269 lines
40 KiB
XML
269 lines
40 KiB
XML
<document id="A066BFF98953C2A3B398235092071FE5" ID-CLB-Dataset="22330" ID-DOI="10.11646/zootaxa.4772.1.5" ID-GBIF-Dataset="b14b0d0d-cf67-4895-965c-12792c622b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3814079" ID-ZooBank="B497198A-08CD-4A21-AE04-14390499853B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834845818" checkinUser="plazi" docAuthor="Polhemus, Dan A." docDate="2020" docId="03FE00115B61FFCEFF38FF51FEA5099A" docLanguage="en" docName="zootaxa.4772.1.5.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enithares kasim Polhemus 2020, new species" docType="treatment" docVersion="8" lastPageNumber="160" masterDocId="FFC778695B7DFFD2FFAFFFC8FFF90E21" masterDocTitle="Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist" masterLastPageNumber="182" masterPageNumber="132" pageNumber="160" updateTime="1698838866946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="182484D22342F3648D9C8AB251935772" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FC4AA010F40385B3BCD41D121D82F4E6">
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<mods:title id="BE3E20EA7D242D21F9CF79C1D89DBF1B">Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist</mods:title>
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</mods:titleInfo>
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<mods:name id="65A944E2AB077271B94327FDFBCC8629" type="personal">
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<mods:role id="1C226C7A020FE92554026ABF5797A317">
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<mods:roleTerm id="659AA60EC5ED6CE657758DEAF8723F0B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F244E86C2A4E979BDAB58D97C5B47EFA">Polhemus, Dan A.</mods:namePart>
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<mods:typeOfResource id="2FAEF6EA97351B5E41DA9C64E22F8F11">text</mods:typeOfResource>
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<mods:titleInfo id="098800970D50BB42B6F4BF94FE4C51C6">
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<mods:title id="7A924ED42FF8AF3F4D0EC304AA2ECE38">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="05CA6EE389103065D622783B54B93340">
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<mods:date id="49DC6D258D2A5C3564D3C171EB85BB89">2020</mods:date>
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<mods:detail id="8F5148E06E8AAFA7EDF8F5A169CF824E" type="pubDate">
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<mods:number id="D15E730D504700C0414FFF959AC5A541">2020-05-07</mods:number>
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</mods:detail>
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<mods:detail id="81CA0E4BF56EA60359335C983AD32145" type="volume">
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<mods:number id="F8FDF3F5D3E02B35AA06EDEBF02A1D7D">4772</mods:number>
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</mods:detail>
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<mods:detail id="CC9832B3FDD0BD04B73A0479921F866E" type="issue">
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<mods:number id="78343324080A47843953CA84DE159137">1</mods:number>
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<mods:start id="1EDDB26CFB3CE1A1FAA5E3F08C29B011">132</mods:start>
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<mods:classification id="18C905188C1D8FC95CD9A0A8169E963A">journal article</mods:classification>
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<mods:identifier id="92A14629A41869C37D1A4E4D5D6E88CE" type="CLB-Dataset">22330</mods:identifier>
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<mods:identifier id="892B8023693E45BBA59F73F506CA8988" type="DOI">10.11646/zootaxa.4772.1.5</mods:identifier>
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<mods:identifier id="AEC08375506B3B17441071995162C477" type="GBIF-Dataset">b14b0d0d-cf67-4895-965c-12792c622b21</mods:identifier>
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<mods:identifier id="1ADAA0AEC3D911C152FB296B39D65AA4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="164FB2958223EBD44B528A7317B335B3" type="Zenodo-Dep">3814079</mods:identifier>
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<mods:identifier id="626841C05A7384AD17D82FD62AB0BB67" type="ZooBank">B497198A-08CD-4A21-AE04-14390499853B</mods:identifier>
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</mods:mods>
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<treatment id="03FE00115B61FFCEFF38FF51FEA5099A" ID-DOI="http://doi.org/10.5281/zenodo.3815538" ID-GBIF-Taxon="163975561" ID-Zenodo-Dep="3815538" LSID="urn:lsid:plazi:treatment:03FE00115B61FFCEFF38FF51FEA5099A" httpUri="http://treatment.plazi.org/id/03FE00115B61FFCEFF38FF51FEA5099A" lastPageNumber="160" pageId="28" pageNumber="160">
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<subSubSection id="C34DE28C5B61FFCEFF38FF51FE0D0E92" box="[151,500,153,180]" pageId="28" pageNumber="160" type="nomenclature">
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<paragraph id="8BE8B1075B61FFCEFF38FF51FE0D0E92" blockId="28.[151,500,153,216]" box="[151,500,153,180]" pageId="28" pageNumber="160">
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<heading id="D0A0066B5B61FFCEFF38FF51FE0D0E92" bold="true" box="[151,500,153,180]" fontSize="11" level="1" pageId="28" pageNumber="160" reason="1">
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<emphasis id="B9236D155B61FFCEFF38FF51FE0D0E92" bold="true" box="[151,500,153,180]" pageId="28" pageNumber="160">
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<taxonomicName id="4C57CA845B61FFCEFF38FF51FEA70E95" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[151,350,153,180]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim" status="sp. nov.">
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<emphasis id="B9236D155B61FFCEFF38FF51FEA70E95" bold="true" box="[151,350,153,180]" italics="true" pageId="28" pageNumber="160">Enithares kasim</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A210D06E5B61FFCEFECAFF52FE0D0E92" box="[357,500,153,180]" pageId="28" pageNumber="160" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34DE28C5B61FFCEFF38FF75FE670EF9" box="[151,414,189,216]" pageId="28" pageNumber="160" type="description">
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||
<paragraph id="8BE8B1075B61FFCEFF38FF75FE670EF9" blockId="28.[151,500,153,216]" box="[151,414,189,216]" pageId="28" pageNumber="160">
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||
(
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<figureCitation id="136CAD825B61FFCEFF30FF75FF160EF9" box="[159,239,189,216]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1014,1039]" captionTargetBox="[151,1436,159,999]" captionTargetId="figure@3.[151,1436,159,1000]" captionTargetPageId="3" captionText="FIGURES 2–7. Enithares species, metaxiphus. 2. E. arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 3. E. bosavi n. sp.; holotype specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 4. E. insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 5. E. kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 6. E. orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Reinduk, CL 7036. 7. E. papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157." figureDoi="http://doi.org/10.5281/zenodo.3814083" httpUri="https://zenodo.org/record/3814083/files/figure.png" pageId="28" pageNumber="160">Figs. 5</figureCitation>
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,
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<figureCitation id="136CAD825B61FFCEFF54FF75FEBD0EF6" box="[251,324,189,216]" captionStart="FIGURES 41–48" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 41–48. Enithares species in the E. megalops group, dorsal habitus photographs. 41–42. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 43–44. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 45–46. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 47–48. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="28" pageNumber="160">45, 46</figureCitation>
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,
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<figureCitation id="136CAD825B61FFCEFEE0FF75FE940EF9" box="[335,365,189,216]" captionStart="FIGURES 49–54" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 49–54. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="28" pageNumber="160">52</figureCitation>
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,
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<figureCitation id="136CAD825B61FFCEFED6FF75FE6F0EF9" box="[377,406,189,216]" captionStart="FIGURE 67" captionStartId="38.[151,250,1916,1941]" captionTargetBox="[158,1430,1097,1882]" captionTargetId="figure@38.[151,1436,1090,1890]" captionTargetPageId="38" captionText="FIGURE 67. Distribution of species in the E. megalops group on New Guinea. Square = E. kasim n. sp.; circles = E. nigra Lansbury; triangles = E. alexis Lansbury; pentagons = E. vulgaris Lansbury." figureDoi="http://doi.org/10.5281/zenodo.3814901" httpUri="https://zenodo.org/record/3814901/files/figure.png" pageId="28" pageNumber="160">67</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34DE28C5B61FFCEFF38FECEFEA70F91" pageId="28" pageNumber="160" type="materials_examined">
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<paragraph id="8BE8B1075B61FFCEFF38FECEFEA70F91" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
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||
<emphasis id="B9236D155B61FFCEFF38FECEFE4C0F01" bold="true" box="[151,437,261,288]" pageId="28" pageNumber="160">Type material examined</emphasis>
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.
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<materialsCitation id="3B3FBB5A5B61FFCEFE6DFECDFBD40F46" ID-GBIF-Occurrence="2610672435" collectingDate="1991-10-02" collectionCode="A, BPBM" collectorName="Polhemus & J. T. Polhemus" country="Indonesia" elevation="15" latitude="-1.2547222" location="New" longLatPrecision="21" longitude="131.23639" pageId="28" pageNumber="160" specimenCount="15" stateProvince="Papua Barat" typeStatus="holotype">
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<typeStatus id="54EC0FA55B61FFCEFE6DFECDFDD40F3E" box="[450,557,261,287]" pageId="28" pageNumber="160" type="holotype">Holotype</typeStatus>
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, male (dissected):
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<collectingCountry id="F340F1975B61FFCEFD54FECDFC6F0F3E" box="[763,918,261,287]" name="Indonesia" pageId="28" pageNumber="160">INDONESIA</collectingCountry>
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||
,
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<collectingRegion id="49937FE55B61FFCEFC0FFECDFB990F01" box="[928,1120,261,288]" country="Indonesia" name="Papua Barat" pageId="28" pageNumber="160">
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||
<emphasis id="B9236D155B61FFCEFC0FFECDFBA00F01" bold="true" box="[928,1113,261,288]" pageId="28" pageNumber="160">Irian Jaya Prov</emphasis>
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||
.
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||
</collectingRegion>
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||
[
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<collectingRegion id="49937FE55B61FFCEFBC3FECEFB000F01" box="[1132,1273,261,288]" country="Indonesia" name="Papua Barat" pageId="28" pageNumber="160">Papua Prov.</collectingRegion>
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||
],
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||
<location id="8E88E7DC5B61FFCEFAA5FECEFAC60F01" LSID="urn:lsid:plazi:treatment:03FE00115B61FFCEFF38FF51FEA5099A:8E88E7DC5B61FFCEFAA5FECEFAC60F01" box="[1290,1343,262,288]" country="Indonesia" latitude="-1.2547222" longLatPrecision="21" longitude="131.23639" name="New" pageId="28" pageNumber="160" stateProvince="Papua Barat">New</location>
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Guinea, pools in muddy lowland rain forest streambed,
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<quantity id="4CAF1CE25B61FFCEFD10FEE1FCF50F62" box="[703,780,297,323]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="28" pageNumber="160" unit="km" value="28.0">28 km</quantity>
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.
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<locationDeviation id="9A5AA8255B61FFCEFCB5FEE2FC4D0F62" box="[794,948,297,324]" location="Kasim" pageId="28" pageNumber="160">NE of Kasim</locationDeviation>
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on pipeline road,
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<specimenCount id="9D517A8E5B61FFCEFB23FEE1FB280F65" box="[1164,1233,297,324]" pageId="28" pageNumber="160" type="generic">
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||
<quantity id="4CAF1CE25B61FFCEFB23FEE1FB340F65" box="[1164,1229,297,324]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="160" unit="m" value="15.0">
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||
<elevation id="007A56345B61FFCEFB23FEE1FB340F65" box="[1164,1229,297,324]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="160" unit="m" value="15.0">15 m</elevation>
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</quantity>
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.
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</specimenCount>
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,
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<date id="FFE997C75B61FFCEFB4FFEE1FA6E0F62" box="[1248,1431,297,323]" pageId="28" pageNumber="160" value="1991-10-02">
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<collectingDate id="EFAD6E2F5B61FFCEFB4FFEE1FA6E0F62" box="[1248,1431,297,323]" pageId="28" pageNumber="160" value="1991-10-02">2 October 1991</collectingDate>
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</date>
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,
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<geoCoordinate id="EE63D7C05B61FFCEFF38FE85FEE20F49" box="[151,283,333,360]" degrees="01" direction="south" minutes="15" orientation="latitude" pageId="28" pageNumber="160" precision="15" seconds="17" value="-1.2547222">01°15′17′′S</geoCoordinate>
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,
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<geoCoordinate id="EE63D7C05B61FFCEFE88FE85FE410F49" box="[295,440,333,360]" degrees="131" direction="east" minutes="14" orientation="longitude" pageId="28" pageNumber="160" precision="15" seconds="11" value="131.23639">131°14′11′′E</geoCoordinate>
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, CL 2630, D.
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<collectionCode id="ED4629C25B61FFCEFDF9FE86FD940F49" box="[598,621,334,360]" pageId="28" pageNumber="160">A</collectionCode>
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||
.
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||
<collectorName id="26A2D4D15B61FFCEFDD7FE85FD1F0F46" box="[632,742,333,359]" pageId="28" pageNumber="160">Polhemus</collectorName>
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||
and
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<collectorName id="26A2D4D15B61FFCEFCB1FE86FC3B0F46" box="[798,962,333,360]" pageId="28" pageNumber="160">J. T. Polhemus</collectorName>
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||
(
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<collectionCode id="ED4629C25B61FFCEFC7EFE85FBDC0F46" box="[977,1061,333,359]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="28" pageNumber="160" type="Museum">BPBM</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B3FBB5A5B61FFCEFB97FE86FBBC0FAD" ID-GBIF-Occurrence="2610672400" collectingDate="1991-10-02" collectionCode="A" collectorName="Polhemus & J. T. Polhemus" country="Indonesia" elevation="15" latitude="-1.2547222" location="New" longLatPrecision="21" longitude="131.23639" pageId="28" pageNumber="160" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
<emphasis id="B9236D155B61FFCEFB97FE86FB530F46" box="[1080,1194,334,359]" italics="true" pageId="28" pageNumber="160">
|
||
<typeStatus id="54EC0FA55B61FFCEFB97FE86FB530F46" box="[1080,1194,334,359]" pageId="28" pageNumber="160" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
:
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<collectingCountry id="F340F1975B61FFCEFB16FE85FAAD0F46" box="[1209,1364,333,359]" name="Indonesia" pageId="28" pageNumber="160">INDONESIA</collectingCountry>
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||
,
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||
<collectingRegion id="49937FE55B61FFCEFACFFE85FEEE0FAD" country="Indonesia" name="Papua Barat" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFACFFE85FEE90FAD" bold="true" pageId="28" pageNumber="160">Irian Jaya Prov</emphasis>
|
||
.
|
||
</collectingRegion>
|
||
[
|
||
<collectingRegion id="49937FE55B61FFCEFE8AFEBAFE4C0FAD" box="[293,437,369,396]" country="Indonesia" name="Papua Barat" pageId="28" pageNumber="160">Papua Prov.</collectingRegion>
|
||
]:
|
||
<specimenCount id="9D517A8E5B61FFCEFE69FEB9FDEC0FAA" box="[454,533,369,395]" pageId="28" pageNumber="160" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D517A8E5B61FFCEFD8EFEB9FD680FAA" box="[545,657,369,396]" pageId="28" pageNumber="160" type="female">5 females</specimenCount>
|
||
, same data as holotype CL 2630, D.
|
||
<collectionCode id="ED4629C25B61FFCEFB81FEBAFBBC0FAD" box="[1070,1093,370,396]" pageId="28" pageNumber="160">A</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3FBB5A5B61FFCEFBFEFEB9FEA30F91" ID-GBIF-Occurrence="2610672560" collectionCode="USNM, BPBM" collectorName="J. T. Polhemus" country="Indonesia" pageId="28" pageNumber="160" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
Polhemus and
|
||
<collectorName id="26A2D4D15B61FFCEFB58FEBAFA650FAA" box="[1271,1436,369,396]" pageId="28" pageNumber="160">J. T. Polhemus</collectorName>
|
||
(
|
||
<collectionCode id="ED4629C25B61FFCEFF30FE5DFF0D0F8E" box="[159,244,405,431]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="160" type="Museum">USNM</collectionCode>
|
||
,
|
||
<collectionCode id="ED4629C25B61FFCEFF50FE5EFEAB0F91" box="[255,338,406,432]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="28" pageNumber="160" type="Museum">BPBM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B61FFCEFF68FE71FC930B5A" pageId="28" pageNumber="160" type="description">
|
||
<paragraph id="8BE8B1075B61FFCEFF68FE71FC1F0FF5" blockId="28.[151,1437,261,1979]" box="[199,998,441,468]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FE71FEA80FF5" bold="true" box="[199,337,441,468]" pageId="28" pageNumber="160">Description</emphasis>
|
||
.
|
||
<emphasis id="B9236D155B61FFCEFECFFE72FE640FF5" bold="true" box="[352,413,442,468]" pageId="28" pageNumber="160">Male</emphasis>
|
||
: Length
|
||
<quantity id="4CAF1CE25B61FFCEFDAEFE71FD9E0FF5" box="[513,615,441,468]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="28" pageNumber="160" unit="mm" value="8.9">8.90 mm</quantity>
|
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, width across pronotum
|
||
<quantity id="4CAF1CE25B61FFCEFCD4FE71FC1B0FF5" box="[891,994,441,468]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="28" pageNumber="160" unit="mm" value="3.2">3.20 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FE15FB960CF1" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FE15FEBB0FD6" box="[199,322,477,503]" italics="true" pageId="28" pageNumber="160">Coloration</emphasis>
|
||
: Ground color dark brown (
|
||
<figureCitation id="136CAD825B61FFCEFDD4FE15FD360FD9" box="[635,719,477,504]" captionStart="FIGURES 41–48" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 41–48. Enithares species in the E. megalops group, dorsal habitus photographs. 41–42. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 43–44. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 45–46. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 47–48. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="28" pageNumber="160">Fig. 45</figureCitation>
|
||
). Vertex and frons creamy white, with diffuse brown centrally at anterior apex of frons. Pronotum with anterior half creamy white, posterior half dark brown. Scutellum dark brownish centrally, transitioning to yellowish-brown along lateral margins. Hemelytra hyaline, with clavus and corium bluish-grey, longitudinally suffused with black along claval suture, inner portion of corium bordering clavus, and along embolium, overall hemelytral coloration appearing darker due to dorsal abdominal color showing through; wing membrane dark fumate brown. Legs yellowish brown, fore and middle tibiae darkened distally, anterior edge of hind femur narrowly margined with dark brown. Venter brown, with coxal plates tan.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FD11FD070D86" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FD11FE4E0CD2" box="[199,439,729,755]" italics="true" pageId="28" pageNumber="160">Structural characters</emphasis>
|
||
: Head broadly rounded anteriorly when viewed dorsally. Head length 0.95; greatest width 2.55, equal to 0.79 pronotal width; anterior width of vertex 0.80, equal to 0.89× head length. Synthlipsis 0.50, about 0.62 anterior width of vertex and clearly shorter than pronotum. Pronotal length along midline 1.10, humeral width 3.20, lateral margins nearly straight, posterior margin weakly sinuate. Dorsal margin of pronotal fovea directed caudad behind eyes (
|
||
<figureCitation id="136CAD825B61FFCEFE2CFCA1FE210DA5" box="[387,472,873,900]" captionStart="FIGURES 41–48" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 41–48. Enithares species in the E. megalops group, dorsal habitus photographs. 41–42. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 43–44. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 45–46. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 47–48. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="28" pageNumber="160">Fig. 45</figureCitation>
|
||
). Nodal furrow curved weakly cephalad, removed by 2.0× its length from membranal suture, length 0.50, distance to membranal suture 1.00.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FC7AFBED0A16" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">Front and hind legs typical for genus, lacking unusual modifications. Middle trochanter angular. Middle femur with sharp subapical tooth bearing a sloping shoulder distally with 3 small, black, peg-like teeth. Lengths of leg segments as follows: fore femur–tibia–tarsal 1–tarsal 2 = 1.35/1.35/0.55/0.30; middle femur–tibia–tarsal 1–tarsal 2 = 2.00/1.70/0.60/0.40; hind femur–tibia–tarsal 1–tarsal 2 = 3.10/2.70/1.10/0.80.</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FB89FB750A7D" blockId="28.[151,1437,261,1979]" box="[199,1164,1089,1116]" pageId="28" pageNumber="160">
|
||
Ventral abdomen with metaxyphus triangular, slightly concave, tip acuminate (
|
||
<figureCitation id="136CAD825B61FFCEFB96FB89FB870A7D" box="[1081,1150,1089,1116]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1014,1039]" captionTargetBox="[151,1436,159,999]" captionTargetId="figure@3.[151,1436,159,1000]" captionTargetPageId="3" captionText="FIGURES 2–7. Enithares species, metaxiphus. 2. E. arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 3. E. bosavi n. sp.; holotype specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 4. E. insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 5. E. kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 6. E. orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Reinduk, CL 7036. 7. E. papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157." figureDoi="http://doi.org/10.5281/zenodo.3814083" httpUri="https://zenodo.org/record/3814083/files/figure.png" pageId="28" pageNumber="160">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FBADFDD10B2E" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FBADFE970A5E" box="[199,366,1125,1151]" italics="true" pageId="28" pageNumber="160">Male genitalia</emphasis>
|
||
when viewed laterally (
|
||
<figureCitation id="136CAD825B61FFCEFD2AFBADFD200AA1" box="[645,729,1125,1152]" captionStart="FIGURES 49–54" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 49–54. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="28" pageNumber="160">Fig. 52</figureCitation>
|
||
) with proctiger angular; pygophore with posteroventral angle forming nearly a right angle bearing a prominent setal tuft, posterior lobe broadly curved along posterior margin, dorsal margin truncate, angled slightly anterad; paramere small, crescent-shaped, reaching only halfway to apex of posterior lobe; lateral arm of basal plate broadly expanded distally, dorsal apex nearly straight; aedeagus semicircular, with numerous transverse folds.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68FAD2FC930B5A" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FAD2FEE70B15" bold="true" box="[199,286,1306,1332]" pageId="28" pageNumber="160">Female</emphasis>
|
||
: Length 8.0 mm., width across pronotum
|
||
<quantity id="4CAF1CE25B61FFCEFD5CFAD1FCA30B15" box="[755,858,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="28" pageNumber="160" unit="mm" value="3.2">3.20 mm</quantity>
|
||
. Similar to male in general structure and coloration, with following exceptions: general coloration yellowish-brown (
|
||
<figureCitation id="136CAD825B61FFCEFCDDFAF5FC3E0B79" box="[882,967,1341,1368]" captionStart="FIGURES 41–48" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 41–48. Enithares species in the E. megalops group, dorsal habitus photographs. 41–42. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 43–44. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 45–46. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 47–48. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="28" pageNumber="160">Fig. 46</figureCitation>
|
||
); upper frons with a pair (1+1) of prominent, longitudinally ovate reddish spots to either side of midline.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B61FFCEFF68FA4EFD860BE2" pageId="28" pageNumber="160" type="etymology">
|
||
<paragraph id="8BE8B1075B61FFCEFF68FA4EFD860BE2" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FA4EFEB10B81" bold="true" box="[199,328,1414,1440]" pageId="28" pageNumber="160">Etymology</emphasis>
|
||
. The name “kasim” is a noun in apposition, and refers to the
|
||
<typeStatus id="54EC0FA55B61FFCEFC47FA4EFBE10B81" box="[1000,1048,1414,1440]" pageId="28" pageNumber="160">type</typeStatus>
|
||
locality at Kasim, near the furthest western point on the island of New
|
||
<collectingCountry id="F340F1975B61FFCEFD89FA61FD820BE2" box="[550,635,1449,1475]" name="Guinea-Bissau" pageId="28" pageNumber="160">Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B61FFCEFF68FA05FEA5099A" pageId="28" pageNumber="160" type="discussion">
|
||
<paragraph id="8BE8B1075B61FFCEFF68FA05FBFD08C5" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68FA05FEBC0BC9" bold="true" box="[199,325,1485,1512]" pageId="28" pageNumber="160">Discussion</emphasis>
|
||
.
|
||
<taxonomicName id="4C57CA845B61FFCEFEFAFA05FDF30BC6" authorityName="Polhemus" authorityYear="2020" box="[341,522,1485,1511]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFEFAFA05FDF30BC6" box="[341,522,1485,1511]" italics="true" pageId="28" pageNumber="160">Enithares kasim</emphasis>
|
||
</taxonomicName>
|
||
is similar in certain respects to
|
||
<taxonomicName id="4C57CA845B61FFCEFCD7FA06FBFD0BC6" authorityName="Lansbury" authorityYear="1968" box="[888,1028,1485,1511]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="megalops">
|
||
<emphasis id="B9236D155B61FFCEFCD7FA06FBFD0BC6" box="[888,1028,1485,1511]" italics="true" pageId="28" pageNumber="160">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
, but is slightly larger and stouter in overall form. The male genitalia will also readily separate the two species. The paramere of
|
||
<taxonomicName id="4C57CA845B61FFCEFB06FA3AFACD082A" authorityName="Lansbury" authorityYear="1968" box="[1193,1332,1521,1547]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="megalops">
|
||
<emphasis id="B9236D155B61FFCEFB06FA3AFACD082A" box="[1193,1332,1521,1547]" italics="true" pageId="28" pageNumber="160">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
is nearly vestigial, being represented by a small, rounded nub, whereas the paramere of
|
||
<taxonomicName id="4C57CA845B61FFCEFBBAF9DEFB8E080E" authorityName="Polhemus" authorityYear="2020" box="[1045,1143,1557,1583]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFBBAF9DEFB8E080E" box="[1045,1143,1557,1583]" italics="true" pageId="28" pageNumber="160">E. kasim</emphasis>
|
||
</taxonomicName>
|
||
is somewhat larger, more obvious, and crescent-shaped (compare
|
||
<figureCitation id="136CAD825B61FFCEFDF4F9F1FD1C0875" box="[603,741,1593,1620]" captionStart="FIGURES 49–54" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 49–54. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="28" pageNumber="160">Figs. 49, 52</figureCitation>
|
||
). The PL of
|
||
<taxonomicName id="4C57CA845B61FFCEFCDCF9F2FC070872" authorityName="Lansbury" authorityYear="1968" box="[883,1022,1593,1619]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="megalops">
|
||
<emphasis id="B9236D155B61FFCEFCDCF9F2FC070872" box="[883,1022,1593,1619]" italics="true" pageId="28" pageNumber="160">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
takes the form of a broadly rounded triangle, whereas that of
|
||
<taxonomicName id="4C57CA845B61FFCEFE07F996FDFE0856" authorityName="Polhemus" authorityYear="2020" box="[424,519,1629,1655]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFE07F996FDFE0856" box="[424,519,1629,1655]" italics="true" pageId="28" pageNumber="160">E. kasim</emphasis>
|
||
</taxonomicName>
|
||
it is bent anteriorly, with a truncate apex (compare
|
||
<figureCitation id="136CAD825B61FFCEFB93F995FB380859" box="[1084,1217,1629,1656]" captionStart="FIGURES 49–54" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 49–54. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="28" pageNumber="160">Figs. 49, 52</figureCitation>
|
||
). Finally, the LABP of
|
||
<taxonomicName id="4C57CA845B61FFCEFF19F94AFEB808BA" authorityName="Lansbury" authorityYear="1968" box="[182,321,1665,1691]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="megalops">
|
||
<emphasis id="B9236D155B61FFCEFF19F94AFEB808BA" box="[182,321,1665,1691]" italics="true" pageId="28" pageNumber="160">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
is gently curved and slightly tapering, with a blunt apex, whereas the LABP in
|
||
<taxonomicName id="4C57CA845B61FFCEFB6BF94AFADC08BA" authorityName="Polhemus" authorityYear="2020" box="[1220,1317,1665,1691]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFB6BF94AFADC08BA" box="[1220,1317,1665,1691]" italics="true" pageId="28" pageNumber="160">E. kasim</emphasis>
|
||
</taxonomicName>
|
||
is broadly expanded distally, with a relatively straight dorsal margin leading to a broadly rounded apex (compare
|
||
<figureCitation id="136CAD825B61FFCEFAA6F96DFA6908E1" box="[1289,1424,1701,1728]" captionStart="FIGURES 49–54" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 49–54. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="28" pageNumber="160">Figs. 49, 52</figureCitation>
|
||
). The mesotrochanter of
|
||
<taxonomicName id="4C57CA845B61FFCEFE33F902FE0508C2" authorityName="Polhemus" authorityYear="2020" box="[412,508,1737,1763]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFE33F902FE0508C2" box="[412,508,1737,1763]" italics="true" pageId="28" pageNumber="160">E. kasim</emphasis>
|
||
</taxonomicName>
|
||
is also more angular than that of
|
||
<taxonomicName id="4C57CA845B61FFCEFCDCF902FC0408C2" authorityName="Lansbury" authorityYear="1968" box="[883,1021,1737,1763]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="megalops">
|
||
<emphasis id="B9236D155B61FFCEFCDCF902FC0408C2" box="[883,1021,1737,1763]" italics="true" pageId="28" pageNumber="160">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B61FFCEFF68F925FEA5099A" blockId="28.[151,1437,261,1979]" pageId="28" pageNumber="160">
|
||
<emphasis id="B9236D155B61FFCEFF68F925FE710929" bold="true" box="[199,392,1773,1800]" pageId="28" pageNumber="160">Ecological notes</emphasis>
|
||
.
|
||
<taxonomicName id="4C57CA845B61FFCEFE38F926FE010926" authorityName="Polhemus" authorityYear="2020" box="[407,504,1773,1799]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="160" phylum="Arthropoda" rank="species" species="kasim">
|
||
<emphasis id="B9236D155B61FFCEFE38F926FE010926" box="[407,504,1773,1799]" italics="true" pageId="28" pageNumber="160">E. kasim</emphasis>
|
||
</taxonomicName>
|
||
is presently known only from the extreme western portion of the Vogelkop Peninsula (
|
||
<figureCitation id="136CAD825B61FFCEFF7DF8D9FEDC090D" box="[210,293,1809,1836]" captionStart="FIGURE 67" captionStartId="38.[151,250,1916,1941]" captionTargetBox="[158,1430,1097,1882]" captionTargetId="figure@38.[151,1436,1090,1890]" captionTargetPageId="38" captionText="FIGURE 67. Distribution of species in the E. megalops group on New Guinea. Square = E. kasim n. sp.; circles = E. nigra Lansbury; triangles = E. alexis Lansbury; pentagons = E. vulgaris Lansbury." figureDoi="http://doi.org/10.5281/zenodo.3814901" httpUri="https://zenodo.org/record/3814901/files/figure.png" pageId="28" pageNumber="160">Fig. 67</figureCitation>
|
||
), occupying the Vogelkop Lowlands area of freshwater endemism (Areas 4) as defined by D.
|
||
<bibRefCitation id="EFC6CCF65B61FFCEFA81F8D9FEBC096E" author="Polhemus, D. A. & Allen, G. A." pageId="28" pageNumber="160" pagination="207 - 245" refId="ref39028" refString="Polhemus, D. A. & Allen, G. A. (2007) Freshwater biogeography of Papua. In: Marshall, A. J. & Beehler, B. M. (Eds.), Ecology of Papua. Part 1. Periplus Editions (HK) Ltd., Singapore, pp. 207 - 245." type="book chapter" year="2007">Polhemus & Allen (2007)</bibRefCitation>
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. The
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<typeStatus id="54EC0FA55B61FFCEFE2EF8FEFE480971" box="[385,433,1846,1872]" pageId="28" pageNumber="160">type</typeStatus>
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locality was a shaded streambed in lowland swamp forest, with scattered pools connected by hyporheic flow. Collections were made in the dry season, when the water is low and aquatic biota concentrates in such remnant pools; during the wet season the channel would be filled, representing a continuous and far more extensive habitat.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |