742 lines
87 KiB
XML
742 lines
87 KiB
XML
<document id="DE6C49AB4C3BE09AD9D271E4E9ED54D5" ID-DOI="10.5281/zenodo.190909" ID-GBIF-Dataset="a1d1a578-4cf6-4dd2-b56d-b288cc17439c" ID-ISSN="1175-5326" ID-Zenodo-Dep="190909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461121462409" checkinUser="plazi" docAuthor="Glaw, Frank, Köhler, Jörn & Vences, Miguel" docDate="2009" docId="03FAF57F117A4170D5BAFB8E4B64345D" docLanguage="en" docName="zt02269p042.pdf" docOrigin="Zootaxa 2269" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Furcifer timoni Glaw, Köhler & Vences, 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="FFC38D07117B4178D52DFFD44F703167" masterDocTitle="A distinctive new species of chameleon of the genus Furcifer (Squamata: Chamaeleonidae) from the Montagne d’Ambre rainforest of northern Madagascar" masterLastPageNumber="42" masterPageNumber="32" pageNumber="33" updateTime="1698596034171" updateUser="plazi">
|
||
<mods:mods id="7C08F532771B5EE52FF10CEA371355AC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5021C28749833FF65C46AA7134DCF200">
|
||
<mods:title id="C95D33B6BF3A971BD7243DE2B10C842F">A distinctive new species of chameleon of the genus Furcifer (Squamata: Chamaeleonidae) from the Montagne d’Ambre rainforest of northern Madagascar</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="CF2B02F087EE78792B46930E2E122249" type="personal">
|
||
<mods:role id="E631ECC0191A4966A87E5426F344E0DC">
|
||
<mods:roleTerm id="50CAC225289D323D2851CDD678D5952E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C7162AA5728BBBA88C73F9CB7FC38E01">Glaw, Frank</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A52DA0C3C1DB2D8DD6915E264EF5CA29" type="personal">
|
||
<mods:role id="D96E5D78B1071A1097D5DE93A0380C86">
|
||
<mods:roleTerm id="701219997CC09B5EFDFAB21AC0F8CD3A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5465C63590412F33F2F94ED84BB3702E">Köhler, Jörn</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C3599F6E27EEC74A9B9F432CA8A85928" type="personal">
|
||
<mods:role id="F659475084499A2D0083A21F35376E22">
|
||
<mods:roleTerm id="C085E89FAC5F7C15D440DEF88AE8588E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="33BFBBADAC450BC818E8B251E01F5EBE">Vences, Miguel</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E63AA9A89143CBB1DE05E924E5E5FEA7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="1C7034F822BB539C6FACCC92D7588BA1" type="host">
|
||
<mods:titleInfo id="EEA73F907BF923C71672276CA89BF0E7">
|
||
<mods:title id="7C76C26038714642F749CCBE646C6300">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="63EA0B35D919D7C5B98DE8D3D0A73A49">
|
||
<mods:date id="FD43A0BD66E3C57E58D2548E63646942">2009</mods:date>
|
||
<mods:detail id="4F627E841B8F8971EF3898F76B760170" type="volume">
|
||
<mods:number id="4C62F5110749737754494F5F107B01D9">2269</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A8FA1C893ED218FB23BF108777C23A82" unit="page">
|
||
<mods:start id="8C9A72701B31D9EC1EA33398ECAB6865">32</mods:start>
|
||
<mods:end id="6B68ED727922059BE09D33E26463CDCD">42</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="96888C52B3B40E709286B2B916047316">journal article</mods:classification>
|
||
<mods:identifier id="725C27684AEA67AB9A6FB0D4297CEC2B" type="DOI">10.5281/zenodo.190909</mods:identifier>
|
||
<mods:identifier id="CDA51640C4351CA848BDC910B0E1F757" type="GBIF-Dataset">a1d1a578-4cf6-4dd2-b56d-b288cc17439c</mods:identifier>
|
||
<mods:identifier id="87A60D555828ED4B291EEC7E38EF8A30" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="8EC971EB6DBBBF94ED565E0E3EEBE9BB" type="Zenodo-Dep">190909</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FAF57F117A4170D5BAFB8E4B64345D" ID-DOI="http://doi.org/10.5281/zenodo.6224500" ID-GBIF-Taxon="119625682" ID-Zenodo-Dep="6224500" LSID="urn:lsid:plazi:treatment:03FAF57F117A4170D5BAFB8E4B64345D" httpUri="http://treatment.plazi.org/id/03FAF57F117A4170D5BAFB8E4B64345D" lastPageId="8" lastPageNumber="40" pageId="1" pageNumber="33">
|
||
<subSubSection id="C34917E2117A4179D5BAFB8E4E7435F0" pageId="1" pageNumber="33" type="nomenclature">
|
||
<paragraph id="8BEC4469117A4179D5BAFB8E4EC73513" blockId="1.[151,439,1114,1175]" box="[151,439,1114,1140]" pageId="1" pageNumber="33">
|
||
<heading id="D0A4F305117A4179D5BAFB8E4EC73513" bold="true" box="[151,439,1114,1140]" fontSize="11" level="1" pageId="1" pageNumber="33" reason="1">
|
||
<emphasis id="B927987B117A4179D5BAFB8E4EC73513" bold="true" box="[151,439,1114,1140]" pageId="1" pageNumber="33">
|
||
<taxonomicName id="4C533FEA117A4179D5BAFB8E4E233513" box="[151,339,1114,1140]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B117A4179D5BAFB8E4E233513" bold="true" box="[151,339,1114,1140]" italics="true" pageId="1" pageNumber="33">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2142500117A4179D477FB8E4EC73513" box="[346,439,1114,1140]" pageId="1" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117A4179D5BAFBA94E7435F0" blockId="1.[151,439,1114,1175]" box="[151,260,1149,1175]" pageId="1" pageNumber="33">
|
||
<figureCitation id="136858EC117A4179D5BAFBA94E7435F0" box="[151,260,1149,1175]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,254,1850,1874]" captionStartId-1="3.[151,261,1406,1430]" captionStartId-2="4.[151,259,813,837]" captionStartId-3="5.[151,254,2012,2036]" captionStartId-4="6.[151,258,977,1001]" captionTargetBox-0="[157,1426,296,1817]" captionTargetBox-1="[251,1332,551,1373]" captionTargetBox-2="[178,1416,203,778]" captionTargetBox-3="[156,1429,719,1984]" captionTargetBox-4="[176,1422,192,939]" captionTargetId-0="figure@2.[157,1431,293,1825]" captionTargetId-1="figure@3.[245,1341,545,1381]" captionTargetId-2="figure@4.[164,1422,197,788]" captionTargetId-3="figure@5.[153,1434,697,1988]" captionTargetId-4="figure@6.[154,1433,158,952]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 in life, (A) lateral view, (B) dorsolateral view of head; and (C) additional male of the same species photographed near the type locality by P. Schönecker (not collected)." captionText-1="FIGURE 2. Drawing of the head of the male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 (by Ruth Kühbandner)." captionText-2="FIGURE 3. Drawings of hemipenis of the male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 (by Ruth Kühbandner): (A) asulcal view, (B) lateral view; (C) sulcal view." captionText-3="FIGURE 4. Gravid female paratype of Furcifer timoni sp. nov., ZSM uncatalogued (FGZC 1884) in life." captionText-4="FIGURE 5. (A) Preserved male holotype of Furcifer bifidus (MNHN 6660) in comparison to (B) preserved male holotype of Furcifer timoni sp. nov. (ZSM 2103 / 2007)." httpUri-0="https://zenodo.org/record/190910/files/figure.png" httpUri-1="https://zenodo.org/record/190911/files/figure.png" httpUri-2="https://zenodo.org/record/190912/files/figure.png" httpUri-3="https://zenodo.org/record/190913/files/figure.png" httpUri-4="https://zenodo.org/record/190914/files/figure.png" pageId="1" pageNumber="33">Figs. 1–5</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34917E2117A4179D5BAFB1E4EFC3700" pageId="1" pageNumber="33" type="materials_examined">
|
||
<paragraph id="8BEC4469117A4179D5BAFB1E4B673455" blockId="1.[151,1437,1226,2026]" pageId="1" pageNumber="33">
|
||
<emphasis id="B927987B117A4179D5BAFB1E4E643583" bold="true" box="[151,276,1226,1252]" pageId="1" pageNumber="33">
|
||
<typeStatus id="54E8FACB117A4179D5BAFB1E4E603583" box="[151,272,1226,1252]" pageId="1" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED42DCAC117A4179D430FB1E4E2B3583" box="[285,347,1226,1252]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="33">ZSM</collectionCode>
|
||
2103/2007 (
|
||
<collectionCode id="ED42DCAC117A4179D4DBFB1E4D353583" box="[502,581,1226,1252]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
1095), adult male, from Montagne d'Ambre National Park (
|
||
<geoCoordinate id="EE6722AE117A4179D007FB1E4AE73583" box="[1322,1431,1226,1252]" direction="east" orientation="longitude" pageId="1" pageNumber="33" precision="925" value="12.5">12°30' E</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6722AE117A4179D5BAFB254F8C346C" box="[151,252,1265,1291]" direction="south" orientation="latitude" pageId="1" pageNumber="33" precision="925" value="-49.183334">49°11' S</geoCoordinate>
|
||
), ca.
|
||
<quantity id="4CABE98C117A4179D410FB254EB4346C" box="[317,452,1265,1291]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="1" pageNumber="33" unit="m" value="775.0" valueMax="800.0" valueMin="750.0">750–800 m</quantity>
|
||
a.s.l., Antsiranana Province, northern
|
||
<collectingCountry id="F34404F9117A4179D6A0FB254B6D346C" box="[909,1053,1265,1291]" name="Madagascar" pageId="1" pageNumber="33">Madagascar</collectingCountry>
|
||
, collected on
|
||
<date id="FFED62A9117A4179D1ECFB254AE7346C" box="[1217,1431,1265,1291]" pageId="1" pageNumber="33" value="2007-02-25">25 February 2007</date>
|
||
, by P. Bora, H. Enting, F. Glaw, A. Knoll, J. Köhler and A. Razafimanantsoa.
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117A4179D5EBFAEA4EFC3700" blockId="1.[151,1437,1226,2026]" pageId="1" pageNumber="33">
|
||
<emphasis id="B927987B117A4179D5EBFAEA4E3B343F" bold="true" box="[198,331,1342,1368]" pageId="1" pageNumber="33">
|
||
<typeStatus id="54E8FACB117A4179D5EBFAEA4E36343F" box="[198,326,1342,1368]" pageId="1" pageNumber="33" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED42DCAC117A4179D47BFAEA4EE2343F" box="[342,402,1342,1368]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="33">ZSM</collectionCode>
|
||
255/2004 (
|
||
<collectionCode id="ED42DCAC117A4179D732FAEA4D1C343F" box="[543,620,1342,1368]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
495),
|
||
<collectionCode id="ED42DCAC117A4179D792FAEA4D8B343F" box="[703,763,1342,1368]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="33">ZSM</collectionCode>
|
||
256/2004 (
|
||
<collectionCode id="ED42DCAC117A4179D6A5FAEA4CA5343F" box="[904,981,1342,1368]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
496),
|
||
<collectionCode id="ED42DCAC117A4179D105FAEA4B09343F" LSID="urn:lsid:biocol.org:col:34613" box="[1064,1145,1342,1368]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="33">ZFMK</collectionCode>
|
||
87585 (originally
|
||
<collectionCode id="ED42DCAC117A4179D04DFAEA4AEC343F" box="[1376,1436,1342,1368]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="33">ZSM</collectionCode>
|
||
257/2004 [
|
||
<collectionCode id="ED42DCAC117A4179D439FAB14E133418" box="[276,355,1381,1407]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
499]),
|
||
<collectionCode id="ED42DCAC117A4179D497FAB14D533418" box="[442,547,1381,1407]" httpUri="http://grbio.org/cool/q3sh-smdb" name="University dAntananarivo, Department de Biologie Animale" pageId="1" pageNumber="33">UADBA</collectionCode>
|
||
uncatalogued (
|
||
<collectionCode id="ED42DCAC117A4179D7F5FAB14C553418" box="[728,805,1381,1407]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
497),
|
||
<collectionCode id="ED42DCAC117A4179D65EFAB14CAC3418" box="[883,988,1381,1407]" httpUri="http://grbio.org/cool/q3sh-smdb" name="University dAntananarivo, Department de Biologie Animale" pageId="1" pageNumber="33">UADBA</collectionCode>
|
||
uncatalogued (
|
||
<collectionCode id="ED42DCAC117A4179D1BDFAB14BAD3418" box="[1168,1245,1381,1407]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
498), five adult and gravid females, from Montagne d'Ambre National Park (same general area as
|
||
<typeStatus id="54E8FACB117A4179D19FFA584A5134C1" box="[1202,1313,1420,1446]" pageId="1" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
locality), Antsiranana Province, northern
|
||
<collectingCountry id="F34404F9117A4179D70EFA664DC934AB" box="[547,697,1458,1484]" name="Madagascar" pageId="1" pageNumber="33">Madagascar</collectingCountry>
|
||
, collected
|
||
<date id="FFED62A9117A4179D613FA664B3C34AB" box="[830,1100,1458,1484]" pageId="1" pageNumber="33" value="2004-02-20" valueMax="2004-02-23" valueMin="2004-02-20">20–23 February 2004</date>
|
||
, by F. Glaw, M. Puente, R. Randrianiaina & A. Razafimanantsoa;
|
||
<collectionCode id="ED42DCAC117A4179D749FA0D4DD13494" box="[612,673,1497,1523]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="33">ZSM</collectionCode>
|
||
uncatalogued (
|
||
<collectionCode id="ED42DCAC117A4179D67AFA0D4CD43494" box="[855,932,1497,1523]" pageId="1" pageNumber="33">FGZC</collectionCode>
|
||
1884) [still in
|
||
<collectionCode id="ED42DCAC117A4179D175FA0D4BB73494" box="[1112,1223,1497,1523]" httpUri="http://grbio.org/cool/q3sh-smdb" name="University dAntananarivo, Department de Biologie Animale" pageId="1" pageNumber="33">UADBA</collectionCode>
|
||
], adult and gravid female, Montagne d'Ambre National Park,
|
||
<quantity id="4CABE98C117A4179D79EF9D44C71377D" box="[691,769,1536,1562]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="33" unit="m" value="850.0">850 m</quantity>
|
||
a.s.l. (similar geographical coordinates as
|
||
<typeStatus id="54E8FACB117A4179D030F9D44AFF377D" box="[1309,1423,1536,1562]" pageId="1" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
), Antsiranana Province, northern
|
||
<collectingCountry id="F34404F9117A4179D723F9F24DEE3727" box="[526,670,1574,1600]" name="Madagascar" pageId="1" pageNumber="33">Madagascar</collectingCountry>
|
||
, collected
|
||
<date id="FFED62A9117A4179D637F9F24C9F3727" box="[794,1007,1574,1600]" pageId="1" pageNumber="33" value="2008-02-26">26 February 2008</date>
|
||
, by N. D'Cruze, F. Glaw, Z. Nagy & A. Razafimanantsoa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34917E2117A417AD5EBF9A74AE73198" lastPageId="2" lastPageNumber="34" pageId="1" pageNumber="33" type="diagnosis">
|
||
<paragraph id="8BEC4469117A417AD5EBF9A74AE73198" blockId="1.[151,1437,1226,2026]" lastBlockId="2.[151,1436,152,255]" lastPageId="2" lastPageNumber="34" pageId="1" pageNumber="33">
|
||
<emphasis id="B927987B117A4179D5EBF9A74E3637EA" bold="true" box="[198,326,1651,1677]" pageId="1" pageNumber="33">Diagnosis.</emphasis>
|
||
The new species differs from all other
|
||
<taxonomicName id="4C533FEA117A4179D63AF9A04C0837EA" box="[791,888,1652,1677]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B927987B117A4179D63AF9A04C0837EA" box="[791,888,1652,1677]" italics="true" pageId="1" pageNumber="33">Furcifer</emphasis>
|
||
</taxonomicName>
|
||
species except
|
||
<taxonomicName id="4C533FEA117A4179D11CF9A04BED37EA" box="[1073,1181,1652,1677]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117A4179D11CF9A04BED37EA" box="[1073,1181,1652,1677]" italics="true" pageId="1" pageNumber="33">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C533FEA117A4179D1F4F9A04A1237EA" box="[1241,1378,1652,1677]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="balteatus">
|
||
<emphasis id="B927987B117A4179D1F4F9A04A1237EA" box="[1241,1378,1652,1677]" italics="true" pageId="1" pageNumber="33">F. balteatus</emphasis>
|
||
</taxonomicName>
|
||
by a whitish lateroventral band that is composed of scales which are arranged in a rosette-like manner. It differs from
|
||
<taxonomicName id="4C533FEA117A4179D5FAF9154E1037BD" box="[215,352,1729,1754]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="balteatus">
|
||
<emphasis id="B927987B117A4179D5FAF9154E1037BD" box="[215,352,1729,1754]" italics="true" pageId="1" pageNumber="33">F. balteatus</emphasis>
|
||
</taxonomicName>
|
||
by smaller size (male SVL
|
||
<quantity id="4CABE98C117A4179D788F9154D8B37BC" box="[677,763,1729,1755]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="33" unit="mm" value="88.0">88 mm</quantity>
|
||
, female SVL
|
||
<quantity id="4CABE98C117A4179D6B1F9154B6A37BC" box="[924,1050,1729,1755]" metricMagnitude="-2" metricUnit="m" metricValue="9.55" metricValueMax="9.6" metricValueMin="9.5" pageId="1" pageNumber="33" unit="mm" value="95.5" valueMax="96.0" valueMin="95.0">95–96 mm</quantity>
|
||
versus male SVL up to
|
||
<quantity id="4CABE98C117A4179D019F9154AE837BC" box="[1332,1432,1729,1755]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="33" unit="mm" value="175.0">175 mm</quantity>
|
||
, female SVL up to
|
||
<quantity id="4CABE98C117A4179D454F93C4E923665" box="[377,482,1768,1794]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="1" pageNumber="33" unit="mm" value="145.0">145 mm</quantity>
|
||
), shorter rostral appendages of the males (
|
||
<quantity id="4CABE98C117A4179D6C5F93C4B5F3665" box="[1000,1071,1768,1794]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="33" unit="mm" value="6.0">6 mm</quantity>
|
||
versus up to
|
||
<quantity id="4CABE98C117A4179D1F9F93C4A5D3665" box="[1236,1325,1768,1794]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="33" unit="mm" value="10.0">10 mm</quantity>
|
||
), general colouration, and the position and extension of an oblique light band on the flanks that runs from the dorsal crest (or slightly below) posterioventrally without reaching the lateroventral band (versus running from the upper flanks to the whitish lateroventral band). It differs from
|
||
<taxonomicName id="4C533FEA117A4179D6A7F8884C8A3612" box="[906,1018,1884,1909]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117A4179D6A7F8884C8A3612" box="[906,1018,1884,1909]" italics="true" pageId="1" pageNumber="33">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
by distinctly smaller size (SVL of adult male
|
||
<quantity id="4CABE98C117A4179D435F8564E1B36FB" box="[280,363,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="33" unit="mm" value="88.0">88 mm</quantity>
|
||
versus up to
|
||
<quantity id="4CABE98C117A4179D72BF8564D1B36FB" box="[518,619,1922,1948]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="33" unit="mm" value="200.0">200 mm</quantity>
|
||
), much shorter rostral appendages of the males (
|
||
<quantity id="4CABE98C117A4179D1B2F8564B9436FB" box="[1183,1252,1922,1948]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="33" unit="mm" value="6.0">6 mm</quantity>
|
||
versus up to
|
||
<quantity id="4CABE98C117A4179D052F8564FBA36A4" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="33" unit="mm" value="24.0">24 mm</quantity>
|
||
,
|
||
<figureCitation id="136858EC117A4179D5FAF87D4E5336A4" box="[215,291,1961,1987]" captionStart="FIGURE 5" captionStartId="6.[151,258,977,1001]" captionTargetBox="[176,1422,192,939]" captionTargetId="figure@6.[154,1433,158,952]" captionTargetPageId="6" captionText="FIGURE 5. (A) Preserved male holotype of Furcifer bifidus (MNHN 6660) in comparison to (B) preserved male holotype of Furcifer timoni sp. nov. (ZSM 2103 / 2007)." httpUri="https://zenodo.org/record/190914/files/figure.png" pageId="1" pageNumber="33">Fig. 5</figureCitation>
|
||
), a lower number of scales (4 vs. 6-8) between nostril and tip of rostral appendage, and by colouration. In addition, males of
|
||
<taxonomicName id="4C533FEA117A4179D71EF8044D9A368E" box="[563,746,2000,2025]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117A4179D71EF8044D9A368E" box="[563,746,2000,2025]" italics="true" pageId="1" pageNumber="33">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
differ from all other
|
||
<taxonomicName id="4C533FEA117A4179D6C0F8044B20368E" box="[1005,1104,2000,2025]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B927987B117A4179D6C0F8044B20368E" box="[1005,1104,2000,2025]" italics="true" pageId="1" pageNumber="33">Furcifer</emphasis>
|
||
</taxonomicName>
|
||
species except those of the
|
||
<taxonomicName id="4C533FEA1179417AD5BAFF4C4E2431D6" box="[151,340,152,177]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B1179417AD5BAFF4C4E2431D6" box="[151,340,152,177]" italics="true" pageId="2" pageNumber="34">Furcifer bifidus</emphasis>
|
||
</taxonomicName>
|
||
group -
|
||
<taxonomicName id="4C533FEA1179417AD490FF4C4D5D31D6" box="[445,557,152,177]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B1179417AD490FF4C4D5D31D6" box="[445,557,152,177]" italics="true" pageId="2" pageNumber="34">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C533FEA1179417AD710FF4C4DBA31D6" box="[573,714,152,177]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="balteatus">
|
||
<emphasis id="B927987B1179417AD710FF4C4DBA31D6" box="[573,714,152,177]" italics="true" pageId="2" pageNumber="34">F. balteatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B927987B1179417AD7F7FF4C4C4C31D6" box="[730,828,152,177]" italics="true" pageId="2" pageNumber="34">F. m i n o r</emphasis>
|
||
(Günther, 1879),
|
||
<taxonomicName id="4C533FEA1179417AD134FF4C4A3A31D5" authority="Gunther, 1890" authorityName="Gunther" authorityYear="1890" box="[1049,1354,152,178]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="willsii">
|
||
<emphasis id="B927987B1179417AD134FF4C4BF131D6" box="[1049,1153,152,177]" italics="true" pageId="2" pageNumber="34">F. willsii</emphasis>
|
||
(Günther, 1890)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C533FEA1179417AD0ABFF4C4D4E31BF" authority="Brygoo & Domergue, 1966" authorityName="Brygoo & Domergue" authorityYear="1966" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="petteri">
|
||
<emphasis id="B927987B1179417AD0ABFF4C4F9431BF" italics="true" pageId="2" pageNumber="34">F. petteri</emphasis>
|
||
(Brygoo & Domergue, 1966)
|
||
</taxonomicName>
|
||
- by the presence of two bony rostral appendages and from adult males of all five species within the
|
||
<taxonomicName id="4C533FEA1179417AD4E7FF314D453199" box="[458,565,229,254]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B1179417AD4E7FF314D453199" box="[458,565,229,254]" italics="true" pageId="2" pageNumber="34">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
group by a shorter length of the rostral appendages (
|
||
<quantity id="4CABE98C1179417AD18CFF314B963198" box="[1185,1254,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="34" unit="mm" value="6.0">6 mm</quantity>
|
||
vs.
|
||
<quantity id="4CABE98C1179417AD038FF314AFA3198" box="[1301,1418,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.4" metricValueMin="0.5" pageId="2" pageNumber="34" unit="mm" value="14.5" valueMax="24.0" valueMin="5.0">5–24 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2C14E11179417AD5BAF8EE4AE03614" httpUri="https://zenodo.org/record/190910/files/figure.png" pageId="2" pageNumber="34" targetBox="[157,1426,296,1817]" targetPageId="2">
|
||
<paragraph id="8BEC44691179417AD5BAF8EE4AE03614" blockId="2.[151,1436,1850,1907]" pageId="2" pageNumber="34">
|
||
<emphasis id="B927987B1179417AD5BAF8EE4E613635" bold="true" box="[151,273,1850,1874]" pageId="2" pageNumber="34">FIGURE 1</emphasis>
|
||
. Male holotype of
|
||
<taxonomicName id="4C533FEA1179417AD4F1F8EE4D0F3636" box="[476,639,1850,1873]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B1179417AD4F1F8EE4D0F3636" box="[476,639,1850,1873]" italics="true" pageId="2" pageNumber="34">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21425001179417AD7A8F8EE4DAB3635" box="[645,731,1850,1874]" pageId="2" pageNumber="34" rank="species">
|
||
<emphasis id="B927987B1179417AD7A8F8EE4DAB3635" bold="true" box="[645,731,1850,1874]" pageId="2" pageNumber="34">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ZSM 2103/2007 in life, (A) lateral view, (B) dorsolateral view of head; and (C) additional male of the same species photographed near the type locality by P. Schönecker (not collected).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34917E21179417FD5EBF8754A683080" lastPageId="7" lastPageNumber="39" pageId="2" pageNumber="34" type="description">
|
||
<paragraph id="8BEC44691179417BD5EBF8754A413080" blockId="2.[151,1436,1953,2019]" lastBlockId="3.[151,1437,152,487]" lastPageId="3" lastPageNumber="35" pageId="2" pageNumber="34">
|
||
<emphasis id="B927987B1179417AD5EBF8754D5636DC" bold="true" box="[198,550,1953,1979]" pageId="2" pageNumber="34">
|
||
Description of the
|
||
<typeStatus id="54E8FACB1179417AD499F8754D5136DC" box="[436,545,1953,1979]" pageId="2" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male, in good condition, both hemipenes completely everted; SVL 88.0 mm (length from tip of rostral appendage to vent 93.0 mm); tail
|
||
<quantity id="4CABE98C1179417AD6E9F81D4B583684" box="[964,1064,1993,2019]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" pageId="2" pageNumber="34" unit="mm" value="121.0">121 mm</quantity>
|
||
; axilla-groin distance
|
||
<quantity id="4CABE98C1179417AD006F81D4AE63684" box="[1323,1430,1993,2019]" metricMagnitude="-2" metricUnit="m" metricValue="4.88" pageId="2" pageNumber="34" unit="mm" value="48.8">48.8 mm</quantity>
|
||
; eye horizontal diameter
|
||
<quantity id="4CABE98C1178417BD49FFF4C4D7F31D5" box="[434,527,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="35" unit="mm" value="8.7">8.7 mm</quantity>
|
||
. Head with two ossified, scaled rostral appendages, length
|
||
<quantity id="4CABE98C1178417BD193FF4C4A6A31D5" box="[1214,1306,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="35" unit="mm" value="5.9">5.9 mm</quantity>
|
||
, projecting beyond the upper lip by
|
||
<quantity id="4CABE98C1178417BD4E1FF6A4D5F31BF" box="[460,559,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="35" unit="mm" value="4.7">4.7 mm</quantity>
|
||
, rostral appendages laterally compressed, almost parallel, but slightly diverging, distance between anterior tips
|
||
<quantity id="4CABE98C1178417BD753FF314DAC3198" box="[638,732,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="3" pageNumber="35" unit="mm" value="9.2">9.2 mm</quantity>
|
||
; rostral crest well developed; supra-orbital crest rounded in lateral view and formed by single, rather smooth row of tubercles; lateral crest moderately distinct, smooth in lateral view; temporal crest recognizable behind eye, almost parallel with lateral crest, both crests fusing posteriorly; parietal crest absent; no traces of occipital lobes; no traces of gular crest. Dorsal crest present, consisting of a single row of 12 pointed tubercles
|
||
<quantity id="4CABE98C1178417BD7F6FE544C6E30FD" box="[731,798,384,410]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="35" unit="mm" value="1.0">1 mm</quantity>
|
||
or less in height on the anterior dorsum; body laterally compressed with homogeneous scalation; no distinct ventral crest; axillary pits not obvious; limbs and tail with homogeneous scalation, tail without dorsal crest of pointed tubercles, feet without tarsal spines.
|
||
</paragraph>
|
||
<caption id="DF2C14E11178417BD5BAFAAA4E5B34DF" httpUri="https://zenodo.org/record/190911/files/figure.png" pageId="3" pageNumber="35" targetBox="[251,1332,551,1373]" targetPageId="3">
|
||
<paragraph id="8BEC44691178417BD5BAFAAA4E5B34DF" blockId="3.[151,1437,1406,1464]" pageId="3" pageNumber="35">
|
||
<emphasis id="B927987B1178417BD5BAFAAA4E6B34F1" bold="true" box="[151,283,1406,1430]" pageId="3" pageNumber="35">FIGURE 2</emphasis>
|
||
. Drawing of the head of the male holotype of
|
||
<taxonomicName id="4C533FEA1178417BD66EFAAB4C8734F1" box="[835,1015,1407,1430]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="35" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B1178417BD66EFAAB4C8734F1" box="[835,1015,1407,1430]" italics="true" pageId="3" pageNumber="35">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21425001178417BD6D2FAAA4B2E34F1" box="[1023,1118,1406,1430]" pageId="3" pageNumber="35" rank="species">
|
||
<emphasis id="B927987B1178417BD6D2FAAA4B2E34F1" bold="true" box="[1023,1118,1406,1430]" pageId="3" pageNumber="35">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ZSM 2103/2007 (by Ruth Kühbandner).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC44691178417BD5E8FA334DF237A5" blockId="3.[151,1437,1511,2001]" pageId="3" pageNumber="35">
|
||
Hemipenis (
|
||
<figureCitation id="136858EC1178417BD475FA334ED03766" box="[344,416,1511,1537]" captionStart="FIGURE 3" captionStartId="4.[151,259,813,837]" captionTargetBox="[178,1416,203,778]" captionTargetId="figure@4.[164,1422,197,788]" captionTargetPageId="4" captionText="FIGURE 3. Drawings of hemipenis of the male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 (by Ruth Kühbandner): (A) asulcal view, (B) lateral view; (C) sulcal view." httpUri="https://zenodo.org/record/190912/files/figure.png" pageId="3" pageNumber="35">Fig. 3</figureCitation>
|
||
) large, total length
|
||
<quantity id="4CABE98C1178417BD7A2FA334D8E3766" box="[655,766,1511,1537]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="3" pageNumber="35" unit="mm" value="17.4">17.4 mm</quantity>
|
||
, morphology strongly differing from that of the closely related species
|
||
<taxonomicName id="4C533FEA1178417BD47CF9DA4EB33740" box="[337,451,1550,1575]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="35" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B1178417BD47CF9DA4EB33740" box="[337,451,1550,1575]" italics="true" pageId="3" pageNumber="35">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C533FEA1178417BD4FEF9DA4D483740" box="[467,568,1550,1575]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="35" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="B927987B1178417BD4FEF9DA4D483740" box="[467,568,1550,1575]" italics="true" pageId="3" pageNumber="35">F. minor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B927987B1178417BD765F9DA4DD53740" box="[584,677,1550,1575]" italics="true" pageId="3" pageNumber="35">F. w i l l s i</emphasis>
|
||
and
|
||
<emphasis id="B927987B1178417BD7C8F9DA4C203740" box="[741,848,1550,1575]" italics="true" pageId="3" pageNumber="35">F. p e t t e r i</emphasis>
|
||
(see drawings and descriptions in
|
||
<bibRefCitation id="EFC239981178417BD1D7F9D94FA43729" author="Ramanantsoa" pageId="3" pageNumber="42" refString="Ramanantsoa, G. A. (1978) Contribution a la connaissance des Chamaleonidae malgaches (Reptilia, Squamata). II. Donnees nouvelles sur Chamaeleo minor A. Gunther, 1879, statut de C. willsii petteri E. - R. Brygoo et Ch. A. Domergue, 1966, etude des hemipenis et de la repartition geographique du groupe bifidus. Bulletin du Museum national d'Histoire Naturelle, Paris, (4) 3, 510, 91 - 103." type="journal article" year="1978">Ramanantsoa 1978</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFC239981178417BD43CF9E04D103729" author="Brygoo" box="[273,608,1588,1614]" pageId="3" pageNumber="41" refString="Brygoo, E. R. & Domergue, C. A. (1969) Chamaeleo balteatus Dum. et Bib. (in C. et A. Dumeril, 1851) n'est pas synonyme de C. bifidus Brongniart, 1800. Description du male. Diagnostic des especes du groupe parsonii. Bulletin du Museum national d'Histoire Naturelle, Paris, (ser. 2), 41, 104 - 116." type="journal article" year="1969">Brygoo & Domergue 1969</bibRefCitation>
|
||
). Truncus covered with calyces, especially in asulcal view, calyces absent around the sulcus spermaticus. Sulcus spermaticus poorly recognizable. Apex bilobed, formed by two big lobes each covered with two rotulae and one field of short papillae; a further field of long papillae on each side at the base of the apex in sulcal view.
|
||
</paragraph>
|
||
<paragraph id="8BEC44691178417BD5E8F91B4E8836B6" blockId="3.[151,1437,1511,2001]" pageId="3" pageNumber="35">
|
||
Colour after almost two years in preservative generally blackish brown, especially on flanks, dorsal surfaces of upper arms and legs and lateral parts of the proximal tail. Throat, sides of head, lower arms, lower legs, distal parts of tail and dorsalmost
|
||
<quantity id="4CABE98C1178417BD74EF8C84DB53651" box="[611,709,1820,1846]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="35" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
of flanks grey to purple. A white band starting at the angle of the mouth runs along the upper lip and fades below the eye. The ventrolateral light band between fore- and hindlimb is faded to brownish but still recognizable. An indistinct whitish midventral line is recognizable. Several blue scales on the dorsal plate of the head are the only colourful remnants of the life colouration. Hemipenis uniformly whitish.
|
||
</paragraph>
|
||
<caption id="DF2C14E1117F417CD5BAFCF94C3B3200" httpUri="https://zenodo.org/record/190912/files/figure.png" pageId="4" pageNumber="36" targetBox="[178,1416,203,778]" targetPageId="4">
|
||
<paragraph id="8BEC4469117F417CD5BAFCF94C3B3200" blockId="4.[151,1436,813,871]" pageId="4" pageNumber="36">
|
||
<emphasis id="B927987B117F417CD5BAFCF94E693222" bold="true" box="[151,281,813,837]" pageId="4" pageNumber="36">FIGURE 3</emphasis>
|
||
. Drawings of hemipenis of the male holotype of
|
||
<taxonomicName id="4C533FEA117F417CD67FFCF94B713223" box="[850,1025,813,836]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B117F417CD67FFCF94B713223" box="[850,1025,813,836]" italics="true" pageId="4" pageNumber="36">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2142500117F417CD124FCF94B153222" box="[1033,1125,813,837]" pageId="4" pageNumber="36" rank="species">
|
||
<emphasis id="B927987B117F417CD124FCF94B153222" bold="true" box="[1033,1125,813,837]" pageId="4" pageNumber="36">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ZSM 2103/2007 (by Ruth Kühbandner): (A) asulcal view, (B) lateral view; (C) sulcal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC4469117F417CD5E8FC414CB635D9" blockId="4.[151,1436,917,1446]" pageId="4" pageNumber="36">
|
||
In life when unstressed (
|
||
<figureCitation id="136858EC117F417CD4C3FC414D4632C8" box="[494,566,917,943]" captionStart="FIGURE 1" captionStartId="2.[151,254,1850,1874]" captionTargetBox="[157,1426,296,1817]" captionTargetId="figure@2.[157,1431,293,1825]" captionTargetPageId="2" captionText="FIGURE 1. Male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 in life, (A) lateral view, (B) dorsolateral view of head; and (C) additional male of the same species photographed near the type locality by P. Schönecker (not collected)." httpUri="https://zenodo.org/record/190910/files/figure.png" pageId="4" pageNumber="36">Fig. 1</figureCitation>
|
||
), dorsal and lateral colouration of head, body, limbs, and tail uniformly lime green, with a whitish-beige oblique stripe on the anterior flanks, starting several millimeters below the seventh dorsal tubercle and ending several millimeters above a whitish lateroventral band of rosette-like scales that runs along the flanks between the insertion of fore- and hindlimbs. A white band along the whole upper lip starting at mouth angle. Eye ball with a characteristic colouration: upper half green, lower half with a light brown outer ring and a blue inner ring, eye opening encircled by a complete, narrow yellowish-orange ring. Lateral crest light brown. Head dorsally with numerous blue scales. Ventral surfaces of head yellowishgreen, body ventrally greyish-green, inner surface of limbs light grey.
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117F417CD5EBFB1E4C7634C1" blockId="4.[151,1436,917,1446]" pageId="4" pageNumber="36">
|
||
<emphasis id="B927987B117F417CD5EBFB1E4D663583" bold="true" box="[198,534,1226,1252]" pageId="4" pageNumber="36">
|
||
Variation of the
|
||
<typeStatus id="54E8FACB117F417CD4BEFB1E4D613583" box="[403,529,1226,1252]" pageId="4" pageNumber="36" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
For measurements and character conditions of the
|
||
<typeStatus id="54E8FACB117F417CD15FFB1E4BD53583" box="[1138,1189,1226,1252]" pageId="4" pageNumber="36">type</typeStatus>
|
||
specimens see
|
||
<tableCitation id="C6D171D2117F417CD076FB1E4FD7346C" captionStart="TABLE 1" captionStartId="4.[151,239,1497,1521]" captionText="TABLE 1. Morphological variation in Furcifer timoni sp. nov. All measurements in mm." pageId="4" pageNumber="36">Table 1</tableCitation>
|
||
. Females of
|
||
<taxonomicName id="4C533FEA117F417CD46FFB254EDD346D" box="[322,429,1265,1290]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117F417CD46FFB254EDD346D" box="[322,429,1265,1290]" italics="true" pageId="4" pageNumber="36">F. timoni</emphasis>
|
||
</taxonomicName>
|
||
lack the rostral appendages of the male and have a distinctly less swollen tail base. Furthermore, the relative tail length of the male
|
||
<typeStatus id="54E8FACB117F417CD7E8FACC4C5C3455" box="[709,812,1304,1330]" pageId="4" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
is longer than in the females (
|
||
<tableCitation id="C6D171D2117F417CD1A4FACC4BAF3455" box="[1161,1247,1304,1330]" captionStart="TABLE 1" captionStartId="4.[151,239,1497,1521]" captionText="TABLE 1. Morphological variation in Furcifer timoni sp. nov. All measurements in mm." pageId="4" pageNumber="36">Table 1</tableCitation>
|
||
) which were all gravid with fully developed, yellowish eggs. The colouration of the female
|
||
<typeStatus id="54E8FACB117F417CD125FAEB4B08343E" box="[1032,1144,1343,1369]" pageId="4" pageNumber="36" type="paratype">paratypes</typeStatus>
|
||
in preservative is almost uniformly blackish. The light ventrolateral bands and the whitish midventral line are more distinct than in the
|
||
<typeStatus id="54E8FACB117F417CD5BAFA584E7234C1" box="[151,258,1420,1446]" pageId="4" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
. Bluish spots are recognizable on the heads.
|
||
</paragraph>
|
||
<caption id="DF2C14E1117F417CD5BAFA0D4B3F3497" box="[151,1103,1497,1521]" pageId="4" pageNumber="36">
|
||
<paragraph id="8BEC4469117F417CD5BAFA0D4B3F3497" blockId="4.[151,1369,1497,1955]" box="[151,1103,1497,1521]" pageId="4" pageNumber="36">
|
||
<emphasis id="B927987B117F417CD5BAFA0D4E793496" bold="true" box="[151,265,1497,1521]" pageId="4" pageNumber="36">TABLE 1.</emphasis>
|
||
Morphological variation in
|
||
<taxonomicName id="4C533FEA117F417CD71BFA0E4DA93496" box="[566,729,1498,1521]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B117F417CD71BFA0E4DA93496" box="[566,729,1498,1521]" italics="true" pageId="4" pageNumber="36">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2142500117F417CD7F2FA0D4C443496" box="[735,820,1497,1521]" pageId="4" pageNumber="36" rank="species">
|
||
<emphasis id="B927987B117F417CD7F2FA0D4C443496" bold="true" box="[735,820,1497,1521]" pageId="4" pageNumber="36">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
All measurements in mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC4469117F417CD4CDF9D84A383744" blockId="4.[151,1369,1497,1955]" box="[480,1352,1548,1571]" pageId="4" pageNumber="36">ZSM 2103/2007 ZSM 255/2004 ZSM 256/2004 ZFMK 87585</paragraph>
|
||
<paragraph id="8BEC4469117F417CD5B2F9E24BAC37AA" blockId="4.[151,1369,1497,1955]" pageId="4" pageNumber="36">
|
||
Status
|
||
<typeStatus id="54E8FACB117F417CD4CDF9E24D4E372A" box="[480,574,1590,1613]" pageId="4" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
<typeStatus id="54E8FACB117F417CD7FFF9E24A60372A" box="[722,1296,1590,1613]" pageId="4" pageNumber="36" type="paratype">paratype paratype paratype</typeStatus>
|
||
Sex male (adult) female (gravid) female (gravid) female (gravid) SVL 88.0 96.0 94.5 95.5 Total length 214 194 195 198
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117F417CD5B2F9354BA6362A" blockId="4.[151,1369,1497,1955]" pageId="4" pageNumber="36">head length 28.7 27.3 26.7 27.5 snout length 20.8 19.6 18.9 19.3 eye diameter 8.7 7.8 7.0 7.6</paragraph>
|
||
<paragraph id="8BEC4469117F417CD5B2F8B54BBF36C4" blockId="4.[151,1369,1497,1955]" pageId="4" pageNumber="36">
|
||
number of dorsal tubercles ca. 12 ca. 10 ca. 12 ca.
|
||
<specimenCount id="9D558FE0117F417CD1F4F8B54FB536C4" pageId="4" pageNumber="36" type="egg">9 egg</specimenCount>
|
||
number in body cavity ---
|
||
<date id="FFED62A9117F417CD7FFF8584BBF36C4" box="[722,1231,1932,1955]" pageId="4" pageNumber="36" value="1913-10-14">14 10 13</date>
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117E417DD5E8FF4C4C6C33CF" blockId="5.[151,1436,152,680]" pageId="5" pageNumber="37">
|
||
During the day the ground colouration of apparently unstressed gravid females in life is green with a pattern of green or brown, mainly transversal bands (
|
||
<figureCitation id="136858EC117E417DD60CFF6A4C1B31BF" box="[801,875,190,216]" captionStart="FIGURE 4" captionStartId="5.[151,254,2012,2036]" captionTargetBox="[156,1429,719,1984]" captionTargetId="figure@5.[153,1434,697,1988]" captionTargetPageId="5" captionText="FIGURE 4. Gravid female paratype of Furcifer timoni sp. nov., ZSM uncatalogued (FGZC 1884) in life." httpUri="https://zenodo.org/record/190913/files/figure.png" pageId="5" pageNumber="37">Fig. 4</figureCitation>
|
||
) on the flanks and sides of the head. Thinner, regularly shaped and regularly spaced transversal bands are also present on tail and legs. A red band starting from middorsum or slightly below runs posterioventrally until the middle of the flanks. The ventrolateral band between fore- and hindlimbs is reddish (
|
||
<figureCitation id="136858EC117E417DD75AFEE64DB1302B" box="[631,705,306,332]" captionStart="FIGURE 4" captionStartId="5.[151,254,2012,2036]" captionTargetBox="[156,1429,719,1984]" captionTargetId="figure@5.[153,1434,697,1988]" captionTargetPageId="5" captionText="FIGURE 4. Gravid female paratype of Furcifer timoni sp. nov., ZSM uncatalogued (FGZC 1884) in life." httpUri="https://zenodo.org/record/190913/files/figure.png" pageId="5" pageNumber="37">Fig. 4</figureCitation>
|
||
). The posterior part of the head is covered by a large more or less triangular red spot which is strongly bordered posteriorly but fades less distinct anteriorly. The head (including the red spot) is covered with many blue scales and similar blue scales, which are partly arranged in longitudinal rows, are scattered along the flanks, decreasing in density posteriorly. There is no distinct white band along the lip, but a whitish spot on the upper lip is recognizable below the eye. The eye ball is similar but less distinctly coloured than in males, usually without distinct blue colour. At night when roosting the body is uniformely green with only indistinct transversal banding pattern, but with distinct diagonal red band on the flanks and triangular red spot on the head. Only few bluish scales are recognizable on the head and no distinctly blue scales on the flanks. The eye ball is brown in the lower half, but without blue colour. When the throat is inflated a reddish reticulated pattern is visible.
|
||
</paragraph>
|
||
<caption id="DF2C14E1117E417DD5BAF8084B8B3694" box="[151,1275,2012,2036]" httpUri="https://zenodo.org/record/190913/files/figure.png" pageId="5" pageNumber="37" targetBox="[156,1429,719,1984]" targetPageId="5">
|
||
<paragraph id="8BEC4469117E417DD5BAF8084B8B3694" blockId="5.[151,1275,2012,2036]" box="[151,1275,2012,2036]" pageId="5" pageNumber="37">
|
||
<emphasis id="B927987B117E417DD5BAF8084E613693" bold="true" box="[151,273,2012,2036]" pageId="5" pageNumber="37">FIGURE 4</emphasis>
|
||
. Gravid female paratype of
|
||
<taxonomicName id="4C533FEA117E417DD713F8094D913693" box="[574,737,2013,2036]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B117E417DD713F8094D913693" box="[574,737,2013,2036]" italics="true" pageId="5" pageNumber="37">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2142500117E417DD7C5F8084C4D3693" box="[744,829,2012,2036]" pageId="5" pageNumber="37" rank="species">
|
||
<emphasis id="B927987B117E417DD7C5F8084C4D3693" bold="true" box="[744,829,2012,2036]" pageId="5" pageNumber="37">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ZSM uncatalogued (FGZC 1884) in life.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C14E1117D417ED5BAFC054D95356B" httpUri="https://zenodo.org/record/190914/files/figure.png" pageId="6" pageNumber="38" targetBox="[176,1422,192,939]" targetPageId="6">
|
||
<paragraph id="8BEC4469117D417ED5BAFC054D95356B" blockId="6.[151,1437,977,1037]" pageId="6" pageNumber="38">
|
||
<emphasis id="B927987B117D417ED5BAFC054E68328E" bold="true" box="[151,280,977,1001]" pageId="6" pageNumber="38">FIGURE 5</emphasis>
|
||
. (A) Preserved male holotype of
|
||
<taxonomicName id="4C533FEA117D417ED7B9FC064C36328E" box="[660,838,978,1001]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117D417ED7B9FC064C36328E" box="[660,838,978,1001]" italics="true" pageId="6" pageNumber="38">Furcifer bifidus</emphasis>
|
||
</taxonomicName>
|
||
(MNHN 6660) in comparison to (B) preserved male holotype of
|
||
<taxonomicName id="4C533FEA117D417ED435FC214ECA356B" box="[280,442,1013,1036]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="B927987B117D417ED435FC214ECA356B" box="[280,442,1013,1036]" italics="true" pageId="6" pageNumber="38">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2142500117D417ED4ECFC214D66356A" box="[449,534,1013,1037]" pageId="6" pageNumber="38" rank="species">
|
||
<emphasis id="B927987B117D417ED4ECFC214D66356A" bold="true" box="[449,534,1013,1037]" pageId="6" pageNumber="38">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(ZSM 2103/2007).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C14E1117D417ED5BAF8074BC2368C" box="[151,1202,2003,2027]" httpUri="https://zenodo.org/record/190915/files/figure.png" pageId="6" pageNumber="38" targetBox="[188,1398,1090,1979]" targetPageId="6">
|
||
<paragraph id="8BEC4469117D417ED5BAF8074BC2368C" blockId="6.[151,1202,2003,2027]" box="[151,1202,2003,2027]" pageId="6" pageNumber="38">
|
||
<emphasis id="B927987B117D417ED5BAF8074E61368C" bold="true" box="[151,273,2003,2027]" pageId="6" pageNumber="38">FIGURE 6</emphasis>
|
||
. Female
|
||
<taxonomicName id="4C533FEA117D417ED459F8004D49368C" box="[372,569,2004,2027]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117D417ED459F8004EBC368C" box="[372,460,2004,2027]" italics="true" pageId="6" pageNumber="38">Furcifer</emphasis>
|
||
cf.
|
||
<emphasis id="B927987B117D417ED4D8F8004D49368C" box="[501,569,2004,2027]" italics="true" pageId="6" pageNumber="38">timoni</emphasis>
|
||
</taxonomicName>
|
||
from Marojejy National Park (Photograph by G. Gomboc).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC4469117C417FD5EBFF434A683080" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
<emphasis id="B927987B117C417FD5EBFF434D1231D6" bold="true" box="[198,610,151,177]" pageId="7" pageNumber="39">Additional photographic records</emphasis>
|
||
. Photographs of one or two males of unknown origin were published in a Japanese chameleon book (
|
||
<bibRefCitation id="EFC23998117C417FD73FFF6A4D9F31BF" author="Kimura" box="[530,751,190,216]" pageId="7" pageNumber="42" refString="Kimura, H. (2003) A complete guide of chameleons. Terra Pro, Terrarium pro book of reptiles & amphibians Vol. 1, 196 pp. (Pisces Publisher)" type="book" year="2003">Kimura 2003: 114</bibRefCitation>
|
||
) and might refer to
|
||
<taxonomicName id="4C533FEA117C417FD6CDFF6B4B3A31BF" box="[992,1098,191,216]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD6CDFF6B4B3A31BF" box="[992,1098,191,216]" italics="true" pageId="7" pageNumber="39">F. timoni</emphasis>
|
||
</taxonomicName>
|
||
. Two additional males from Montagne d'Ambre were photographed by H.-P. Berghof and P. Schönecker (e.g.,
|
||
<figureCitation id="136858EC117C417FD141FF314BDF3198" box="[1132,1199,229,255]" captionStart="FIGURE 1" captionStartId="2.[151,254,1850,1874]" captionTargetBox="[157,1426,296,1817]" captionTargetId="figure@2.[157,1431,293,1825]" captionTargetPageId="2" captionText="FIGURE 1. Male holotype of Furcifer timoni sp. nov., ZSM 2103 / 2007 in life, (A) lateral view, (B) dorsolateral view of head; and (C) additional male of the same species photographed near the type locality by P. Schönecker (not collected)." httpUri="https://zenodo.org/record/190910/files/figure.png" pageId="7" pageNumber="39">Fig. 1</figureCitation>
|
||
c), respectively. All these photographs strongly resemble the
|
||
<typeStatus id="54E8FACB117C417FD75AFED84DAE3041" box="[631,734,268,294]" pageId="7" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C533FEA117C417FD628FED84CC83042" box="[773,952,268,293]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD628FED84CC83042" box="[773,952,268,293]" italics="true" pageId="7" pageNumber="39">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
in morphology and principal colouration and strongly confirm that it represents a distinctive new species and not a subadult or aberrant specimen of the green colour morph of
|
||
<taxonomicName id="4C533FEA117C417FD48FFE8D4D2A3015" box="[418,602,345,370]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117C417FD48FFE8D4D2A3015" box="[418,602,345,370]" italics="true" pageId="7" pageNumber="39">Furcifer bifidus</emphasis>
|
||
</taxonomicName>
|
||
as shown in
|
||
<bibRefCitation id="EFC23998117C417FD7DDFE8D4B4E3014" author="Martin" box="[752,1086,345,371]" pageId="7" pageNumber="42" refString="Martin, J. & Wolfe, A. (1992) Masters of disguise: A natural history of chameleons. Facts on file, New York, 176 pp." type="book" year="1992">Martin & Wolfe (1992: 139)</bibRefCitation>
|
||
. Similarly, the photographs of females taken from the pet trade (
|
||
<bibRefCitation id="EFC23998117C417FD714FE544DAF30FD" author="Kimura" box="[569,735,384,410]" pageId="7" pageNumber="42" refString="Kimura, H. (2003) A complete guide of chameleons. Terra Pro, Terrarium pro book of reptiles & amphibians Vol. 1, 196 pp. (Pisces Publisher)" type="book" year="2003">Kimura 2003</bibRefCitation>
|
||
and own photographs of a specimen taken in the 1990s) resemble those from Montagne d'Ambre. Unstressed gravid females appear to be largely green, with a network pattern of brown bands which are principally arranged similarly among different females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34917E2117C417FD5EBFE274FAC3353" pageId="7" pageNumber="39" type="etymology">
|
||
<paragraph id="8BEC4469117C417FD5EBFE274FAC3353" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
<emphasis id="B927987B117C417FD5EBFE274E23336A" bold="true" box="[198,339,499,525]" pageId="7" pageNumber="39">Etymology.</emphasis>
|
||
The senior author wishes to dedicate this new chameleon species to his son Timon Robert Glaw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34917E2117C417FD5EBFD944D87372F" pageId="7" pageNumber="39" type="distribution">
|
||
<paragraph id="8BEC4469117C417FD5EBFD944F82354C" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
<emphasis id="B927987B117C417FD5EBFD944D2E333D" bold="true" box="[198,606,576,602]" pageId="7" pageNumber="39">Distribution, habitat and habits.</emphasis>
|
||
<taxonomicName id="4C533FEA117C417FD74AFD954C6B333D" box="[615,795,577,602]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD74AFD954C6B333D" box="[615,795,577,602]" italics="true" pageId="7" pageNumber="39">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Montagne d’Ambre National Park, between 750 and
|
||
<quantity id="4CABE98C117C417FD49FFDBC4E8E33E5" box="[434,510,616,642]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="39" unit="m" value="900.0">900 m</quantity>
|
||
altitude. The discovery of this distinctive new
|
||
<taxonomicName id="4C533FEA117C417FD114FDBC4BEC33E6" box="[1081,1180,616,641]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="genus">
|
||
<emphasis id="B927987B117C417FD114FDBC4BEC33E6" box="[1081,1180,616,641]" italics="true" pageId="7" pageNumber="39">Furcifer</emphasis>
|
||
</taxonomicName>
|
||
species in Montagne d’Ambre was very surprising since this area has been repeatedly and intensively surveyed for reptiles over many years (
|
||
<bibRefCitation id="EFC23998117C417FD410FD614D7F33A8" author="Mocquard" box="[317,527,693,719]" pageId="7" pageNumber="42" refString="Mocquard, M. F. (1895) Sur les reptiles recueillis a Madagascar par M. M. Alluaud et Belly. Bulletin de la Societe Philomatique de Paris, ser. 8, 7, 112 - 136." type="journal article" year="1895">Mocquard 1895</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC23998117C417FD733FD614C6833A8" author="Ramanantsoa" box="[542,792,693,719]" pageId="7" pageNumber="42" refString="Ramanantsoa, G. A. (1974) Connaissance des Cameleonides communs de la Province de Diego-Suarez par la Population Paysanne. Bulletin de l'Academie Malgache, 51 (1973), 147 - 149." type="journal article" year="1974">Ramanantsoa 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC23998117C417FD60AFD614C8233A8" author="Andreone" box="[807,1010,693,719]" pageId="7" pageNumber="41" refString="Andreone, F. (1991) Reptile field records from Malagasy rainforests. Bollettino del Museo Regionale di Scienze Naturali di Torino, 9, 437 - 460." type="journal article" year="1991">Andreone 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC23998117C417FD12CFD614BAF33A8" author="Kauffmann" box="[1025,1247,693,719]" pageId="7" pageNumber="42" refString="Kauffmann, J. (1994) Veldwaarnemingen aan Calumma brevicornis en C. oshaughnessyi ambreensis op Madagascar. Lacerta, 53, 2 - 10." type="journal article" year="1994">Kauffmann 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC23998117C417FD1C3FD614E283391" author="Raxworthy" pageId="7" pageNumber="42" refString="Raxworthy, C. J. & Nussbaum, R. A. (1994) A rainforest survey of amphibians, reptiles and small mammals at Montagne d'Ambre, Madagascar. Biological Conservation, 69, 65 - 73." type="journal article" year="1994">Raxworthy & Nussbaum 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC23998117C417FD44EFD084D103391" author="Glaw" box="[355,608,732,758]" pageId="7" pageNumber="41" refString="Glaw, F. & Vences, M. (1994) A fieldguide to the amphibians and reptiles of Madagascar, 2 nd edition. Vences & Glaw Verlag, Koln, 480 pp. + 48 colour pp." type="book" year="1994">Glaw & Vences 1994</bibRefCitation>
|
||
; D'Cruze
|
||
<emphasis id="B927987B117C417FD7FEFD084C7D3392" box="[723,781,732,757]" italics="true" pageId="7" pageNumber="39">et al.</emphasis>
|
||
2008). Apart from the
|
||
<typeStatus id="54E8FACB117C417FD136FD084B3E3391" box="[1051,1102,732,758]" pageId="7" pageNumber="39">type</typeStatus>
|
||
locality the species possibly occurs in the Marojejy National Park (
|
||
<figureCitation id="136858EC117C417FD74DFCD64DD7327B" box="[608,679,770,796]" captionStart="FIGURE 6" captionStartId="6.[151,254,2003,2027]" captionTargetBox="[188,1398,1090,1979]" captionTargetId="figure@6.[188,1398,1090,1979]" captionTargetPageId="6" captionText="FIGURE 6. Female Furcifer cf. timoni from Marojejy National Park (Photograph by G. Gomboc)." httpUri="https://zenodo.org/record/190915/files/figure.png" pageId="7" pageNumber="39">Fig. 6</figureCitation>
|
||
) as well, since photographs of a female taken by G. Gomboc at Marojejy (between Camp 1 and Camp 2) resemble females of
|
||
<taxonomicName id="4C533FEA117C417FD6A0FCFD4C893225" box="[909,1017,809,834]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD6A0FCFD4C893225" box="[909,1017,809,834]" italics="true" pageId="7" pageNumber="39">F. timoni</emphasis>
|
||
</taxonomicName>
|
||
and might represent this or a very closely related species. It differs from the studied
|
||
<typeStatus id="54E8FACB117C417FD7CFFC844C65320D" box="[738,789,848,874]" pageId="7" pageNumber="39">type</typeStatus>
|
||
specimens (see
|
||
<tableCitation id="C6D171D2117C417FD6F8FC844B5A320D" box="[981,1066,848,874]" captionStart="TABLE 1" captionStartId="4.[151,239,1497,1521]" captionText="TABLE 1. Morphological variation in Furcifer timoni sp. nov. All measurements in mm." pageId="7" pageNumber="39">Table 1</tableCitation>
|
||
) by a dorsal crest composed of more (ca. 20) tubercles (see
|
||
<figureCitation id="136858EC117C417FD4CEFCA24D5A32F7" box="[483,554,886,912]" captionStart="FIGURE 6" captionStartId="6.[151,254,2003,2027]" captionTargetBox="[188,1398,1090,1979]" captionTargetId="figure@6.[188,1398,1090,1979]" captionTargetPageId="6" captionText="FIGURE 6. Female Furcifer cf. timoni from Marojejy National Park (Photograph by G. Gomboc)." httpUri="https://zenodo.org/record/190915/files/figure.png" pageId="7" pageNumber="39">Fig. 6</figureCitation>
|
||
). Similar to the situation at Montagne d'Ambre, all previous herpetological surveys in the Marojejy National Park since 1968 (see summary in
|
||
<bibRefCitation id="EFC23998117C417FD694FC494B9832D0" author="Raselimanana" box="[953,1256,925,951]" pageId="7" pageNumber="42" refString="Raselimanana, A. P., Raxworthy, C. J. & Nussbaum, R. A. (2000) Herpetofaunal species diversity and elevational distribution within the Parc National de Marojejy, Madagascar. In: A floral and faunal inventory of the Parc National de Marojejy: with reference to elevational variation (S. M. Goodman ed.). Fieldiana Zoology (new series), 97, 157 - 174." type="journal article" year="2000">
|
||
Raselimanana
|
||
<emphasis id="B927987B117C417FD14AFC494BD332D1" box="[1127,1187,925,950]" italics="true" pageId="7" pageNumber="39">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
) did not reveal any record of a chameleon resembling
|
||
<emphasis id="B927987B117C417FD770FC104DBD32BA" box="[605,717,964,989]" italics="true" pageId="7" pageNumber="39">
|
||
<taxonomicName id="4C533FEA117C417FD770FC104DB832BA" box="[605,712,964,989]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">F. timoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
indicating that both the Montagne d'Ambre and the Marojejy populations had been overlooked, possibly due to them occupying a cryptic niche hidden high up in the tree canopy.
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117C417FD5E8FBEC4D9E345D" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
At Montagne d'Ambre,
|
||
<taxonomicName id="4C533FEA117C417FD4CFFBEC4DE83536" box="[482,664,1080,1105]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD4CFFBEC4DE83536" box="[482,664,1080,1105]" italics="true" pageId="7" pageNumber="39">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
was exclusively found in mid-altitude primary rainforest during the rainy season, in close syntopy with
|
||
<taxonomicName id="4C533FEA117C417FD75DFB8B4C58351F" box="[624,808,1119,1144]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="petteri">
|
||
<emphasis id="B927987B117C417FD75DFB8B4C58351F" box="[624,808,1119,1144]" italics="true" pageId="7" pageNumber="39">Furcifer petteri</emphasis>
|
||
</taxonomicName>
|
||
. All specimens we captured in the years
|
||
<date id="FFED62A9117C417FD03BFB8A4AEC351F" box="[1302,1436,1118,1144]" pageId="7" pageNumber="39" value="2004" valueMax="2008">2004–2008</date>
|
||
were found in the period
|
||
<date id="FFED62A9117C417FD490FB514D0835F8" box="[445,632,1157,1183]" pageId="7" pageNumber="39" value="1920-02-26">20–26 February</date>
|
||
, roosting at night in the vegetation. The only male seen and captured during the same period was captured from a position more than
|
||
<quantity id="4CABE98C117C417FD68EFB784CA235A1" box="[931,978,1196,1222]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="39" unit="m" value="3.0">3 m</quantity>
|
||
above the ground. In contrast, gravid females were relatively common in late February, usually sitting on branches ca.
|
||
<quantity id="4CABE98C117C417FD171FB064BCE358B" box="[1116,1214,1234,1260]" metricMagnitude="0" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="7" pageNumber="39" unit="m" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 m</quantity>
|
||
above the ground. We suspect that they were just descending from the trees to bury their eggs into the ground. Data on egg numbers of preserved females are given in
|
||
<tableCitation id="C6D171D2117C417FD7BDFAF44D98345D" box="[656,744,1312,1338]" captionStart="TABLE 1" captionStartId="4.[151,239,1497,1521]" captionText="TABLE 1. Morphological variation in Furcifer timoni sp. nov. All measurements in mm." pageId="7" pageNumber="39">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117C417FD5EBFA924D87372F" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
<emphasis id="B927987B117C417FD5EBFA924EEA3407" bold="true" box="[198,410,1350,1376]" pageId="7" pageNumber="39">Available names.</emphasis>
|
||
Among the
|
||
<collectingCountry id="F34404F9117C417FD703FA924DD03407" box="[558,672,1350,1376]" name="Madagascar" pageId="7" pageNumber="39">Malagasy</collectingCountry>
|
||
chameleons with paired bony rostral appendages in males, there is only one taxon,
|
||
<taxonomicName id="4C533FEA117C417FD45CFAB94D0134E1" box="[369,625,1389,1414]" class="Reptilia" family="Chamaeleonidae" genus="Dicranosaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifurca">
|
||
<emphasis id="B927987B117C417FD45CFAB94D0134E1" box="[369,625,1389,1414]" italics="true" pageId="7" pageNumber="39">Dicranosaura bifurca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C533FEA117C417FD754FAB94CB634E0" authority="Gray, 1864" authorityName="Gray" authorityYear="1864" box="[633,966,1389,1415]" pageId="7" pageNumber="39" rank="variety" variety="crassicornis">
|
||
var.
|
||
<emphasis id="B927987B117C417FD781FAB94C4B34E1" box="[684,827,1389,1414]" italics="true" pageId="7" pageNumber="39">crassicornis</emphasis>
|
||
<bibRefCitation id="EFC23998117C417FD66EFAB94CB634E0" author="Gray" box="[835,966,1389,1415]" pageId="7" pageNumber="41" refString="Gray, J. E. (1864) Revision of the genera and species of Chamaeleonidae, with the description of some new species. Proceedings of the Zoological Society of London, 30, 465 - 477." type="journal article" year="1864">Gray, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
that needs to be discussed as a possible earlier name for
|
||
<taxonomicName id="4C533FEA117C417FD476FA404D7F34CA" box="[347,527,1428,1453]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B117C417FD476FA404D7F34CA" box="[347,527,1428,1453]" italics="true" pageId="7" pageNumber="39">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
(the other synonyms of
|
||
<taxonomicName id="4C533FEA117C417FD61DFA404C9934CA" box="[816,1001,1428,1453]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117C417FD61DFA404C9934CA" box="[816,1001,1428,1453]" italics="true" pageId="7" pageNumber="39">Furcifer bifidus</emphasis>
|
||
</taxonomicName>
|
||
are replacement names due to errors and therefore not available).
|
||
<bibRefCitation id="EFC23998117C417FD4CBFA6E4DE334B3" author="Brygoo" box="[486,659,1466,1492]" pageId="7" pageNumber="41" refString="Brygoo, E. R. (1971) Reptiles Sauriens Chamaeleonidae - Genre Chamaeleo. Faune de Madagascar, 33, 1 - 318." type="journal article" year="1971">Brygoo (1971)</bibRefCitation>
|
||
considered the possibility that
|
||
<emphasis id="B927987B117C417FD6D0FA6F4BFC34B3" box="[1021,1164,1467,1492]" italics="true" pageId="7" pageNumber="39">crassicornis</emphasis>
|
||
could be a synonym of
|
||
<taxonomicName id="4C533FEA117C417FD5BAFA354E5A349D" box="[151,298,1505,1530]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="balteatus">
|
||
<emphasis id="B927987B117C417FD5BAFA354E5A349D" box="[151,298,1505,1530]" italics="true" pageId="7" pageNumber="39">F. balteatus</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C533FEA117C417FD4E9FA354D49349D" box="[452,569,1505,1530]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B117C417FD4E9FA354D49349D" box="[452,569,1505,1530]" italics="true" pageId="7" pageNumber="39">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
, but the
|
||
<typeStatus id="54E8FACB117C417FD786FA354D91349C" box="[683,737,1505,1531]" pageId="7" pageNumber="39">type</typeStatus>
|
||
specimens were apparently never identified or studied. Unfortunately, the original description of this taxon is very short, imprecise and difficult to interpret and we here repeat the whole description (
|
||
<bibRefCitation id="EFC23998117C417FD706F9FA4D9B372F" author="Gray" box="[555,747,1582,1608]" pageId="7" pageNumber="41" refString="Gray, J. E. (1864) Revision of the genera and species of Chamaeleonidae, with the description of some new species. Proceedings of the Zoological Society of London, 30, 465 - 477." type="journal article" year="1864">Gray 1864: 479</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34917E2117C4170D5BAF9814B64345D" lastPageId="8" lastPageNumber="40" pageId="7" pageNumber="39" type="description">
|
||
<paragraph id="8BEC4469117C417FD5BAF9814EC33708" blockId="7.[151,1437,151,2034]" box="[151,435,1621,1647]" pageId="7" pageNumber="39">
|
||
"
|
||
<taxonomicName id="4C533FEA117C417FD5B2F9814E173709" box="[159,359,1621,1647]" pageId="7" pageNumber="42" rank="variety" variety="crassicornis">
|
||
Var.
|
||
<emphasis id="B927987B117C417FD5F5F9814E173709" box="[216,359,1621,1646]" italics="true" pageId="7" pageNumber="39">crassicornis</emphasis>
|
||
</taxonomicName>
|
||
B. M.
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117C417FD5EBF9A84D223784" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
One of the males, with the horns only partly developed, has them very thick and trigonal at the base, so as nearly to reach across the nose. In another young male, about the same size, they are compressed and far apart at the base, as in the
|
||
<typeStatus id="54E8FACB117C417FD4A7F91D4ECD3784" box="[394,445,1737,1763]" pageId="7" pageNumber="39">type</typeStatus>
|
||
specimens."
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117C417FD5EBF9244B5036AC" blockId="7.[151,1437,151,2034]" pageId="7" pageNumber="39">
|
||
Colin McCarthy (in litt.,
|
||
<date id="FFED62A9117C417FD4C6F9244C6C366D" box="[491,796,1776,1802]" pageId="7" pageNumber="39" value="2009-07-22" valueMax="2009-08-24" valueMin="2009-07-22">22 July and 24 Aug. 2009</date>
|
||
) kindly provided us the following information: "The situation regarding the
|
||
<typeStatus id="54E8FACB117C417FD4CAF8C24D553657" box="[487,549,1814,1840]" pageId="7" pageNumber="39">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C533FEA117C417FD761F8C34C3B3657" box="[588,843,1815,1840]" class="Reptilia" family="Chamaeleonidae" genus="Dicranosaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifurca">
|
||
<emphasis id="B927987B117C417FD761F8C34C3B3657" box="[588,843,1815,1840]" italics="true" pageId="7" pageNumber="39">Dicranosaura bifurca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C533FEA117C417FD67EF8C24BEF3657" authority="Gray, 1864" authorityName="Gray" authorityYear="1864" box="[851,1183,1814,1840]" pageId="7" pageNumber="39" rank="variety" variety="crassicornis">
|
||
var.
|
||
<emphasis id="B927987B117C417FD6A8F8C34B643657" box="[901,1044,1815,1840]" italics="true" pageId="7" pageNumber="39">crassicornis</emphasis>
|
||
<bibRefCitation id="EFC23998117C417FD131F8C24BEF3657" author="Gray" box="[1052,1183,1814,1840]" pageId="7" pageNumber="41" refString="Gray, J. E. (1864) Revision of the genera and species of Chamaeleonidae, with the description of some new species. Proceedings of the Zoological Society of London, 30, 465 - 477." type="journal article" year="1864">Gray, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
is unclear. We do not appear to have distinguished the
|
||
<typeStatus id="54E8FACB117C417FD731F8E94D2A3630" box="[540,602,1853,1879]" pageId="7" pageNumber="39">types</typeStatus>
|
||
of this variety in our collection. What we have are the four specimens of
|
||
<emphasis id="B927987B117C417FD594F8B04EC9361A" box="[185,441,1892,1917]" italics="true" pageId="7" pageNumber="39">
|
||
'
|
||
<taxonomicName id="4C533FEA117C417FD590F8B04EC9361A" box="[189,441,1892,1917]" class="Reptilia" family="Chamaeleonidae" genus="Chamaeleo" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="bifurcus">Chamaeleo bifurcus</taxonomicName>
|
||
</emphasis>
|
||
' mentioned in Gray's (1845) lizard catalogue to which I assume could be added BMNH 1863.4.16.1 which appears to have been the only other specimen of this taxon added to the collections before Gray's paper of 1864. So the relevant specimens for consideration are:
|
||
</paragraph>
|
||
<paragraph id="8BEC4469117C4170D5BAF80C4C3131D5" blockId="7.[151,1437,151,2034]" lastBlockId="8.[151,1437,152,1338]" lastPageId="8" lastPageNumber="40" pageId="7" pageNumber="39">
|
||
BMNH xxv.12a
|
||
<collectingCountry id="F34404F9117C417FD470F80C4E9D3695" box="[349,493,2008,2034]" name="Madagascar" pageId="7" pageNumber="39">Madagascar</collectingCountry>
|
||
[SVL:
|
||
<quantity id="4CABE98C117C417FD765F80C4DDD3695" box="[584,685,2008,2034]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="39" unit="mm" value="170.0">170 mm</quantity>
|
||
, tail length:
|
||
<quantity id="4CABE98C117C417FD669F80C4CDA3695" box="[836,938,2008,2034]" metricMagnitude="-1" metricUnit="m" metricValue="2.33" pageId="7" pageNumber="39" unit="mm" value="233.0">233 mm</quantity>
|
||
, rostral appendage (length, ventral edge):
|
||
<quantity id="4CABE98C11734170D5BAFF4C4E7231D5" box="[151,258,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="8" pageNumber="40" unit="mm" value="26.2">26.2 mm</quantity>
|
||
, rostral appendage length (projection):
|
||
<quantity id="4CABE98C11734170D7E3FF4C4C4A31D5" box="[718,826,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="8" pageNumber="40" unit="mm" value="23.4">23.4 mm</quantity>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFF6A4CBD3198" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
BMNH xxv.12b
|
||
<collectingCountry id="F34404F911734170D470FF6A4E9B31BF" box="[349,491,190,216]" name="Madagascar" pageId="8" pageNumber="40">Madagascar</collectingCountry>
|
||
/ Pres: J.E. Gray [SVL:
|
||
<quantity id="4CABE98C11734170D623FF6A4C1531BF" box="[782,869,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="8" pageNumber="40" unit="mm" value="97.0">97 mm</quantity>
|
||
, tail length:
|
||
<quantity id="4CABE98C11734170D6D4FF6A4B2D31BF" box="[1017,1117,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="8" pageNumber="40" unit="mm" value="119.0">119 mm</quantity>
|
||
, rostral appendage (length, ventral edge):
|
||
<quantity id="4CABE98C11734170D46DFF314EED3198" box="[320,413,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="40" unit="mm" value="5.7">5.7 mm</quantity>
|
||
, rostral appendage length (projection):
|
||
<quantity id="4CABE98C11734170D644FF314CB63198" box="[873,966,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="40" unit="mm" value="5.4">5.4 mm</quantity>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFED84CBD302B" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
BMNH xxv.12c
|
||
<collectingCountry id="F34404F911734170D474FED84E963041" box="[345,486,268,294]" name="Madagascar" pageId="8" pageNumber="40">Madagascar</collectingCountry>
|
||
/ Pres: J.E. Gray [SVL:
|
||
<quantity id="4CABE98C11734170D62BFED84C193041" box="[774,873,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="40" unit="mm" value="105.0">105 mm</quantity>
|
||
, tail length:
|
||
<quantity id="4CABE98C11734170D6D6FED84B2F3041" box="[1019,1119,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.28" pageId="8" pageNumber="40" unit="mm" value="128.0">128 mm</quantity>
|
||
, rostral appendage (length, ventral edge):
|
||
<quantity id="4CABE98C11734170D46DFEE64EED302B" box="[320,413,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="40" unit="mm" value="6.6">6.6 mm</quantity>
|
||
, rostral appendage length (projection):
|
||
<quantity id="4CABE98C11734170D644FEE64CB6302B" box="[873,966,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="8" pageNumber="40" unit="mm" value="6.1">6.1 mm</quantity>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFE8D4BE030FD" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
BMNH RR1935.12.8.1 (formerly xxv.12d)
|
||
<collectingCountry id="F34404F911734170D7B8FE8D4C523014" box="[661,802,345,371]" name="Madagascar" pageId="8" pageNumber="40">Madagascar</collectingCountry>
|
||
/ Pres: J.E. Gray [SVL:
|
||
<quantity id="4CABE98C11734170D16DFE8D4BD43014" box="[1088,1188,345,371]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="40" unit="mm" value="125.0">125 mm</quantity>
|
||
, tail length:
|
||
<quantity id="4CABE98C11734170D019FE8D4AE83014" box="[1332,1432,345,371]" metricMagnitude="-1" metricUnit="m" metricValue="1.7399999999999998" pageId="8" pageNumber="40" unit="mm" value="174.0">174 mm</quantity>
|
||
, rostral appendage (length, ventral edge): n/a, rostral appendage length (projection): n/a
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFE724CA03080" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
BMNH 1863.4.16.1
|
||
<collectingCountry id="F34404F911734170D4A7FE724D6730A7" box="[394,535,422,448]" name="Madagascar" pageId="8" pageNumber="40">Madagascar</collectingCountry>
|
||
/ Purch: Mr Stevens [SVL:
|
||
<quantity id="4CABE98C11734170D649FE724CB930A7" box="[868,969,422,448]" metricMagnitude="-1" metricUnit="m" metricValue="1.47" pageId="8" pageNumber="40" unit="mm" value="147.0">147 mm</quantity>
|
||
, tail length:
|
||
<quantity id="4CABE98C11734170D176FE724BB030A7" box="[1115,1216,422,448]" metricMagnitude="-1" metricUnit="m" metricValue="2.12" pageId="8" pageNumber="40" unit="mm" value="212.0">212 mm</quantity>
|
||
, rostral appendage (length, ventral edge): n/a?, rostral appendage length (projection): n/a]
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFE204C4333E5" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
Just looking at the specimens, at least one of the possible candidates for
|
||
<emphasis id="B927987B11734170D12FFE204BE4336A" box="[1026,1172,500,525]" italics="true" pageId="8" pageNumber="40">crassicornis</emphasis>
|
||
appears to be BMNH 1863.4.16.1. This has very short horns and I think that a very faded label on the jar may say
|
||
<emphasis id="B927987B11734170D1C1FDCE4AFA3354" box="[1260,1418,538,563]" italics="true" pageId="8" pageNumber="40">'crassicornis'</emphasis>
|
||
- though it is difficult to be sure. However this specimen is rather larger than BMNH xxv.12b and BMNH xxv.12c (which are relatively close in size to each other).
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5BAFD5A4CCB33A8" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
I think we can rule out BMNH xxv.12a since, as
|
||
<bibRefCitation id="EFC2399811734170D7CCFD5A4C1033CF" author="Gray" box="[737,864,654,680]" pageId="8" pageNumber="41" refString="Gray, J. E. (1845) Catalogue of the specimens of lizards in the British Museum. London." type="book" year="1845">Gray 1845</bibRefCitation>
|
||
states, it has 'elongated nose horns' and BMNH RR1935.12.8.1 (formerly xxv.12d) is Gray's 'female without horns'."
|
||
</paragraph>
|
||
<paragraph id="8BEC446911734170D5EBFD084B64345D" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="40">
|
||
We fully agree with this interpretation and after examination of these specimens in the Natural History Museum on
|
||
<date id="FFED62A911734170D41DFCD64D6F327B" box="[304,543,770,796]" pageId="8" pageNumber="40" value="2009-09-30">30 September 2009</date>
|
||
we consider only the three specimens (BMNH xxv.12b [subadult male], BMNH xxv.12c [subadult male], and BMNH 1863.4.16.1. (formerly xxv.12d) [probably adult male] as
|
||
<typeStatus id="54E8FACB11734170D5BAFC844E73320D" box="[151,259,848,874]" pageId="8" pageNumber="40" type="syntype">syntypes</typeStatus>
|
||
. To remedy the uncertainty in the usage of this name, we hereby designate BMNH 1863.4.16.1 as
|
||
<typeStatus id="54E8FACB11734170D5BAFCA24E7532F7" box="[151,261,886,912]" pageId="8" pageNumber="40" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C533FEA11734170D403FCA34D4232F7" box="[302,562,887,912]" class="Reptilia" family="Chamaeleonidae" genus="Dicranosaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="bifurca">
|
||
<emphasis id="B927987B11734170D403FCA34D4232F7" box="[302,562,887,912]" italics="true" pageId="8" pageNumber="40">Dicranosaura bifurca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C533FEA11734170D716FCA24CE232F7" authority="Gray, 1864" authorityName="Gray" authorityYear="1864" box="[571,914,886,912]" pageId="8" pageNumber="40" rank="variety" variety="crassicornis">
|
||
var.
|
||
<emphasis id="B927987B11734170D742FCA34C7132F7" box="[623,769,887,912]" italics="true" pageId="8" pageNumber="40">crassicornis</emphasis>
|
||
<bibRefCitation id="EFC2399811734170D627FCA24CE232F7" author="Gray" box="[778,914,886,912]" pageId="8" pageNumber="41" refString="Gray, J. E. (1864) Revision of the genera and species of Chamaeleonidae, with the description of some new species. Proceedings of the Zoological Society of London, 30, 465 - 477." type="journal article" year="1864">Gray, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
. This probably adult male (total length
|
||
<quantity id="4CABE98C11734170D042FCA24FBA32D0" metricMagnitude="-1" metricUnit="m" metricValue="3.59" pageId="8" pageNumber="40" unit="mm" value="359.0">359 mm</quantity>
|
||
, tail base moderately swollen), apparently labelled as "
|
||
<emphasis id="B927987B11734170D6A7FC494B5532D1" box="[906,1061,925,950]" italics="true" pageId="8" pageNumber="40">crassicornis</emphasis>
|
||
" has very short and probably malformed rostral appendages, but otherwise agrees with males of
|
||
<taxonomicName id="4C533FEA11734170D694FC104B0432BA" box="[953,1140,964,989]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B11734170D694FC104B0432BA" box="[953,1140,964,989]" italics="true" pageId="8" pageNumber="40">Furcifer bifidus</emphasis>
|
||
</taxonomicName>
|
||
. It differs from the male
|
||
<typeStatus id="54E8FACB11734170D5BAFC3E4F8E3563" box="[151,254,1002,1028]" pageId="8" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C533FEA11734170D408FC3E4EA83564" box="[293,472,1002,1027]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="timoni">
|
||
<emphasis id="B927987B11734170D408FC3E4EA83564" box="[293,472,1002,1027]" italics="true" pageId="8" pageNumber="40">Furcifer timoni</emphasis>
|
||
</taxonomicName>
|
||
by much larger size (
|
||
<quantity id="4CABE98C11734170D7FAFC3E4C4A3563" box="[727,826,1002,1028]" metricMagnitude="-1" metricUnit="m" metricValue="1.47" pageId="8" pageNumber="40" unit="mm" value="147.0">147 mm</quantity>
|
||
vs.
|
||
<quantity id="4CABE98C11734170D647FC3E4CCD3563" box="[874,957,1002,1028]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="40" unit="mm" value="88.0">88 mm</quantity>
|
||
SVL, total length
|
||
<quantity id="4CABE98C11734170D1B5FC3E4B893563" box="[1176,1273,1002,1028]" metricMagnitude="-1" metricUnit="m" metricValue="3.59" pageId="8" pageNumber="40" unit="mm" value="359.0">359 mm</quantity>
|
||
vs.
|
||
<quantity id="4CABE98C11734170D007FC3E4AFF3563" box="[1322,1423,1002,1028]" metricMagnitude="-1" metricUnit="m" metricValue="2.1399999999999997" pageId="8" pageNumber="40" unit="mm" value="214.0">214 mm</quantity>
|
||
), more tubercles of the dorsal crest (ca. 25 vs. ca. 12) and a broader ventrolateral band of rossette-like scales (maximum width
|
||
<quantity id="4CABE98C11734170D445FBEC4EB23535" box="[360,450,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="40" unit="mm" value="4.8">4.8 mm</quantity>
|
||
vs.
|
||
<quantity id="4CABE98C11734170D4DCFBEC4D203535" box="[497,592,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="40" unit="mm" value="2.2">2.2 mm</quantity>
|
||
). The remaining two
|
||
<typeStatus id="54E8FACB11734170D664FBEC4C823535" box="[841,1010,1080,1106]" pageId="8" pageNumber="40" type="paralectotype">paralectotypes</typeStatus>
|
||
are smaller than the
|
||
<typeStatus id="54E8FACB11734170D1CBFBEC4A253535" box="[1254,1365,1080,1106]" pageId="8" pageNumber="40" type="lectotype">lectotype</typeStatus>
|
||
. They superficially resemble the
|
||
<typeStatus id="54E8FACB11734170D4CDFB8A4D3C351F" box="[480,588,1118,1144]" pageId="8" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="B927987B11734170D75AFB8B4D91351F" box="[631,737,1119,1144]" italics="true" pageId="8" pageNumber="40">F. t i m o n i</emphasis>
|
||
in size and general morphology, but differ by a broader ventrolateral band of rosette-like scales (maximum width
|
||
<quantity id="4CABE98C11734170D679FB514C9035F8" box="[852,992,1157,1183]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="8" pageNumber="40" unit="mm" value="3.75" valueMax="3.8" valueMin="3.7">3.7-3.8 mm</quantity>
|
||
vs.
|
||
<quantity id="4CABE98C11734170D13EFB514B0135F8" box="[1043,1137,1157,1183]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="40" unit="mm" value="2.2">2.2 mm</quantity>
|
||
) and the scalation of the rostral appendage which consists of more scales than in
|
||
<emphasis id="B927987B11734170D66DFB784CD735A2" box="[832,935,1196,1221]" italics="true" pageId="8" pageNumber="40">F. t i m o n i</emphasis>
|
||
(7 vs. 4 scales between nostril and tip of rostral appendage). A higher number (6-8) of scales between nostril and tip of rostral appendage is also typical for adult (BMNH xxv.12a, drawings in
|
||
<bibRefCitation id="EFC2399811734170D74BFB2D4C733474" author="Brygoo" box="[614,771,1273,1299]" pageId="8" pageNumber="41" refString="Brygoo, E. R. (1971) Reptiles Sauriens Chamaeleonidae - Genre Chamaeleo. Faune de Madagascar, 33, 1 - 318." type="journal article" year="1971">Brygoo 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC2399811734170D622FB2D4C3B3474" author="Brygoo" box="[783,843,1273,1299]" pageId="8" pageNumber="41" refString="Brygoo, E. R. (1978) Reptiles Sauriens Chamaeleonidae - Genre Brookesia et complement pour le genre Chamaeleo. Faune de Madagascar, 47, 1 - 173." type="journal article" year="1978">1978</bibRefCitation>
|
||
) and subadult (ZMB 7551) males of
|
||
<taxonomicName id="4C533FEA11734170D1D3FB2D4A1B3475" box="[1278,1387,1273,1298]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B11734170D1D3FB2D4A1B3475" box="[1278,1387,1273,1298]" italics="true" pageId="8" pageNumber="40">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
and we therefore consider the two
|
||
<typeStatus id="54E8FACB11734170D4D7FAF44DD3345D" box="[506,675,1312,1338]" pageId="8" pageNumber="40" type="paralectotype">paralectotypes</typeStatus>
|
||
as subadult males of
|
||
<taxonomicName id="4C533FEA11734170D68CFAF44B7D345E" box="[929,1037,1312,1337]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="bifidus">
|
||
<emphasis id="B927987B11734170D68CFAF44B7D345E" box="[929,1037,1312,1337]" italics="true" pageId="8" pageNumber="40">F. bifidus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |