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<document id="4F9595673E7F76519A0B8EAF2322580B" ID-DOI="http://dx.doi.org/10.3897/zookeys.473.8659" ID-GBIF-Dataset="f0fbc7da-005d-4414-b66d-ca97ccf32a02" ID-PMC="PMC4304041" ID-Pensoft-Pub="1313-2970-473-1" ID-PubMed="25632253" ID-ZooBank="02943D336D534CB6A6BD47526EC80C67" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-473-1" ModsDocOrigin="ZooKeys 473" ModsDocTitle="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)" checkinTime="1451244823217" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Ablett, Jonathan, Lương, Hao Văn, Fred Naggs, &amp; Asami, Takahiro" docDate="2015" docId="E4A4895E498AF7B643C19D5206003CE5" docLanguage="en" docName="ZooKeys 473: 1-118" docOrigin="ZooKeys 473" docSource="http://dx.doi.org/10.3897/zookeys.473.8659" docTitle="Gudeodiscus (Gudeodiscus) messageri subsp. messageri Gude 1909" docType="treatment" docVersion="5" lastPageNumber="25" masterDocId="145AFF9BD02C0945FFA62A78FF974356" masterDocTitle="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)" masterLastPageNumber="118" masterPageNumber="1" pageNumber="24" updateTime="1701314957421" updateUser="plazi">
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<mods:title id="10AAA1E94E29D27A1C7F4A3D615483D7">Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)</mods:title>
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<mods:name id="2C781ED4BE8370D3441E4A6059138756" type="personal">
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<mods:roleTerm id="DC94293CFF18D6BCC3AAC198F48A79E3">Author</mods:roleTerm>
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<mods:namePart id="5A9E8B7E1CF6B5EA327636C4DDC93674">Pall-Gergely, Barna</mods:namePart>
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<mods:namePart id="EFBF2C087BA2793168DFD6389692A755">Hunyadi, Andras</mods:namePart>
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<mods:namePart id="431F6CDB4EA882BC5A5F953831640509">Ablett, Jonathan</mods:namePart>
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<mods:namePart id="57890C635E362E917878F7B9A65FEE84">Lương, Hao Văn</mods:namePart>
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<mods:namePart id="6FFD2AB228ECD6E4111F24303CF32B57">Fred Naggs,</mods:namePart>
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</mods:role>
<mods:namePart id="E119E30A3FA231B4854A4BBCA1BE08E7">Asami, Takahiro</mods:namePart>
</mods:name>
<mods:typeOfResource id="74815A22A6FED63B1E12D8414B480559">text</mods:typeOfResource>
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<mods:title id="83937264AA26836430826F519572DCB0">ZooKeys</mods:title>
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<mods:date id="AAB57A93D43196AF2B9EE98EC0F51009">2015</mods:date>
<mods:detail id="2085A187E443B41271636116382E2BD7" type="volume">
<mods:number id="47A56AFDD5002655817901DFB4B42B74">473</mods:number>
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<mods:classification id="403AA504758BB276081CED8A018DC246">journal article</mods:classification>
<mods:identifier id="28836BEBB7893FB1CB6D56CB5D1936C5" type="DOI">http://dx.doi.org/10.3897/zookeys.473.8659</mods:identifier>
<mods:identifier id="C6C41B69670CB811199651A59288A7D8" type="Pensoft-Pub">1313-2970-473-1</mods:identifier>
<mods:identifier id="A19A3DED94C10F07766C19384CDCDDF7" type="ZooBank">02943D336D534CB6A6BD47526EC80C67</mods:identifier>
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<treatment id="E4A4895E498AF7B643C19D5206003CE5" ID-GBIF-Taxon="152057796" LSID="urn:lsid:plazi:treatment:E4A4895E498AF7B643C19D5206003CE5" httpUri="http://treatment.plazi.org/id/E4A4895E498AF7B643C19D5206003CE5" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="18C95F20140E0EAB2303EF13BEA94D99" pageId="23" pageNumber="24" type="multiple">
<paragraph id="E8DC01FD5BBBAAC22EF58CD531D9FEFA" pageId="23" pageNumber="24">Taxon classification Animalia Stylommatophora Plectopylidae</paragraph>
</subSubSection>
<subSubSection id="E46C3FD6872526670F16A1EF06C224C9" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="11B61EBA2D8AAC401D8AC23219E7DD8A" pageId="23" pageNumber="24">
<taxonomicName id="299C8C3C352EE9F6F4F78085474CFA41" ID-CoL="7H7PK" authority="Gude, 1909" authorityName="Gude" authorityYear="1909" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus (Gudeodiscus) messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="messageri" subGenus="Gudeodiscus" subSpecies="messageri">Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909)</taxonomicName>
Figures 5
<normalizedToken id="39202A891993E3812A0BDDBCBE9F3D9F" originalValue="F–G">F-G</normalizedToken>
, 9E, 12
<normalizedToken id="4B3579746C03B4C1294AAF1E49A968FB" originalValue="N–Q">N-Q</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="AF39C5F07D9AE28C7E07062D8C220348" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="45A9ACBF34F3F3C904DA1F94E1BB1546" pageId="23" pageNumber="24">
<taxonomicName id="11BD1D56290D867F141B6FD8BD52B01D" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="messageri">Plectopylis messageri</taxonomicName>
Gude 1909, Proceedings of the Malacological Society of London, 8: 214-215., Plate 9, Figs 4, 4
<normalizedToken id="BB44408313634C35DF4D86E776BFF6E1" originalValue="a–b">a-b</normalizedToken>
[
<normalizedToken id="1E23C5791E35589BAC2AE45B88B80A57" originalValue="“Moung-Hum”">&quot;Moung-Hum&quot;</normalizedToken>
, &quot;Nat-Son, Pac-Kha, and Trinh-Tuong&quot;].
</paragraph>
<paragraph id="82D9F23C967F68012E97F4B57F510CC9" pageId="23" pageNumber="24">
<taxonomicName id="E72DF8977DFDF7345F037C2723F318BF" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="messageri">Gudeodiscus messageri</taxonomicName>
, -
<normalizedToken id="98D1E14C9295AD05B86D4A282BCE1C85" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi 2013, Archiv
<normalizedToken id="8F079C82418334807639292773FE9DEE" originalValue="für">fuer</normalizedToken>
Molluskenkunde 142 (1): 8.
</paragraph>
</subSubSection>
<subSubSection id="22844C775A2D9BC5BB6A02EEC8300F6E" pageId="23" pageNumber="24" type="types examined">
<paragraph id="6849581AB8178FD752C1145CB1FBB815" pageId="23" pageNumber="24">Types examined.</paragraph>
<paragraph id="D11EFDD9457AB6AE49E2B682DAEE418D" pageId="23" pageNumber="24">
Tonkin, Muong-Hum, leg. Messager, NHMUK 1922.8.29.53 (holotype of
<taxonomicName id="73139AAE6D614059BA0C831FBCA02390" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="messageri">Plectopylis messageri</taxonomicName>
, Figure 5F), Tonkin, Nat-Son, leg. Messager, NHMUK 1922.8.29.54 (holotype of
<taxonomicName id="224678DB5B7416E6C85DB14DFFD82341" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Euthyneura messageri var. minor" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="messageri" variety="minor">messageri var. minor</taxonomicName>
, Figure 5G).
</paragraph>
</subSubSection>
<subSubSection id="30A2EBD9C59B2093C0FDDEAAAFC1964A" pageId="23" pageNumber="24" type="museum material examined">
<paragraph id="54AA5DCA9B4D13BB9A8BA1C9971DBB13" pageId="23" pageNumber="24">Museum material examined.</paragraph>
<paragraph id="5D47C07B322A690400814D296CEFBF04" pageId="23" pageNumber="24">
Tonkin, coll. Dosch ex Rolle ex Messager, SMF 172088/4; Tonkin, coll. Dosch ex Rolle, SMF 172076/2; Tonkin, Trinh-Tuong, coll. Dosch ex Rolle, SMF 172086/4; Tonkin, Drinch-Tuom (Trinh-Thuong?), coll. Jaeckel ex Messager, SMF 207675/3; Tonkin, alw.
<normalizedToken id="9DB3FE237C5330AE95E21A7A3D557EF0" originalValue="Müller">Mueller</normalizedToken>
, coll. Kaltenbach, SMF 294867/2; Tonkin, Gia-Phu, coll. Dosch ex Rolle, SMF 172089/4; Tonkin, Muong-Bo, coll. Dosch ex Rolle, SMF 172087/4; Tonkin, Muong-Kong, coll. Pfeiffer, K. L. ex Naschloss (?) ex Rolle, January 1938, SMF 102820/1; Tonkin, coll. Dosch ex Rolle ex Messager, SMF 182088/4; Tonkin, Ba-Nat (?), NHMSB 131/200, 122812-122813/2;
<normalizedToken id="0844478B3A79794734B68975F6979860" originalValue="Pakhé">Pakhe</normalizedToken>
, leg. Messager, MNHN-IM-2012-2129/9; Muong-Hum, leg. Messager, MNHN-IM-2012-2134/15; Nat-Son, Trinh-Thuong, leg. Messager, MNHN-IM-2012-2136/16 (&quot;var.
<taxonomicName id="CB1D3FD7D15B0AF2AC4C4F7D31A1DB8F" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura minor var. minor" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="minor" variety="minor">minor</taxonomicName>
&quot;); Muong-Kong, leg. Messager, MNHN-IM-2012-2137/4; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2142/2+4jb; Muong-Hum, leg. Messager, MNHN-IM-2012-2131/5; Muong-Hum, leg. Messager, MNHN-IM-2012-2143/3; Muong-Hum, leg. Messager, MNHN-IM-2012-2145/74;
<normalizedToken id="288EB273EBEA5C78F5301BA9EE3F1B85" originalValue="Pakhé">Pakhe</normalizedToken>
, leg. Messager, MNHN-IM-2012-2149/1; Pac-Kha (
<normalizedToken id="F3D77477263A5E3B1F86086E770C793A" originalValue="Pakhé">Pakhe</normalizedToken>
), leg. Messager, MNHN-IM-2012-2151/10; Nat-Son, leg. Messager, MNHN-IM-2012-2154/6; Muong-Kong, leg. Messager, MNHN-IM-2012-2159/1; Nat-Son, leg. Messager, MNHN-IM-2012-2162/29; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2163/20; Nat-Son, leg. Messager, MNHN-IM-2012-2165/8+25jb; Bac-Kan, leg. Messager, MNHN-IM-2012-2166/6; Bac-Kan, leg. Messager, MNHN-IM-2012-2172/4; Muong-Hum, leg. Messager, MNHN-IM-2012-2173/3; Muong-Hum, leg. Messager, MNHN-IM-2012-2183/4;
<normalizedToken id="00F285FA0D68827711879A0998F83815" originalValue="Pakhé">Pakhe</normalizedToken>
, leg. Messager, MNHN-IM-2012-2184/1; Long-Ping, leg. Messager, MNHN-IM-2012-2186/8; Muong-Hum, leg. Messager, MNHN-IM-2012-2188/8; Bac-Kan, leg. Messager, MNHN-IM-2012-2194/3; Muong-Hum, leg. Messager, MNHN-IM-2012-2196/4; Nat-Son, leg. Messager, MNHN-IM-2012-2198/2; Nat-Son, leg. Messager, MNHN-IM-2012-2199/2; Tonkin, leg. Messager, MNHN-IM-2012-2202/1; Trinh Thuong, leg. Messager, MNHN-IM-2012-2205/12; Muong-Kong, leg. Messager, MNHN-IM-2012-2479/1; Bac-Kan, leg. Messager, MNHN-IM-2012-2475/10; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2472/16; Cao-Bang, leg. Messager, MNHN-IM-2012-2471/1; Tonkin,
<normalizedToken id="52115422592F961D7359E326B6741716" originalValue="Pakhé">Pakhe</normalizedToken>
, leg. Messager, MNHN-IM-2012-2458/7; Long-Ping, leg. Messager, MNHN-IM-2012-2457/23; label not readable, leg. Messager, MNHN-IM-2012-2449/2; Bac-Kan, leg. Messager, MNHN-IM-2012-2403/1; Trinh-Thuong, coll. Levazzari, 1929, MNHN-IM-2012-2408/9; Tonkin, coll. Staadt, 1969, MNHN-IM-2012-2411/3; Nat-Son, coll. Letellier, 1949, MNHN-IM-2012-2414/2; Pac-Kha, coll. Letellier, 1949, MNHN-IM-2012-2415/2; Gia-Phu, MNHN-IM-2012-2418/3; Trinh-Thuong, coll. Lavezzari, 1929, MNHN-IM-2012-2419/10; Tonkin, leg. Messager, MNHN-IM-2012-2425/3; Gia-Phu, leg. Messager, MNHN-IM-2012-2215/33; Muong-Hum, leg. Messager, MNHN-IM-2012-2216/3; Long-Ping, leg. Messager, MNHN-IM-2012-2217/9; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2219/12; Col de Nuages, leg. Messager, MNHN-IM-2012-2221/1; Trinh Tuong, leg. Messager, MNHN-IM-2012-2223/2; Tonkin, leg. Messager, MNHN-IM-2012-2225/4; Tonkin, leg. Messager, MNHN-IM-2012-2230/1; Long-Phai, leg. Messager, 1901, MNHN-IM-2012-2237/2; Muang-Kong, leg. Messager, MNHN-IM-2012-2242/3; Nat-Son, coll. Staadt, 1969, MNHN-IM-2012-2282/1; Tonkin, leg. M. Balansa, 1889 July, MNHN-IM-2012-2296/10;
<normalizedToken id="AE3F2C6B93BC60B6D01D97F920164B1D" originalValue="Pakhé">Pakhe</normalizedToken>
, leg. Messager, MNHN-IM-2012-2339/1; Bac-Kan, leg. Messager, MNHN-IM-2012-2315/1; Gia-Phu, leg. Messager, MNHN-IM-2012-2364/2; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2379/1; Trinh-Thuong, leg. Messager, MNHN-IM-2012-2394/1; Tonkin, Pac-Kha, NHMUK1916.3.16.15/1; Tonkin, Pac-Kha, coll. Kennard, A.S. ex auct. (Gude), NHMUK 20130620.2/1; Tonkin, Muong-Hum, coll. Biggs, H.E.J. ex Gygngell, 1930, Acc. no. 2258, NHMUK 20130626/2; Tonkin, Gia-Phu, coll. Kennard, A.S. ex auct. (Gude), NHMUK 20130627/2; Tonkin, Muong-Kong, coll. Salisbury ex Beddome, NHMUK 20130628/2; Tonkin, Muong-Kong, 31/3/09, NHMUK 20130629/3; Tonkin, Muong-Hum, 5/1/09, NHMUK 20130630/3; Tonkin, Pac-Kha, 3/11/08, NHMUK 20130631/2 (&quot;var.
<taxonomicName id="F3514ED2261AF96D2F68202A9CD7F018" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura minor var. minor" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="minor" variety="minor">minor</taxonomicName>
&quot;); Tonkin, 5/1/09, NHMUK 20130632/3; Tonkin, Muong-Hum, coll. Preston, NHMUK 20130633/2; Tonkin, Muong-Bo, 3/11/08, NHMUK 20130634/2 (&quot;var.
<taxonomicName id="BE56DFBAD914293AE8ED319E822B574D" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura major var. major" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="major" variety="major">major</taxonomicName>
&quot;); Tonkin,
<normalizedToken id="8B805604CBE5AC8B59E19593DC1F8B53" originalValue="That-Khé">That-Khe</normalizedToken>
, coll. Salisbury ex Beddome, NHMUK 20130635/1; Tonkin, Muong-Hum, coll. Kennard, NHMUK 20130636/1; Tonkin, Muong-Hum, NHMUK 1909.3.17.29-31/3; Tonkin, Muong-Hum, NHMUK 1916.3.16.16-18/3; Tonkin, Pac-Kha, NHMUK 1909.3.17.32-34/3 (&quot;var.
<taxonomicName id="81EBEE0BE5875BBA559319E5BEDF8643" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura minor var. minor" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="minor" variety="minor">minor</taxonomicName>
&quot;); Tonkin, Pac-Kha, NHMUK 1909.3.17.24-25/2; Tonkin, Muong-Bo, NHMUK 1909.3.17.35-36/2 (&quot;var.
<taxonomicName id="6947429A75CC4419A9C64F641AB29846" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura major var. major" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="major" variety="major">major</taxonomicName>
&quot;); Tonkin, Gia-Phee, coll.
<normalizedToken id="26303333DF37BA013744597419B361FC" originalValue="Rušnov">Rusnov</normalizedToken>
ex Rolle ex Messager, NHMW 92576/1; Tonkin, Trisch-Tuong, coll. Edlauer ex Werner, NHMW 75000/E/7983/2; Tonkin, Muong-Hum, coll. Oberwimmer ex Wagner ex Messager, NHMW 92573/2; Tonkin, Bac-Kan, coll. Oberwimmer, NHMW 71640/O/14028/1; Tonkin, Long-Ping, 3000 m, coll. Oberwimmer ex Wagner ex Messager, NHMW 92572/1; Tonkin, Muong Hum, coll. Rosen ex Messager, NHMW 71640/O/9476/2; Tonkin, Trinh-Tua (?), coll. Rolle, NHMW 92574/2; Tonkin, Ban-Tao, coll.
<normalizedToken id="91127BF8782906B84B637A346649E57E" originalValue="Rušnov">Rusnov</normalizedToken>
ex Blume, NHMW 92575/1; Tonkin, Muong Kong, NHMW 71640/O/46293/2; Tonkin, Nat-Son, coll. Rosen ex Messager, NHMW 71640/O/9477/1; Tonkin, Trisch Tuong, coll.
<normalizedToken id="0E85AE69095C6045412838FC90D92A8F" originalValue="Rušnov">Rusnov</normalizedToken>
ex Rolle ex Messager, NHMW 92578/2; Tonkin, Ban-Lao, coll. Rolle, NHMW 92577/1; Tonkin, Bac-Kan, coll. Oberwimmer, NHMW 92567; Tonkin, Bac-Kan, coll. Oberwimmer, NHMW 103353/1; Tonkin, Nat-Son, coll.
<normalizedToken id="F3C7B43F9ADB9FE53CF1BE260A3DECC6" originalValue="Rušnov">Rusnov</normalizedToken>
ex Messager, NHMW 103355/1; Vietnam/132, Lao Cai Province, Cox-Xan, 400 m, leg.
<normalizedToken id="18F25673BFA11864E6AE892FE1C4CA88" originalValue="Topál">Topal</normalizedToken>
&amp;
<normalizedToken id="4AB43E9CC4C5DE7B98D452E00B4570C9" originalValue="Matskási">Matskasi</normalizedToken>
, 27.11.1971., VA/10.
</paragraph>
</subSubSection>
<subSubSection id="12E60F7053C4C6D09EB0A5ED9B8D438E" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="5EA5CDAB57267D0A845729EC5FC7D40E" pageId="23" pageNumber="24">Diagnosis.</paragraph>
<paragraph id="DB5180A0D6427E148C4360E2F62CB2FD" pageId="23" pageNumber="24">
At least one shell was opened of every larger samples. Anterior lamella normal (not dissolved into small denticles); lower parietal plica does not extend beyond the anterior lamella in the anterior direction (Figures 12
<normalizedToken id="E3C1491F4FA21B3B798B06C67E957F64" originalValue="N–Q">N-Q</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="8FF4BAFADA8E24CC9449FE349103015E" pageId="23" pageNumber="24" type="measurements">
<paragraph id="54E8C226CB71035C50425A469BC9C4D8" pageId="23" pageNumber="24">Measurements</paragraph>
<paragraph id="510DF3E5E86F9B557B7907E1B33BC237" pageId="23" pageNumber="24">(in mm). D = 12.75-18.5 (according to the original description).</paragraph>
</subSubSection>
<subSubSection id="6CADFABCE0437DCF824EBEDEE4C52E7D" pageId="23" pageNumber="24" type="differential diagnosis">
<paragraph id="8F1E2E29A9F300A719099932FA74037F" pageId="23" pageNumber="24">Differential diagnosis.</paragraph>
<paragraph id="BC0EBBBA04D6F5C874EBDF46D8510524" pageId="23" pageNumber="24">
<taxonomicName id="658775EC448911921C65E0A83037C42C" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="messageri" subSpecies="messageri">Gudeodiscus messageri messageri</taxonomicName>
inhabits northern Vietnam and in many museum samples it is mixed with
<taxonomicName id="3C879FA1247740498BE5B20FA923A050" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis gouldingi" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="gouldingi">Plectopylis gouldingi</taxonomicName>
or
<taxonomicName id="5D09DD4221C0141B43248BA4F7D4710E" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis fallax" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="fallax">Plectopylis fallax</taxonomicName>
(synonyms of
<taxonomicName id="C04F62CDF360EF74D4627915FA716C24" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
). These two forms have flat shells with a sharp and angled callus, and sometimes with an apertural denticle. Also, the aperture of
<taxonomicName id="CB343865080D20F757B75151EDB5AF3E" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="messageri">Gudeodiscus messageri</taxonomicName>
is rather rounded, whereas it is rather elongated in those populations of
<taxonomicName id="ACDF7716C2EF5CE73A5521EE17974427" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
(Figures 9D:
<taxonomicName id="98DE5F4D71D08BC342BCB1675A699905" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura phlyarius" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="phlyarius">phlyarius</taxonomicName>
, Figure 9E:
<taxonomicName id="27FA6004AB87F2045E15AE5B31770E87" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura messageri" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="messageri">messageri</taxonomicName>
). This allows
<taxonomicName id="E4C6173D766E7107958C649453E1145D" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="messageri">Gudeodiscus messageri</taxonomicName>
and
<taxonomicName id="C66C1073E50ECBFCC5DC30DBE4847317" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
to be distinguished without breaking the shell. The lower parietal plica, which does not extend beyond the anterior lamella in the anterior direction, is characteristic of
<taxonomicName id="3D475130315462D5A277BF1DB00400C9" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="messageri" subSpecies="messageri">Gudeodiscus messageri messageri</taxonomicName>
(see also Remarks), but almost always extends in &quot;
<taxonomicName id="80059CED1EDED49AFF751317F304549B" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis fallax" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="fallax">Plectopylis fallax</taxonomicName>
&quot; and &quot;
<taxonomicName id="5AF90ABE683CA5BF4BE6EF043C3A7BF4" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis gouldingi" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="gouldingi">Plectopylis gouldingi</taxonomicName>
&quot;. &quot;
<taxonomicName id="E13A7D879B706845755F81CD8D522DB8" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis verecunda" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="verecunda">Plectopylis verecunda</taxonomicName>
&quot; (synonym of
<taxonomicName id="0E44CA4F1FD93311BC8AC8510AF447EF" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
) and typical
<taxonomicName id="70BAE6BBCFF44F13D7BEF1CB4288CB50" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus phlyarius" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="phlyarius">Gudeodiscus phlyarius</taxonomicName>
always have a strong apertural fold. Moreover, the lower parietal plica of the latter usually extends beyond the anterior lamella in the anterior direction. For comparison with
<taxonomicName id="D2734D4FD03BE471B6969D5E877EF789" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus messageri" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="messageri" subSpecies="raheemi">Gudeodiscus messageri raheemi</taxonomicName>
ssp. n., see there.
</paragraph>
</subSubSection>
<subSubSection id="9B8AB31F46C1668FDE5B792F26479089" pageId="23" pageNumber="24" type="intrasubpecific diversity">
<paragraph id="FC2691D1548C40BA11BEA669F5035AF2" pageId="23" pageNumber="24">Intrasubpecific diversity.</paragraph>
<paragraph id="8960F7FC59066CB51D26AC482D779D0C" pageId="23" pageNumber="24">Low; the shell size, and the relationship between the lower parietal plica and the anterior lamella show some variability (see remarks). The shell and aperture shape are stable characters.</paragraph>
</subSubSection>
<subSubSection id="00C3398585A941C9E15585A05501E29B" pageId="23" pageNumber="24" type="distribution">
<paragraph id="B95A4F03EBF247CB6E105E58383E6F83" pageId="23" pageNumber="24">Distribution</paragraph>
<paragraph id="CAC7D1D1DBAFE9ECB5F68BCCA7A82989" pageId="23" pageNumber="24">(see Figure 43): Only museum material was available for study, which suggested that this species is located along the Chinese (Yunnan) border.</paragraph>
</subSubSection>
<subSubSection id="6B117003AF3DCAA31B80A382ECB9B1A5" pageId="24" pageNumber="25" type="remarks">
<paragraph id="982275216AC025B08C43ED36FC76B9EA" pageId="24" pageNumber="25">
<pageBreakToken id="61FC44BED34EED271F56AD39AC76E505" pageId="24" pageNumber="25" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="6AA4E3E3E3058E9B6BFB04236F54BC49" pageId="24" pageNumber="25">In one sample (MNHN-IM-2012-2215) a specimen had longer lower plica which extended beyond the anterior lamella in the anterior direction.</paragraph>
</subSubSection>
</treatment>
</document>
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