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<document id="6FA268096BF501CDE19568F0A5C9B96F" ID-CLB-Dataset="21443" ID-DOI="10.11646/zootaxa.4816.3.4" ID-GBIF-Dataset="8de569a5-ae44-4994-8313-57163e9a4514" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954447" ID-ZooBank="FA112B5F-E75E-4E6F-8E4C-E0D87674FCCA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339868132" checkinUser="plazi" docAuthor="Ahyong, Shane T. &amp; Ng, Peter K. L." docDate="2020" docId="03B087FADC16FFC7C798FECFFB122D86" docLanguage="en" docName="zootaxa.4816.3.4.pdf" docOrigin="Zootaxa 4816 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Buergeres Ng &amp; Manning 2003" docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FF89FF82DC1EFFCEC70FFFABFFCA2C29" masterDocTitle="New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="349" masterPageNumber="333" pageNumber="341" updateTime="1698843768168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="172595A288AB2A48742798C94F2024A7">New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="035E0EE14E9802D99CF1B8896A7AF6C1">Ahyong, Shane T.</mods:namePart>
<mods:affiliation id="30DE2BDF45F216D097D876B77619AF24">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia, and School of Biological, Earth &amp; Environmental Sciences, University of New South Wales, NSW 2052, Australia.</mods:affiliation>
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<mods:namePart id="C9E76A48ABB27F38503E2348D181F2ED">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="968B956C0A7BAF0D60ECFA69F8397B68">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:title id="534C63ABF1D17E36A19B2401CB4777F4">Zootaxa</mods:title>
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<mods:date id="AC7E809BCA829E088AE5617D2355FFB4">2020</mods:date>
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<treatment id="03B087FADC16FFC7C798FECFFB122D86" ID-DOI="http://doi.org/10.5281/zenodo.4330016" ID-GBIF-Taxon="165857112" ID-Zenodo-Dep="4330016" LSID="urn:lsid:plazi:treatment:03B087FADC16FFC7C798FECFFB122D86" httpUri="http://treatment.plazi.org/id/03B087FADC16FFC7C798FECFFB122D86" lastPageId="9" lastPageNumber="341" pageId="8" pageNumber="341">
<subSubSection id="C3036567DC16FFC6C798FECFFEF42D57" box="[151,318,356,382]" pageId="8" pageNumber="341" type="nomenclature">
<paragraph id="8BA636ECDC16FFC6C798FECFFEF42D57" blockId="8.[151,318,356,418]" box="[151,318,356,382]" pageId="8" pageNumber="341">
<heading id="D0EE8180DC16FFC6C798FECFFEF42D57" bold="true" box="[151,318,356,382]" fontSize="11" level="1" pageId="8" pageNumber="341" reason="1">
<emphasis id="B96DEAFEDC16FFC6C798FECFFEF42D57" bold="true" box="[151,318,356,382]" pageId="8" pageNumber="341">
<taxonomicName id="4C194D6FDC16FFC6C798FECFFED92D57" ID-CoL="7NVB5" authorityName="Ng &amp; Manning" authorityYear="2003" box="[151,275,356,382]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC16FFC6C798FECFFED92D57" bold="true" box="[151,275,356,382]" italics="true" pageId="8" pageNumber="341">Buergeres</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC16FFC6C798FE23FF212D8B" box="[151,235,391,418]" pageId="8" pageNumber="341" type="description">
<paragraph id="8BA636ECDC16FFC6C798FE23FF212D8B" blockId="8.[151,318,356,418]" box="[151,235,391,418]" pageId="8" pageNumber="341">
(
<figureCitation id="13222A69DC16FFC6C7AFFE23FF282D8B" box="[160,226,391,418]" captionStart="FIGURE 5" captionStartId="9.[151,250,1855,1880]" captionTargetBox="[153,1433,458,1826]" captionTargetId="figure@9.[151,1436,453,1830]" captionTargetPageId="9" captionText="FIGURE 5. Buergeres sp., male, cl 5.3 mm, cw 5.6 mm, Manga Point, Bohol, Visayas, Philippines, ZRC 2018.0784:A, dorsal habitus; B, anterior cephalothorax; C, right maxilliped 3; D, right chela; EH, right P25; I, anterior thoracic sternum; J, pleon; KL, left G1, sternal and pleonal views, respectively; MN, left G2, sternal and pleonal views, respectively. Scales:A = 2.0 mm; B, DH, J = 1.0 mm; C, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954457" httpUri="https://zenodo.org/record/3954457/files/figure.png" pageId="8" pageNumber="341">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC16FFC6C798FE64FE142E1B" pageId="8" pageNumber="341" type="materials_examined">
<paragraph id="8BA636ECDC16FFC6C798FE64FE142E1B" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">
<emphasis id="B96DEAFEDC16FFC6C798FE64FEB02DC3" bold="true" box="[151,378,463,490]" pageId="8" pageNumber="341">Material examined</emphasis>
.
<materialsCitation id="3B713CB1DC16FFC6C686FE7BFE102E1B" ID-GBIF-Occurrence="2816749302" collectingDate="2004-06-17" collectionCode="ZRC" country="Philippines" elevation="5" latitude="9.701667" location="Manga Point" longLatPrecision="129" longitude="123.855" municipality="Visayas" pageId="8" pageNumber="341" specimenCode="ZRC 2018.0784" specimenCount="1" specimenCount-male="1" stateProvince="Bohol">
<specimenCode id="DBBF9E97DC16FFC6C686FE7BFD8B2DC3" box="[393,577,464,490]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="341">ZRC 2018.0784</specimenCode>
,
<specimenCount id="9D1FFD65DC16FFC6C543FE7BFD522DC3" box="[588,664,464,490]" pageId="8" pageNumber="341" type="male">1 male</specimenCount>
(cl
<quantity id="4CE19B09DC16FFC6C5CAFE64FCD42DC3" box="[709,798,463,490]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="341" unit="mm" value="5.3">5.3 mm</quantity>
, cw
<quantity id="4CE19B09DC16FFC6C45DFE64FC672DC3" box="[850,941,463,490]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="8" pageNumber="341" unit="mm" value="5.6">5.6 mm</quantity>
),
<location id="8EC66037DC16FFC6C4CFFE7BFB992DC3" LSID="urn:lsid:plazi:treatment:03B087FADC16FFC7C798FECFFB122D86:8EC66037DC16FFC6C4CFFE7BFB992DC3" box="[960,1107,464,490]" country="Philippines" latitude="9.701667" longLatPrecision="129" longitude="123.855" municipality="Visayas" name="Manga Point" pageId="8" pageNumber="341" stateProvince="Bohol">Manga Point</location>
,
<collectingRegion id="49DDF80EDC16FFC6C350FE7BFB6C2DC3" box="[1119,1190,464,490]" country="Philippines" name="Bohol" pageId="8" pageNumber="341">Bohol</collectingRegion>
,
<collectingMunicipality id="6BC2AC96DC16FFC6C3BDFE7BFAC62DC3" box="[1202,1292,464,490]" pageId="8" pageNumber="341">Visayas</collectingMunicipality>
,
<collectingCountry id="F30E767CDC16FFC6C218FE7BFA522DC3" box="[1303,1432,464,490]" name="Philippines" pageId="8" pageNumber="341">Philippines</collectingCountry>
,
<geoCoordinate id="EE2D502BDC16FFC6C798FE5FFEDA2E27" box="[151,272,500,526]" degrees="09" direction="north" minutes="42.1" orientation="latitude" pageId="8" pageNumber="341" precision="92" value="9.701667">09°42.1N</geoCoordinate>
,
<geoCoordinate id="EE2D502BDC16FFC6C611FE58FE6B2E27" box="[286,417,499,526]" degrees="123" direction="east" minutes="51.3" orientation="longitude" pageId="8" pageNumber="341" precision="92" value="123.855">123°51.3E</geoCoordinate>
,
<quantity id="4CE19B09DC16FFC6C6BFFE5FFE362E27" box="[432,508,500,526]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="8" pageNumber="341" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="0034D1DFDC16FFC6C6BFFE5FFE362E27" box="[432,508,500,526]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="8" pageNumber="341" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">01 m</elevation>
</quantity>
, mangrove and mixed intertidal habitat, from
<taxonomicName id="4C194D6FDC16FFC6C31BFE5FFAC22E24" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[1044,1288,500,526]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="8" pageNumber="341" phylum="Echinodermata" rank="species" species="impatiens">
<emphasis id="B96DEAFEDC16FFC6C31BFE5FFAC22E24" box="[1044,1288,500,526]" italics="true" pageId="8" pageNumber="341">Holothuria impatiens</emphasis>
</taxonomicName>
, PANGLAO 2004 stn M24,
<date id="FFA7102CDC16FFC6C64EFDB3FE102E1B" box="[321,474,536,562]" pageId="8" pageNumber="341" value="2004-06-17">
<collectingDate id="EFE3E9C4DC16FFC6C64EFDB3FE102E1B" box="[321,474,536,562]" pageId="8" pageNumber="341" value="2004-06-17">17 June 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC16FFC6C7C8FD90FAEC2A9B" pageId="8" pageNumber="341" type="description">
<paragraph id="8BA636ECDC16FFC6C7C8FD90FE3F2F23" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">
<emphasis id="B96DEAFEDC16FFC6C7C8FD90FE9B2E7F" bold="true" box="[199,337,571,598]" pageId="8" pageNumber="341">Description</emphasis>
. MALE: Carapace rounded, high, vaulted, slightly wider than long, widest near midlength. Front produced, anterior margin concave in dorsal view. Anterolateral margins poorly defined. Dorsum smooth, glabrous, regions scarcely indicated centrally; dorsal midline convex in profile. Epistome with narrow triangular interantennular septum; median buccal margin with obtuse, triangular median point. Antennular sinus of slightly larger than orbit; antennules folded slightly obliquely. Antennal articles 1 and 2 fused to epistome. Eyes visible in dorsal view, filling orbit, cornea pigmented.
</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8FCBFFD712FCB" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">Maxilliped 3 ischiomerus length twice width; inner proximal margin convex; inner margin with blunt, obtuse angle slightly distal to midlength, margin on either side weakly concave; outer margin strongly convex. Carpus half propodus length. Propodus spatulate, gently tapering in distal half, apex rounded, dorsally and distally setose, length exceeding twice dactylus length. Dactylus sub-clavate, widest distal to midlength, apex rounded, distally setose, inserted at propodal midlength, apex not reaching end of propodus. Exopod inner margin straight, outer margin convex; flagellum with 2 articles, distally setose.</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8FC47FB792893" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">Chelipeds symmetrical from left to right. Dactylus and pollex relatively straight, crossing distally, without gape. Dactylus dorsal margin convex, smooth, glabrous; occlusal margin with blunt triangular tooth proximal to midlength, otherwise straight, sparsely setose. Pollex occlusal margin with 2 blunt triangular teeth proximally, remaining margin weakly crenulate, straight, sparsely setose, minutely denticulate; with fringe of setae on inner ventral margin extending onto lower inner palm surface. Propodus palm outer surface smooth, glabrous; dorsal margin 1.3× height, 1.1× length of dactylus; ventral margin sinuous. Carpus unarmed, inner margin setose.</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8FB6FFC37299F" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">
Pereopods 25 (walking legs 14) symmetrical from left to right, unarmed; relative lengths in decreasing order: P3
<emphasis id="B96DEAFEDC16FFC6C7B3FB43FF052928" box="[188,207,1256,1281]" italics="true" pageId="8" pageNumber="341">=</emphasis>
P4
<emphasis id="B96DEAFEDC16FFC6C7F3FB43FEC52928" box="[252,271,1256,1281]" italics="true" pageId="8" pageNumber="341">&gt;</emphasis>
P2
<emphasis id="B96DEAFEDC16FFC6C634FB43FE842928" box="[315,334,1256,1281]" italics="true" pageId="8" pageNumber="341">&gt;</emphasis>
P5. P3 merus length 0.5× pcl. P24 similar; merus extensor margin broadly convex, extensor and flexor margins sparsely setose; carpus glabrous; dactylus and propodus flexor margins setose; dactyli subequal, stout, falcate, evenly arcuate, distally spiniform, length 0.4× propodus length; basis anterior surface smooth, not granulate. P5 merus tapering distally, distinctly shorter than P24 merus, extensor and flexor margins margins straight; propodus margins parallel, flexor margin setose; dactylus slender, weakly curved, almost straight, flexor margin setose but unarmed, 0.8× propodus length, 1.3× P24 dactylus length.
</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8FA6BFD8529D7" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">Thoracic sternum anterior margin concave medially; sternites 13 indistinguishably fused; sterno-pleonal cavity reaching level of sternite 3/4 suture.</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8F9A3FDF02A6F" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">Pleon slender, margins evenly converging distally; comprising 6 free somites and telson, widest at somites 13; telson as wide as long, apex rounded.</paragraph>
<paragraph id="8BA636ECDC16FFC6C7C8F9FBFAEC2A9B" blockId="8.[151,1437,463,2038]" pageId="8" pageNumber="341">Gonopod 1 elongate, slender, gently arcuate, dorsoventrally compressed, about ¾ length of pleon, tapering to convex apex with short, obliquely directed spiniform tip; margins strongly setose. Gonopod 2 endopod simple, about ¼ length of gonopod 1; exopod short, slender, about half length of endopod. Pleopods 35 absent.</paragraph>
</subSubSection>
<subSubSection id="C3036567DC16FFC6C7C8F917FCA42AFE" box="[199,878,1723,1751]" pageId="8" pageNumber="341" type="biology_ecology">
<paragraph id="8BA636ECDC16FFC6C7C8F917FCA42AFE" blockId="8.[151,1437,463,2038]" box="[199,878,1723,1751]" pageId="8" pageNumber="341">
<emphasis id="B96DEAFEDC16FFC6C7C8F917FF352AFF" bold="true" box="[199,255,1724,1750]" pageId="8" pageNumber="341">Host</emphasis>
.
<taxonomicName id="4C194D6FDC16FFC6C602F917FCA02AFE" authority="(Forskal, 1775)" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[269,874,1723,1751]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="8" pageNumber="341" phylum="Echinodermata" rank="species" species="impatiens">
<emphasis id="B96DEAFEDC16FFC6C602F917FE342AFC" box="[269,510,1724,1750]" italics="true" pageId="8" pageNumber="341">Holothuria impatiens</emphasis>
(Forskål, 1775) (Holothuriidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036567DC16FFC7C7C8F94BFB122D86" lastPageId="9" lastPageNumber="342" pageId="8" pageNumber="341" type="discussion">
<paragraph id="8BA636ECDC16FFC7C7C8F94BFB122D86" blockId="8.[151,1437,463,2038]" lastBlockId="9.[151,1437,153,431]" lastPageId="9" lastPageNumber="342" pageId="8" pageNumber="341">
<emphasis id="B96DEAFEDC16FFC6C7C8F94BFEFE2AD3" bold="true" box="[199,308,1760,1786]" pageId="8" pageNumber="341">Remarks</emphasis>
. Prior to the present study, knowledge of male
<taxonomicName id="4C194D6FDC16FFC6C45DF94BFC092AD0" authorityName="Ng &amp; Manning" authorityYear="2003" box="[850,963,1760,1785]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC16FFC6C45DF94BFC092AD0" box="[850,963,1760,1785]" italics="true" pageId="8" pageNumber="341">Buergeres</emphasis>
</taxonomicName>
has been derived from a single specimen of
<taxonomicName id="4C194D6FDC16FFC6C7B9F8AFFE972B37" authorityName="Chopra" authorityYear="1931" box="[182,349,1796,1822]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="deccanensis">
<emphasis id="B96DEAFEDC16FFC6C7B9F8AFFE972B37" box="[182,349,1796,1822]" italics="true" pageId="8" pageNumber="341">B. deccanensis</emphasis>
</taxonomicName>
reported by Chopra (1931) (although without figures or description of the gonopods). We report here a male of an unidentified species of
<taxonomicName id="4C194D6FDC16FFC6C568F883FD122B68" authorityName="Ng &amp; Manning" authorityYear="2003" box="[615,728,1832,1857]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC16FFC6C568F883FD122B68" box="[615,728,1832,1857]" italics="true" pageId="8" pageNumber="341">Buergeres</emphasis>
</taxonomicName>
from the
<collectingCountry id="F30E767CDC16FFC6C448F883FC0F2B6B" box="[839,965,1832,1858]" name="Philippines" pageId="8" pageNumber="341">Philippines</collectingCountry>
(
<figureCitation id="13222A69DC16FFC6C4DAF883FBD32B6B" box="[981,1049,1831,1858]" captionStart="FIGURE 5" captionStartId="9.[151,250,1855,1880]" captionTargetBox="[153,1433,458,1826]" captionTargetId="figure@9.[151,1436,453,1830]" captionTargetPageId="9" captionText="FIGURE 5. Buergeres sp., male, cl 5.3 mm, cw 5.6 mm, Manga Point, Bohol, Visayas, Philippines, ZRC 2018.0784:A, dorsal habitus; B, anterior cephalothorax; C, right maxilliped 3; D, right chela; EH, right P25; I, anterior thoracic sternum; J, pleon; KL, left G1, sternal and pleonal views, respectively; MN, left G2, sternal and pleonal views, respectively. Scales:A = 2.0 mm; B, DH, J = 1.0 mm; C, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954457" httpUri="https://zenodo.org/record/3954457/files/figure.png" pageId="8" pageNumber="341">Fig. 5</figureCitation>
). It corresponds well to
<taxonomicName id="4C194D6FDC16FFC6C224F883FA562B68" authorityName="Ng &amp; Manning" authorityYear="2003" box="[1323,1436,1832,1857]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC16FFC6C224F883FA562B68" box="[1323,1436,1832,1857]" italics="true" pageId="8" pageNumber="341">Buergeres</emphasis>
</taxonomicName>
in maxilliped 3 morphology and in the subequal P24 dactyli that are all shorter and more strongly curved than the straight, elongate P5 dactylus. Based on what is known of
<taxonomicName id="4C194D6FDC16FFC6C433F8DBFC2E2BA3" authorityName="Chopra" authorityYear="1931" box="[828,996,1904,1930]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="deccanensis">
<emphasis id="B96DEAFEDC16FFC6C433F8DBFC2E2BA3" box="[828,996,1904,1930]" italics="true" pageId="8" pageNumber="341">B. deccanensis</emphasis>
</taxonomicName>
, the P5 dactylus is essentially straight in both sexes, but proportionally shorter in males; the present male reflects such a condition. The Philippine male differs from male
<taxonomicName id="4C194D6FDC16FFC6C66CF813FDC32BFB" authorityName="Chopra" authorityYear="1931" box="[355,521,1976,2002]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="deccanensis">
<emphasis id="B96DEAFEDC16FFC6C66CF813FDC32BFB" box="[355,521,1976,2002]" italics="true" pageId="8" pageNumber="341">B. deccanensis</emphasis>
</taxonomicName>
in carapace shape (rounded versus subquadrate), and given the locality, might be referable to either
<taxonomicName id="4C194D6FDC16FFC6C669F877FDC22BDF" authorityName="Semper" authorityYear="1880" box="[358,520,2012,2038]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="holothuriae">
<emphasis id="B96DEAFEDC16FFC6C669F877FDC22BDF" box="[358,520,2012,2038]" italics="true" pageId="8" pageNumber="341">B. holothuriae</emphasis>
</taxonomicName>
or
<taxonomicName id="4C194D6FDC16FFC6C522F877FD792BDC" authorityName="Burger" authorityYear="1895" box="[557,691,2012,2037]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B96DEAFEDC16FFC6C522F877FD792BDC" box="[557,691,2012,2037]" italics="true" pageId="8" pageNumber="341">B. ortmanni</emphasis>
</taxonomicName>
, both of which occur in the
<collectingCountry id="F30E767CDC16FFC6C4FFF877FBA42BDF" box="[1008,1134,2012,2038]" name="Philippines" pageId="8" pageNumber="341">Philippines</collectingCountry>
(
<bibRefCitation id="EF884B1DDC16FFC6C373F877FAC72BDF" author="Burger, O." box="[1148,1293,2011,2038]" pageId="8" pageNumber="341" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger 1895</bibRefCitation>
; Ng &amp; Man- ning 2003). The specific identity of the male cannot be confirmed at present because it was not associated with a known female, but we believe that it is probably
<taxonomicName id="4C194D6FDC17FFC7C5BEFF15FCFC2CFE" authorityName="Burger" authorityYear="1895" box="[689,822,190,215]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B96DEAFEDC17FFC7C5BEFF15FCFC2CFE" box="[689,822,190,215]" italics="true" pageId="9" pageNumber="342">B. ortmanni</emphasis>
</taxonomicName>
: the outer surface of the cheliped palm is glabrous as in female
<taxonomicName id="4C194D6FDC17FFC7C7E5FF49FEBA2CD2" authorityName="Burger" authorityYear="1895" box="[234,368,226,251]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B96DEAFEDC17FFC7C7E5FF49FEBA2CD2" box="[234,368,226,251]" italics="true" pageId="9" pageNumber="342">B. ortmanni</emphasis>
</taxonomicName>
(
<figureCitation id="13222A69DC17FFC7C670FF4AFE122CD2" box="[383,472,224,251]" captionStart="FIGURE 5" captionStartId="9.[151,250,1855,1880]" captionTargetBox="[153,1433,458,1826]" captionTargetId="figure@9.[151,1436,453,1830]" captionTargetPageId="9" captionText="FIGURE 5. Buergeres sp., male, cl 5.3 mm, cw 5.6 mm, Manga Point, Bohol, Visayas, Philippines, ZRC 2018.0784:A, dorsal habitus; B, anterior cephalothorax; C, right maxilliped 3; D, right chela; EH, right P25; I, anterior thoracic sternum; J, pleon; KL, left G1, sternal and pleonal views, respectively; MN, left G2, sternal and pleonal views, respectively. Scales:A = 2.0 mm; B, DH, J = 1.0 mm; C, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954457" httpUri="https://zenodo.org/record/3954457/files/figure.png" pageId="9" pageNumber="342">Fig. 5D</figureCitation>
;
<bibRefCitation id="EF884B1DDC17FFC7C6EBFF4AFD182CD2" author="Ng, P. K. L. &amp; Manning, R. B." box="[484,722,225,251]" pageId="9" pageNumber="342" pagination="901 - 919" refId="ref11058" refString="Ng, P. K. L. &amp; Manning, R. B. (2003) On two new genera of pea crabs parasitic in holothurians (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the Indo West Pacific, with notes on allied genera. Proceedings of the Biological Society of Washington, 116, 901 - 919." type="journal article" year="2003">Ng &amp; Manning 2003</bibRefCitation>
: fig. 4C), rather than tomentose as in female
<taxonomicName id="4C194D6FDC17FFC7C3C7FF49FAA02CD2" authorityName="Semper" authorityYear="1880" box="[1224,1386,225,251]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="holothuriae">
<emphasis id="B96DEAFEDC17FFC7C3C7FF49FAA02CD2" box="[1224,1386,225,251]" italics="true" pageId="9" pageNumber="342">B. holothuriae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF884B1DDC17FFC7C276FF4AFE962D36" author="Ng, P. K. L. &amp; Manning, R. B." pageId="9" pageNumber="342" pagination="901 - 919" refId="ref11058" refString="Ng, P. K. L. &amp; Manning, R. B. (2003) On two new genera of pea crabs parasitic in holothurians (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the Indo West Pacific, with notes on allied genera. Proceedings of the Biological Society of Washington, 116, 901 - 919." type="journal article" year="2003">Ng &amp; Manning 2003</bibRefCitation>
: fig. 5D). Despite the unconfirmed identification, the present male is figured here to document its morphology, especially that of the gonopods. As in known males of
<taxonomicName id="4C194D6FDC17FFC7C494FE82FBDC2D6A" authorityName="P.K.L.Ng &amp; R.B.Manning" authorityYear="2003" box="[923,1046,297,323]" class="Malacostraca" family="Pinnotheridae" genus="Holotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC17FFC7C494FE82FBDC2D6A" box="[923,1046,297,323]" italics="true" pageId="9" pageNumber="342">Holotheres</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF884B1DDC17FFC7C35BFE82FA8E2D6A" author="Ng, P. K. L. &amp; Manning, R. B." box="[1108,1348,297,323]" pageId="9" pageNumber="342" pagination="901 - 919" refId="ref11058" refString="Ng, P. K. L. &amp; Manning, R. B. (2003) On two new genera of pea crabs parasitic in holothurians (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the Indo West Pacific, with notes on allied genera. Proceedings of the Biological Society of Washington, 116, 901 - 919." type="journal article" year="2003">Ng &amp; Manning 2003</bibRefCitation>
: fig. 1I, 2H), the G1 of the present male is broadly curved, with a short, slender, obliquely directed apex (albeit proportionally shorter than in
<taxonomicName id="4C194D6FDC17FFC7C67DFEDAFE272DA2" authorityName="P.K.L.Ng &amp; R.B.Manning" authorityYear="2003" box="[370,493,369,395]" class="Malacostraca" family="Pinnotheridae" genus="Holotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC17FFC7C67DFEDAFE272DA2" box="[370,493,369,395]" italics="true" pageId="9" pageNumber="342">Holotheres</emphasis>
</taxonomicName>
) (
<figureCitation id="13222A69DC17FFC7C509FEDAFD972DA2" box="[518,605,368,395]" captionStart="FIGURE 5" captionStartId="9.[151,250,1855,1880]" captionTargetBox="[153,1433,458,1826]" captionTargetId="figure@9.[151,1436,453,1830]" captionTargetPageId="9" captionText="FIGURE 5. Buergeres sp., male, cl 5.3 mm, cw 5.6 mm, Manga Point, Bohol, Visayas, Philippines, ZRC 2018.0784:A, dorsal habitus; B, anterior cephalothorax; C, right maxilliped 3; D, right chela; EH, right P25; I, anterior thoracic sternum; J, pleon; KL, left G1, sternal and pleonal views, respectively; MN, left G2, sternal and pleonal views, respectively. Scales:A = 2.0 mm; B, DH, J = 1.0 mm; C, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954457" httpUri="https://zenodo.org/record/3954457/files/figure.png" pageId="9" pageNumber="342">Fig. 5L</figureCitation>
). As in many other pinnotherids, the G2 of the male
<taxonomicName id="4C194D6FDC17FFC7C3A3FED9FAD72DA2" authorityName="Ng &amp; Manning" authorityYear="2003" box="[1196,1309,370,395]" class="Malacostraca" family="Pinnotheridae" genus="Buergeres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFEDC17FFC7C3A3FED9FAD72DA2" box="[1196,1309,370,395]" italics="true" pageId="9" pageNumber="342">Buergeres</emphasis>
</taxonomicName>
has a short but distinct exopod (
<figureCitation id="13222A69DC17FFC7C672FE3EFE372D86" box="[381,509,404,431]" captionStart="FIGURE 5" captionStartId="9.[151,250,1855,1880]" captionTargetBox="[153,1433,458,1826]" captionTargetId="figure@9.[151,1436,453,1830]" captionTargetPageId="9" captionText="FIGURE 5. Buergeres sp., male, cl 5.3 mm, cw 5.6 mm, Manga Point, Bohol, Visayas, Philippines, ZRC 2018.0784:A, dorsal habitus; B, anterior cephalothorax; C, right maxilliped 3; D, right chela; EH, right P25; I, anterior thoracic sternum; J, pleon; KL, left G1, sternal and pleonal views, respectively; MN, left G2, sternal and pleonal views, respectively. Scales:A = 2.0 mm; B, DH, J = 1.0 mm; C, KN = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954457" httpUri="https://zenodo.org/record/3954457/files/figure.png" pageId="9" pageNumber="342">Fig. 5M, N</figureCitation>
;
<bibRefCitation id="EF884B1DDC17FFC7C507FE3EFD2E2D86" author="Ahyong, S. T. &amp; Komai, T. &amp; Watanabe, T." box="[520,740,405,431]" pageId="9" pageNumber="342" pagination="35 - 47" refId="ref8937" refString="Ahyong, S. T., Komai, T. &amp; Watanabe, T. (2012) First Viridotheres Manning, 1996, from Japan, with a key to the species (Decapoda, Brachyura, Pinnotheridae). In: Komatsu, H., Okuno, J. &amp; Fukuoka, K. (Eds.), Studies in Malacostraca: a homage to Masatsune Takeda. Crustaceana Monographs 17. Brill, Leiden, pp. 35 - 47. https: // doi. org / 10.1163 / 9789004202894 _ 003" type="book chapter" year="2012">
Ahyong
<emphasis id="B96DEAFEDC17FFC7C566FE3DFD6B2D86" box="[617,673,405,431]" italics="true" pageId="9" pageNumber="342">et al.</emphasis>
2012
</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC17FFC7C5E0FE3EFC6C2D86" author="Ng, P. K. L. &amp; Ho, P. - H." box="[751,934,405,431]" pageId="9" pageNumber="342" pagination="229 - 241" refId="ref10826" refString="Ng, P. K. L. &amp; Ho, P. - H. (2016 a) Orthotheres baoyu, a new species of pea crab (Crustacea: Brachyura: Pinnotheridae) associated with abalones from Tungsha Island, Taiwan; with notes on the genus. Raffles Bulletin of Zoology, 64, 229 - 241." type="journal article" year="2016">Ng &amp; Ho 2016a</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC17FFC7C4BEFE3EFBEE2D86" author="Ng, P. K. L." box="[945,1060,405,431]" pageId="9" pageNumber="342" pagination="474 - 485" refId="ref10601" refString="Ng, P. K. L. (2018 b) Arcotheres placunicola, a new species of pea crab (Crustacea: Brachyura: Pinnotheridae) from the windowpane shell, Placuna ephippium Philipsson, 1788 (Placunidae) in Singapore. Raffles Bulletin of Zoology, 66, 474 - 485." type="journal article" year="2018">Ng 2018b</bibRefCitation>
;
<bibRefCitation id="EF884B1DDC17FFC7C321FE3EFB062D86" author="Ahyong, S. T." box="[1070,1228,405,431]" pageId="9" pageNumber="342" pagination="107 - 118" refId="ref8819" refString="Ahyong, S. T. (2019) First Indonesian Viridotheres Manning, 1996, and redescription of male Afropinnotheres dofleini Manning, 1993, from South Africa (Decapoda, Pinnotheridae). Crustaceana, 92, 107 - 118. https: // doi. org / 10.1163 / 15685403 - 00003855" type="journal article" year="2019">Ahyong 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>