorsal to
<emphasis id="B2ECE8451F42FF9D19CBF9B9FC3E44B6" box="[968,1000,1648,1671]" italics="true" pageId="73" pageNumber="73">Eb</emphasis>
<subScript id="1C1C36121F42FF9D19E4F9B3FC2744B9" attach="left" box="[999,1009,1658,1672]" fontSize="6" pageId="73" pageNumber="73">2</subScript>
.
</paragraph>
<paragraph id="802734571F42FF9E18D8F946FF134373" blockId="73.[701,1228,1472,1759]" lastBlockId="74.[93,620,213,1759]" lastPageId="74" lastPageNumber="74" pageId="73" pageNumber="73">
<emphasis id="B2ECE8451F42FF9D18D8F946FCEE4495" box="[731,824,1679,1700]" italics="true" pageId="73" pageNumber="73">Tergites</emphasis>
: Tergites IVI, surfaces finely granular, more coarsely and densely so in ♂. Tergite VII tetracarinate, paired DL carinae restricted to posterior two-thirds of segment, paired DSM carinae to posterior half; surfaces more coarsely and densely granular in posterior half.
</paragraph>
<paragraph id="802734571F41FF9E1A79FE82FE91401D" blockId="74.[93,620,213,1759]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E1A79FE82FF664351" box="[122,176,331,352]" italics="true" pageId="74" pageNumber="74">Legs</emphasis>
: Femur and patella, prolateral surfaces finely granular, retrolateral surfaces smooth. Femur, VI and EM carinae complete in ♂, less developed in
<specimenCount id="969EFFDE1F41FF9E1B6AFE56FEAB4388" box="[361,381,415,441]" pageId="74" pageNumber="74" type="female"></specimenCount>
; DE and DI carinae vestigial. Patella acarinate. Telotarsi, proand retroventral rows of spiniform macrosetae with following counts on leg I, 1/1; II, 2/2; III and IV, 3/3.
</paragraph>
<paragraph id="802734571F41FF9E1A79FDFFFF7F40B7" blockId="74.[93,620,213,1759]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E1A79FDFFFF0E407A" box="[122,216,566,587]" italics="true" pageId="74" pageNumber="74">Pectines</emphasis>
: Pectinal tooth count: ♂, 2024 (
<geoCoordinate id="E5AC52901F41FF9E1A65FD9DFF604059" box="[102,182,594,617]" degrees="5" direction="north" minutes="34" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.5666666">
<emphasis id="B2ECE8451F41FF9E1A65FD9DFFA24058" box="[102,116,596,617]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 34
</geoCoordinate>
, mode 5 22);
<specimenCount id="969EFFDE1F41FF9E1B5AFD99FEB8405B" box="[345,366,592,618]" pageId="74" pageNumber="74" type="female"></specimenCount>
, 1922 (
<geoCoordinate id="E5AC52901F41FF9E1BCCFD9DFDC94059" box="[463,543,594,617]" degrees="5" direction="north" minutes="24" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.4">
<emphasis id="B2ECE8451F41FF9E1BCCFD9DFE0B4058" box="[463,477,596,617]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 24
</geoCoordinate>
, mode 5 20).
</paragraph>
<paragraph id="802734571F41FF9E1A79FD46FF194128" blockId="74.[93,620,213,1759]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E1A79FD46FF084095" box="[122,222,655,676]" italics="true" pageId="74" pageNumber="74">Sternites</emphasis>
: Sternites IIIVI, surfaces smooth (
<specimenCount id="969EFFDE1F41FF9E1A65FD60FFAC40F2" box="[102,122,681,707]" pageId="74" pageNumber="74" type="female"></specimenCount>
) or matt in posterior three-quarters (♂); spiracles small, narrow. Sternite VII, surface smooth (
<specimenCount id="969EFFDE1F41FF9E1AC5FD2AFF0D40CC" box="[198,219,739,765]" pageId="74" pageNumber="74" type="female"></specimenCount>
) or finely granular (♂) (fig. 19C), acarinate.
</paragraph>
<paragraph id="802734571F41FF9E1A79FCEBFCC343AD" blockId="74.[93,620,213,1759]" lastBlockId="74.[667,1194,213,1478]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E1A79FCEBFF244106" box="[122,242,802,823]" italics="true" pageId="74" pageNumber="74">Metasoma</emphasis>
: Segment I, DL carinae complete; ML carinae complete, posterior granules more prominent than adjacent granulation; one pair of ML macrosetae; LIM carinae reduced to few granules in posterior third; VL carinae obsolete; VSM carinae absent (fig. 19C); two pairs of VL and VSM macrosetae. Segments I and II, ventral intercarinal surfaces finely granular (♂). Segments IIIV, surfaces between DL and ML carinae finely and sparsely granular, granulation decreasing posteriorly. Segments II and III, DL carinae complete; one pair of DL macrosetae in 14
<emphasis id="B2ECE8451F41FF9E1B8EFB57FE734682" box="[397,421,1182,1203]" italics="true" pageId="74" pageNumber="74">%</emphasis>
and 50
<emphasis id="B2ECE8451F41FF9E1822FB57FDEF4682" box="[545,569,1182,1203]" italics="true" pageId="74" pageNumber="74">%</emphasis>
of specimens, respectively; ML carinae as for segment I; LIM carinae reduced to few granules posteriorly on segment II, restricted to posterior third (♂) or absent (
<specimenCount id="969EFFDE1F41FF9E1820FAD9FDEE471B" box="[547,568,1296,1322]" pageId="74" pageNumber="74" type="female"></specimenCount>
) on segment III; VL carinae as for segment I but less developed; VSM carinae absent; three pairs of VL and VSM macrosetae. Segment IV, DL carinae complete, granular; one pair of DL macrosetae; ML carinae complete, granular; LIM carinae absent; VL carinae as for segment III (absent in some subadult and juvenile specimens); VSM carinae absent; usually three pairs of VL and VSM macrosetae. Segment V, length/width ratio: ♂, 2.182.44 (
<geoCoordinate id="E5AC52901F41FF9E1811F99CFDB0445B" box="[530,614,1619,1642]" degrees="5" direction="north" minutes="6" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.1">
<emphasis id="B2ECE8451F41FF9E1811F99CFDF6445B" box="[530,544,1621,1642]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 6
</geoCoordinate>
, mean 5 2.31),
<specimenCount id="969EFFDE1F41FF9E1B13F9A7FEF344B9" box="[272,293,1646,1672]" pageId="74" pageNumber="74" type="female"></specimenCount>
, 2.012.20 (
<geoCoordinate id="E5AC52901F41FF9E1BB5F9BBFE2B44B6" box="[438,509,1648,1671]" degrees="5" direction="north" minutes="4" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.0666666">
<emphasis id="B2ECE8451F41FF9E1BB5F9BBFE1244B6" box="[438,452,1650,1671]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 4
</geoCoordinate>
, mean 5 2.10); DL carinae complete, granular; lateral surfaces sparsely granular near DL carinae (fig. 20D); VL carinae complete; VSM carinae reduced to few granules medially; VM carina complete, obscured by surface granulation in posterior half; usually three pairs of VL and VSM macrosetae; two pairs of macrosetae along posterior margin; ventral intercarinal surfaces, granulation decreasing anteriorly.
</paragraph>
<paragraph id="802734571F41FF9E18BBFE6AFB97405A" blockId="74.[667,1194,213,1478]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E18BBFE6AFCD5438B" box="[696,771,419,442]" italics="true" pageId="74" pageNumber="74">Telson</emphasis>
: Length/height ratio: ♂, 3.594.00 (
<geoCoordinate id="E5AC52901F41FF9E18A7FE0BFD3B43E6" box="[676,749,449,472]" degrees="5" direction="north" minutes="9" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.15">
<emphasis id="B2ECE8451F41FF9E18A7FE0BFD6443E6" box="[676,690,450,471]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 9
</geoCoordinate>
, mean 5 3.76);
<specimenCount id="969EFFDE1F41FF9E19B0FE76FC1E43E8" box="[947,968,447,473]" pageId="74" pageNumber="74" type="female"></specimenCount>
, 2.903.17 (
<geoCoordinate id="E5AC52901F41FF9E1E59FE0BFB7243E9" box="[1114,1188,449,472]" degrees="5" direction="north" minutes="6" orientation="latitude" pageId="74" pageNumber="74" precision="925" value="5.1">
<emphasis id="B2ECE8451F41FF9E1E59FE0BFBBE43E6" box="[1114,1128,450,471]" italics="true" pageId="74" pageNumber="74">n</emphasis>
5 6
</geoCoordinate>
, mean 5 3.03). Vesicle slightly elongated (♂, fig. 23F) or globose (
<specimenCount id="969EFFDE1F41FF9E19B2FE33FC134025" box="[945,965,506,532]" pageId="74" pageNumber="74" type="female"></specimenCount>
, fig. 24D); dorsal surface smooth, flat or slightly concave, gland not apparent (♂); ventral surface sparsely granular. Aculeus short and curved.
</paragraph>
<paragraph id="802734571F41FF9E18BBFDBAFCE141FF" blockId="74.[667,1194,213,1478]" pageId="74" pageNumber="74">
<emphasis id="B2ECE8451F41FF9E18BBFDBAFC4840BB" box="[696,926,627,650]" italics="true" pageId="74" pageNumber="74">Hemispermatophore</emphasis>
: Apex approximately half the length of lamina, distal border subtriangular and slightly curved to ventral margin; distal crest curved like ventral margin, with one or two small folds in 80
<emphasis id="B2ECE8451F41FF9E1E93FD23FB7E40CE" box="[1168,1192,746,767]" italics="true" pageId="74" pageNumber="74">%</emphasis>
of specimens. Frontal crest elongated; basal part oblique; distal part parallel to ventral margin of lamina, lateral projections complete, slightly undulated, and larger than basal part. Basal lobe, terminal process extending to constriction of frontal crest (fig. 40D, E).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F41FF9E18BBFC1CFBA94652" pageId="74" pageNumber="74" type="distribution">
<paragraph id="802734571F41FF9E18BBFC1CFBA94652" blockId="74.[667,1194,213,1478]" pageId="74" pageNumber="74">
DISTRIBUTION:
<taxonomicName id="47984FD41F41FF9E1985FC1FFD04463A" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="74" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F41FF9E1985FC1FFD04463A" italics="true" pageId="74" pageNumber="74">Orobothriurus quewerukana</emphasis>
</taxonomicName>
is endemic to the western slopes of the Andes, at
<quantity id="476099B21F41FF9E1932FBDBFC0A4616" box="[817,988,1042,1063]" metricMagnitude="3" metricUnit="m" metricValue="3.2769999999999997" metricValueMax="3.732" metricValueMin="2.822" pageId="74" pageNumber="74" unit="m" value="3277.0" valueMax="3732.0" valueMin="2822.0">28223732 m</quantity>
, in the
<collectingRegion id="425CFAB51F41FF9E1E63FBDBFB7F4616" box="[1120,1193,1042,1063]" country="Peru" name="Tacna" pageId="74" pageNumber="74">Tacna</collectingRegion>
Department of southern
<collectingCountry id="F88F74C71F41FF9E19C8FBF9FBD74674" box="[971,1025,1072,1093]" name="Peru" pageId="74" pageNumber="74">Peru</collectingCountry>
and the
<collectingRegion id="425CFAB51F41FF9E1E76FBF9FCEA4652" country="Chile" name="Tarapaca" pageId="74" pageNumber="74">Tarapacá Region</collectingRegion>
of northern
<collectingCountry id="F88F74C71F41FF9E19D6FB84FBC44652" box="[981,1042,1101,1123]" name="Chile" pageId="74" pageNumber="74">Chile</collectingCountry>
(fig. 53).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F41FF9E18BBFBA0FB8047F4" pageId="74" pageNumber="74" type="biology_ecology">
<paragraph id="802734571F41FF9E18BBFBA0FB8047F4" blockId="74.[667,1194,213,1478]" pageId="74" pageNumber="74">
ECOLOGY: The area inhabited by
<taxonomicName id="47984FD41F41FF9E1E8FFBA2FCFC46AE" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="74" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F41FF9E1E8FFBA2FCFC46AE" italics="true" pageId="74" pageNumber="74">O. quewerukana</emphasis>
</taxonomicName>
belongs to the Serrania Esteparia ecoregion (
<bibRefCitation id="E40949A61F41FF9E1919FB6FFC70468D" author="Brack, A." box="[794,934,1190,1212]" pageId="74" pageNumber="74" pagination="175 - 319" refId="ref42164" refString="Brack, A. 1986. Ecologia de un pais complejo. In Gran geografia del Peru. Naturaleza y Hombre 2, 175 - 319. Barcelona: Manfer-Juan Mejia Baca." type="journal article" year="1986">Brack, 1986</bibRefCitation>
) or Estepa Altoandina botanical region (
<bibRefCitation id="E40949A61F41FF9E1959FB0DFC2B46EB" author="Gajardo, R." box="[858,1021,1220,1242]" pageId="74" pageNumber="74" refId="ref42994" refString="Gajardo, R. 1993. La vegetacion natural de Chile, clasificacion y distribucion geografica. Santiago de Chile: Editorial Universitaria, 165 pp." type="book" year="1993">Gajardo, 1993</bibRefCitation>
), the vegetation of which is characterised by abundant cacti and some shrubs, like
<taxonomicName id="47984FD41F41FF9E1976FB37FC324724" box="[885,996,1278,1301]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="74" pageNumber="74" phylum="Tracheophyta" rank="genus">
<emphasis id="B2ECE8451F41FF9E1976FB37FC324724" box="[885,996,1278,1301]" italics="true" pageId="74" pageNumber="74">Baccharis</emphasis>
</taxonomicName>
L.,
<taxonomicName id="47984FD41F41FF9E1E23FB36FBAD4725" box="[1056,1147,1279,1300]" class="Magnoliopsida" family="Fabaceae" genus="Lupinus" kingdom="Plantae" order="Fabales" pageId="74" pageNumber="74" phylum="Tracheophyta" rank="genus">
<emphasis id="B2ECE8451F41FF9E1E23FB36FBAD4725" box="[1056,1147,1279,1300]" italics="true" pageId="74" pageNumber="74">Lupinus</emphasis>
</taxonomicName>
L., and
<taxonomicName id="47984FD41F41FF9E18DAFAD4FCE64703" box="[729,816,1309,1330]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="74" pageNumber="74" phylum="Tracheophyta" rank="genus">
<emphasis id="B2ECE8451F41FF9E18DAFAD4FCE64703" box="[729,816,1309,1330]" italics="true" pageId="74" pageNumber="74">Senecio</emphasis>
</taxonomicName>
L. Specimens were collected under stones during the day and with UV light detection at night.
<taxonomicName id="47984FD41F41FF9E19C9FA9EFCDF47BA" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="74" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F41FF9E19C9FA9EFCDF47BA" italics="true" pageId="74" pageNumber="74">Orobothriurus quewerukana</emphasis>
</taxonomicName>
is sympatric with another bothriurid,
<taxonomicName id="47984FD41F41FF9E1908FA5BFCBF47F4" authority="Ochoa and Acosta, 2002" authorityName="Ochoa and Acosta" authorityYear="2002" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="74" phylum="Arthropoda" rank="species" species="quiscapata">
<emphasis id="B2ECE8451F41FF9E1908FA5BFB9B4799" box="[779,1101,1426,1449]" italics="true" pageId="74" pageNumber="74">Brachistosternus quiscapata</emphasis>
<bibRefCitation id="E40949A61F41FF9E1E5FFA5BFCBF47F4" author="Acosta, L. E. &amp; J. A. Ochoa" pageId="74" pageNumber="74" pagination="15 - 23" refId="ref42068" refString="Acosta, L. E., and J. A. Ochoa. 2002. Lista de los escorpiones bolivia nos (Chelicerata: Scorpiones), con notas sobre su distribucion. Revista de la Sociedad Argentina de Entomologia 61 (3 - 4): 15 - 23." type="journal article" year="2002">Ochoa and Acosta, 2002</bibRefCitation>
</taxonomicName>
, at Palca and Putre.
</paragraph>
</subSubSection>
</treatment>
</document>