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<document id="5907F970884F178A7A309316A3281E25" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:2A491D5E-45DB-4D83-8137-AB1EA6B49523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893275264" checkinUser="felipe" docAuthor="Engel, Michael S. &amp; Packer, Laurence" docDate="2013" docId="05237E4C5906614ECC51FC0AF0E923AB" docLanguage="en" docName="JMelittology.2013.10.1-7.pdf" docOrigin="Journal of Melittology 2013 (10)" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Chiasmognathus scythicus Engel &amp; Packer 2013, new species" docType="treatment" docUuid="F5C60D49-59D0-4772-A771-E77B712E8823" docUuidSource="ZooBank" docVersion="1" lastPageNumber="7" masterDocId="F91A063459076148CD1FFFEBF1412212" masterDocTitle="A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1722899446385" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="4AB8B912EC116FAF528AB03901C79614">A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="2BEEE03DEC5CF454AC813FD30B70DEDE">Engel, Michael S.</mods:namePart>
<mods:affiliation id="484BF2E08AD3032C70CEBA38284E430B">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu).</mods:affiliation>
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<mods:namePart id="55C18365356DB413B9701099C1FAB5B3">Packer, Laurence</mods:namePart>
<mods:affiliation id="DAB5DD18507F721A982D5A0B66D5ABA7">Department of Biology, York University, 4700 Keele Street, Toronto, Ontario M 3 J 1 P 3, Canada (xeromelissa @ mail. com).</mods:affiliation>
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<mods:title id="E71F1F01C95FA2C687B167FF82D42767">Journal of Melittology</mods:title>
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<mods:date id="1915D8804140D2A9FE46AD889991B9C4">2013</mods:date>
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<mods:number id="AE5A34440CEE490648C499999928EC8F">2013-05-07</mods:number>
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<subSubSection id="C5909CD159066149CC51FC0AF571260B" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8D35CF5A59066149CC51FC0AF2A321EB" blockId="1.[256,1072,993,1081]" box="[334,994,993,1017]" pageId="1" pageNumber="2">
<heading id="D67D783659066149CC51FC0AF2A321EB" box="[334,994,993,1017]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="4A8AB4D959066149CC51FC0AF20B21EB" authority="Engel &amp; Packer" authorityName="Engel &amp; Packer" authorityYear="2013" box="[334,842,993,1017]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="scythicus" status="sp. nov.">
<emphasis id="BFFE134859066149CC51FC0AF3CC21EB" bold="true" box="[334,653,993,1017]" italics="true" pageId="1" pageNumber="2">Chiasmognathus scythicus</emphasis>
Engel &amp; Packer
</taxonomicName>
,
<taxonomicNameLabel id="A4CDAE3359066149CE4AFC09F2A321EB" box="[853,994,993,1017]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</heading>
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ZooBank
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<subSubSection id="C5909CD159066149CF4AFBCAF39A262B" box="[597,731,1057,1081]" pageId="1" pageNumber="2" type="description">
<paragraph id="8D35CF5A59066149CF4AFBCAF39A262B" blockId="1.[256,1072,993,1081]" box="[597,731,1057,1081]" pageId="1" pageNumber="2">
(
<figureCitation id="15B1D3DF59066149CF41FBCAF393262B" box="[606,722,1057,1081]" captionStart-0="Figures 13" captionStart-1="Figures 48" captionStart-2="Figures 911" captionStartId-0="2.[160,241,1409,1431]" captionStartId-1="3.[161,242,1224,1246]" captionStartId-2="4.[160,241,1410,1432]" captionTargetBox-0="[163,1137,241,1385]" captionTargetBox-1="[194,1081,244,1200]" captionTargetBox-2="[172,1135,246,1379]" captionTargetId-0="figure-110@2.[160,1168,224,1400]" captionTargetId-1="figure-173@3.[161,1169,224,1217]" captionTargetId-2="figure-128@4.[160,1168,224,1400]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figures 13. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." captionText-1="Figures 48. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." captionText-2="Figures 911. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." pageId="1" pageNumber="2">Figs. 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C5909CD159066149CDCFFB8AF5512448" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8D35CF5A59066149CDCFFB8AF5512448" blockId="1.[160,1169,1121,1786]" pageId="1" pageNumber="2">
DIAGNOSIS: Superficially the new species resembles
<taxonomicName id="4A8AB4D959066149CE35FB89F291266B" baseAuthorityName="Popov" baseAuthorityYear="1937" box="[810,976,1121,1145]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149CE35FB89F291266B" box="[810,976,1121,1145]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
but differs most notably by the shape of the hidden metasomal sterna (
<figureCitation id="15B1D3DF59066149CE3BFB6AF2CF268B" box="[804,910,1153,1177]" captionStart="Figures 48" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 48. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." pageId="1" pageNumber="2">Figs. 4, 5</figureCitation>
:
<emphasis id="BFFE134859066149CE85FB6AF2ED268B" box="[922,940,1153,1177]" italics="true" pageId="1" pageNumber="2">cf</emphasis>
.
<bibRefCitation id="E91BB2AB59066149CEA5FB69F50C268B" author="Popov, V. B." box="[954,1101,1153,1177]" pageId="1" pageNumber="2" pagination="10 - 14" refId="ref2932" refString="Popov, V. B. 1937. Zur Kenntnis der Bienengattung Parammobatodes Popov (Hymenoptera, Apoidea). Konowia 16 (1): 10 - 14." type="journal article" year="1937">Popov, 1937</bibRefCitation>
): seventh metasomal sternum broader apically before process, giving disc a rectangular or trapezoidal shape (more strongly narrowed apically and thereby triangular in
<taxonomicName id="4A8AB4D959066149C967FB29F06626EB" baseAuthorityName="Popov" baseAuthorityYear="1937" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149C967FB29F06626EB" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
), apical process glabrous (with short setae in
<taxonomicName id="4A8AB4D959066149CE55FB09F2B026EB" baseAuthorityName="Popov" baseAuthorityYear="1937" box="[842,1009,1249,1273]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149CE55FB09F2B026EB" box="[842,1009,1249,1273]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
); eighth sternum with discs reduced laterally and deeply concave basolaterally (discs broad laterally in
<taxonomicName id="4A8AB4D959066149CDEBFAC9F0DD272B" baseAuthorityName="Popov" baseAuthorityYear="1937" box="[244,412,1313,1337]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149CDEBFAC9F0DD272B" box="[244,412,1313,1337]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
), and lateral apodemes broad and rounded apically (narrowed with acutely rounded proximal angle and pointed apical angle in
<taxonomicName id="4A8AB4D959066149CEBDFAA9F509274B" baseAuthorityName="Popov" baseAuthorityYear="1937" box="[930,1096,1345,1369]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149CEBDFAA9F509274B" box="[930,1096,1345,1369]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
). The dorsal division between the gonocoxae is much deeper (extending half the length of the gonocoxae) than in
<taxonomicName id="4A8AB4D959066149CCA9FA69F31C278B" baseAuthorityName="Popov" baseAuthorityYear="1937" box="[438,605,1409,1433]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="BFFE134859066149CCA9FA69F31C278B" box="[438,605,1409,1433]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName>
(approximately ¼ the length according to
<bibRefCitation id="E91BB2AB59066149C947FA69F04627AB" author="Popov, V. B." pageId="1" pageNumber="2" pagination="10 - 14" refId="ref2932" refString="Popov, V. B. 1937. Zur Kenntnis der Bienengattung Parammobatodes Popov (Hymenoptera, Apoidea). Konowia 16 (1): 10 - 14." type="journal article" year="1937">Popov, 1937</bibRefCitation>
: fig. 2) resulting in more prominent dorsoapical lobes (
<figureCitation id="15B1D3DF59066149CEB5FA4AF2AE27AB" box="[938,1007,1441,1465]" captionStart="Figures 48" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 48. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." pageId="1" pageNumber="2">Fig. 8</figureCitation>
). The sterna more closely resemble the shape observed in
<taxonomicName id="4A8AB4D959066149CFA2FA29F54C27CB" authority="(Warncke, 1983)" baseAuthorityName="Warncke" baseAuthorityYear="1983" box="[701,1037,1473,1497]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="orientanus">
<emphasis id="BFFE134859066149CFA2FA29F21027CB" box="[701,849,1474,1497]" italics="true" pageId="1" pageNumber="2">C. orientanus</emphasis>
(
<bibRefCitation id="E91BB2AB59066149CE40FA2AF54727CB" author="Warncke, K." box="[863,1030,1473,1497]" pageId="1" pageNumber="2" pagination="261 - 347" refId="ref3063" refString="Warncke, K. 1983. Zur Kenntnis der Bienengattung Pasites Jurine, 1807, in der Westpalaarktis (Hymenoptera, Apidae, Nomadinae). Entomofauna 4 (21): 261 - 347." type="journal article" year="1983">Warncke, 1983</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F59D8FCA59066149C949FA2AF19F27EB" name="Bulgaria" pageId="1" pageNumber="2">Bulgaria</collectingCountry>
, Crete,
<collectingCountry id="F59D8FCA59066149CC28FA0AF0D027EB" box="[311,401,1505,1529]" name="Cyprus" pageId="1" pageNumber="2">Cyprus</collectingCountry>
,
<collectingCountry id="F59D8FCA59066149CC82FA0AF0B227EB" box="[413,499,1505,1529]" name="Turkey" pageId="1" pageNumber="2">Turkey</collectingCountry>
, and
<collectingCountry id="F59D8FCA59066149CF2CFA0AF33527EB" box="[563,628,1505,1529]" name="Israel" pageId="1" pageNumber="2">Israel</collectingCountry>
(but likely spread throughout the Levant) (
<emphasis id="BFFE134859066149C968FA0AF5C827EB" box="[1143,1161,1505,1529]" italics="true" pageId="1" pageNumber="2">cf</emphasis>
.
<bibRefCitation id="E91BB2AB59066149CDBFF9EAF006240B" author="Warncke, K." box="[160,327,1537,1561]" pageId="1" pageNumber="2" pagination="261 - 347" refId="ref3063" refString="Warncke, K. 1983. Zur Kenntnis der Bienengattung Pasites Jurine, 1807, in der Westpalaarktis (Hymenoptera, Apidae, Nomadinae). Entomofauna 4 (21): 261 - 347." type="journal article" year="1983">Warncke, 1983</bibRefCitation>
), but differ in the shorter lateral apodemes of the eighth sternum, the disc more developed laterally, and more constricted apical process beyond the disc (broadly tapering from lateral apodemes to apical process in
<taxonomicName id="4A8AB4D959066149CE71F9A9F540244B" baseAuthorityName="Warncke" baseAuthorityYear="1983" box="[878,1025,1602,1625]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="orientanus">
<emphasis id="BFFE134859066149CE71F9A9F540244B" box="[878,1025,1602,1625]" italics="true" pageId="1" pageNumber="2">C. orientanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C5909CD15906614DCDCFF98AF2F0262B" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2" type="description">
<paragraph id="8D35CF5A5906614BCDCFF98AF3162797" blockId="1.[160,1169,1121,1786]" lastBlockId="3.[160,1175,1357,1797]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
DESCRIPTION:
<specimenCount id="9B8C04D359066149CC6FF989F0C5246E" box="[368,388,1634,1660]" count="1" pageId="1" pageNumber="2" type="male"></specimenCount>
: Total body length
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(
<quantity id="4A7262BF59066149CFFDF98AF2C92468" box="[738,904,1633,1658]" metricMagnitude="-3" metricUnit="m" metricValue="2.505" metricValueMax="2.87" metricValueMin="2.14" pageId="1" pageNumber="2" unit="mm" value="2.505" valueMax="2.87" valueMin="2.14">2.142.87 mm</quantity>
); forewing length
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(
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). Head wider than long, width
<quantity id="4A7262BF59066149CE0FF96AF236248B" box="[784,887,1665,1689]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="2" unit="mm" value="0.86">0.86 mm</quantity>
(
<quantity id="4A7262BF59066149CE97F96AF56E2488" box="[904,1071,1665,1690]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="1" pageNumber="2" unit="mm" value="0.885" valueMax="0.95" valueMin="0.82">0.820.95 mm</quantity>
), length
<quantity id="4A7262BF59066149CDBFF94AF04524AB" box="[160,260,1697,1721]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="2" unit="mm" value="0.7">0.70 mm</quantity>
(
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); inner margins of compound eyes straight, convergent below; apex of clypeus at lower tangent of compound eyes; ocelli above upper tangent of compound eyes, ocellar triangle prominent, slightly swollen above curvature of head, vertex weakly concave between lateral ocelli; clypeus weakly convex, nearly flat, apicolateral corners of clypeus with small patches of tightly packed, elongate, curved setae; malar space vestigial posteriorly, 0.7X diameter of median ocellus anteriorly; mandibles simple, crossing in repose but not covering labrum; frontal line distinct from just below lower tangent of antennal toruli to median ocellus, raised below, impressed above. Mesoscutum with median and parapsidal lines weakly impressed, median line extending to just before mesoscutal midlength. Intertegular distance (
<emphasis id="BFFE13485905614AC975F957F5C324C1" box="[1130,1154,1724,1747]" italics="true" pageId="2" pageNumber="3">i.e</emphasis>
., distance between inner margins of tegulae)
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(
<quantity id="4A7262BF5905614ACE02F937F28524E6" box="[797,964,1756,1780]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="6.0" metricValueMin="4.3" pageId="2" pageNumber="3" unit="mm" value="0.515" valueMax="0.6" valueMin="0.43">0.430.60 mm</quantity>
). Forewing marginal cell broadly truncate; both m-cu crossveins entering second submarginal cell. Terminalia as depicted in figures 48.
</paragraph>
<caption id="D9F59FD25905614ACDBFFA6AF31D27A6" pageId="2" pageNumber="3" startId="2.[160,241,1409,1431]" targetBox="[163,1137,241,1385]" targetPageId="2" targetType="figure">
<paragraph id="8D35CF5A5905614ACDBFFA6AF31D27A6" blockId="2.[160,1124,1409,1460]" pageId="2" pageNumber="3">
<emphasis id="BFFE13485905614ACDBFFA6AF0632784" bold="true" box="[160,290,1409,1431]" pageId="2" pageNumber="3">Figures 13.</emphasis>
Photomicrographs of male of
<taxonomicName id="4A8AB4D95905614ACF7CFA6AF2242784" authorityName="Engel &amp; Packer" authorityYear="2013" box="[611,869,1409,1430]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="scythicus" status="sp. nov.">
<emphasis id="BFFE13485905614ACF7CFA6AF2242784" box="[611,869,1409,1430]" italics="true" pageId="2" pageNumber="3">Chiasmognathus scythicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4CDAE335905614ACE6EFA6AF2B22785" box="[881,1011,1409,1431]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="BFFE13485905614AC91DFA6AF5542784" bold="true" box="[1026,1045,1409,1430]" pageId="2" pageNumber="3">1.</emphasis>
Lateral habitus.
<emphasis id="BFFE13485905614ACC1EFA75F05227A1" bold="true" box="[257,275,1438,1459]" pageId="2" pageNumber="3">2.</emphasis>
Dorsal habitus.
<emphasis id="BFFE13485905614ACCD9FA75F09927A1" bold="true" box="[454,472,1438,1459]" pageId="2" pageNumber="3">3.</emphasis>
Facial view.
</paragraph>
</caption>
<caption id="D9F59FD25904614BCDBEFB23F33B270A" pageId="3" pageNumber="4" startId="3.[161,242,1224,1246]" targetBox="[194,1081,244,1200]" targetPageId="3" targetType="figure">
<paragraph id="8D35CF5A5904614BCDBEFB23F33B270A" blockId="3.[161,1143,1224,1304]" pageId="3" pageNumber="4">
<emphasis id="BFFE13485904614BCDBEFB23F06226CC" bold="true" box="[161,291,1224,1246]" pageId="3" pageNumber="4">Figures 48.</emphasis>
Male terminalia of
<taxonomicName id="4A8AB4D95904614BCCEEFB22F3B526CC" authorityName="Engel &amp; Packer" authorityYear="2013" box="[497,756,1225,1246]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scythicus" status="sp. nov.">
<emphasis id="BFFE13485904614BCCEEFB22F3B526CC" box="[497,756,1225,1246]" italics="true" pageId="3" pageNumber="4">Chiasmognathus scythicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4CDAE335904614BCE1FFB22F2C326CD" box="[768,898,1225,1247]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="BFFE13485904614BCE8EFB22F2E226CC" bold="true" box="[913,931,1225,1246]" pageId="3" pageNumber="4">4.</emphasis>
Seventh metasomal sternum.
<emphasis id="BFFE13485904614BCC14FB0DF05C26E9" bold="true" box="[267,285,1254,1275]" pageId="3" pageNumber="4">5.</emphasis>
Eighth metasomal sternum.
<emphasis id="BFFE13485904614BCF4AFB0EF32626E8" bold="true" box="[597,615,1253,1274]" pageId="3" pageNumber="4">6.</emphasis>
Genital capsule, ventral view.
<emphasis id="BFFE13485904614BCEACFB0DF28426E9" bold="true" box="[947,965,1254,1275]" pageId="3" pageNumber="4">7.</emphasis>
Genital capsule, lateral view.
<emphasis id="BFFE13485904614BCC30FAE9F0032705" bold="true" box="[303,322,1282,1303]" pageId="3" pageNumber="4">8.</emphasis>
Genital capsule, dorsal view.
</paragraph>
</caption>
<paragraph id="8D35CF5A5904614CCDCFFA66F53F24B2" blockId="3.[160,1175,1357,1797]" lastBlockId="4.[160,1168,1512,1792]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Integument generally shining (
<figureCitation id="15B1D3DF5904614BCF56FA66F3F727B7" box="[585,694,1421,1445]" captionStart="Figures 13" captionStartId="2.[160,241,1409,1431]" captionTargetBox="[163,1137,241,1385]" captionTargetId="figure-110@2.[160,1168,224,1400]" captionTargetPageId="2" captionText="Figures 13. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." pageId="3" pageNumber="4">Figs. 13</figureCitation>
). Labrum with punctures irregular in size and spacing, surface smooth; clypeus with small punctures separated by 0.52 times a puncture width centrally, punctures closer laterally, with a few scattered larger punctures, apex impunctate; face and vertex with small punctures separated by 0.252 times a puncture width (in a few places where punctures are particularly close they can be nearly contiguous), integument between punctures smooth, punctures on vertex posterior to ocelli somewhat weaker; punctures on gena and postgena separated by 0.52 times a puncture width, integument otherwise smooth and shining. Mesoscutum sparsely punctate on central disc, punctures small and separated by 14.5 times a puncture width, punctures closer on margins, integument otherwise smooth and shining; mesoscutellum with punctures slightly larger and deeper than those of mesoscutum, separated by 0.52 times a puncture width; metanotum with punctures separated by a puncture width; pleura with small punctures separated by 0.52.5 times a puncture width, integument otherwise smooth; propodeum with short and narrow basal area coarsely imbricate and impunctate, otherwise integument with punctures separated by less than or equal to a puncture width except medially around pit on posterior surface smooth. Metasomal terga and sterna faintly and finely imbricate, with fine, scattered punctures (T1 and T2 almost devoid of sculpture in some specimens).
</paragraph>
<caption id="D9F59FD25903614CCDBFFA69F33527A7" pageId="4" pageNumber="5" startId="4.[160,241,1410,1432]" targetBox="[172,1135,246,1379]" targetPageId="4" targetType="figure">
<paragraph id="8D35CF5A5903614CCDBFFA69F33527A7" blockId="4.[160,1156,1410,1461]" pageId="4" pageNumber="5">
<emphasis id="BFFE13485903614CCDBFFA69F06F2785" bold="true" box="[160,302,1410,1432]" pageId="4" pageNumber="5">Figures 911.</emphasis>
Photomicrographs of female of
<taxonomicName id="4A8AB4D95903614CCF9DFA69F2C42785" authorityName="Engel &amp; Packer" authorityYear="2013" box="[642,901,1410,1431]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="scythicus" status="sp. nov.">
<emphasis id="BFFE13485903614CCF9DFA69F2C42785" box="[642,901,1410,1431]" italics="true" pageId="4" pageNumber="5">Chiasmognathus scythicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4CDAE335903614CCE8EFA68F553278A" box="[913,1042,1410,1432]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="BFFE13485903614CC93DFA69F5752785" bold="true" box="[1058,1076,1410,1431]" pageId="4" pageNumber="5">9.</emphasis>
Lateral habitus.
<emphasis id="BFFE13485903614CCC1EFA74F05E27A6" bold="true" box="[257,287,1439,1460]" pageId="4" pageNumber="5">10.</emphasis>
Dorsal habitus.
<emphasis id="BFFE13485903614CCCCDFA74F0B127A6" bold="true" box="[466,496,1439,1460]" pageId="4" pageNumber="5">11.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="8D35CF5A5903614DCDCFF942F2AD234B" blockId="4.[160,1168,1512,1792]" lastBlockId="5.[160,1169,225,1786]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Integument of head and mesosoma black to dark brown and shining except yellowish to reddish brown on mandibles (which have reddish apices), yellowish to reddish brown on labrum, light brown to brown on labiomaxillary complex, light brown to yellow on scape and pedicel, light yellow brown on flagellum, translucent amber on tegula, and light brown on legs except lighter on tarsi and at femorotibial and tibiobasitarsal joints. Wing veins amber except C and Sc+R darker; membranes hyaline. Metasoma dark chestnut brown; apical impressed areas lighter in color.</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFE8AF31120C8" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">Pubescence silvery white. Head with numerous, fine, appressed to subappressed plumose setae, such setae nearly obscuring integument of face around and below level of antennal toruli, and intermingled with a few suberect to erect finer, simple setae; appressed plumose setae present on gena. Setae of mesosoma like those of head although more sparse centrally on mesoscutum and mesoscutellum; setae similar to those of gena on pleura (although longer and more diffuse to sparse centrally on mesepisternum), metanotum, and dorsolateral portions of propodeum, obscuring underlying integument especially on metanotum. Metasoma with sparse, erect to suberect, short simple setae, without prominent apical fasciae; first metasomal tergum with small, weak apicolateral patches of appressed to subappressed plumose setae (rubbed off in some specimens); succeeding terga with similar patches although often more diffuse or smaller than those of first tergum.</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFD09F210214B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">
<specimenCount id="9B8C04D35902614DCDCFFD09F1A520EE" box="[208,228,738,764]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
: As described for male except in usual gender differences (
<bibRefCitation id="E91BB2AB5902614DCEA7FD0AF50520EB" author="Engel, M. S." box="[952,1092,737,761]" pageId="5" pageNumber="6" pagination="113 - 121" refId="ref2665" refString="Engel, M. S. 2006. A new genus of minute ammobatine bees (Hymenoptera: Apidae). Acta Entomologica Slovenica 14 (2): 113 - 121." type="journal article" year="2006">Engel, 2006</bibRefCitation>
,
<bibRefCitation id="E91BB2AB5902614DC94DFD0AF5C820EB" author="Engel, M. S." box="[1106,1161,737,761]" pageId="5" pageNumber="6" pagination="237 - 247" refId="ref2820" refString="Engel, M. S. 2009. The bee genus Chiasmognathus in the Arabian Peninsula (Hymenoptera: Apidae). Fauna of Arabia 24: 237 - 247." type="journal article" year="2009">2009</bibRefCitation>
) and as follows: Total body length
<quantity id="4A7262BF5902614DCF32FCEAF38E210B" box="[557,719,769,793]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.77" metricValueMin="2.13" pageId="5" pageNumber="6" unit="mm" value="2.45" valueMax="2.77" valueMin="2.13">2.132.77 mm</quantity>
; forewing length 1.602.00 mm. Head wider than long, width
<quantity id="4A7262BF5902614DCCDEFCCAF327212B" box="[449,614,801,825]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" metricValueMax="9.9" metricValueMin="7.9" pageId="5" pageNumber="6" unit="mm" value="0.89" valueMax="0.99" valueMin="0.79">0.790.99 mm</quantity>
, length
<quantity id="4A7262BF5902614DCFD6FCCAF22E212B" box="[713,879,801,825]" metricMagnitude="-4" metricUnit="m" metricValue="6.8500000000000005" metricValueMax="8.100000000000001" metricValueMin="5.6000000000000005" pageId="5" pageNumber="6" unit="mm" value="0.685" valueMax="0.81" valueMin="0.56">0.560.81 mm</quantity>
. Ocellar triangle not as prominent as in male. Intertegular distance
<quantity id="4A7262BF5902614DCFB5FCAAF20C214B" box="[682,845,833,857]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="5" pageNumber="6" unit="mm" value="0.525" valueMax="0.6" valueMin="0.45">0.450.60 mm</quantity>
.
</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFC8AF350218B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">Sculpturing as in male except punctures of mesoscutellum closer, separated by 0.51.5 times a puncture width.</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFC4AF305260B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">
Coloration as in male except sometimes lighter brown throughout on those areas where male is black, especially face below antennae (
<figureCitation id="15B1D3DF5902614DCE5FFC2AF2F921CB" box="[832,952,961,985]" captionStart="Figures 911" captionStartId="4.[160,241,1410,1432]" captionTargetBox="[172,1135,246,1379]" captionTargetId="figure-128@4.[160,1168,224,1400]" captionTargetPageId="4" captionText="Figures 911. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." pageId="5" pageNumber="6">Figs. 911</figureCitation>
), and nearly black around ocellar triangle regardless of background color of integument (lighter colors likely due to fading of integument).
</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFBCAF2F0262B" blockId="5.[160,1169,225,1786]" box="[208,945,1057,1081]" pageId="5" pageNumber="6">Pubescence more dense on clypeus and lower paraocular area.</paragraph>
</subSubSection>
<subSubSection id="C5909CD15902614DCDCFFBAAF2B5240B" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8D35CF5A5902614DCDCFFBAAF05026CB" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">
<materialsCitation id="3DE2C5075902614DCDCFFBAAF05026CB" collectingDate="1988-07-30" collectionCode="KASC" country="Kazakhstan" county="Tauqum Sands" elevation="550" latitude="44.0343" location="Kazenas" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="holotype">
<typeStatus id="523171F85902614DCDCFFBAAF005264B" box="[208,324,1089,1113]" pageId="5" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9B8C04D35902614DCC4EFBA9F024264E" box="[337,357,1090,1116]" count="1" pageId="5" pageNumber="6" type="male"></specimenCount>
,
<collectingCountry id="F59D8FCA5902614DCC6CFBAAF340264B" box="[371,513,1089,1113]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
:
<collectingRegion id="4F4E01B85902614DCF15FBAAF324264B" box="[522,613,1089,1113]" country="Kazakhstan" name="Almaty Oblysy" pageId="5" pageNumber="6">Almaty</collectingRegion>
,
<collectingCounty id="6454B7D65902614DCF6FFBA9F263264B" box="[624,802,1089,1113]" pageId="5" pageNumber="6">Taūqum Sands</collectingCounty>
,
<quantity id="4A7262BF5902614DCE32FBAAF235264B" box="[813,884,1089,1113]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="km" value="20.0">20 km</quantity>
[North] of
<collectingMunicipality id="6D5155205902614DCEE7FBAAF51C264B" box="[1016,1117,1089,1113]" pageId="5" pageNumber="6">Aydarly</collectingMunicipality>
,
<quantity id="4A7262BF5902614DC977FBAAF1FB266B" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="m" value="550.0">
<elevation id="06A728695902614DC977FBAAF1FB266B" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="m" value="550.0">550 m</elevation>
</quantity>
,
<geoCoordinate id="E8BEA99D5902614DCDDAFB8AF07F266B" box="[197,318,1121,1145]" degrees="44.0343" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="44.0343">44.0343°N</geoCoordinate>
,
<geoCoordinate id="E8BEA99D5902614DCC56FB8AF0FF266B" box="[329,446,1121,1145]" degrees="79.5184" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="79.5184">79.5184°E</geoCoordinate>
,
<date id="F934E99A5902614DCCD7FB8AF302266B" box="[456,579,1121,1145]" pageId="5" pageNumber="6" value="1988-07-30">
<collectingDate id="E97010725902614DCCD7FB8AF302266B" box="[456,579,1121,1145]" pageId="5" pageNumber="6" value="1988-07-30">30.vii.1988</collectingDate>
</date>
[
<date id="F934E99A5902614DCF4FFB8AF3A2266B" box="[592,739,1121,1145]" pageId="5" pageNumber="6" value="1988-07-30">
<collectingDate id="E97010725902614DCF4FFB8AF3A2266B" box="[592,739,1121,1145]" pageId="5" pageNumber="6" value="1988-07-30">30 July 1988</collectingDate>
</date>
],
<location id="885599815902614DCFEAFB89F219266B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCFEAFB89F219266B" box="[757,856,1122,1145]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Kazenas" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Kazenas</location>
(
<collectionCode id="EB9B579F5902614DCE79FB8AF2F4266B" box="[870,949,1121,1145]" pageId="5" pageNumber="6">KASC</collectionCode>
).
<location id="885599815902614DCED3FB8AF2B8266B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCED3FB8AF2B8266B" box="[972,1017,1121,1145]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="The" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">The</location>
type locality is in southeastern
<collectingCountry id="F59D8FCA5902614DCC67FB6AF342268B" box="[376,515,1153,1177]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
east of the
<location id="885599815902614DCF94FB6AF2C6268B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCF94FB6AF2C6268B" box="[651,903,1153,1177]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Kapshagay Reservoir" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Kapshagay Reservoir</location>
and around the
<location id="885599815902614DC953FB6AF0CA26AB" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DC953FB6AF0CA26AB" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Altyn Emel National Park" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Altyn Emel National Park</location>
, and generally part of the xeric to mountainous ecoregion of
<collectingCountry id="F59D8FCA5902614DC979FB49F04C26CB" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFB0AF2B5240B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">
<materialsCitation id="3DE2C5075902614DCDCFFB0AF03E270B" collectingDate="1988-07-30" collectionCode="KASC, KUEC, PCYU" country="Kazakhstan" county="Tauqum Sands" elevation="550" latitude="44.0343" location="Kazenas" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" pageId="5" pageNumber="6" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<typeStatus id="523171F85902614DCDCFFB0AF00B26EB" box="[208,330,1249,1273]" pageId="5" pageNumber="6" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9B8C04D35902614DCC48FB0AF0CC26EE" box="[343,397,1249,1276]" count="1" pageId="5" pageNumber="6" type="male">1♂♂</specimenCount>
,
<specimenCount id="9B8C04D35902614DCC85FB0AF09126EE" box="[410,464,1249,1276]" count="4" pageId="5" pageNumber="6" type="female">4♀♀</specimenCount>
, same data as holotype (
<collectionCode id="EB9B579F5902614DCFECFB0AF27F26EB" box="[755,830,1249,1273]" pageId="5" pageNumber="6">KASC</collectionCode>
except
<specimenCount id="9B8C04D35902614DCE88FB09F55926EB" box="[919,1048,1249,1273]" count="1" pageId="5" pageNumber="6" type="female">one female</specimenCount>
at each of
<collectionCode id="EB9B579F5902614DCDBFFAEAF1AA270B" box="[160,235,1281,1305]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="5" pageNumber="6">KUEC</collectionCode>
and
<collectionCode id="EB9B579F5902614DCC35FAEAF036270B" box="[298,375,1281,1305]" country="Canada" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="5" pageNumber="6" type="University or college">PCYU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE2C5075902614DCC94FAEAF272272B" collectingDate="1988-07-24" collectingDateMax="1998-07-24" collectingDateMin="1988-07-24" collectionCode="KASC, KUEC, PCYU" country="Kazakhstan" county="Tauqum Sands" elevation="550" latitude="44.0343" location="Kazenas" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" pageId="5" pageNumber="6" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="9B8C04D35902614DCC94FAEAF080270E" box="[395,449,1281,1308]" count="2" pageId="5" pageNumber="6" type="male">2♂♂</specimenCount>
,
<specimenCount id="9B8C04D35902614DCCCFFAEAF0B0270E" box="[464,497,1281,1308]" count="1" pageId="5" pageNumber="6" type="female">1♀</specimenCount>
, same data as holotype except
<date id="F934E99A5902614DCE76FAEAF2A4270B" box="[873,997,1281,1305]" pageId="5" pageNumber="6" value="1988-07-24">
<collectingDate id="E97010725902614DCE76FAEAF2A4270B" box="[873,997,1281,1305]" pageId="5" pageNumber="6" value="1988-07-24">24.vii.1988</collectingDate>
</date>
[
<date id="F934E99A5902614DCEEBFAEAF5CB270B" box="[1012,1162,1281,1305]" pageId="5" pageNumber="6" value="1998-07-24">
<collectingDate id="E97010725902614DCEEBFAEAF5CB270B" box="[1012,1162,1281,1305]" pageId="5" pageNumber="6" value="1998-07-24">24 July 1998</collectingDate>
</date>
] (female at
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<specimenCount id="9B8C04D35902614DCC73FAC9F096272B" box="[364,471,1313,1337]" count="1" pageId="5" pageNumber="6" type="male">one male</specimenCount>
each at of
<collectionCode id="EB9B579F5902614DCF48FACAF3E3272B" box="[599,674,1313,1337]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="5" pageNumber="6">KUEC</collectionCode>
and
<collectionCode id="EB9B579F5902614DCFC2FACAF26B272B" box="[733,810,1313,1337]" country="Canada" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="5" pageNumber="6" type="University or college">PCYU</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="9B8C04D35902614DCE21FACAF21E272E" box="[830,863,1313,1340]" count="1" pageId="5" pageNumber="6" type="male">1♂</specimenCount>
,
<collectingCountry id="F59D8FCA5902614DCE72FACAF2BA272B" box="[877,1019,1313,1337]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
:
<collectingRegion id="4F4E01B85902614DC91BFACAF51E272B" box="[1028,1119,1313,1337]" country="Kazakhstan" name="Almaty Oblysy" pageId="5" pageNumber="6">Almaty</collectingRegion>
,
<collectingMunicipality id="6D5155205902614DC974FACAF1A3274B" pageId="5" pageNumber="6">Ilie River</collectingMunicipality>
,
<quantity id="4A7262BF5902614DCDF1FAAAF077274B" box="[238,310,1345,1369]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="km" value="30.0">30 km</quantity>
S [South-Southeast] of
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,
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,
<geoCoordinate id="E8BEA99D5902614DCE30FAAAF2E2274B" box="[815,931,1345,1369]" degrees="79.8064" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="79.8064">79.8064°E</geoCoordinate>
,
<quantity id="4A7262BF5902614DCEB1FAAAF2B9274B" box="[942,1016,1345,1369]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="600.0">
<elevation id="06A728695902614DCEB1FAAAF2B9274B" box="[942,1016,1345,1369]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="F934E99A5902614DC91BFAAAF533274B" box="[1028,1138,1345,1369]" pageId="5" pageNumber="6" value="1993-07-06">
<collectingDate id="E97010725902614DC91BFAAAF533274B" box="[1028,1138,1345,1369]" pageId="5" pageNumber="6" value="1993-07-06">6.vii.1993</collectingDate>
</date>
[
<date id="F934E99A5902614DC99EFAAAF050276B" pageId="5" pageNumber="6" value="1993-07-06">
<collectingDate id="E97010725902614DC99EFAAAF050276B" pageId="5" pageNumber="6" value="1993-07-06">6 July 1993</collectingDate>
</date>
],
<collectorName id="207FAA8C5902614DCC3BFA89F0C6276B" box="[292,391,1378,1401]" pageId="5" pageNumber="6">Kazenas</collectorName>
(
<collectionCode id="EB9B579F5902614DCC89FA8AF0A5276B" box="[406,484,1377,1401]" pageId="5" pageNumber="6">KASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE2C5075902614DCCE7FA8AF556278B" collectingDate="1997-06-13" collectionCode="SEMC" collectorName="Kazenas" country="Kazakhstan" latitude="43.5474" location="Borandrusy" longLatPrecision="6" longitude="78.0766" municipality="Chilik" pageId="5" pageNumber="6" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="9B8C04D35902614DCCE7FA8AF358276E" box="[504,537,1377,1404]" count="1" pageId="5" pageNumber="6" type="male">1♂</specimenCount>
,
<specimenCount id="9B8C04D35902614DCF38FA8AF308276E" box="[551,585,1377,1404]" count="1" pageId="5" pageNumber="6" type="female">1♀</specimenCount>
,
<collectingCountry id="F59D8FCA5902614DCF48FA8AF3A4276B" box="[599,741,1377,1401]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
:
<collectingRegion id="4F4E01B85902614DCFF0FA8AF20B276B" box="[751,842,1377,1401]" country="Kazakhstan" name="Almaty Oblysy" pageId="5" pageNumber="6">Almaty</collectingRegion>
,
<quantity id="4A7262BF5902614DCE48FA8AF2DE276B" box="[855,927,1377,1401]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="km" value="30.0">30 km</quantity>
[East] of
<collectingMunicipality id="6D5155205902614DC90EFA8AF51D276B" box="[1041,1116,1377,1401]" pageId="5" pageNumber="6">Chilik</collectingMunicipality>
,
<location id="885599815902614DC977FA89F04E278B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DC977FA89F04E278B" country="Kazakhstan" latitude="43.5474" longLatPrecision="6" longitude="78.0766" municipality="Chilik" name="Borandrusy" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Borandrusy</location>
,
<geoCoordinate id="E8BEA99D5902614DCC05FA6AF0D5278B" box="[282,404,1409,1433]" degrees="43.5474" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="43.5474">43.5474°N</geoCoordinate>
,
<geoCoordinate id="E8BEA99D5902614DCC80FA6AF355278B" box="[415,532,1409,1433]" degrees="78.0766" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="78.0766">78.0766°E</geoCoordinate>
,
<date id="F934E99A5902614DCF00FA6AF3D2278B" box="[543,659,1409,1433]" pageId="5" pageNumber="6" value="1997-06-13">
<collectingDate id="E97010725902614DCF00FA6AF3D2278B" box="[543,659,1409,1433]" pageId="5" pageNumber="6" value="1997-06-13">13.vi.1997</collectingDate>
</date>
[
<date id="F934E99A5902614DCFBEFA6AF27A278B" box="[673,827,1409,1433]" pageId="5" pageNumber="6" value="1997-06-13">
<collectingDate id="E97010725902614DCFBEFA6AF27A278B" box="[673,827,1409,1433]" pageId="5" pageNumber="6" value="1997-06-13">13 June 1997</collectingDate>
</date>
],
<collectorName id="207FAA8C5902614DCE52FA69F2F1278B" box="[845,944,1410,1433]" pageId="5" pageNumber="6">Kazenas</collectorName>
(
<collectionCode id="EB9B579F5902614DCEA0FA6AF54E278B" box="[959,1039,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="6" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
. The type series is in less than desirable condition, with many of the specimens poorly mounted, glued to cards so that various portions are embedded and obscured (and the glue has deteriorated into a crystalline matrix rendering it virtually impossible to separate specimens).
<materialsCitation id="3DE2C5075902614DCC27F9EAF3AE240B" box="[312,751,1537,1561]" collectionCode="KASC" country="Kazakhstan" location="July" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="9B8C04D35902614DCC27F9EAF0E8240B" box="[312,425,1537,1561]" count="1" pageId="5" pageNumber="6" type="male">One male</specimenCount>
collected 24
<location id="885599815902614DCF5FF9EAF331240B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCF5FF9EAF331240B" box="[576,624,1537,1561]" country="Kazakhstan" name="July" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">July</location>
(in
<collectionCode id="EB9B579F5902614DCF82F9EAF3A9240B" box="[669,744,1537,1561]" pageId="5" pageNumber="6">KASC</collectionCode>
)
</materialsCitation>
has been decapitated.
</paragraph>
</subSubSection>
<subSubSection id="C5909CD15902614ECDCFF9CAF319230B" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8D35CF5A5902614ECDCFF9CAF319230B" blockId="5.[160,1169,225,1786]" lastBlockId="6.[160,1168,225,441]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">VARIATION: There is variation in metasomal coloration among the available specimens. For example, in some males the first and other terga are lighter in their apical two-thirds, resulting in a somewhat banded appearance. Females seem to vary in the degree to which the first tergum is red and the degree to which it is differentiated in color from the succeeding terga. The degree to which pubescence is developed on the metasoma is variable (perhaps due to the degree to which individuals are worn), with patches of appressed to subappressed plumose setae apicolaterally on the terga somewhat larger and more dense in some individuals. As noted above, there is variation in the degree of sculpture on T1 and T2.</paragraph>
</subSubSection>
<subSubSection id="C5909CD15901614ECDCFFECAF0E923AB" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8D35CF5A5901614ECDCFFECAF0E923AB" blockId="6.[160,1168,225,441]" pageId="6" pageNumber="7">
ETYMOLOGY: The specific epithet is a reference to the ancient region of Scythia (8th Century BC to 2nd Century AD), which encompassed much of modern-day
<collectingCountry id="F59D8FCA5901614EC932FEAAF190236B" name="Kazakhstan" pageId="6" pageNumber="7">Kazakhstan</collectingCountry>
(particularly the Kazakh steppe and Saka tigrakhaude) as well as bordering regions in Central Asia (Sakastan, Ponto-Caspian steppe) and easternmost Europe (Sarmatia, Scythia Minor).
</paragraph>
</subSubSection>
</treatment>
</document>