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<mods:title id="896897505590DB99E98107E7C681B5C2">Systematic Revision Of The Andean Scorpion Genus Orobothriurus Maury, 1976 (Bothriuridae), With Discussion Of The Altitude Record For Scorpions</mods:title>
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<taxonomicName id="47984FD41F3FFFE01A8FFC89FDEB4167" ID-CoL="757VR" authority="Acosta, 2001" authorityName="Acosta" authorityYear="2001" box="[140,573,832,855]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01A8FFC89FE4D4167" box="[140,411,832,855]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
<bibRefCitation id="E40949A61F3FFFE01BA7FC88FDEB4167" author="Acosta, L. E. &amp; J. A. Ochoa" box="[420,573,833,854]" pageId="52" pageNumber="52" pagination="203 - 214" refId="ref41988" refString="Acosta, L. E., and J. A. Ochoa. 2001. Two new species of Orobothriurus Maury 1976, from Argentina and Peru, with comments on the systematics of the genus (Scorpiones, Bothriuridae). In V. Fet and P. A. Selden (editors), Scorpions 2001. In memoriam Gary A. Polis: 203 - 214. Burnham Beeches, Bucks: British Arachnological Society." type="book chapter" year="2001">Acosta, 2001</bibRefCitation>
</taxonomicName>
<figureCitation id="18A328D21F3FFFE01AF6FC96FEA44144" box="[245,370,863,885]" captionStart="Fig" captionStartId="51.[150,188,1625,1644]" captionTargetBox="[170,1186,211,1594]" captionTargetId="figure-1@51.[170,1187,210,1594]" captionTargetPageId="51" captionText="Fig. 36. Orobothriurus Maury, 1976, sinistral hemispermatophore, ectal aspect showing lamina. AC. Orobothriurus compagnuccii, n. sp., ♂ holotype, ♂ paratypes (MACN-Ar). D, E. Orobothriurus grismadoi Ojanguren Affilastro et al., 2009, ♂ paratypes (MACN-Ar). FH. Orobothriurus famatina Acosta, 2001, ♂ (MACN-Ar). I, J. Orobothriurus alticola (Pocock, 1899), ♂ (MACN-Ar). KO. Orobothriurus calchaqui, n. sp., ♂ paratypes (MACN-Ar). Scale bar 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611232" httpUri="https://zenodo.org/record/4611232/files/figure.png" pageId="52" pageNumber="52">Figures 36</figureCitation>
FH, 54
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<treatmentCitation id="013912461F3FFFE01A5EFC42FE2441AE" author="Maury, E. A." box="[93,498,907,928]" page="17" pageId="52" pageNumber="52" year="1976">
<taxonomicName id="47984FD41F3FFFE01A5EFC42FE2441AE" ID-CoL="6T87R" authority=": Maury 1976: 17" authorityName=": Maury" authorityPageNumber="17" authorityYear="1976" box="[93,498,907,928]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="B2ECE8451F3FFFE01A5EFC42FEE941AE" box="[93,319,907,927]" italics="true" pageId="52" pageNumber="52">Orobothriurus alticola</emphasis>
:
<bibRefCitation id="E40949A61F3FFFE01B4FFC44FE2441AE" author="Maury, E. A." box="[332,498,908,928]" pageId="52" pageNumber="52" pagination="14 - 25" refId="ref44084" refString="Maury, E. A. 1976. Escorpiones y escorpionismo en el Peru V. Orobothriurus, un nuevo genero de escorpiones altoandinos (Bothriuridae). Revista Peruana de Entomologia 18 (1): 14 - 25." type="journal article" year="1976">Maury 1976: 17</bibRefCitation>
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(part);
<treatmentCitation id="013912461F3FFFE01841FC45FF08418B" author="Cei, J. M." page="660" pageId="52" pageNumber="52" year="1982">
<bibRefCitation id="E40949A61F3FFFE01841FC45FF08418B" author="Cei, J. M." pageId="52" pageNumber="52" pagination="643 - 672" refId="ref42502" refString="Cei, J. M. 1982. Aspetti geo-biogeografici inediti della Sierra di Famatina, il piu elevato massiccio d'America del Sud dopo le cordigliere andine (Argentina centro-occidentale). L'Universo (Instituto Geografico Militare, Firenze) 62 (4): 643 - 672." type="journal article" year="1982">Cei, 1982: 660</bibRefCitation>
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, 669 [misidentification].
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<emphasis id="B2ECE8451F3FFFE01A5EFC0BFE8F41E7" box="[93,345,962,982]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
Acosta
</taxonomicName>
<emphasis id="B2ECE8451F3FFFE01BC6FC0AFE0E41E4" box="[453,472,963,981]" italics="true" pageId="52" pageNumber="52">in</emphasis>
<treatmentCitation id="013912461F3FFFE01BE9FC0AFEAA41C1" author="Acosta, L. E. &amp; J. A. Ochoa" page="205" pageId="52" pageNumber="52" year="2001">
<bibRefCitation id="E40949A61F3FFFE01BE9FC0AFEAA41C1" author="Acosta, L. E. &amp; J. A. Ochoa" pageId="52" pageNumber="52" pagination="203 - 214" refId="ref41988" refString="Acosta, L. E., and J. A. Ochoa. 2001. Two new species of Orobothriurus Maury 1976, from Argentina and Peru, with comments on the systematics of the genus (Scorpiones, Bothriuridae). In V. Fet and P. A. Selden (editors), Scorpions 2001. In memoriam Gary A. Polis: 203 - 214. Burnham Beeches, Bucks: British Arachnological Society." type="book chapter" year="2001">Acosta and Ochoa, 2001: 205211</bibRefCitation>
</treatmentCitation>
, figs. 414;
<treatmentCitation id="013912461F3FFFE01823FC17FF3E463D" author="Ochoa, J. A." page="43" pageId="52" pageNumber="52" year="2004">
<bibRefCitation id="E40949A61F3FFFE01823FC17FF3E463D" author="Ochoa, J. A." pageId="52" pageNumber="52" pagination="43 - 73" refId="ref44598" refString="Ochoa, J. A. 2004 a. Filogenia del genero Orobothriurus y descripcion de un nuevo genero de Bothriuridae (Scorpiones). Revista Iberica de Aracnologia 9: 43 - 73." type="journal article" year="2004">Ochoa, 2004a: 43</bibRefCitation>
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, 52, 55, 73,
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;
<treatmentCitation id="013912461F3FFFE01BFEFC30FE5B4616" author="Ojanguren Affilastro, A. A." page="177" pageId="52" pageNumber="52" year="2005">
<bibRefCitation id="E40949A61F3FFFE01BFEFC30FE5B4616" author="Ojanguren Affilastro, A. A." pageId="52" pageNumber="52" pagination="75 - 241" refId="ref45124" refString="Ojanguren Affilastro, A. A. 2005. Estudio monografico de los escorpiones de la Republica Argentina. Revista Iberica de Aracnologia 11: 75 - 241." type="journal article" year="2005">Ojanguren Affilastro, 2005: 177181</bibRefCitation>
</treatmentCitation>
, figs. 458466, 477, 488, 663, table 15.
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<subSubSection id="C88267DC1F3FFFE01A79FB90FDBA44EE" pageId="52" pageNumber="52" type="materials_examined">
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TYPE MATERIAL:
<emphasis id="B2ECE8451F3FFFE01B48FB92FF1E46BF" bold="true" pageId="52" pageNumber="52">
<collectingCountry id="F88F74C71F3FFFE01B48FB92FE214640" box="[331,503,1115,1137]" name="Argentina" pageId="52" pageNumber="52">ARGENTINA</collectingCountry>
:
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:
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Department
</collectorName>
</emphasis>
:
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♂ (MACN-Ar 9932),
<collectingMunicipality id="6043AE2D1F3FFFE01B45FB5FFDE6469D" box="[326,560,1174,1196]" pageId="52" pageNumber="52">Sierra de Famatina</collectingMunicipality>
,
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canyon, path to
<collectorName id="2D6D51811F3FFFE01B62FB7DFDD446F8" box="[353,514,1204,1226]" pageId="52" pageNumber="52">Mina El Oro</collectorName>
, 29
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069S 67
<emphasis id="B2ECE8451F3FFFE01A7BFB18FF5546D7" bold="true" box="[120,131,1233,1254]" pageId="52" pageNumber="52">°</emphasis>
429W,
<quantity id="476099B21F3FFFE01AD7FB18FEF946D6" box="[212,303,1233,1255]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="52" pageNumber="52" unit="m" value="2450.0">
<elevation id="0BB5D3641F3FFFE01AD7FB18FEF946D6" box="[212,303,1233,1255]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="52" pageNumber="52" unit="m" value="2450.0">2450 m</elevation>
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,
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<collectingDate id="E462EB7F1F3FFFE01B3CFB18FE6746D6" box="[319,433,1233,1255]" pageId="52" pageNumber="52" value="1998-12-06">6.xii.1998</collectingDate>
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,
<collectorName id="2D6D51811F3FFFE01BC3FB1BFDEF46D7" box="[448,569,1233,1255]" pageId="52" pageNumber="52">L. Acosta</collectorName>
,
<collectorName id="2D6D51811F3FFFE0184BFB1BFF784735" pageId="52" pageNumber="52">M. Acosta</collectorName>
and G.
<collectorName id="2D6D51811F3FFFE01B2CFB26FE454734" box="[303,403,1263,1285]" pageId="52" pageNumber="52">Repossi.</collectorName>
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<typeStatus id="5F238AF51F3FFFE01BA7FB26FDCC4735" box="[420,538,1263,1284]" pageId="52" pageNumber="52" type="paratype">Paratypes</typeStatus>
: same data, 1 ♂,
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(
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), 1 ♂ (
<specimenCode id="D03E9C2C1F3FFFE01BD7FAC4FD9F4713" box="[468,585,1292,1314]" collectionCode="CDA" country="Canada" name="Canadian Department of Agriculture" pageId="52" pageNumber="52">CDA 051</specimenCode>
), 1 ♂ (
<collectionCode id="E689AC921F3FFFE01A87FAE3FF324771" box="[132,228,1322,1344]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="52" pageNumber="52">MHNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30F03E0A1F3FFFE01AFAFAE3FE39476F" ID-GBIF-Occurrence="3053988375" collectingDate="1951-04" collectorName="R. Gonzalez Amorio" country="Argentina" location="Famatina" pageId="52" pageNumber="52" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja" typeStatus="holotype">
<location id="8547628C1F3FFFE01AFAFAE3FEBB4771" LSID="urn:lsid:plazi:treatment:083185411F3FFFE01A8FFC89FC274423:8547628C1F3FFFE01AFAFAE3FEBB4771" box="[249,365,1322,1344]" country="Argentina" name="Famatina" pageId="52" pageNumber="52" stateProvince="La Rioja">Famatina</location>
,
<date id="F42612971F3FFFE01B7BFAE3FE184771" box="[376,462,1322,1344]" pageId="52" pageNumber="52" value="1951-04">
<collectingDate id="E462EB7F1F3FFFE01B7BFAE3FE184771" box="[376,462,1322,1344]" pageId="52" pageNumber="52" value="1951-04">iv.1951</collectingDate>
</date>
,
<collectorName id="2D6D51811F3FFFE01BD9FAE2FF6A476F" pageId="52" pageNumber="52">R. González Amorío</collectorName>
,
<specimenCount id="969EFFDE1F3FFFE01ACCFA81FED4476E" box="[207,258,1349,1375]" pageId="52" pageNumber="52" type="female">1 ♀</specimenCount>
(MACN-Ar 6844)
</materialsCitation>
;
<materialsCitation id="30F03E0A1F3FFFE01800FA81FEE347A8" ID-GBIF-Occurrence="3053988386" collectingDate="1956-02" collectorName="Mina El Oro &amp; M. E. Galiano" country="Argentina" location="Chilecito" pageId="52" pageNumber="52" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja" typeStatus="holotype">
<collectorName id="2D6D51811F3FFFE01800FA81FF58474B" pageId="52" pageNumber="52">Mina El Oro</collectorName>
,
<location id="8547628C1F3FFFE01AA3FAACFEDA474A" LSID="urn:lsid:plazi:treatment:083185411F3FFFE01A8FFC89FC274423:8547628C1F3FFFE01AA3FAACFEDA474A" box="[160,268,1381,1403]" country="Argentina" name="Chilecito" pageId="52" pageNumber="52" stateProvince="La Rioja">Chilecito</location>
,
<date id="F42612971F3FFFE01B1DFAACFEBB474A" box="[286,365,1381,1403]" pageId="52" pageNumber="52" value="1956-02">
<collectingDate id="E462EB7F1F3FFFE01B1DFAACFEBB474A" box="[286,365,1381,1403]" pageId="52" pageNumber="52" value="1956-02">ii.1956</collectingDate>
</date>
,
<collectorName id="2D6D51811F3FFFE01B7CFAAFFDFC474A" box="[383,554,1381,1403]" pageId="52" pageNumber="52">M.E. Galiano</collectorName>
,
<specimenCount id="969EFFDE1F3FFFE01838FAACFDBD474D" box="[571,619,1378,1404]" pageId="52" pageNumber="52" type="female">1 ♀</specimenCount>
(MACN-Ar 6843)
</materialsCitation>
;
<materialsCitation id="30F03E0A1F3FFFE01B41FA4AFF18443E" ID-GBIF-Occurrence="3053988349" collectingDate="1998-12-05" collectionCode="MHNC" collectorName="Acosta &amp; M. Acosta &amp; G. Repossi" country="Argentina" elevation="3060" location="Mina La Mejicana" municipality="Sierra de Famatina" pageId="52" pageNumber="52" specimenCode="CDA 050" specimenCount="3" specimenCount-female="3" stateProvince="La Rioja" typeStatus="holotype">
<collectingMunicipality id="6043AE2D1F3FFFE01B41FA4AFDFF47A8" box="[322,553,1411,1433]" pageId="52" pageNumber="52">Sierra de Famatina</collectingMunicipality>
, path to
<location id="8547628C1F3FFFE01A7DFA69FE884787" LSID="urn:lsid:plazi:treatment:083185411F3FFFE01A8FFC89FC274423:8547628C1F3FFFE01A7DFA69FE884787" box="[126,350,1440,1462]" country="Argentina" municipality="Sierra de Famatina" name="Mina La Mejicana" pageId="52" pageNumber="52" stateProvince="La Rioja">Mina La Mejicana</location>
,
<quantity id="476099B21F3FFFE01B68FA69FE124787" box="[363,452,1440,1462]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="52" pageNumber="52" unit="m" value="3060.0">
<elevation id="0BB5D3641F3FFFE01B68FA69FE124787" box="[363,452,1440,1462]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="52" pageNumber="52" unit="m" value="3060.0">3060 m</elevation>
</quantity>
,
<date id="F42612971F3FFFE01BD1FA69FD924787" box="[466,580,1440,1462]" pageId="52" pageNumber="52" value="1998-12-05">
<collectingDate id="E462EB7F1F3FFFE01BD1FA69FD924787" box="[466,580,1440,1462]" pageId="52" pageNumber="52" value="1998-12-05">5.xii.1998</collectingDate>
</date>
, L.
<collectorName id="2D6D51811F3FFFE01A5EFA77FF6747E2" box="[93,177,1470,1491]" pageId="52" pageNumber="52">Acosta</collectorName>
,
<collectorName id="2D6D51811F3FFFE01AC6FA76FE9D47E2" box="[197,331,1470,1492]" pageId="52" pageNumber="52">M. Acosta</collectorName>
, and
<collectorName id="2D6D51811F3FFFE01B99FA77FDF347E5" box="[410,549,1470,1492]" pageId="52" pageNumber="52">G. Repossi</collectorName>
,
<specimenCount id="969EFFDE1F3FFFE0183AFA77FDBD47E4" box="[569,619,1467,1493]" pageId="52" pageNumber="52" type="female">1 ♀</specimenCount>
(
<specimenCode id="D03E9C2C1F3FFFE01A66FA15FF3547C0" box="[101,227,1499,1521]" collectionCode="CDA" country="Canada" name="Canadian Department of Agriculture" pageId="52" pageNumber="52">CDA 050</specimenCode>
),
<specimenCount id="969EFFDE1F3FFFE01B03FA15FEE247C3" box="[256,308,1496,1522]" pageId="52" pageNumber="52" type="female">1 ♀</specimenCount>
(MACN-Ar 9933),
<specimenCount id="969EFFDE1F3FFFE0183BFA15FDBD47C3" box="[568,619,1496,1522]" pageId="52" pageNumber="52" type="female">1 ♀</specimenCount>
(
<collectionCode id="E689AC921F3FFFE01A66FA30FF10443E" box="[101,198,1529,1551]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="52" pageNumber="52">MHNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="802734571F3FFFE01A79F9DFFDBA44EE" blockId="52.[93,620,1113,1760]" pageId="52" pageNumber="52">
NEW RECORDS:
<emphasis id="B2ECE8451F3FFFE01B44F9DEFF1E447A" bold="true" pageId="52" pageNumber="52">
<collectingCountry id="F88F74C71F3FFFE01B44F9DEFE24441C" box="[327,498,1559,1581]" name="Argentina" pageId="52" pageNumber="52">ARGENTINA</collectingCountry>
:
<collectingRegion id="425CFAB51F3FFFE01801F9D1FF15447A" country="Argentina" name="La Rioja" pageId="52" pageNumber="52">La Rioja Province</collectingRegion>
:
</emphasis>
<emphasis id="B2ECE8451F3FFFE01ADAF9FFFE09447A" box="[217,479,1590,1611]" italics="true" pageId="52" pageNumber="52">
<taxonomicName id="47984FD41F3FFFE01ADAF9FFFE90447A" authorityName="Acosta" authorityYear="2001" box="[217,326,1590,1611]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">Famatina</taxonomicName>
Department
</emphasis>
: Sierra de
<taxonomicName id="47984FD41F3FFFE01A5EF99AFE054458" authority=", Cuevas de Noronha" authorityName="Cuevas de Noronha" box="[93,467,1619,1641]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">Famatina, Cuevas de Noronha</taxonomicName>
(path to La Mejicana mine), 28
<emphasis id="B2ECE8451F3FFFE01B5CF9B9FEBC44B4" bold="true" box="[351,362,1648,1669]" pageId="52" pageNumber="52">°</emphasis>
559310
<geoCoordinate id="E5AC52901F3FFFE01BB1F9B9FDD644B4" box="[434,512,1648,1670]" degrees="67" direction="south" orientation="latitude" pageId="52" pageNumber="52" precision="55555" value="-67.0">
S 67
<emphasis id="B2ECE8451F3FFFE01BF6F9B9FDD644B4" bold="true" box="[501,512,1648,1669]" pageId="52" pageNumber="52">°</emphasis>
</geoCoordinate>
409230W,
<quantity id="476099B21F3FFFE01A5EF947FF634492" box="[93,181,1678,1700]" metricMagnitude="3" metricUnit="m" metricValue="2.846" pageId="52" pageNumber="52" unit="m" value="2846.0">2846 m</quantity>
,
<date id="F42612971F3FFFE01AC1F947FEFA4495" box="[194,300,1678,1700]" pageId="52" pageNumber="52" value="2006-01-29">29.i.2006</date>
, L. Piacentini, L. Compagnucci, and A. Ojanguren Affilastro, 2 ♂,
<specimenCount id="969EFFDE1F3FFFE01839F962FDB544F3" box="[570,611,1704,1730]" pageId="52" pageNumber="52" type="female">6 ♀</specimenCount>
,
<specimenCount id="969EFFDE1F3FFFE01A5EF900FF7C44EE" box="[93,170,1737,1759]" pageId="52" pageNumber="52" type="generic">12 juv.</specimenCount>
(MACN-Ar),
<specimenCount id="969EFFDE1F3FFFE01B51F900FEAD44D1" box="[338,379,1734,1760]" pageId="52" pageNumber="52" type="female">2 ♀</specimenCount>
(
<collectionCode id="E689AC921F3FFFE01B8BF900FE3044EF" box="[392,486,1737,1758]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="52" pageNumber="52">AMNH</collectionCode>
[LP 5846]).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F3FFFE018BBFF1AFB7F4680" pageId="52" pageNumber="52" type="diagnosis">
<paragraph id="802734571F3FFFE018BBFF1AFB7F4680" blockId="52.[667,1194,211,1554]" pageId="52" pageNumber="52">
DIAGNOSIS:
<taxonomicName id="47984FD41F3FFFE0195EFF1DFBAB42DB" authorityName="Acosta" authorityYear="2001" box="[861,1149,212,235]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE0195EFF1DFBAB42DB" box="[861,1149,212,235]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
</taxonomicName>
is most closely related to
<taxonomicName id="47984FD41F3FFFE019B5FF3BFB9E4339" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[950,1096,241,264]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B2ECE8451F3FFFE019B5FF3BFB9E4339" box="[950,1096,241,264]" italics="true" pageId="52" pageNumber="52">O. calchaqui</emphasis>
</taxonomicName>
(fig. 5), from which it differs in the following respects.
<taxonomicName id="47984FD41F3FFFE01912FEE5FBFD4373" authorityName="Acosta" authorityYear="2001" box="[785,1067,300,323]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01912FEE5FBFD4373" box="[785,1067,300,323]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
</taxonomicName>
possesses one ML macroseta on metasomal segment IV and only 10
<emphasis id="B2ECE8451F3FFFE0195AFEAEFCA7434D" box="[857,881,359,380]" italics="true" pageId="52" pageNumber="52">%</emphasis>
of the specimens possess one DL macroseta on segment III (absent in 90
<emphasis id="B2ECE8451F3FFFE018B5FE6BFD184386" box="[694,718,418,439]" italics="true" pageId="52" pageNumber="52">%</emphasis>
), whereas
<taxonomicName id="47984FD41F3FFFE0194DFE6BFC084389" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[846,990,417,440]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B2ECE8451F3FFFE0194DFE6BFC084389" box="[846,990,417,440]" italics="true" pageId="52" pageNumber="52">O. calchaqui</emphasis>
</taxonomicName>
possesses two or three macrosetae on segment IV and 70
<emphasis id="B2ECE8451F3FFFE01E6CFE76FB5143E5" box="[1135,1159,447,468]" italics="true" pageId="52" pageNumber="52">%</emphasis>
of the specimens possess one DL macroseta on segment III (absent in 30
<emphasis id="B2ECE8451F3FFFE019C9FE33FC34403E" box="[970,994,506,527]" italics="true" pageId="52" pageNumber="52">%</emphasis>
). There are also differences in the trichobothrial pattern. Pedipalpal chela trichobothria
<emphasis id="B2ECE8451F3FFFE01E1FFDFAFBE1407B" box="[1052,1079,563,586]" italics="true" pageId="52" pageNumber="52">db</emphasis>
and
<emphasis id="B2ECE8451F3FFFE01E80FDFCFB48407B" box="[1155,1182,565,586]" italics="true" pageId="52" pageNumber="52">Et</emphasis>
<subScript id="1C1C36121F3FFFE01E9CFDF7FB7F407D" attach="left" box="[1183,1193,574,588]" fontSize="6" pageId="52" pageNumber="52">3</subScript>
are located in the same axis as, or slightly distal to
<emphasis id="B2ECE8451F3FFFE0191EFDA6FC9540B5" box="[797,835,623,644]" italics="true" pageId="52" pageNumber="52">Est</emphasis>
in 70
<emphasis id="B2ECE8451F3FFFE0199CFDA6FC6140B5" box="[927,951,623,644]" italics="true" pageId="52" pageNumber="52">%</emphasis>
and 30
<emphasis id="B2ECE8451F3FFFE01E2AFDA6FB9740B5" box="[1065,1089,623,644]" italics="true" pageId="52" pageNumber="52">%</emphasis>
of the specimens of
<taxonomicName id="47984FD41F3FFFE01940FD45FC044090" authorityName="Acosta" authorityYear="2001" box="[835,978,652,673]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01940FD45FC044090" box="[835,978,652,673]" italics="true" pageId="52" pageNumber="52">O. famatina</emphasis>
</taxonomicName>
, respectively, but always located proximal to
<emphasis id="B2ECE8451F3FFFE019E0FD63FBDF408E" box="[995,1033,682,703]" italics="true" pageId="52" pageNumber="52">Est</emphasis>
in
<taxonomicName id="47984FD41F3FFFE01E31FD60FD6B40ED" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B2ECE8451F3FFFE01E31FD60FD6B40ED" italics="true" pageId="52" pageNumber="52">O. calchaqui</emphasis>
</taxonomicName>
. Pedipalp patella trichobothrium
<emphasis id="B2ECE8451F3FFFE01E5CFD0EFB5640ED" box="[1119,1152,711,732]" italics="true" pageId="52" pageNumber="52">em</emphasis>
<subScript id="1C1C36121F3FFFE01E83FD19FB5C40EF" attach="left" box="[1152,1162,720,734]" fontSize="6" pageId="52" pageNumber="52">2</subScript>
is located in the same axis as, or distal to
<emphasis id="B2ECE8451F3FFFE01E63FD2DFB5740C8" box="[1120,1153,740,761]" italics="true" pageId="52" pageNumber="52">em</emphasis>
<quantity id="476099B21F3FFFE01E82FD24FB7F40CB" box="[1153,1193,740,763]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="52" pageNumber="52" unit="in" value="1.0">
<subScript id="1C1C36121F3FFFE01E82FD24FB5D40CA" attach="left" box="[1153,1163,749,763]" fontSize="6" pageId="52" pageNumber="52">1</subScript>
in
</quantity>
70
<emphasis id="B2ECE8451F3FFFE018B5FCC8FD184127" box="[694,718,769,790]" italics="true" pageId="52" pageNumber="52">%</emphasis>
of the specimens of
<taxonomicName id="47984FD41F3FFFE019DEFCC8FBBA4127" authorityName="Acosta" authorityYear="2001" box="[989,1132,769,790]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE019DEFCC8FBBA4127" box="[989,1132,769,790]" italics="true" pageId="52" pageNumber="52">O. famatina</emphasis>
</taxonomicName>
, but proximal to
<emphasis id="B2ECE8451F3FFFE01933FCD6FC874105" box="[816,849,799,820]" italics="true" pageId="52" pageNumber="52">em</emphasis>
<quantity id="476099B21F3FFFE01952FCE1FCAA4105" box="[849,892,798,822]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="52" pageNumber="52" unit="in" value="1.0">
<subScript id="1C1C36121F3FFFE01952FCE1FC8D4107" attach="left" box="[849,859,808,822]" fontSize="6" pageId="52" pageNumber="52">1</subScript>
in
</quantity>
80
<emphasis id="B2ECE8451F3FFFE019A3FCD7FC6E4102" box="[928,952,798,819]" italics="true" pageId="52" pageNumber="52">%</emphasis>
of the specimens of
<taxonomicName id="47984FD41F3FFFE01898FCF5FCFD4163" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[667,811,827,850]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B2ECE8451F3FFFE01898FCF5FCFD4163" box="[667,811,827,850]" italics="true" pageId="52" pageNumber="52">O. calchaqui</emphasis>
</taxonomicName>
.
<taxonomicName id="47984FD41F3FFFE01938FCF2FB9C4161" authorityName="Acosta" authorityYear="2001" box="[827,1098,827,850]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01938FCF2FB9C4161" box="[827,1098,827,850]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
</taxonomicName>
may be differentiated from
<taxonomicName id="47984FD41F3FFFE0198EFC90FB9D415F" authorityName="Ochoa, Ojanguren Affilastro, Mattoni &amp; Prendini" authorityYear="2011" box="[909,1099,857,878]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="compagnucci">
<emphasis id="B2ECE8451F3FFFE0198EFC90FB9D415F" box="[909,1099,857,878]" italics="true" pageId="52" pageNumber="52">O. compagnucci</emphasis>
</taxonomicName>
by the more elongated apex of the hemispermatophore lamina: the apex comprises 66.6 68.48
<emphasis id="B2ECE8451F3FFFE018DAFC78FD2741F7" box="[729,753,945,966]" italics="true" pageId="52" pageNumber="52">%</emphasis>
(
<geoCoordinate id="E5AC52901F3FFFE01909FC78FCB641F7" box="[778,864,943,966]" degrees="5" direction="north" minutes="3" orientation="latitude" pageId="52" pageNumber="52" precision="925" value="5.05">
<emphasis id="B2ECE8451F3FFFE01909FC78FCCE41F7" box="[778,792,945,966]" italics="true" pageId="52" pageNumber="52">n</emphasis>
5 3
</geoCoordinate>
, mean 5 67.02
<emphasis id="B2ECE8451F3FFFE01E29FC78FB9441F7" box="[1066,1090,945,966]" italics="true" pageId="52" pageNumber="52">%</emphasis>
) of the lamina in
<taxonomicName id="47984FD41F3FFFE01928FC07FC1741D2" authorityName="Acosta" authorityYear="2001" box="[811,961,974,995]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01928FC07FC1741D2" box="[811,961,974,995]" italics="true" pageId="52" pageNumber="52">O. famatina</emphasis>
</taxonomicName>
(fig. 36FH) and 52.17
<emphasis id="B2ECE8451F3FFFE018DAFC22FD274631" box="[729,753,1003,1024]" italics="true" pageId="52" pageNumber="52">%</emphasis>
54.54
<emphasis id="B2ECE8451F3FFFE0193FFC22FC824631" box="[828,852,1003,1024]" italics="true" pageId="52" pageNumber="52">%</emphasis>
(
<geoCoordinate id="E5AC52901F3FFFE0196BFC25FC664631" box="[872,944,1002,1025]" degrees="5" direction="north" minutes="3" orientation="latitude" pageId="52" pageNumber="52" precision="925" value="5.05">
<emphasis id="B2ECE8451F3FFFE0196BFC25FCA04630" box="[872,886,1004,1025]" italics="true" pageId="52" pageNumber="52">n</emphasis>
5 3
</geoCoordinate>
; mean 5 52.98
<emphasis id="B2ECE8451F3FFFE01E64FC22FBA94631" box="[1127,1151,1003,1024]" italics="true" pageId="52" pageNumber="52">%</emphasis>
) in
<taxonomicName id="47984FD41F3FFFE01898FBC0FC8D462F" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[667,859,1033,1054]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="compagnuccii">
<emphasis id="B2ECE8451F3FFFE01898FBC0FC8D462F" box="[667,859,1033,1054]" italics="true" pageId="52" pageNumber="52">O. compagnuccii</emphasis>
</taxonomicName>
(fig. 36AC). The two species can also be separated by the VL and VSM carinae of metasomal segment I and sternite VII in the male, which are well developed in
<taxonomicName id="47984FD41F3FFFE0193AFBB7FC1246A3" authorityName="Acosta" authorityYear="2001" box="[825,964,1149,1171]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE0193AFBB7FC1246A3" box="[825,964,1149,1171]" italics="true" pageId="52" pageNumber="52">O. famatina</emphasis>
</taxonomicName>
, and weakly developed to absent in
<taxonomicName id="47984FD41F3FFFE0196DFB52FBFD4681" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[878,1067,1179,1200]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="compagnuccii">
<emphasis id="B2ECE8451F3FFFE0196DFB52FBFD4681" box="[878,1067,1179,1200]" italics="true" pageId="52" pageNumber="52">O. compagnuccii</emphasis>
</taxonomicName>
(fig. 18F).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F3FFFE018BBFB7EFB7F4775" pageId="52" pageNumber="52" type="distribution">
<paragraph id="802734571F3FFFE018BBFB7EFB7F4775" blockId="52.[667,1194,211,1554]" pageId="52" pageNumber="52">
DISTRIBUTION:
<taxonomicName id="47984FD41F3FFFE01974FB71FB5C46FF" authorityName="Acosta" authorityYear="2001" box="[887,1162,1208,1231]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F3FFFE01974FB71FB5C46FF" box="[887,1162,1208,1231]" italics="true" pageId="52" pageNumber="52">Orobothriurus famatina</emphasis>
</taxonomicName>
is known only from high altitudes (
<quantity id="476099B21F3FFFE01E60FB1FFD254738" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="52" pageNumber="52" unit="m" value="2850.0" valueMax="3200.0" valueMin="2500.0">2500 3200 m</quantity>
) of the Sierra Famatina, a mountain range close to the Andes, in
<collectingRegion id="425CFAB51F3FFFE019FDFAD8FD024775" country="Argentina" name="La Rioja" pageId="52" pageNumber="52">La Rioja Province</collectingRegion>
of central-western
<collectingCountry id="F88F74C71F3FFFE019A3FAE7FBD84775" box="[928,1038,1326,1348]" name="Argentina" pageId="52" pageNumber="52">Argentina</collectingCountry>
(figs. 3G, 54).
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F3FFFE018BBFA83FC274423" pageId="52" pageNumber="52" type="biology_ecology">
<paragraph id="802734571F3FFFE018BBFA83FC274423" blockId="52.[667,1194,211,1554]" pageId="52" pageNumber="52">
ECOLOGY: This species inhabits an area of grassland and shrub steppe that belongs to the Prepuna and Altoandina phytogeographic provinces (
<bibRefCitation id="E40949A61F3FFFE01949FA6DFB734788" author="Cabrera, A. L. &amp; A. W. Willink" box="[842,1189,1444,1466]" pageId="52" pageNumber="52" refId="ref42425" refString="Cabrera, A. L., and A. W. Willink. 1980. Biogeografia de America Latina. Serie de Biologia, OEA Monographs vol. 13, 122 pp." type="book" year="1980">Cabrera and Willink, 1980</bibRefCitation>
; fig. 3G). It occurs in sympatry with another bothriurid,
<taxonomicName id="47984FD41F3FFFE0192EFA17FB7F47C5" authority="Acosta, 1997" authorityName="Acosta" authorityYear="1997" box="[813,1193,1502,1525]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="52" pageNumber="52" phylum="Arthropoda" rank="species" species="olaen">
<emphasis id="B2ECE8451F3FFFE0192EFA17FC2247C4" box="[813,1012,1502,1525]" italics="true" pageId="52" pageNumber="52">Bothriurus olaen</emphasis>
<bibRefCitation id="E40949A61F3FFFE01E04FA16FB7F47C5" author="Acosta, L. E." box="[1031,1193,1503,1524]" pageId="52" pageNumber="52" pagination="1 - 8" refId="ref41706" refString="Acosta, L. E. 1997. Descripcion de Bothriurus olaen, nueva especie de escorpion de Argentina central (Scorpiones: Bothriuridae). Revue Arachnologique 12 (1): 1 - 8." type="journal article" year="1997">Acosta, 1997</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E40949A61F3FFFE018A1FA35FC334423" author="Ojanguren Affilastro, A. A." box="[674,997,1532,1554]" pageId="52" pageNumber="52" pagination="75 - 241" refId="ref45124" refString="Ojanguren Affilastro, A. A. 2005. Estudio monografico de los escorpiones de la Republica Argentina. Revista Iberica de Aracnologia 11: 75 - 241." type="journal article" year="2005">Ojanguren Affilastro, 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>