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<document id="E7E807F2BCA4925FABD1C8C971EC4A47" ID-DOI="10.11646/zootaxa.5380.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10216220" ID-ZooBank="6B51DC3F-67C4-4833-92C7-3387297985A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701246344783" checkinUser="plazi" docAuthor="Gouvea, Bruno, Valente, Roberta M. &amp; Asenjo, Angélico" docDate="2023" docId="03B087E32145F257FF29FB2CA8EC1408" docLanguage="en" docName="zootaxa.5380.2.2.pdf" docOrigin="Zootaxa 5380 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5380.2.2/52383" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxarthrius spiculatus Gouvea, Valente &amp; Asenjo 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="144" masterDocId="FF89FF9B2147F25DFFBEFFA3AA3D156D" masterDocTitle="A new species from the eastern Amazon rainforest with lectotype designations and key for Brazilian Oxarthrius Reitter, 1883 (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="150" masterPageNumber="134" pageNumber="136" updateTime="1701246758839" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8706E66E107B8449AE3CEF8E76C38EF5">A new species from the eastern Amazon rainforest with lectotype designations and key for Brazilian Oxarthrius Reitter, 1883 (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="952A8DDEC8A644E7243D8C8814C5B05E">Gouvea, Bruno</mods:namePart>
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<mods:affiliation id="FE04C6D8B3721D8B3B8C0E8732B38245">Universidade Federal do Pará, Instituto de Ciências Biológicas, Laboratório de Invertebrados, Belém, Pará, Brazil. brunoleandro 555 @ gmail. com; https: // orcid. org / 0009 - 0002 - 1540 - 9303</mods:affiliation>
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<mods:namePart id="DFDA78472FB119273457B8D84B126CD4">Valente, Roberta M.</mods:namePart>
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<mods:affiliation id="0F2F5D22F69004C385ED0CCAE5D30B51">Universidade Federal do Pará, Instituto de Ciências Biológicas, Laboratório de Invertebrados, Belém, Pará, Brazil. roberta. mvalente @ gmail. com; https: // orcid. org / 0000 - 0002 - 8249 - 5937</mods:affiliation>
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<mods:namePart id="79A4C2108209FBC099870FA1EB587FBC">Asenjo, Angélico</mods:namePart>
<mods:affiliation id="D86123ED5712CDB4754B50E2C25CA535">Instituto Tecnológico VALE, R. Boaventura da Silva 955, Nazaré, 66055 - 090 Belém, PA. Brazil. &amp; Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Jesús María, Lima 14, Perú.</mods:affiliation>
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<taxonomicName id="4C194D762145F25FFF29FB2CA8D211C4" authority="Gouvea, Valente &amp; Asenjo" authorityName="Gouvea, Valente &amp; Asenjo" authorityYear="2023" box="[151,751,1167,1194]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72145F25FFF29FB2CABA311C7" bold="true" box="[151,414,1167,1194]" italics="true" pageId="2" pageNumber="136">Oxarthrius spiculatus</emphasis>
Gouvea, Valente &amp; Asenjo
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, 
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(
<figureCitation id="13222A702145F25FFF21FB74AB3F119F" box="[159,258,1239,1266]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1685,1709]" captionStartId-1="5.[151,250,1878,1902]" captionStartId-2="6.[151,250,1489,1513]" captionStartId-3="7.[151,250,1848,1872]" captionStartId-4="8.[151,250,1487,1511]" captionTargetBox-0="[151,1435,164,1659]" captionTargetBox-1="[151,1435,191,1852]" captionTargetBox-2="[152,1436,179,1463]" captionTargetBox-3="[152,1435,180,1821]" captionTargetBox-4="[154,1433,181,1462]" captionTargetId-0="figure-20@3.[151,1435,164,1659]" captionTargetId-1="figure-20@5.[151,1435,179,1852]" captionTargetId-2="figure-19@6.[152,1436,179,1463]" captionTargetId-3="figure-20@7.[151,1436,179,1822]" captionTargetId-4="figure-19@8.[154,1433,181,1462]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." captionText-1="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." captionText-2="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." captionText-3="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." captionText-4="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi-0="http://doi.org/10.5281/zenodo.10216222" figureDoi-1="http://doi.org/10.5281/zenodo.10216226" figureDoi-2="http://doi.org/10.5281/zenodo.10216228" figureDoi-3="http://doi.org/10.5281/zenodo.10216234" figureDoi-4="http://doi.org/10.5281/zenodo.10216236" httpUri-0="https://zenodo.org/record/10216222/files/figure.png" httpUri-1="https://zenodo.org/record/10216226/files/figure.png" httpUri-2="https://zenodo.org/record/10216228/files/figure.png" httpUri-3="https://zenodo.org/record/10216234/files/figure.png" httpUri-4="https://zenodo.org/record/10216236/files/figure.png" pageId="2" pageNumber="136">Figs 1–5</figureCitation>
)
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<emphasis id="B96DEAE72145F25FFF29FABCAB031054" bold="true" box="[151,318,1311,1337]" pageId="2" pageNumber="136">Type material</emphasis>
(
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, 
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).
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. 
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: 
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:
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, labeled [dissected]: “STP0599 / BRAZIL: Paragominas, / Mineradora Hydro. CP, / 
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, 
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[without other data]” “ 
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[red card] / 
<taxonomicName id="4C194D762145F25FFC65FAC4AE6B10EC" authorityName="Reitter" authorityYear="1883" box="[987,1110,1383,1409]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="genus">
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/ 
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<emphasis id="B96DEAE72145F25FFBD3FAC4AEE210EC" box="[1133,1247,1383,1409]" italics="true" pageId="2" pageNumber="136">spiculatus</emphasis>
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/ Desig. Gouvea 
<emphasis id="B96DEAE72145F25FFF4BFA2FAB1B10C8" box="[245,294,1419,1445]" italics="true" pageId="2" pageNumber="136">et al</emphasis>
. 2023”; (
<collectionCode id="ED08AE302145F25FFE31FA28ABDE10C8" box="[399,483,1419,1445]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
).
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<materialsCitation id="3B713CA82145F25FFF79FA0CA89B10A4" box="[199,678,1454,1482]" country="Brazil" location="Para" pageId="2" pageNumber="136" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Para" typeStatus="paratype">
<emphasis id="B96DEAE72145F25FFF79FA0CAB6110A4" bold="true" box="[199,348,1455,1481]" pageId="2" pageNumber="136">
<typeStatus id="54A288572145F25FFF79FA0CAB0210A4" box="[199,319,1455,1481]" pageId="2" pageNumber="136" type="paratype">Paratypes</typeStatus>
(
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♂♂, 
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<emphasis id="B96DEAE72145F25FFE69FA0CA89B10A4" bold="true" box="[471,678,1454,1481]" pageId="2" pageNumber="136">
). 
<collectingCountry id="F30E76652145F25FFE53FA0CA86010A4" box="[493,605,1455,1481]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: 
<collectingRegion id="49DDF8172145F25FFDD6FA0DA89B10A4" box="[616,678,1454,1481]" country="Brazil" name="Para" pageId="2" pageNumber="136">Pará</collectingRegion>
</emphasis>
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<emphasis id="B96DEAE72145F25FFD1BFA0DA89710A4" bold="true" box="[677,682,1454,1481]" pageId="2" pageNumber="136">:</emphasis>
<materialsCitation id="3B713CA82145F25FFD0EFA13A99C1080" collectionCode="MPEG" country="Brazil" latitude="-3.2575" longLatPrecision="21" longitude="-47.70778" pageId="2" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="paratype">
<specimenCount id="9D1FFD7C2145F25FFD0EFA13A8FA10A7" box="[688,711,1456,1482]" count="1" pageId="2" pageNumber="136" type="male">♂</specimenCount>
, “STP0266 / BRAZIL: Paragominas, / Mineradora Hydro. AL10, 
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. / Winkler [without other data]” (
<collectionCode id="ED08AE302145F25FFCFBFA70A9A41080" box="[837,921,1491,1517]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79FA5BAE321358" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79FA5BAE311358" collectionCode="MPEG" country="Brazil" latitude="-3.2386112" location="BRAZIL" longLatPrecision="21" longitude="-47.698612" pageId="2" pageNumber="136" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D1FFD7C2145F25FFF79FA5BAAE3137F" box="[199,222,1528,1554]" count="1" pageId="2" pageNumber="136" type="male">♂</specimenCount>
, [specimen for Scanning Electron Micrographs]; “STP0269 / 
<collectingCountry id="F30E76652145F25FFC23FA54AE3A137C" box="[925,1031,1527,1553]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: Paragominas, / Mineradora Hydro. AL3, / 
<geoCoordinate id="EE2D50322145F25FFF59F9B8AB4E1358" box="[231,371,1563,1589]" degrees="03" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="19" value="-3.2386112">S 03°14’19”</geoCoordinate>
<geoCoordinate id="EE2D50322145F25FFEC7F9BFA820135B" box="[377,541,1563,1590]" degrees="047" direction="west" minutes="41" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="55" value="-47.698612">W 047°41’55”</geoCoordinate>
/ 20-VI[June]-18[2018]. Winkler” (
<collectionCode id="ED08AE302145F25FFC11F9B8AE3E1358" box="[943,1027,1563,1589]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F9E3AB571310" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79F9E3AB5B1310" collectionCode="UFPA" country="Brazil" latitude="-3.2386112" location="BRAZIL" longLatPrecision="21" longitude="-47.698612" pageId="2" pageNumber="136" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D1FFD7C2145F25FFF79F9E3AAE31337" box="[199,222,1600,1626]" count="1" pageId="2" pageNumber="136" type="male">♂</specimenCount>
, [dissected]; “STP0275 / 
<collectingCountry id="F30E76652145F25FFDBEF99CA8571334" box="[512,618,1599,1625]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: Paragominas, / Mineradora Hydro. AL3, / 
<geoCoordinate id="EE2D50322145F25FFBEAF99CAEDC1334" box="[1108,1249,1599,1625]" degrees="03" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="19" value="-3.2386112">S 03°14’19”</geoCoordinate>
<geoCoordinate id="EE2D50322145F25FFB56F9E3AFB01337" box="[1256,1421,1599,1626]" degrees="047" direction="west" minutes="41" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="55" value="-47.698612">W 047°41’55”</geoCoordinate>
/ Winkler.” (
<collectionCode id="ED08AE302145F25FFEACF9C0AB631310" box="[274,350,1635,1661]" pageId="2" pageNumber="136">UFPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F92BA86113A8" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79F92BA86513A8" collectionCode="MPEG" country="Brazil" latitude="-3.2619445" location="BRAZIL" longLatPrecision="21" longitude="-47.701946" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1FFD7C2145F25FFF79F92BAAE113CF" box="[199,220,1672,1698]" count="1" pageId="2" pageNumber="136" type="female">♀</specimenCount>
, [dissected]; “STP0273 / 
<collectingCountry id="F30E76652145F25FFE40F924A85513CC" box="[510,616,1671,1697]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: Paragominas, / Mineradora Hydro. AL9, / 
<geoCoordinate id="EE2D50322145F25FFBF1F924AEE613CF" box="[1103,1243,1671,1698]" degrees="03" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="43" value="-3.2619445">S 03°15’43”</geoCoordinate>
<geoCoordinate id="EE2D50322145F25FFB5CF92BAFB413CC" box="[1250,1417,1671,1698]" degrees="047" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="07" value="-47.701946">W 047°42’07”</geoCoordinate>
. / 7-V[May]-18[2018]. Winkler” (
<collectionCode id="ED08AE302145F25FFE42F908A86D13A8" box="[508,592,1707,1733]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F973A8BF1260" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79F973A8421260" collectionCode="MPEG" country="Brazil" county="Winkler" latitude="-3.2619445" location="Paragominas" longLatPrecision="21" longitude="-47.701946" municipality="June" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1FFD7C2145F25FFF79F973AAE11387" box="[199,220,1744,1770]" count="1" pageId="2" pageNumber="136" type="female">♀</specimenCount>
, [MEV] “STP0274 / 
<collectingCountry id="F30E76652145F25FFE6AF96CA8031384" box="[468,574,1743,1769]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: 
<location id="8EC6602E2145F25FFDF4F96CA8E31384" LSID="urn:lsid:plazi:treatment:03B087E32145F257FF29FB2CA8EC1408:8EC6602E2145F25FFDF4F96CA8E31384" box="[586,734,1743,1769]" country="Brazil" county="Winkler" latitude="-3.2619445" longLatPrecision="21" longitude="-47.701946" municipality="June" name="Paragominas" pageId="2" pageNumber="136">Paragominas</location>
, / Mineradora Hydro. / 
<location id="8EC6602E2145F25FFC50F96CAE301384" LSID="urn:lsid:plazi:treatment:03B087E32145F257FF29FB2CA8EC1408:8EC6602E2145F25FFC50F96CAE301384" box="[1006,1037,1743,1769]" country="Brazil" county="Winkler" latitude="-3.2619445" longLatPrecision="21" longitude="-47.701946" municipality="June" name="Al" pageId="2" pageNumber="136">Al</location>
9(4), 
<geoCoordinate id="EE2D50322145F25FFBF7F96CAEEB1387" box="[1097,1238,1743,1770]" degrees="03" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="43" value="-3.2619445">S 03°15’43”</geoCoordinate>
/ 
<geoCoordinate id="EE2D50322145F25FFB51F973AFA41384" box="[1263,1433,1743,1770]" degrees="047" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="07" value="-47.701946">W 047°42’07”</geoCoordinate>
. 29-VI[
<collectingMunicipality id="6BC2AC8F2145F25FFF5BF950AB221260" box="[229,287,1779,1805]" pageId="2" pageNumber="136">June</collectingMunicipality>
]-18[2018]. / 
<collectingCounty id="62C74E792145F25FFE12F950A8341260" box="[428,521,1779,1805]" pageId="2" pageNumber="136">Winkler</collectingCounty>
” (
<collectionCode id="ED08AE302145F25FFD9CF950A84B1260" box="[546,630,1779,1805]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F8BBA8601238" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79F8BBA8641238" collectionCode="MPEG" country="Brazil" county="Winkler" latitude="-3.2619445" location="Paragominas" longLatPrecision="21" longitude="-47.701946" municipality="June" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1FFD7C2145F25FFF79F8BBAAE1125F" box="[199,220,1816,1842]" count="1" pageId="2" pageNumber="136" type="female">♀</specimenCount>
, “STP0267 / 
<collectingCountry id="F30E76652145F25FFE3FF8B4ABD6125C" box="[385,491,1815,1841]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: 
<location id="8EC6602E2145F25FFE44F8B4A8B3125C" LSID="urn:lsid:plazi:treatment:03B087E32145F257FF29FB2CA8EC1408:8EC6602E2145F25FFE44F8B4A8B3125C" box="[506,654,1815,1841]" country="Brazil" county="Winkler" latitude="-3.2619445" longLatPrecision="21" longitude="-47.701946" municipality="June" name="Paragominas" pageId="2" pageNumber="136">Paragominas</location>
, / Mineradora Hydro. / AL9(2), 
<geoCoordinate id="EE2D50322145F25FFBB1F8B4AEA3125F" box="[1039,1182,1815,1842]" degrees="03" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="43" value="-3.2619445">S 03°15’43”</geoCoordinate>
/ 
<geoCoordinate id="EE2D50322145F25FFB02F8BBAF55125C" box="[1212,1384,1815,1842]" degrees="047" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="07" value="-47.701946">W 047°42’07”</geoCoordinate>
. 20- VI[
<collectingMunicipality id="6BC2AC8F2145F25FFF01F898AAC41238" box="[191,249,1851,1877]" pageId="2" pageNumber="136">June</collectingMunicipality>
]-18[2018]. / 
<collectingCounty id="62C74E792145F25FFE39F898ABD91238" box="[391,484,1851,1877]" pageId="2" pageNumber="136">Winkler</collectingCounty>
” (
<collectionCode id="ED08AE302145F25FFE43F898A86C1238" box="[509,593,1851,1877]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="136">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F8C3A86912F0" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
<materialsCitation id="3B713CA82145F25FFF79F8C3A86D12F0" collectionCode="UFPA" country="Brazil" latitude="-3.2619445" location="BRAZIL" longLatPrecision="21" longitude="-47.701946" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1FFD7C2145F25FFF79F8C3AAE11217" box="[199,220,1888,1914]" count="1" pageId="2" pageNumber="136" type="female">♀</specimenCount>
, [dissected] “STP0271 / 
<collectingCountry id="F30E76652145F25FFE46F8FCA85F1214" box="[504,610,1887,1913]" name="Brazil" pageId="2" pageNumber="136">BRAZIL</collectingCountry>
: Paragominas, / Mineradora Hydro. AL9, / 
<geoCoordinate id="EE2D50322145F25FFBF3F8FCAEE41217" box="[1101,1241,1887,1914]" degrees="03" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="136" precision="15" seconds="43" value="-3.2619445">S 03°15’43”</geoCoordinate>
<geoCoordinate id="EE2D50322145F25FFB5FF8C3AFB41214" box="[1249,1417,1887,1914]" degrees="047" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="136" precision="15" seconds="07" value="-47.701946">W 047°42’07”</geoCoordinate>
. / 7-V[May]-18[2018]. Winkler” (
<collectionCode id="ED08AE302145F25FFE42F820A87512F0" box="[508,584,1923,1949]" pageId="2" pageNumber="136">UFPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636F52145F25FFF79F804AADE1288" blockId="2.[151,1437,1311,2021]" pageId="2" pageNumber="136">
All 
<typeStatus id="54A288572145F25FFF4AF80BAB6212AF" box="[244,351,1960,1986]" pageId="2" pageNumber="136" type="paratype">paratypes</typeStatus>
with label “ 
<typeStatus id="54A288572145F25FFE55F804A84412AC" box="[491,633,1959,1985]" pageId="2" pageNumber="136" type="paratype">PARATYPE</typeStatus>
[yellow card] / 
<taxonomicName id="4C194D762145F25FFC8DF804A99312AC" authorityName="Reitter" authorityYear="1883" box="[819,942,1959,1985]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE72145F25FFC8DF804A99312AC" box="[819,942,1959,1985]" italics="true" pageId="2" pageNumber="136">Oxarthrius</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C194D762145F25FFC79F804AE0412AC" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[967,1081,1959,1985]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72145F25FFC79F804AE0412AC" box="[967,1081,1959,1985]" italics="true" pageId="2" pageNumber="136">spiculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E579C2145F25FFBFCF80BAEA712AF" box="[1090,1178,1960,1986]" pageId="2" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
/ Desig. Gouvea et al. 2023”.
</paragraph>
</subSubSection>
<caption id="DF66667D2144F25EFF29F936AF1813BF" ID-DOI="http://doi.org/10.5281/zenodo.10216222" ID-Zenodo-Dep="10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="3" pageNumber="137" startId="3.[151,250,1685,1709]" targetBox="[151,1435,164,1659]" targetPageId="3" targetType="figure">
<paragraph id="8BA636F52144F25EFF29F936AF1813BF" blockId="3.[151,1436,1685,1746]" pageId="3" pageNumber="137">
<emphasis id="B96DEAE72144F25EFF29F936AB3313C0" bold="true" box="[151,270,1685,1709]" pageId="3" pageNumber="137">FIGURE 1</emphasis>
. 
<taxonomicName id="4C194D762144F25EFEA5F936ABC513C0" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[283,504,1685,1709]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72144F25EFEA5F936ABC513C0" box="[283,504,1685,1709]" italics="true" pageId="3" pageNumber="137">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72144F25EFE41F935A86D13C3" bold="true" box="[511,592,1686,1710]" pageId="3" pageNumber="137">
<taxonomicNameLabel id="A25E579C2144F25EFE41F935A86D13C3" box="[511,592,1686,1710]" pageId="3" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) 
<emphasis id="B96DEAE72144F25EFA99F936AF7D13C0" bold="true" box="[1319,1344,1685,1709]" pageId="3" pageNumber="137">A.</emphasis>
Habitus, dorsal view. 
<emphasis id="B96DEAE72144F25EFEA6F919AB1213BF" bold="true" box="[280,303,1722,1746]" pageId="3" pageNumber="137">B.</emphasis>
Habitus, dorsal view. 
<emphasis id="B96DEAE72144F25EFDAFF91AA81713BC" bold="true" box="[529,554,1721,1745]" pageId="3" pageNumber="137">C.</emphasis>
Metaventrite. 
<emphasis id="B96DEAE72144F25EFD00F919A8EA13BF" bold="true" box="[702,727,1722,1746]" pageId="3" pageNumber="137">D.</emphasis>
Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D).
</paragraph>
</caption>
<subSubSection id="C303657E2144F25EFF79F95CAE6E12EB" pageId="3" pageNumber="137" type="diagnosis">
<paragraph id="8BA636F52144F25EFF79F95CAE6E12EB" blockId="3.[151,1437,1791,2034]" pageId="3" pageNumber="137">
<emphasis id="B96DEAE72144F25EFF79F95CAB7F1274" bold="true" box="[199,322,1791,1817]" pageId="3" pageNumber="137">Diagnosis.</emphasis>
<taxonomicName id="4C194D762144F25EFEF5F8A3A87C1277" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[331,577,1792,1818]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72144F25EFEF5F8A3A87C1277" box="[331,577,1792,1818]" italics="true" pageId="3" pageNumber="137">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72144F25EFDF5F8A3A89A1277" bold="true" box="[587,679,1792,1818]" pageId="3" pageNumber="137">
<taxonomicNameLabel id="A25E579C2144F25EFDF5F8A3A89A1277" box="[587,679,1792,1818]" pageId="3" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more similar to 
<taxonomicName id="4C194D762144F25EFC39F8A3AE101277" box="[903,1069,1792,1818]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE72144F25EFC39F8A3A9A61277" box="[903,923,1792,1818]" italics="true" pageId="3" pageNumber="137">O</emphasis>
. 
<emphasis id="B96DEAE72144F25EFC12F8A3AE101277" box="[940,1069,1792,1818]" italics="true" pageId="3" pageNumber="137">anthicoides</emphasis>
</taxonomicName>
due to the rough surface of the pronotum. However, 
<taxonomicName id="4C194D762144F25EFE3BF887A8451253" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[389,632,1828,1854]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72144F25EFE3BF887A8451253" box="[389,632,1828,1854]" italics="true" pageId="3" pageNumber="137">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72144F25EFDC1F887A8E51253" bold="true" box="[639,728,1828,1854]" pageId="3" pageNumber="137">
<taxonomicNameLabel id="A25E579C2144F25EFDC1F887A8E51253" box="[639,728,1828,1854]" pageId="3" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the thin vertexal sulcus [vs] the lack of carinae on the head, and by pronotum lacking lateral longitudinal sulci [lls]. In 
<taxonomicName id="4C194D762144F25EFBB4F8EBAE90120F" box="[1034,1197,1864,1890]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE72144F25EFBB4F8EBAE23120F" box="[1034,1054,1864,1890]" italics="true" pageId="3" pageNumber="137">O</emphasis>
. 
<emphasis id="B96DEAE72144F25EFB92F8EBAE90120F" box="[1068,1197,1864,1890]" italics="true" pageId="3" pageNumber="137">anthicoides</emphasis>
</taxonomicName>
the vertexal sulcus is large and carinate, and the lateral longitudinal sulci [lls] are evident on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C303657E2144F257FF79F82CA8141594" lastPageId="10" lastPageNumber="144" pageId="3" pageNumber="137" type="description">
<paragraph id="8BA636F52144F25EFF79F82CAE44129F" blockId="3.[151,1437,1791,2034]" pageId="3" pageNumber="137">
<emphasis id="B96DEAE72144F25EFF79F82CAB6412C4" bold="true" box="[199,345,1935,1961]" pageId="3" pageNumber="137">Description.</emphasis>
<typeStatus id="54A288572144F25EFEDEF833ABF512C7" box="[352,456,1936,1962]" pageId="3" pageNumber="137" type="holotype">Holotype</typeStatus>
male, body length: 2.81 (
<figureCitation id="13222A702144F25EFD56F833A97312C7" box="[744,846,1936,1962]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1685,1709]" captionStartId-1="5.[151,250,1878,1902]" captionTargetBox-0="[151,1435,164,1659]" captionTargetBox-1="[151,1435,191,1852]" captionTargetId-0="figure-20@3.[151,1435,164,1659]" captionTargetId-1="figure-20@5.[151,1435,179,1852]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." captionText-1="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi-0="http://doi.org/10.5281/zenodo.10216222" figureDoi-1="http://doi.org/10.5281/zenodo.10216226" httpUri-0="https://zenodo.org/record/10216222/files/figure.png" httpUri-1="https://zenodo.org/record/10216226/files/figure.png" pageId="3" pageNumber="137">Figs 1–2</figureCitation>
). Head and pronotum dark reddish-brown, tegument densely rough. Antennae, legs, elytra and abdomen light reddish brown, tegument of elytra and abdomen shiny and polished. Tarsi and mouthparts paler. Body covered with sparse long setae. (
<figureCitation id="13222A702144F25EFC55F87BAE6D129F" box="[1003,1104,2008,2034]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1685,1709]" captionStartId-1="5.[151,250,1878,1902]" captionStartId-2="6.[151,250,1489,1513]" captionTargetBox-0="[151,1435,164,1659]" captionTargetBox-1="[151,1435,191,1852]" captionTargetBox-2="[152,1436,179,1463]" captionTargetId-0="figure-20@3.[151,1435,164,1659]" captionTargetId-1="figure-20@5.[151,1435,179,1852]" captionTargetId-2="figure-19@6.[152,1436,179,1463]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." captionText-1="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." captionText-2="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi-0="http://doi.org/10.5281/zenodo.10216222" figureDoi-1="http://doi.org/10.5281/zenodo.10216226" figureDoi-2="http://doi.org/10.5281/zenodo.10216228" httpUri-0="https://zenodo.org/record/10216222/files/figure.png" httpUri-1="https://zenodo.org/record/10216226/files/figure.png" httpUri-2="https://zenodo.org/record/10216228/files/figure.png" pageId="3" pageNumber="137">Figs 1–3</figureCitation>
, 
<figureCitation id="13222A702144F25EFBE5F874AE56129F" box="[1115,1131,2007,2034]" captionStart="FIGURE 5" captionStartId="8.[151,250,1487,1511]" captionTargetBox="[154,1433,181,1462]" captionTargetId="figure-19@8.[154,1433,181,1462]" captionTargetPageId="8" captionText="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="3" pageNumber="137">5</figureCitation>
).
</paragraph>
<paragraph id="8BA636F52143F259FF79FF34AE7C179C" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Head: slightly oval (length 0.32: width 0.41) with small antennal tubercles at lateral margin of frontal ridge. Antennal insertions of head not visible in dorsal view. Eyes prominent, situated at posterior half of head with sparse interfacetgal setae between ommatidia, genal area small and rounded. Neck strongly constricted (neck width 0.20). Head with two deep vertexal foveae [dorsal tentorial pits] [vf] connected by a slightly distinguished U-shaped vertexal sulcus [vs] that extends to a depression between antennal tubercles (
<figureCitation id="13222A702143F259FBB0FE84AE51142C" box="[1038,1132,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">Figs 2B</figureCitation>
, 
<figureCitation id="13222A702143F259FBC4FE84AEA2142C" box="[1146,1183,295,321]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">3A</figureCitation>
). Median longitudinal carina extending from base of constricted neck to anterior margin of head (
<figureCitation id="13222A702143F259FBBBFEE8AE5F1408" box="[1029,1122,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3A</figureCitation>
). Ocular-mandibular carina present, extending from anterior margin of eyes to base of clypeus. Maxillary palpi four-segmented pubescent, distal segment longer and wider than others with unscarred external face. Gular carina absent. Gular foveae [gf] present in posterior region of head. Antenna thick and long (length 0.92) with antennal club formed by last three antennomeres, antennal length exceed their middle body length (
<figureCitation id="13222A702143F259FCC8FE78A9E81498" box="[886,981,475,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1A</figureCitation>
, 
<figureCitation id="13222A702143F259FC5CFE78AE161498" box="[994,1067,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A–D</figureCitation>
). Scape subcylindrical (
<figureCitation id="13222A702143F259FA87FE78AFAA1498" box="[1337,1431,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">Figs 2A</figureCitation>
, 
<figureCitation id="13222A702143F259FF29FE5CAA861774" box="[151,187,511,537]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">3A</figureCitation>
, 
<figureCitation id="13222A702143F259FF79FE5DAAD71774" box="[199,234,510,537]" captionStart="FIGURE 5" captionStartId="8.[151,250,1487,1511]" captionTargetBox="[154,1433,181,1462]" captionTargetId="figure-19@8.[154,1433,181,1462]" captionTargetPageId="8" captionText="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="4" pageNumber="138">5B</figureCitation>
), longer than wide (length 0.14: width 0.08), base curved, apex notched at dorsal and ventral view. Rounded pedicel, thinner and shorter than scape (length 0.06: width 0.06). Segment 3–7 slightly longer than wide (
<figureCitation id="13222A702143F259FA84FD80AFA51750" box="[1338,1432,547,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1A</figureCitation>
, 
<figureCitation id="13222A702143F259FF29FDE4AADC170C" box="[151,225,583,609]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A–D</figureCitation>
): 3 (length 0.05: width 0.04); 4 (length 0.06: width 0.05); 5 (length 0.07: width 0.05); 6 (length 0.07: width 0.05); 7 (length 0.07: width 0.06). Segment 8 (length 0.07: width 0.06). Segment 9–11 forming distinctive antennal club (
<figureCitation id="13222A702143F259FF66FD2CAB0B17C4" box="[216,310,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1A</figureCitation>
, 
<figureCitation id="13222A702143F259FEFDFD2CABB017C4" box="[323,397,655,681]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A–D</figureCitation>
): 9 subcylindrical, straight angles (length 0.09: width 0.07); 10 subquadrate, straight angles (length 0.08: width 0.09); 11 fusiform, straight angles at base and obtuse at apex (length 0.19: width 0.09). All antennal segments covered with microsetae, segments 9–11 with microsetae denser.
</paragraph>
<paragraph id="8BA636F52143F259FF79FD58AFBD117C" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Thorax: Cordiform shape, similar in length (0.34) and width (0.34), edges of anterior half very tumescent, edge of posterior half tapering posteriorly; basal margin almost right; posterolateral angles almost right angle. Integument densely rough almost like wave pattern, with posterior half less rough (
<figureCitation id="13222A702143F259FC77FCE0AE1E1630" box="[969,1059,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3B</figureCitation>
). Pair of conspicuous basolateral spines next to one deep lateral antebasal foveae [laf] on external side (
<figureCitation id="13222A702143F259FC16FCC4AE3D16EC" box="[936,1024,871,897]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3B</figureCitation>
). Outer basolateral foveae [oblf] and inner basolateral foveae [iblf] at basal margin present (
<figureCitation id="13222A702143F259FD44FC28A96C16C8" box="[762,849,907,933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3B</figureCitation>
). Prosternum (
<figureCitation id="13222A702143F259FC48FC28AE7316C8" box="[1014,1102,907,933]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">Fig. 2A</figureCitation>
) with anteroprosternal foveae [apsf], lateral procoxal foveae present [lpcf] proepimeral foveae [pef] all present. Mesoventrite (
<figureCitation id="13222A702143F259FB6EFC0CAF1616A4" box="[1232,1323,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1C</figureCitation>
, 
<figureCitation id="13222A702143F259FA86FC0CAF6116A4" box="[1336,1372,943,969]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A</figureCitation>
) with pair of lateral mesosternal foveae [lmsf] and pair of lateral mesocoxal foveae [lmcf]. Metaventrite with pair of lateral metasternal foveae [lmtf] and median oval excavation (
<figureCitation id="13222A702143F259FCBFFC54A961117C" box="[769,860,1015,1041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1C</figureCitation>
, 
<figureCitation id="13222A702143F259FCD7FC54A9B4117C" box="[873,905,1015,1041]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2E</figureCitation>
) impressed by pattern of transverse grooves.
</paragraph>
<paragraph id="8BA636F52143F259FF79FBB8A80711CC" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Elytra (
<figureCitation id="13222A702143F259FEA5FBB8AB441158" box="[283,377,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1A</figureCitation>
, 
<figureCitation id="13222A702143F259FE38FBB8AB971158" box="[390,426,1051,1077]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">3C</figureCitation>
): wider than long (EL: 0.53; EW: 0.60), slightly wider than pronotum, sides rounded and gradually widening from the anterior to the posterior region, anterior margin straight, posterior margin sinuous. Humeri slightly prominently. Each elytron with one deep basal elytral fovea [bef] near scutellum (
<figureCitation id="13222A702143F259FB61FBC0AF051110" box="[1247,1336,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3C</figureCitation>
). Sutural stria [ss] complete, evident (
<figureCitation id="13222A702143F259FE6AFB24A81011CC" box="[468,557,1159,1185]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BA636F52143F259FF79FB08A8EB1014" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Legs: Thin and long. Procoxae and mesocoxae (
<figureCitation id="13222A702143F259FD5FFB08A90611A8" box="[737,827,1195,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1C</figureCitation>
, 
<figureCitation id="13222A702143F259FCF6FB08A95111A8" box="[840,876,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A</figureCitation>
) conical, prominent, contiguous; metacoxae (
<figureCitation id="13222A702143F259FAD4FB08AA841184" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1C</figureCitation>
, 
<figureCitation id="13222A702143F259FF74FB6CAB221184" box="[202,287,1231,1257]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">2A, 2E</figureCitation>
) transversely elongated, slightly prominent, separated. Protrochanter and mesotrochanther smaller, triangular shape; metatrochanter bigger, with minute black spine on the apex (
<figureCitation id="13222A702143F259FC48FB50AE6D1060" box="[1014,1104,1267,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Figs 1C</figureCitation>
, 
<figureCitation id="13222A702143F259FBE2FB51AEBF1060" box="[1116,1154,1266,1293]" captionStart="FIGURE 5" captionStartId="8.[151,250,1487,1511]" captionTargetBox="[154,1433,181,1462]" captionTargetId="figure-19@8.[154,1433,181,1462]" captionTargetPageId="8" captionText="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="4" pageNumber="138">5A</figureCitation>
). Femora thickening near the apex, mesofemur with acute and prominent spine on basal half, also one median crest posteriorly to the spine (
<figureCitation id="13222A702143F259FF21FA98AAC41038" box="[159,249,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="4" pageNumber="138">Fig. 1C</figureCitation>
). Tibiae elongated, slim, narrow. Tarsi 3-segmented, first tarsomere small, the last two elongated.Tarsal claws simple, with other minute claw on each tarsi.
</paragraph>
<paragraph id="8BA636F52143F259FF79FA20AE3F13D0" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Abdomen: Rounded (length 0.67), constricted on base. At dorsal view, paratergites visible only in tergite IV; intersegmental membranes with minute rectangular sclerites. Tergite IV (
<figureCitation id="13222A702143F259FC73FA04AE1910AC" box="[973,1060,1447,1473]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="4" pageNumber="138">Fig. 3D</figureCitation>
) (length 0.20: width 0.56) longer than tergite V, with pair of basolateral foveae [blf] and pair of mediobasal foveae [mbf]. Tergite V (length 0.08: width 0.48) and VI (length 0.06: width 0.41) very short. Tergite VII (length 0.13: width 0.31) longer than tergite V– VI and shorter than tergite IV, transverse, angles slightly converging from the base to the apex, base with projections that extends below tergite VI. Tergite VIII large, U-shaped (
<figureCitation id="13222A702143F259FC91F994A9B5133C" box="[815,904,1591,1617]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="4" pageNumber="138">Fig. 4D</figureCitation>
), concave centrally on anterior. Sternite III very short, with one deep mediobasal foveae [mbf] evident on the center. Sternite IV longer than III. Sternites V-VII short. Sternite VIII (
<figureCitation id="13222A702143F259FEC3F9DCABEF13F4" box="[381,466,1663,1689]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="4" pageNumber="138">Fig. 4F</figureCitation>
) large, transverse, slightly sinuous on anterior margin, protrude centrally on posterior margin, with large medial depression and subapical curved keel (
<figureCitation id="13222A702143F259FCD0F900A9F513D0" box="[878,968,1699,1725]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="4" pageNumber="138">Figs 2E</figureCitation>
, 
<figureCitation id="13222A702143F259FC6AF900A9C813D0" box="[980,1013,1699,1725]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="4" pageNumber="138">4F</figureCitation>
).
</paragraph>
<paragraph id="8BA636F52143F259FF79F964AB391220" blockId="4.[151,1437,151,1869]" pageId="4" pageNumber="138">
Aedeagus: Slightly asymmetrical (length 
<quantity id="4CE19B102143F259FD26F964A8CF138C" box="[664,754,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="138" unit="mm" value="0.3">0.3 mm</quantity>
). Median lobe bulbous; apex almost straight with one central plate (cp) divided into two short opposite branches (
<figureCitation id="13222A702143F259FD60F948A9561268" box="[734,875,1771,1797]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="4" pageNumber="138">Figs 4A, 4C</figureCitation>
), ventrally with two poorly defined spots of setae, median pointed projection (mpp) ventrally facing on each side of aedeagus, discrete (
<figureCitation id="13222A702143F259FBE4F8ACAED41244" box="[1114,1257,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="4" pageNumber="138">Figs 4A, 4B</figureCitation>
), parameres not observed.
</paragraph>
<caption id="DF66667D2142F258FF29F8F5A89912DA" ID-DOI="http://doi.org/10.5281/zenodo.10216226" ID-Zenodo-Dep="10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="5" pageNumber="139" startId="5.[151,250,1878,1902]" targetBox="[151,1435,191,1852]" targetPageId="5" targetType="figure">
<paragraph id="8BA636F52142F258FF29F8F5A89912DA" blockId="5.[151,1436,1878,1975]" pageId="5" pageNumber="139">
<emphasis id="B96DEAE72142F258FF29F8F5AB2F1203" bold="true" box="[151,274,1878,1902]" pageId="5" pageNumber="139">FIGURE 2.</emphasis>
<taxonomicName id="4C194D762142F258FEA9F8F5ABCE1203" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[279,499,1878,1902]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72142F258FEA9F8F5ABCE1203" box="[279,499,1878,1902]" italics="true" pageId="5" pageNumber="139">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72142F258FE46F8F4A87A1202" bold="true" box="[504,583,1879,1903]" pageId="5" pageNumber="139">
<taxonomicNameLabel id="A25E579C2142F258FE46F8F4A87A1202" box="[504,583,1879,1903]" pageId="5" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A-E. Male holotype - STP0599; F. Female paratype – STP0267) 
<emphasis id="B96DEAE72142F258FB63F8F5AECB1203" bold="true" box="[1245,1270,1878,1902]" pageId="5" pageNumber="139">A.</emphasis>
Habitus, ventral view. 
<emphasis id="B96DEAE72142F258FF6AF8D8AAD612FE" bold="true" box="[212,235,1915,1939]" pageId="5" pageNumber="139">B.</emphasis>
Head, dorsal view. 
<emphasis id="B96DEAE72142F258FE06F8D9ABED12FF" bold="true" box="[440,464,1914,1938]" pageId="5" pageNumber="139">C.</emphasis>
Habitus, lateral view. 
<emphasis id="B96DEAE72142F258FD09F8D8A8ED12FE" bold="true" box="[695,720,1915,1939]" pageId="5" pageNumber="139">D.</emphasis>
Antennae. 
<emphasis id="B96DEAE72142F258FCFAF8D8A96612FE" bold="true" box="[836,859,1915,1939]" pageId="5" pageNumber="139">E.</emphasis>
Abdomen ventral view. 
<emphasis id="B96DEAE72142F258FBE9F8D8AE5712FE" bold="true" box="[1111,1130,1915,1939]" pageId="5" pageNumber="139">F.</emphasis>
Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D).
</paragraph>
</caption>
<caption id="DF66667D2141F25BFF29FA72A8851338" ID-DOI="http://doi.org/10.5281/zenodo.10216228" ID-Zenodo-Dep="10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="6" pageNumber="140" startId="6.[151,250,1489,1513]" targetBox="[152,1436,179,1463]" targetPageId="6" targetType="figure">
<paragraph id="8BA636F52141F25BFF29FA72A8851338" blockId="6.[151,1436,1489,1621]" pageId="6" pageNumber="140">
<emphasis id="B96DEAE72141F25BFF29FA72AB2B1084" bold="true" box="[151,278,1489,1513]" pageId="6" pageNumber="140">FIGURE 3.</emphasis>
<taxonomicName id="4C194D762141F25BFEA1FA72ABC21084" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[287,511,1489,1513]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72141F25BFEA1FA72ABC21084" box="[287,511,1489,1513]" italics="true" pageId="6" pageNumber="140">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72141F25BFDB6FA71A8661087" bold="true" box="[520,603,1490,1514]" pageId="6" pageNumber="140">
<taxonomicNameLabel id="A25E579C2141F25BFDB6FA71A8661087" box="[520,603,1490,1514]" pageId="6" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male- STP0269. 
<emphasis id="B96DEAE72141F25BFCC2FA72A9A81084" bold="true" box="[892,917,1489,1513]" pageId="6" pageNumber="140">A.</emphasis>
Head, dorsal view. 
<emphasis id="B96DEAE72141F25BFBD7FA71AEBD1087" bold="true" box="[1129,1152,1490,1514]" pageId="6" pageNumber="140">B.</emphasis>
Pronotum, dorsal view. 
<emphasis id="B96DEAE72141F25BFA3AFA72AFA11084" bold="true" box="[1412,1436,1489,1513]" pageId="6" pageNumber="140">C.</emphasis>
Elytra, dorsal view. 
<emphasis id="B96DEAE72141F25BFEDDFA55AB411363" bold="true" box="[355,380,1526,1550]" pageId="6" pageNumber="140">D.</emphasis>
Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae.
</paragraph>
</caption>
<paragraph id="8BA636F52141F25BFF79F927AB72138B" blockId="6.[151,1437,1667,2018]" pageId="6" pageNumber="140">
Pubescence: Thin and long setae [ts] are the most common and found all over the body (
<figureCitation id="13222A702141F25BFB21F927AF1B13F3" box="[1183,1318,1667,1694]" captionStart="FIGURE 5" captionStartId="8.[151,250,1487,1511]" captionTargetBox="[154,1433,181,1462]" captionTargetId="figure-19@8.[154,1433,181,1462]" captionTargetPageId="8" captionText="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="6" pageNumber="140">Fig. 5B, 5D</figureCitation>
), flattened setae [fs] present on the anterior edge of the head (
<figureCitation id="13222A702141F25BFD77F90BA92313AF" box="[713,798,1703,1730]" captionStart="FIGURE 5" captionStartId="8.[151,250,1487,1511]" captionTargetBox="[154,1433,181,1462]" captionTargetId="figure-19@8.[154,1433,181,1462]" captionTargetPageId="8" captionText="FIGURE 5. Oxarthrius spiculatus sp. nov., paratype male-STP0269. A. metatrochanter. B. Antennal scape. C. Head setae. D. Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="6" pageNumber="140">Fig. 5C</figureCitation>
) and dorsal surface of the abdomen, hollow setae present on the antennae.
</paragraph>
<paragraph id="8BA636F52141F25BFF79F953A87E128F" blockId="6.[151,1437,1667,2018]" pageId="6" pageNumber="140">
<emphasis id="B96DEAE72141F25BFF79F953AB261267" box="[199,283,1776,1802]" italics="true" pageId="6" pageNumber="140">Female</emphasis>
. Similar to male, except for: mesofemora (
<figureCitation id="13222A702141F25BFCB2F953A95B1267" box="[780,870,1776,1802]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="6" pageNumber="140">Fig. 1D</figureCitation>
), without spine and median crest; metatrochanter without any spine (
<figureCitation id="13222A702141F25BFED0F8B7ABFB1243" box="[366,454,1812,1838]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="6" pageNumber="140">Fig. 1D</figureCitation>
); metaventrite with median longitudinal sulcus (
<figureCitation id="13222A702141F25BFC59F8B7AE071243" box="[999,1082,1812,1838]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="6" pageNumber="140">Fig. 2F</figureCitation>
) instead of excavation as in the 
<emphasis id="B96DEAE72141F25BFF29F89BAAF0123F" box="[151,205,1848,1874]" italics="true" pageId="6" pageNumber="140">male</emphasis>
(
<figureCitation id="13222A702141F25BFF60F89BAB00123F" box="[222,317,1848,1874]" captionStart="FIGURE 1" captionStartId="3.[151,250,1685,1709]" captionTargetBox="[151,1435,164,1659]" captionTargetId="figure-20@3.[151,1435,164,1659]" captionTargetPageId="3" captionText="FIGURE 1. Oxarthrius spiculatus sp. nov. (A, C. Male holotype – STP0599; B. D. Female paratype – STP0267) A. Habitus, dorsal view. B. Habitus, dorsal view. C. Metaventrite. D. Metaventrite. Scale bars = 1 mm (A, B); 0.25 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.10216222" httpUri="https://zenodo.org/record/10216222/files/figure.png" pageId="6" pageNumber="140">Figs 1D</figureCitation>
, 
<figureCitation id="13222A702141F25BFEF5F89BAB53123F" box="[331,366,1848,1874]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype – STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="6" pageNumber="140">2E</figureCitation>
). Tergite VIII (
<figureCitation id="13222A702141F25BFDA5F89BA84F123F" box="[539,626,1848,1874]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="6" pageNumber="140">Fig. 4E</figureCitation>
) trapezoidal, straight on anterior margin, rounded on posterior margin. Sternite VIII (
<figureCitation id="13222A702141F25BFE8CF8FFABB4121B" box="[306,393,1884,1910]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="6" pageNumber="140">Fig. 4G</figureCitation>
) large, rounded, slightly sinuous on posterior margin, no keel, no depression. Genital complex (
<figureCitation id="13222A702141F25BFF21F823AAC412F7" box="[159,249,1920,1946]" captionStart="FIGURE 4" captionStartId="7.[151,250,1848,1872]" captionTargetBox="[152,1435,180,1821]" captionTargetId="figure-20@7.[151,1436,179,1822]" captionTargetPageId="7" captionText="FIGURE 4. Oxarthrius spiculatus sp. nov. (A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) A. Aedeagus, ventral view. B. Aedeagus, lateral view. C. Aedeagus, dorsal view. D. Tergite VIII. E. Tergite VIII. F. Sternite VIII. G. Sternite VIII. H. Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection." figureDoi="http://doi.org/10.5281/zenodo.10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="6" pageNumber="140">Fig. 4H</figureCitation>
) trapezoidal, weak sclerotized; posterior margin rounded, projecting laterally, narrowed; anterior margin membranous; median region with one evident transversal curved carina, centrally with one pair of transversal sclerites, slightly elongated anteriorly.
</paragraph>
<caption id="DF66667D2140F25AFF29F89BAB9412D1" ID-DOI="http://doi.org/10.5281/zenodo.10216234" ID-Zenodo-Dep="10216234" httpUri="https://zenodo.org/record/10216234/files/figure.png" pageId="7" pageNumber="141" startId="7.[151,250,1848,1872]" targetBox="[152,1435,180,1821]" targetPageId="7" targetType="figure">
<paragraph id="8BA636F52140F25AFF29F89BAB9412D1" blockId="7.[151,1437,1848,1980]" pageId="7" pageNumber="141">
<emphasis id="B96DEAE72140F25AFF29F89BAB2B123D" bold="true" box="[151,278,1848,1872]" pageId="7" pageNumber="141">FIGURE 4.</emphasis>
<taxonomicName id="4C194D762140F25AFEA1F89BABC3123D" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[287,510,1848,1872]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="141" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72140F25AFEA1F89BABC3123D" box="[287,510,1848,1872]" italics="true" pageId="7" pageNumber="141">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72140F25AFDB9F89BA867123D" bold="true" box="[519,602,1848,1872]" pageId="7" pageNumber="141">
<taxonomicNameLabel id="A25E579C2140F25AFDB9F89BA867123D" box="[519,602,1848,1872]" pageId="7" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A–D, F. Male holotype – STP0599; E, G, H. Female paratype - STP0273) 
<emphasis id="B96DEAE72140F25AFA3AF89BAFA1123D" bold="true" box="[1412,1436,1848,1872]" pageId="7" pageNumber="141">A.</emphasis>
Aedeagus, ventral view. 
<emphasis id="B96DEAE72140F25AFE2CF8FFAB941219" bold="true" box="[402,425,1884,1908]" pageId="7" pageNumber="141">B.</emphasis>
Aedeagus, lateral view. 
<emphasis id="B96DEAE72140F25AFD1CF8FFA8861219" bold="true" box="[674,699,1884,1908]" pageId="7" pageNumber="141">C.</emphasis>
Aedeagus, dorsal view. 
<emphasis id="B96DEAE72140F25AFC0DF8FFA9F61219" bold="true" box="[947,971,1884,1908]" pageId="7" pageNumber="141">D.</emphasis>
Tergite VIII. 
<emphasis id="B96DEAE72140F25AFBE9F8FFAE521219" bold="true" box="[1111,1135,1884,1908]" pageId="7" pageNumber="141">E.</emphasis>
Tergite VIII. 
<emphasis id="B96DEAE72140F25AFB45F8FFAF331219" bold="true" box="[1275,1294,1884,1908]" pageId="7" pageNumber="141">F.</emphasis>
Sternite VIII. 
<emphasis id="B96DEAE72140F25AFF29F823AA8C12F5" bold="true" box="[151,177,1920,1944]" pageId="7" pageNumber="141">G.</emphasis>
Sternite VIII. 
<emphasis id="B96DEAE72140F25AFEF9F823AB5F12F5" bold="true" box="[327,354,1920,1944]" pageId="7" pageNumber="141">H.</emphasis>
Genital complex. Scale bars = 0.1 mm (A–C, H); 0.25 mm (D–G). Abbreviations: cp, central plate; mpp, median pointed projection.
</paragraph>
</caption>
<caption id="DF66667D214FF255FF29FA6CABDC1342" ID-DOI="http://doi.org/10.5281/zenodo.10216236" ID-Zenodo-Dep="10216236" httpUri="https://zenodo.org/record/10216236/files/figure.png" pageId="8" pageNumber="142" startId="8.[151,250,1487,1511]" targetBox="[154,1433,181,1462]" targetPageId="8" targetType="figure">
<paragraph id="8BA636F5214FF255FF29FA6CABDC1342" blockId="8.[151,1437,1487,1583]" pageId="8" pageNumber="142">
<emphasis id="B96DEAE7214FF255FF29FA6CAB28108A" bold="true" box="[151,277,1487,1511]" pageId="8" pageNumber="142">FIGURE 5.</emphasis>
<taxonomicName id="4C194D76214FF255FEA0FA6CABC3108A" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[286,510,1487,1511]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="142" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE7214FF255FEA0FA6CABC3108A" box="[286,510,1487,1511]" italics="true" pageId="8" pageNumber="142">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE7214FF255FDB8FA6CA864108A" bold="true" box="[518,601,1487,1511]" pageId="8" pageNumber="142">
<taxonomicNameLabel id="A25E579C214FF255FDB8FA6CA864108A" box="[518,601,1487,1511]" pageId="8" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male-STP0269. 
<emphasis id="B96DEAE7214FF255FCCEFA6CA9B4108A" bold="true" box="[880,905,1487,1511]" pageId="8" pageNumber="142">A.</emphasis>
metatrochanter. 
<emphasis id="B96DEAE7214FF255FB89FA6CAE72108A" bold="true" box="[1079,1103,1487,1511]" pageId="8" pageNumber="142">B.</emphasis>
Antennal scape. 
<emphasis id="B96DEAE7214FF255FABDFA6CAF21108A" bold="true" box="[1283,1308,1487,1511]" pageId="8" pageNumber="142">C.</emphasis>
Head setae. 
<emphasis id="B96DEAE7214FF255FF29FA50AA8D1366" bold="true" box="[151,176,1523,1547]" pageId="8" pageNumber="142">D.</emphasis>
Scutelum. Scale bars = 50 μm (A, D); 20 μm (B, C). Abbreviations: ts, thin setae; fs, flattened setae; iblf, inner basolateral foveae; bef, basal elytral foveae.
</paragraph>
</caption>
<paragraph id="8BA636F5214FF255FF79F9FEA80E1246" blockId="8.[151,1437,1629,2016]" pageId="8" pageNumber="142">
<emphasis id="B96DEAE7214FF255FF79F9FEA87F131A" bold="true" box="[199,578,1629,1655]" pageId="8" pageNumber="142">Study area and sampling notes.</emphasis>
All specimens of 
<taxonomicName id="4C194D76214FF255FCAFF9FDA99A131A" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[785,935,1629,1656]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="142" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214FF255FCAFF9FDA99A131A" box="[785,935,1629,1656]" italics="true" pageId="8" pageNumber="142">O. spiculatus</emphasis>
</taxonomicName>
were collected in the area of the Mineração Paragominas S.A. Company of the Norsk Hydro ASA, located approximately 
<quantity id="4CE19B10214FF255FB93F922AE4513F6" box="[1069,1144,1665,1691]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="142" unit="km" value="70.0">70 km</quantity>
(coordinates: 3˚15’53”S, 47˚423’06”W) from the municipal headquarters of Paragominas, state of 
<collectingRegion id="49DDF817214FF255FC59F906AE2113D2" box="[999,1052,1701,1727]" country="Brazil" name="Para" pageId="8" pageNumber="142">Pará</collectingRegion>
, in the eastern Brazilian Amazon (
<figureCitation id="13222A70214FF255FF21F96AAADB138E" box="[159,230,1737,1763]" captionStart="FIGURE 6" captionStartId="9.[151,250,1457,1481]" captionTargetBox="[167,1426,529,1424]" captionTargetId="figure-146@9.[154,1434,522,1430]" captionTargetPageId="9" captionText="FIGURE 6. Bauxite mining area, southwestern portion of Paragominas Municipality, Pará, Brazil. (Map data: Google © 2023 CNES/Airbus, Maxar Technologies)" figureDoi="http://doi.org/10.5281/zenodo.10216238" httpUri="https://zenodo.org/record/10216238/files/figure.png" pageId="8" pageNumber="142">Fig. 6</figureCitation>
). Paragominas represents one of the largest bauxite reserves of the World, having estimated reserves of 1 billion tonnes (
<bibRefCitation id="EF884B04214FF255FEFFF94EA868126A" author="Veredict Research" box="[321,597,1773,1799]" pageId="8" pageNumber="142" refId="ref10562" refString="Veredict Research. (2008) Mining Technology. Paragominas Bauxite Mine - Brazil. Available from: https: // www. miningtechnology. com / projects / paragominas / (accessed 29 Jul. 2023)" type="url" year="2008">Veredict Research 2008</bibRefCitation>
). Its main soils are Yellow Latosols, Yellow Argisols, Plinthsols, Gleysols and Neosols (
<bibRefCitation id="EF884B04214FF255FE91F8B2A81A1246" author="Rodrigues, T. E. &amp; Silva, R. C. &amp; Silva, J. M. L. &amp; Oliveira Junior, R. C. &amp; Gama, J. R. N. &amp; Valente, M. A." box="[303,551,1809,1836]" pageId="8" pageNumber="142" refId="ref10233" refString="Rodrigues, T. E., Silva, R. C., Silva, J. M. L., Oliveira Junior, R. C., Gama, J. R. N. &amp; Valente, M. A. (2003) Caracterizacao e classificacao dos solos do municipio de Paragominas, Estado do Para. Embrapa Amazonia Oriental, Belem." type="book" year="2003">
Rodrigues 
<emphasis id="B96DEAE7214FF255FE15F8B1ABE11246" box="[427,476,1809,1835]" italics="true" pageId="8" pageNumber="142">et al</emphasis>
. 2003
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636F5214FF254FF79F895AADD1594" blockId="8.[151,1437,1629,2016]" lastBlockId="9.[151,1437,150,465]" lastPageId="9" lastPageNumber="143" pageId="8" pageNumber="142">
In this region the climate is tropical wet and dry (Aw 
<typeStatus id="54A28857214FF255FC87F895A951123D" box="[825,876,1846,1872]" pageId="8" pageNumber="142">type</typeStatus>
, Köppen-Geiger’s Climate classification), with a pronounced dry season occurring from June to November besides a wetter season (from February to May) and a transition season (from December to January) (
<bibRefCitation id="EF884B04214FF255FD0DF8DEA97E12F5" author="Bastos, T. X. &amp; Pacheco, N. A. &amp; Figueiredo, R. D. O. &amp; Silva, G." box="[691,835,1917,1944]" pageId="8" pageNumber="142" refId="ref8392" refString="Bastos, T. X., Pacheco, N. A., Figueiredo, R. D. O. &amp; Silva, G. (2005) Caracteristicas agroclimaticas do municipio de Paragominas. Embrapa." type="book" year="2005">Bastos 2005</bibRefCitation>
; 
<bibRefCitation id="EF884B04214FF255FCEFF8DEAE0312FA" author="Dubreuil, V. &amp; Fante, K. P. &amp; Planchon, O. &amp; Neto, J. L. S. A." box="[849,1086,1917,1944]" pageId="8" pageNumber="142" refId="ref8750" refString="Dubreuil, V., Fante, K. P., Planchon, O. &amp; Neto, J. L. S. A. (2018) Os tipos de climas anuais no Brasil: uma aplicacao da classificacao de Koppen de 1961 a 2015. Confins. Revista franco-brasilera de geografia, (37). https: // doi. org / 10.4000 / confins. 15738" type="book" year="2018">
Dubreuil 
<emphasis id="B96DEAE7214FF255FC00F8DDA9CC12FA" box="[958,1009,1917,1943]" italics="true" pageId="8" pageNumber="142">et al</emphasis>
. 2018
</bibRefCitation>
). The precipitation varies a lot throughout the year, ranging from 
<quantity id="4CE19B10214FF255FDA4F802A85412D1" box="[538,617,1953,1980]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="142" unit="mm" value="28.0">28 mm</quantity>
(August and September) to 
<quantity id="4CE19B10214FF255FC1BF802AE3F12D1" box="[933,1026,1953,1980]" metricMagnitude="-1" metricUnit="m" metricValue="3.57" pageId="8" pageNumber="142" unit="mm" value="357.0">357 mm</quantity>
(March; average annual 
<quantity id="4CE19B10214FF255FAA4F802AFAD12D6" box="[1306,1424,1953,1979]" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="142" unit="mm" value="1800.0">1,800 mm</quantity>
), whereas relative humidity (76% to 85%; annual average 81%) and air temperature (25.6°C to 27°C; annual average 26.3°C) are relatively stable throughout the year (
<bibRefCitation id="EF884B04214EF254FD60FF34A94F15DC" author="Bastos, T. X. &amp; Pacheco, N. A. &amp; Figueiredo, R. D. O. &amp; Silva, G." box="[734,882,150,177]" pageId="9" pageNumber="143" refId="ref8392" refString="Bastos, T. X., Pacheco, N. A., Figueiredo, R. D. O. &amp; Silva, G. (2005) Caracteristicas agroclimaticas do municipio de Paragominas. Embrapa." type="book" year="2005">Bastos 2005</bibRefCitation>
). The predominant vegetation is dense Amazon Rainforest showing different levels of degradation and deforestation, resulting in a landscape mosaic (
<bibRefCitation id="EF884B04214EF254FAB4FF18AAEE1594" author="Uhl, C. &amp; Vieira, I. C. G." pageId="9" pageNumber="143" pagination="98 - 106" refId="ref10505" refString="Uhl, C. &amp; Vieira, I. C. G. (1989) Ecological impacts of selective logging in the Brazilian Amazon: a case study from the Paragominas region of the state of Para. Biotropica, 21, 98 - 106. https: // doi. org / 10.2307 / 2388700" type="journal article" year="1989">Uhl &amp; Vieira 1989</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636F5214EF254FF79FEA0A9D914BC" blockId="9.[151,1437,150,465]" pageId="9" pageNumber="143">
In the study area, the mining activity due to bauxite extraction has changed the landscape due to total vegetation suppression through total topsoil and soil removal. The bauxite mined areas are later reforested with native species according to the Degraded Area Rehabilitation Plan (PRAD) which incorporates three different systems of rehabilitation implemented by the Mineração Paragominas: “nucleação”, spontaneous natural regeneration and planted forest species (
<bibRefCitation id="EF884B04214EF254FE20FE30A8BD14C0" author="Martins, W. B. R. &amp; Ferreira, G. C. &amp; de Assis Oliveira, F." box="[414,640,403,429]" pageId="9" pageNumber="143" pagination="37 - 48" refId="ref9231" refString="Martins, W. B. R., Ferreira, G. C. &amp; de Assis Oliveira, F. (2018) Deposicao de serapilheira e nutrientes em areas de mineracao submetidas a metodos de restauracao florestal em Paragominas, Para. Floresta, 48 (1), 37 - 48. https: // doi. org / 10.5380 / rf. v 48 i 1.49288" type="journal article" year="2018">
Martins 
<emphasis id="B96DEAE7214EF254FDBFFE30A80814C0" box="[513,565,403,429]" italics="true" pageId="9" pageNumber="143">et al</emphasis>
. 2018
</bibRefCitation>
; 
<bibRefCitation id="EF884B04214EF254FD30FE30A9BB14C0" author="Cerqueira, R. M. &amp; Jardim, M. A. G. &amp; Silva Junior, L. L. M. &amp; Paixao, L. P. &amp; Martins, M. B." box="[654,902,403,429]" pageId="9" pageNumber="143" pagination="22 - 41" refId="ref8527" refString="Cerqueira, R. M., Jardim, M. A. G., Silva Junior, L. L. M., Paixao, L. P. &amp; Martins, M. B. (2021) Fitossociologia do estrato arboreo em floresta nativa e em areas do programa de recuperacao de areas degradadas sob influencia da mineracao, Paragominas, Para, Brasil. Nature and Conservation, 14 (3), 22 - 41. https: // doi. org / 10.6008 / CBPC 2318 - 2881.2021.003.0002" type="journal article" year="2021">
Cerqueira 
<emphasis id="B96DEAE7214EF254FCB8FE30A90714C0" box="[774,826,403,429]" italics="true" pageId="9" pageNumber="143">et al</emphasis>
. 2021
</bibRefCitation>
). Therefore, the landscape of the study area is mainly composed of Forest, bauxite mining areas and PRAD areas (
<figureCitation id="13222A70214EF254FC2FFE14A9E814BC" box="[913,981,439,465]" captionStart="FIGURE 6" captionStartId="9.[151,250,1457,1481]" captionTargetBox="[167,1426,529,1424]" captionTargetId="figure-146@9.[154,1434,522,1430]" captionTargetPageId="9" captionText="FIGURE 6. Bauxite mining area, southwestern portion of Paragominas Municipality, Pará, Brazil. (Map data: Google © 2023 CNES/Airbus, Maxar Technologies)" figureDoi="http://doi.org/10.5281/zenodo.10216238" httpUri="https://zenodo.org/record/10216238/files/figure.png" pageId="9" pageNumber="143">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF66667D214EF254FF29FA12A8351080" ID-DOI="http://doi.org/10.5281/zenodo.10216238" ID-Zenodo-Dep="10216238" httpUri="https://zenodo.org/record/10216238/files/figure.png" pageId="9" pageNumber="143" startId="9.[151,250,1457,1481]" targetBox="[167,1426,529,1424]" targetPageId="9" targetType="figure">
<paragraph id="8BA636F5214EF254FF29FA12A8351080" blockId="9.[151,1437,1457,1517]" pageId="9" pageNumber="143">
<emphasis id="B96DEAE7214EF254FF29FA12AB2E10A4" bold="true" box="[151,275,1457,1481]" pageId="9" pageNumber="143">FIGURE 6.</emphasis>
Bauxite mining area, southwestern portion of Paragominas Municipality, Pará, Brazil. (Map data: Google © 2023 CNES/Airbus, Maxar Technologies)
</paragraph>
</caption>
<paragraph id="8BA636F5214EF254FF79F9B8A92113A8" blockId="9.[151,1437,1563,2022]" pageId="9" pageNumber="143">
The bauxite mining areas were not sampled. The field collections occurred between 2017 and 2019: 
<date id="FFA71035214EF254FA98F9B8AB5B1337" pageId="9" pageNumber="143" value="2017-11" valueMax="2017-12-31" valueMin="2017-11-00">November to December 2017</date>
(dry season to transition season), 
<date id="FFA71035214EF254FD5DF9E3A96B1337" box="[739,854,1600,1626]" pageId="9" pageNumber="143" value="2018-05">May 2018</date>
(wet season), 
<date id="FFA71035214EF254FC49F9E3AE541337" box="[1015,1129,1600,1626]" pageId="9" pageNumber="143" value="2018-06">June 2018</date>
(dry season), and 
<date id="FFA71035214EF254FA89F99CAAF21313" pageId="9" pageNumber="143" value="2019-02">February 2019</date>
(wet season). The sampling points were randomly scattered in the study area to represent its environmental variability, with about 60% of the points in forest. In the PRAD areas, sampling points were spread out close to the edge of the forest. The sampling unit was not established.
</paragraph>
<paragraph id="8BA636F5214EF257FF79F96CA8141594" blockId="9.[151,1437,1563,2022]" lastBlockId="10.[151,1436,150,357]" lastPageId="10" lastPageNumber="144" pageId="9" pageNumber="143">
All specimens of 
<taxonomicName id="4C194D76214EF254FE2FF96CA8311384" authorityName="Reitter" authorityYear="1883" box="[401,524,1743,1769]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214EF254FE2FF96CA8311384" box="[401,524,1743,1769]" italics="true" pageId="9" pageNumber="143">Oxarthrius</emphasis>
</taxonomicName>
collected belong to 
<taxonomicName id="4C194D76214EF254FD4BF973A9B71384" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[757,906,1743,1770]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="143" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214EF254FD4BF973A9B71384" box="[757,906,1743,1770]" italics="true" pageId="9" pageNumber="143">O. spiculatus</emphasis>
</taxonomicName>
. They were recorded exclusively in the forest, at elevations from 
<quantity id="4CE19B10214EF254FECEF957AB8D1263" box="[368,432,1780,1806]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="143" unit="m" value="100.0">100m</quantity>
to 
<quantity id="4CE19B10214EF254FE67F957A8211263" box="[473,540,1780,1806]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="143" unit="m" value="140.0">140m</quantity>
, between the end of the wet season (
<date id="FFA71035214EF254FC77F957AE7F1263" box="[969,1090,1780,1806]" pageId="9" pageNumber="143" value="2018-05">May 2018</date>
) and the beginning of the dry season (
<date id="FFA71035214EF254FF4CF8BBAB50125F" box="[242,365,1816,1842]" pageId="9" pageNumber="143" value="2018-06">June 2018</date>
). Different sampling methods were used for the collections: Classical Light Trap (white cloth, with mercury-vapor or blacklight lamps), Winkler extractor, Yellow Pan Trap (or Moericke), Butterfly Sweep Net, Sweep Net, Quadrate Beating Sheet or white Umbrella, and active search under decaying logs, leaves, flowers and logs. However, all specimens of 
<taxonomicName id="4C194D76214EF254FDBCF827A8AB12F0" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[514,662,1923,1950]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="143" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214EF254FDBCF827A8AB12F0" box="[514,662,1923,1950]" italics="true" pageId="9" pageNumber="143">O. spiculatus</emphasis>
</taxonomicName>
were gathered exclusively from Winkler extractor samples. Here, the Winkler samples consisted of litter with some part of the topsoil. Each Winkler litter sample were put in coarsemesh bags suspended inside a large closed cloth bag suspended over a collection 100 ml bottle containing 75 ml of 80% ethanol, and left to dry at controlled temperature of 25ºc for three days. We believe that 
<taxonomicName id="4C194D76214DF257FB72FF34AF5C15DC" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[1228,1377,151,177]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="144" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214DF257FB72FF34AF5C15DC" box="[1228,1377,151,177]" italics="true" pageId="10" pageNumber="144">O. spiculatus</emphasis>
</taxonomicName>
lives as a ground-dwelling rove beetle living in the leaf litter of the Amazon forest. It seems to be unable to colonize the local PRAD areas during this study.
</paragraph>
</subSubSection>
<subSubSection id="C303657E214DF257FF79FEA1A8CC1470" box="[199,753,258,285]" pageId="10" pageNumber="144" type="distribution">
<paragraph id="8BA636F5214DF257FF79FEA1A8CC1470" blockId="10.[151,1436,150,357]" box="[199,753,258,285]" pageId="10" pageNumber="144">
<emphasis id="B96DEAE7214DF257FF79FEA1AB5C1471" bold="true" box="[199,353,258,284]" pageId="10" pageNumber="144">Distribution.</emphasis>
Only known from the 
<typeStatus id="54A28857214DF257FDDEFEA0A8AD1470" box="[608,656,259,285]" pageId="10" pageNumber="144">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C303657E214DF257FF79FE84A8EC1408" pageId="10" pageNumber="144" type="etymology">
<paragraph id="8BA636F5214DF257FF79FE84A8EC1408" blockId="10.[151,1436,150,357]" pageId="10" pageNumber="144">
<emphasis id="B96DEAE7214DF257FF79FE84AB73142C" bold="true" box="[199,334,295,321]" pageId="10" pageNumber="144">Etymology.</emphasis>
The specific epithet refers to the Latin term 
<taxonomicName id="4C194D76214DF257FCEAFE84A9FB142C" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[852,966,295,321]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="144" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214DF257FCEAFE84A9FB142C" box="[852,966,295,321]" italics="true" pageId="10" pageNumber="144">spiculatus</emphasis>
</taxonomicName>
, meaning equipped with small spines, in reference to the spines on the male metatrochanter.
</paragraph>
</subSubSection>
</treatment>
</document>