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<mods:title id="B8A109460D95DEE461C9276FA0F5CA09">A revision of the taxonomy and distribution of Archispirostreptus Silvestri 1895 (Diplopoda, Spirostreptida, Spirostreptidae), and description of a new spirostreptid genus with three new species</mods:title>
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<mods:namePart id="0F14054C134D6859215F98DF1D70B581">Barraclough, David</mods:namePart>
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<treatment id="080E878B1C6C770EFF144AD52A597870" ID-DOI="http://doi.org/10.5281/zenodo.6207853" ID-GBIF-Taxon="119394063" ID-Zenodo-Dep="6207853" LSID="urn:lsid:plazi:treatment:080E878B1C6C770EFF144AD52A597870" httpUri="http://treatment.plazi.org/id/080E878B1C6C770EFF144AD52A597870" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13">
<subSubSection id="C8BD65161C6C7713FF144AD5288E7CDF" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8018369D1C6C7713FF144AD52A327CF2" blockId="12.[151,615,1451,1512]" box="[151,615,1451,1477]" pageId="12" pageNumber="13">
<heading id="DB5081F11C6C7713FF144AD52A327CF2" bold="true" box="[151,615,1451,1477]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="47A74D1E1C6C7713FF144AD52A327CF2" ID-CoL="G9Q6" authority="Peters 1855" authorityName="Peters" authorityYear="1855" box="[151,615,1451,1477]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6C7713FF144AD52A327CF2" bold="true" box="[151,615,1451,1477]" pageId="12" pageNumber="13">
<emphasis id="B2D3EA8F1C6C7713FF144AD529E87CF2" bold="true" box="[151,445,1451,1477]" italics="true" pageId="12" pageNumber="13">Archispirostreptus gigas</emphasis>
(
<bibRefCitation id="E4364B6C1C6C7713FE484AD52A0A7CF2" author="Peters" box="[459,607,1451,1477]" pageId="12" pageNumber="13" refString="Peters, W. (1855) Uber die Myriapoden im allgemeinen und insbesondere uber die in Mossambique beobachteten Arten dieser Familie. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 75, 75 - 79." type="journal article" year="1855">Peters 1855</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8018369D1C6C7713FF144AB0288E7CDF" blockId="12.[151,615,1451,1512]" box="[151,219,1486,1512]" pageId="12" pageNumber="13">
<figureCitation id="189C2A181C6C7713FF144AB0288E7CDF" box="[151,219,1486,1512]" captionStart="FIGURE 5 a, b" captionStartId="15.[151,255,1982,2006]" captionTargetBox="[200,1394,233,1944]" captionTargetId="figure@15.[194,1399,194,1965]" captionTargetPageId="15" captionText="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C6C7713FF14496D2A4F7F9D" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8018369D1C6C7713FF14496D2AF77F1D" blockId="12.[151,1436,1555,1706]" box="[151,674,1555,1578]" pageId="12" pageNumber="13">
<taxonomicName id="47A74D1E1C6C7713FF14496D29BF7F1D" ID-CoL="4Z8SV" authority="Peters 1855" authorityName="Peters" authorityYear="1855" box="[151,490,1555,1578]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6C7713FF14496D29317F1D" box="[151,356,1555,1578]" italics="true" pageId="12" pageNumber="13">Spirostreptus gigas</emphasis>
<bibRefCitation id="E4364B6C1C6C7713FEE9496D29BF7F1D" author="Peters" box="[362,490,1555,1578]" pageId="12" pageNumber="13" refString="Peters, W. (1855) Uber die Myriapoden im allgemeinen und insbesondere uber die in Mossambique beobachteten Arten dieser Familie. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 75, 75 - 79." type="journal article" year="1855">Peters 1855</bibRefCitation>
</taxonomicName>
, p. 536, figs 12.
</paragraph>
<paragraph id="8018369D1C6C7713FF14494D296B7F5D" blockId="12.[151,1436,1555,1706]" pageId="12" pageNumber="13">
<taxonomicName id="47A74D1E1C6C7713FF14494D2A167F7D" ID-CoL="G9Q6" authority="Silvestri 1895" authorityName="Silvestri" authorityYear="1895" box="[151,579,1587,1610]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6C7713FF14494D29C97F7D" box="[151,412,1587,1610]" italics="true" pageId="12" pageNumber="13">Archispirostreptus gigas</emphasis>
:
<bibRefCitation id="E4364B6C1C6C7713FE29494D2A167F7D" author="Silvestri" box="[426,579,1587,1610]" pageId="12" pageNumber="13" refString="Silvestri, F. (1895) Chilopodi e diplopodi. Annali del Museo Civico di Storia Naturale di Genova, (2) 15, 479 - 490. Genova." type="journal article" year="1895">Silvestri 1895</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="E4364B6C1C6C7713FDCC494D2AB87F7D" author="Schubart" box="[591,749,1587,1610]" pageId="12" pageNumber="13" refString="Schubart, O. (1951) Uber einige Diplopoden vom Bergmassiv Air in der Sud-Sahara, gesammelt von L. Chopard und A. Villiers im Jahre 1947. Bulletin de I'Institut Francaise d'Afrique Noire, 13, 116 - 125." type="journal article" year="1951">Schubart 1951</bibRefCitation>
;
<bibRefCitation id="E4364B6C1C6C7713FD7A494D2BCD7F7D" author="Hoffman" box="[761,920,1587,1610]" pageId="12" pageNumber="13" refString="Hoffman, R. L. (1965) A new species in the diplopod genus Archispirostreptus from south western Arabia. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 3 (51), 17 - 23." type="journal article" year="1965">Hoffman 1965</bibRefCitation>
;
<bibRefCitation id="E4364B6C1C6C7713FC26494D2CF87F7D" author="Krabbe" box="[933,1197,1587,1610]" pageId="12" pageNumber="13" refString="Krabbe, E. &amp; Enghoff, H. (1978) A new species of Archispirostreptus from Kenya (Diplopoda: Spirostreptida, Spirostreptidae). Entomologica Scandinavica, 9, 247 - 249." type="journal article" year="1978">Krabbe &amp; Enghoff 1978</bibRefCitation>
;
<bibRefCitation id="E4364B6C1C6C7713FB3A494D2D1D7F7D" author="Krabbe" box="[1209,1352,1587,1610]" pageId="12" pageNumber="13" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
;
<bibRefCitation id="E4364B6C1C6C7713FAD0494D28AA7F5D" author="Hamer" pageId="12" pageNumber="13" refString="Hamer, M. (1998) Checklist of Southern African millipedes. Annals of the Natal Museum, 39, 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
,
<bibRefCitation id="E4364B6C1C6C7713FE8A492D296B7F5D" author="Hamer" box="[265,318,1619,1642]" pageId="12" pageNumber="13" refString="Hamer, M. (1999) An illustrated key to the spirostreptidian (Diplopoda: Spirostreptida) genera of Southern Africa. Annals of the Natal Museum, 40, 1 - 22." type="journal article" year="1999">1999</bibRefCitation>
</paragraph>
<paragraph id="8018369D1C6C7713FF14490D2A957FBD" blockId="12.[151,1436,1555,1706]" box="[151,704,1651,1674]" pageId="12" pageNumber="13">
<taxonomicName id="47A74D1E1C6C7713FF14490D2A797FBD" ID-CoL="3H8V8" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[151,556,1651,1674]" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6C7713FF14490D29C67FBD" box="[151,403,1651,1674]" italics="true" pageId="12" pageNumber="13">Graphidostreptus gigas</emphasis>
:
<bibRefCitation id="E4364B6C1C6C7713FE23490D2A797FBD" author="Attems" box="[416,556,1651,1674]" pageId="12" pageNumber="13" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25, 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="E4364B6C1C6C7713FDB4490D2A957FBD" author="Attems" box="[567,704,1651,1674]" pageId="12" pageNumber="13" refString="Attems, C. (1928) The Myriapoda of South Africa. Annals of the South African Museum, 26, 1 - 431." type="journal article" year="1928">Attems 1928</bibRefCitation>
</paragraph>
<paragraph id="8018369D1C6C7713FF1449ED2A4F7F9D" blockId="12.[151,1436,1555,1706]" box="[151,538,1683,1706]" pageId="12" pageNumber="13">
Further synonymy in
<bibRefCitation id="E4364B6C1C6C7713FEFE49ED2A4F7F9D" author="Krabbe" box="[381,538,1683,1706]" pageId="12" pageNumber="13" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe (1982)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C6C7713FF1449AD2A5B7E6A" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8018369D1C6C7713FF1449AD2D247FDA" blockId="12.[151,1437,1747,2035]" box="[151,1393,1747,1773]" pageId="12" pageNumber="13">
<emphasis id="B2D3EA8F1C6C7713FF1449AD29117FDA" bold="true" box="[151,324,1747,1773]" pageId="12" pageNumber="13">
<typeStatus id="5F1C883F1C6C7713FF1449AD28877FDA" box="[151,210,1747,1773]" pageId="12" pageNumber="13">Type</typeStatus>
material
</emphasis>
(not examined):
<collectingCountry id="F8B0760D1C6C7713FD8D49AD2A887FDA" box="[526,733,1747,1773]" name="Mozambique" pageId="12" pageNumber="13">MOZAMBIQUE</collectingCountry>
: 1 3, Tete.The location of the
<typeStatus id="5F1C883F1C6C7713FBC249AD2C267FDA" box="[1089,1139,1747,1773]" pageId="12" pageNumber="13">type</typeStatus>
material is unknown.
</paragraph>
<paragraph id="8018369D1C6C7713FF4649862A5B7E6A" blockId="12.[151,1437,1747,2035]" pageId="12" pageNumber="13">
<emphasis id="B2D3EA8F1C6C7713FF4649862A137E25" bold="true" box="[197,582,1784,1810]" pageId="12" pageNumber="13">Additional material examined:</emphasis>
<collectingCountry id="F8B0760D1C6C7713FDCE49872AE07E24" box="[589,693,1785,1811]" name="Kenya" pageId="12" pageNumber="13">KENYA</collectingCountry>
: 1 3, 2 Ƥ, Tsavo, Taita Discovery Center [
<geoCoordinate id="E593505A1C6C7713FB2A49872D4D7E24" box="[1193,1304,1785,1811]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="-3.4166667">03° 25' S</geoCoordinate>
,
<geoCoordinate id="E593505A1C6C7713FAA549872DC57E24" box="[1318,1424,1785,1811]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="38.766666">38° 46'E</geoCoordinate>
],
<date id="F419105D1C6C7713FF14486029447E0F" box="[151,273,1822,1848]" pageId="12" pageNumber="13" value="2000-12-06">6.xii.2000</date>
, Van den Spiegel D. (
<collectionCode id="E6B6AE581C6C7713FD9148602A3F7E0F" box="[530,618,1822,1848]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="12" pageNumber="13">MRAC</collectionCode>
20254); 1 3, 1 Ƥ, Ngaia Forest [
<geoCoordinate id="E593505A1C6C7713FC7548602C097E0F" box="[1014,1116,1822,1848]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="0.4">0° 24' N</geoCoordinate>
,
<geoCoordinate id="E593505A1C6C7713FBEA48602C817E0F" box="[1129,1236,1822,1848]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="38.033333">38° 02'E</geoCoordinate>
],
<date id="F419105D1C6C7713FB6B48602D347E0F" box="[1256,1377,1822,1848]" pageId="12" pageNumber="13" value="2002-12-03">3.xii.2002</date>
, Van den Spiegel D. (
<collectionCode id="E6B6AE581C6C7713FED0483D29FE7E6A" box="[339,427,1859,1885]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="12" pageNumber="13">MRAC</collectionCode>
20764).
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C6C7713FF4648162B9E7EC4" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8018369D1C6C7713FF4648162B9E7EC4" blockId="12.[151,1437,1747,2035]" pageId="12" pageNumber="13">
<emphasis id="B2D3EA8F1C6C7713FF46481629187EB5" bold="true" box="[197,333,1896,1922]" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Lateral metaplical process short and broad with a short terminal spine; lateral metaplical process at acute angle with apical metaplical process; lateral and medial edges of apical metaplical process convex proximally; lateral metaplical process darker proximally than distally; apical metaplical process darker laterally and proximally than medially or distally (
<figureCitation id="189C2A181C6C7713FCB448A72BE87EC4" box="[823,957,2009,2035]" captionStart="FIGURE 5 a, b" captionStartId="15.[151,255,1982,2006]" captionTargetBox="[200,1394,233,1944]" captionTargetId="figure@15.[194,1399,194,1965]" captionTargetPageId="15" captionText="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="12" pageNumber="13">Figs 5a, 5b</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D4D866151C6D7712FF1448D929537EE9" httpUri="https://zenodo.org/record/197295/files/figure.png" pageId="13" pageNumber="14" targetBox="[377,1219,200,1936]" targetPageId="13">
<paragraph id="8018369D1C6D7712FF1448D929537EE9" blockId="13.[151,1437,1959,2014]" pageId="13" pageNumber="14">
<emphasis id="B2D3EA8F1C6D7712FF1448D929187E88" bold="true" box="[151,333,1959,1983]" pageId="13" pageNumber="14">FIGURE 4a, b.</emphasis>
<taxonomicName id="47A74D1E1C6D7712FED648D62AD67E88" box="[341,643,1960,1983]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dodsoni">
<emphasis id="B2D3EA8F1C6D7712FED648D62AD67E88" box="[341,643,1960,1983]" italics="true" pageId="13" pageNumber="14">Archispirostreptus dodsoni</emphasis>
</taxonomicName>
(BMNH 1897.11.10.87).
<emphasis id="B2D3EA8F1C6D7712FC2948D92BEA7E88" bold="true" box="[938,959,1959,1983]" pageId="13" pageNumber="14">a,</emphasis>
oral view of gonopods;
<emphasis id="B2D3EA8F1C6D7712FB5B48D92CBA7E88" bold="true" box="[1240,1263,1959,1983]" pageId="13" pageNumber="14">b,</emphasis>
aboral view of gonopods.
</paragraph>
</caption>
<caption id="D4D866151C6E7711FF144CA1289C7D21" httpUri="https://zenodo.org/record/197296/files/figure.png" pageId="14" pageNumber="15" targetBox="[155,1432,191,963]" targetPageId="14">
<paragraph id="8018369D1C6E7711FF144CA1289C7D21" blockId="14.[151,1436,991,1046]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF144CA129727AC0" bold="true" box="[151,295,991,1015]" pageId="14" pageNumber="15">FIGURE 4c.</emphasis>
<taxonomicName id="47A74D1E1C6E7711FEAC4CA12A057AC1" box="[303,592,991,1014]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dodsoni">
<emphasis id="B2D3EA8F1C6E7711FEAC4CA12A057AC1" box="[303,592,991,1014]" italics="true" pageId="14" pageNumber="15">Archispirostreptus dodsoni</emphasis>
</taxonomicName>
(BMNH 1897.11.10.87).
<emphasis id="B2D3EA8F1C6E7711FCE84CA12B2B7AC0" bold="true" box="[875,894,991,1015]" pageId="14" pageNumber="15">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs.
</paragraph>
</caption>
<subSubSection id="C8BD65161C6E7711FF464B412B0E7FCB" pageId="14" pageNumber="15" type="description">
<paragraph id="8018369D1C6E7711FF464B412C227D6E" blockId="14.[151,1437,1087,2012]" box="[197,1143,1087,1113]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464B41290A7D6E" bold="true" box="[197,351,1087,1113]" pageId="14" pageNumber="15">Description:</emphasis>
<emphasis id="B2D3EA8F1C6E7711FEE44B4129C37D6F" box="[359,406,1087,1112]" italics="true" pageId="14" pageNumber="15">Size</emphasis>
: Body length
<quantity id="475F9B781C6E7711FDBF4B412A8E7D6E" box="[572,731,1087,1113]" metricMagnitude="-1" metricUnit="m" metricValue="2.19" metricValueMax="2.6" metricValueMin="1.78" pageId="14" pageNumber="15" unit="mm" value="219.0" valueMax="260.0" valueMin="178.0">178260 mm</quantity>
; maximum body width
<quantity id="475F9B781C6E7711FC724B412C267D6E" box="[1009,1139,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="14" pageNumber="15" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">1519 mm</quantity>
.
</paragraph>
<paragraph id="8018369D1C6E7711FF464B1B2A737D49" blockId="14.[151,1437,1087,2012]" box="[197,550,1124,1150]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464B1B29907D49" box="[197,453,1125,1150]" italics="true" pageId="14" pageNumber="15">Number of body rings</emphasis>
: 6270.
</paragraph>
<paragraph id="8018369D1C6E7711FF464BF429267D93" blockId="14.[151,1437,1087,2012]" box="[197,371,1162,1188]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464BF4294D7D94" box="[197,280,1162,1187]" italics="true" pageId="14" pageNumber="15">Colour</emphasis>
: Black.
</paragraph>
<paragraph id="8018369D1C6E7711FF464BD12D187DFE" blockId="14.[151,1437,1087,2012]" box="[197,1357,1199,1225]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464BD1294E7DFF" box="[197,283,1199,1224]" italics="true" pageId="14" pageNumber="15">Collum</emphasis>
: Rounded without anterior lobe (
<figureCitation id="189C2A181C6E7711FD224BD12AA27DFE" box="[673,759,1199,1225]" captionStart-0="FIGURE 5 a, b" captionStart-1="FIGURE 5 c, d" captionStartId-0="15.[151,255,1982,2006]" captionStartId-1="16.[151,256,1953,1977]" captionTargetBox-0="[200,1394,233,1944]" captionTargetBox-1="[286,1299,211,1926]" captionTargetId-0="figure@15.[194,1399,194,1965]" captionTargetId-1="figure@16.[280,1307,194,1932]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." captionText-1="FIGURE 5 c, d. Archispirostreptus gigas (MRAC 20764). c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri-0="https://zenodo.org/record/197297/files/figure.png" httpUri-1="https://zenodo.org/record/197298/files/figure.png" pageId="14" pageNumber="15">Fig. 5c</figureCitation>
), with 23 complete striae and an incomplete stria.
</paragraph>
<paragraph id="8018369D1C6E7711FF464BA629567C20" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464BA62A807DC6" box="[197,725,1237,1265]" italics="true" pageId="14" pageNumber="15">Pre-femoral process of 1st pair of male legs</emphasis>
: Proximally wide with laterally deflected apical extension (
<figureCitation id="189C2A181C6E7711FF1C4B8328A07C20" box="[159,245,1277,1303]" captionStart-0="FIGURE 5 a, b" captionStart-1="FIGURE 5 c, d" captionStartId-0="15.[151,255,1982,2006]" captionStartId-1="16.[151,256,1953,1977]" captionTargetBox-0="[200,1394,233,1944]" captionTargetBox-1="[286,1299,211,1926]" captionTargetId-0="figure@15.[194,1399,194,1965]" captionTargetId-1="figure@16.[280,1307,194,1932]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." captionText-1="FIGURE 5 c, d. Archispirostreptus gigas (MRAC 20764). c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri-0="https://zenodo.org/record/197297/files/figure.png" httpUri-1="https://zenodo.org/record/197298/files/figure.png" pageId="14" pageNumber="15">Fig. 5c</figureCitation>
).
</paragraph>
<paragraph id="8018369D1C6E7711FF464A5C2BF97C56" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464A5C29677C0C" box="[197,306,1314,1339]" italics="true" pageId="14" pageNumber="15">Gonopod</emphasis>
: (
<figureCitation id="189C2A181C6E7711FEC94A5C299B7C0B" box="[330,462,1314,1340]" captionStart="FIGURE 5 a, b" captionStartId="15.[151,255,1982,2006]" captionTargetBox="[200,1394,233,1944]" captionTargetId="figure@15.[194,1399,194,1965]" captionTargetPageId="15" captionText="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="14" pageNumber="15">Figs 5a, 5b</figureCitation>
) Sternum broad, short and apically rounded; apex of sternum not reaching level of paracoxite apex; paracoxite rounded and fused tightly to metaplica.
</paragraph>
<paragraph id="8018369D1C6E7711FF464A132B3B7C9B" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">Proximal proplica with raised lateral and medial edges; apically proplicae rounded, overlapping lateral metaplical process proximally and with an apical medial lobe.</paragraph>
<paragraph id="8018369D1C6E7711FF464AC92B027CE6" blockId="14.[151,1437,1087,2012]" box="[197,855,1463,1489]" pageId="14" pageNumber="15">Proximal medial edges of oral fold of metaplicae raised.</paragraph>
<paragraph id="8018369D1C6E7711FF464AA329667F2B" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">
Proximal half of opposite telocoxites touching medially, further apart distal to lateral metaplical process (
<figureCitation id="189C2A181C6E7711FF1C497C29707F2B" box="[159,293,1538,1564]" captionStart="FIGURE 5 a, b" captionStartId="15.[151,255,1982,2006]" captionTargetBox="[200,1394,233,1944]" captionTargetId="figure@15.[194,1399,194,1965]" captionTargetPageId="15" captionText="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="14" pageNumber="15">Figs 5a, 5b</figureCitation>
).
</paragraph>
<paragraph id="8018369D1C6E7711FF4649592CCC7F50" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">Lateral metaplical processes short and broad, crescent-shaped and tapering distally, and may have a small terminal spine. Lateral metaplical process at acute angle with apical metaplical process.</paragraph>
<paragraph id="8018369D1C6E7711FF46490C29FE7F86" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">Lateral and medial edges of apical metaplical process convex proximally; apical metaplical process broad and opaque proximally.</paragraph>
<paragraph id="8018369D1C6E7711FF4649C32B657FE0" blockId="14.[151,1437,1087,2012]" box="[197,816,1725,1751]" pageId="14" pageNumber="15">
Aborally, medial edge of metaplicae raised (
<figureCitation id="189C2A181C6E7711FD4849C32B777FE0" box="[715,802,1725,1751]" captionStart="FIGURE 5 a, b" captionStartId="15.[151,255,1982,2006]" captionTargetBox="[200,1394,233,1944]" captionTargetId="figure@15.[194,1399,194,1965]" captionTargetPageId="15" captionText="FIGURE 5 a, b. Archispirostreptus gigas (MRAC 20764). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="14" pageNumber="15">Fig. 5b</figureCitation>
).
</paragraph>
<paragraph id="8018369D1C6E7711FF46499C2B0E7FCB" blockId="14.[151,1437,1087,2012]" box="[197,859,1762,1788]" pageId="14" pageNumber="15">Apex of antetorsal process extends past paracoxite apex.</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C6E7711FF46487928BF7E70" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8018369D1C6E7711FF46487928BF7E70" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF464879293E7E16" bold="true" box="[197,363,1799,1825]" pageId="14" pageNumber="15">Distribution:</emphasis>
Known from
<collectingCountry id="F8B0760D1C6E7711FD9048792AE47E16" box="[531,689,1799,1825]" name="Mozambique" pageId="14" pageNumber="15">Mozambique</collectingCountry>
,
<collectingCountry id="F8B0760D1C6E7711FD3D48792B0D7E16" box="[702,856,1799,1825]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
,
<collectingCountry id="F8B0760D1C6E7711FCE648792B9A7E16" box="[869,975,1799,1825]" name="Tanzania" pageId="14" pageNumber="15">Zanzibar</collectingCountry>
,
<collectingCountry id="F8B0760D1C6E7711FC5848792C6B7E16" box="[987,1086,1799,1825]" name="Somalia" pageId="14" pageNumber="15">Somalia</collectingCountry>
,
<collectingCountry id="F8B0760D1C6E7711FBC848792CE17E16" box="[1099,1204,1799,1825]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
(
<bibRefCitation id="E4364B6C1C6E7711FB4648792D347E16" author="Krabbe" box="[1221,1377,1799,1825]" pageId="14" pageNumber="15" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
) and
<collectingCountry id="F8B0760D1C6E7711FF14485328B37E70" box="[151,230,1837,1863]" name="Kenya" pageId="14" pageNumber="15">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C6E770EFF46482C2A597870" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8018369D1C6E7711FF46482C28887EEB" blockId="14.[151,1437,1087,2012]" pageId="14" pageNumber="15">
<emphasis id="B2D3EA8F1C6E7711FF46482C29177E5B" bold="true" box="[197,322,1874,1900]" pageId="14" pageNumber="15">Remarks:</emphasis>
The opposite telocoxites of the gonopods in
<taxonomicName id="47A74D1E1C6E7711FCE7482C2B927E5C" box="[868,967,1874,1899]" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6E7711FCE7482C2B927E5C" box="[868,967,1874,1899]" italics="true" pageId="14" pageNumber="15">A. gigas</emphasis>
</taxonomicName>
are separated at the level of the lateral metaplical processes as in
<taxonomicName id="47A74D1E1C6E7711FE4848062A067EA6" box="[459,595,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phillipsii">
<emphasis id="B2D3EA8F1C6E7711FE4848062A067EA6" box="[459,595,1912,1937]" italics="true" pageId="14" pageNumber="15">A. phillipsii</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C6E7711FDE248062AB77EA6" box="[609,738,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C6E7711FDE248062AB77EA6" box="[609,738,1912,1937]" italics="true" pageId="14" pageNumber="15">A. lugubris</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C6E7711FD7248062B277EA6" box="[753,882,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B2D3EA8F1C6E7711FD7248062B277EA6" box="[753,882,1912,1937]" italics="true" pageId="14" pageNumber="15">A. syriacus</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C6E7711FC0248062C7B7EA6" box="[897,1070,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tumuliporus">
<emphasis id="B2D3EA8F1C6E7711FC0248062C7B7EA6" box="[897,1070,1912,1937]" italics="true" pageId="14" pageNumber="15">A. tumuliporus</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C6E7711FBBF48062CF97EA6" box="[1084,1196,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="smithii">
<emphasis id="B2D3EA8F1C6E7711FBBF48062CF97EA6" box="[1084,1196,1912,1937]" italics="true" pageId="14" pageNumber="15">A. smithii</emphasis>
</taxonomicName>
and
<taxonomicName id="47A74D1E1C6E7711FB6748062D347EA6" box="[1252,1377,1912,1937]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dodsoni">
<emphasis id="B2D3EA8F1C6E7711FB6748062D347EA6" box="[1252,1377,1912,1937]" italics="true" pageId="14" pageNumber="15">A. dodsoni</emphasis>
</taxonomicName>
. The convex shapes of the lateral and medial edges of the apical metaplical process of the gonopod are unique to
<taxonomicName id="47A74D1E1C6E7711FA0048E328837EEC" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6E7711FA0048E32DC07E81" box="[1411,1429,1949,1974]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.
<emphasis id="B2D3EA8F1C6E7711FF1448BC28837EEC" box="[151,214,1986,2011]" italics="true" pageId="14" pageNumber="15">gigas</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="D4D866151C6F7710FF1448C02D1F7EE2" box="[151,1354,1982,2006]" httpUri="https://zenodo.org/record/197297/files/figure.png" pageId="15" pageNumber="16" targetBox="[200,1394,233,1944]" targetPageId="15">
<paragraph id="8018369D1C6F7710FF1448C02D1F7EE2" blockId="15.[151,1354,1982,2006]" box="[151,1354,1982,2006]" pageId="15" pageNumber="16">
<emphasis id="B2D3EA8F1C6F7710FF1448C029177EE1" bold="true" box="[151,322,1982,2006]" pageId="15" pageNumber="16">FIGURE 5a, b.</emphasis>
<taxonomicName id="47A74D1E1C6F7710FECA48C02A1C7EE2" box="[329,585,1982,2005]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C6F7710FECA48C02A1C7EE2" box="[329,585,1982,2005]" italics="true" pageId="15" pageNumber="16">Archispirostreptus gigas</emphasis>
</taxonomicName>
(MRAC 20764).
<emphasis id="B2D3EA8F1C6F7710FC8448C02B4E7EE1" bold="true" box="[775,795,1982,2006]" pageId="15" pageNumber="16">a,</emphasis>
oral view of gonopods;
<emphasis id="B2D3EA8F1C6F7710FB9D48C02C617EE1" bold="true" box="[1054,1076,1982,2006]" pageId="15" pageNumber="16">b,</emphasis>
aboral view of gonopods.
</paragraph>
</caption>
<caption id="D4D866151C70770FFF1448DF29DC7EEA" httpUri="https://zenodo.org/record/197298/files/figure.png" pageId="16" pageNumber="17" targetBox="[286,1299,211,1926]" targetPageId="16">
<paragraph id="8018369D1C70770FFF1448DF29DC7EEA" blockId="16.[151,1436,1953,2013]" pageId="16" pageNumber="17">
<emphasis id="B2D3EA8F1C70770FFF1448DF29117E8E" bold="true" box="[151,324,1953,1977]" pageId="16" pageNumber="17">FIGURE 5c, d.</emphasis>
<taxonomicName id="47A74D1E1C70770FFECF48DC2A047E8E" box="[332,593,1954,1977]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C70770FFECF48DC2A047E8E" box="[332,593,1954,1977]" italics="true" pageId="16" pageNumber="17">Archispirostreptus gigas</emphasis>
</taxonomicName>
(MRAC 20764).
<emphasis id="B2D3EA8F1C70770FFC9748DF2B727E8E" bold="true" box="[788,807,1953,1977]" pageId="16" pageNumber="17">c,</emphasis>
oral view of prefemoral lobe of first pair of male legs;
<emphasis id="B2D3EA8F1C70770FFA0548DF2DC97E8E" bold="true" box="[1414,1436,1953,1977]" pageId="16" pageNumber="17">d,</emphasis>
lateral view of collum.
</paragraph>
</caption>
<paragraph id="8018369D1C71770EFF464FE62A8379CB" blockId="17.[151,1436,152,327]" pageId="17" pageNumber="18">
Considering where
<taxonomicName id="47A74D1E1C71770EFE344FE62A497986" box="[439,540,152,177]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C71770EFE344FE62A497986" box="[439,540,152,177]" italics="true" pageId="17" pageNumber="18">A. gigas</emphasis>
</taxonomicName>
has been collected and the distance between the localities, the currently known distribution may be a collecting artifact. The known distribution suggests that it is more widespread along the east coast of Africa than further inland.
</paragraph>
<paragraph id="8018369D1C71770EFF464E762A597870" blockId="17.[151,1436,152,327]" pageId="17" pageNumber="18">
The
<typeStatus id="5F1C883F1C71770EFF7F4E7629657815" box="[252,304,264,290]" pageId="17" pageNumber="18">type</typeStatus>
is missing from the ZMH collection where it was said to be housed (
<bibRefCitation id="E4364B6C1C71770EFBFD4E762D767815" author="Krabbe" box="[1150,1315,264,290]" pageId="17" pageNumber="18" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
); this was confirmed by museum curators.
</paragraph>
</subSubSection>
</treatment>
</document>