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<document id="F5198779FA27FB033A27042F3B58B7A6" ID-DOI="10.11646/zootaxa.3852.5.1" ID-GBIF-Dataset="089d3fcb-fa57-4f3e-8ba0-d8c86d970c81" ID-ISSN="1175-5326" ID-Zenodo-Dep="229686" ID-ZooBank="B3987891-E380-4525-9CB6-4177BD38716F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460886016237" checkinUser="plazi" docAuthor="Ale-Rocha, Rosaly &amp; Freitas-Silva, Rafael Augusto Pinheiro De" docDate="2014" docId="03FA1303BA36FFF3F8EEFB1C89EFF860" docLanguage="en" docName="zt03852p539.pdf" docOrigin="Zootaxa 3852 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oropezella Collin 1926" docType="treatment" docVersion="9" lastPageNumber="503" masterDocId="FFC36B7BBA34FFF1F879FFB5883AFF95" masterDocTitle="New species of Oropezella Collin (Diptera, Hybotidae, Ocydromiinae) from Brazil and Costa Rica, with comments on the relationships among species-groups" masterLastPageNumber="539" masterPageNumber="501" pageNumber="503" updateTime="1698359118705" updateUser="plazi">
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<mods:title id="9D6EDD140478B91EB8D0C6FB477D0F7F">New species of Oropezella Collin (Diptera, Hybotidae, Ocydromiinae) from Brazil and Costa Rica, with comments on the relationships among species-groups</mods:title>
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<mods:namePart id="3D34883A9168EB173997588EBCFB6970">Ale-Rocha, Rosaly</mods:namePart>
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<mods:namePart id="19CBC2A7A40B16C59F316E12FF8A65B1">Freitas-Silva, Rafael Augusto Pinheiro De</mods:namePart>
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<mods:date id="C590BC5F57B0571FCC02D276C8008B65">2014</mods:date>
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<mods:number id="887FAC904DE8E3BEDD327B29476850B7">3852</mods:number>
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<treatment id="03FA1303BA36FFF3F8EEFB1C89EFF860" ID-DOI="http://doi.org/10.5281/zenodo.5689858" ID-GBIF-Taxon="119586517" ID-Zenodo-Dep="5689858" LSID="urn:lsid:plazi:treatment:03FA1303BA36FFF3F8EEFB1C89EFF860" httpUri="http://treatment.plazi.org/id/03FA1303BA36FFF3F8EEFB1C89EFF860" lastPageNumber="503" pageId="2" pageNumber="503">
<subSubSection id="C349F19EBA36FFF3F8EEFB1C898DFB56" box="[151,439,1193,1219]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="8BECA215BA36FFF3F8EEFB1C898DFB56" blockId="2.[151,439,1193,1219]" box="[151,439,1193,1219]" pageId="2" pageNumber="503">
<heading id="D0A41579BA36FFF3F8EEFB1C898DFB56" bold="true" box="[151,439,1193,1219]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<taxonomicName id="4C53D996BA36FFF3F8EEFB1C898DFB56" ID-CoL="8YQKJ" ID-ENA="369870" authority="Collin, 1926" authorityName="Collin" authorityYear="1926" box="[151,439,1193,1219]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F8EEFB1C898DFB56" bold="true" box="[151,439,1193,1219]" pageId="2" pageNumber="503">
<emphasis id="B9277E07BA36FFF3F8EEFB1C8923FB56" bold="true" box="[151,281,1193,1219]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
<bibRefCitation id="EFC2DFE4BA36FFF3F959FB1C898DFB56" author="Collin" box="[288,439,1193,1219]" pageId="2" pageNumber="503" refString="Collin, J. E. (1926) Notes on the Empididae (Diptera) with additions and corrections to the British List. Entomologist´s Monthly Magazine, 62, 213 - 219." type="journal article" year="1926">Collin, 1926</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349F19EBA36FFF3F8EEFB458CCFFAD6" pageId="2" pageNumber="503" type="reference_group">
<paragraph id="8BECA215BA36FFF3F8EEFB458993FAB1" blockId="2.[151,1436,1264,1347]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="AB43853BBA36FFF3F8EEFB458993FAB1" pageId="2" pageNumber="503">
<taxonomicName id="4C53D996BA36FFF3F8EEFB4589F4FA93" ID-CoL="8YQKJ" authority="Collin, 1926: 214" authorityName="Collin" authorityPageNumber="214" authorityYear="1926" box="[151,462,1264,1286]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F8EEFB45893DFA93" box="[151,263,1264,1286]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
<treatmentCitation id="0AF28404BA36FFF3F968FB4589F4FA93" author="Collin" box="[273,462,1264,1286]" page="214" pageId="2" pageNumber="503" year="1926">
<bibRefCitation id="EFC2DFE4BA36FFF3F968FB4589ADFA93" author="Collin" box="[273,407,1264,1286]" pageId="2" pageNumber="503" refString="Collin, J. E. (1926) Notes on the Empididae (Diptera) with additions and corrections to the British List. Entomologist´s Monthly Magazine, 62, 213 - 219." type="journal article" year="1926">Collin, 1926</bibRefCitation>
: 214
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AF28404BA36FFF3F9A4FB448AF4FA93" author="Yang" box="[477,718,1264,1286]" page="334" pageId="2" pageNumber="503" year="2007">
<bibRefCitation id="EFC2DFE4BA36FFF3F9A4FB448AA3FA93" author="Yang" box="[477,665,1264,1286]" pageId="2" pageNumber="503" refString="Yang, D., Zhang, K., Yao, G. &amp; Zhang, J. (2007) World catalog of Empididae (Insecta: Diptera). China Agricultural University Press, Beijing, 599 pp." type="book" year="2007">
Yang
<emphasis id="B9277E07BA36FFF3FA65FB448A77FA93" box="[540,589,1264,1286]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
., 2007
</bibRefCitation>
: 334
</treatmentCitation>
335 (catalogue).
<typeStatus id="54E81CB7BA36FFF3FBF7FB448BFAFA93" box="[910,960,1265,1286]" pageId="2" pageNumber="503">Type</typeStatus>
species:
<taxonomicName id="4C53D996BA36FFF3FC5FFB448DA6FA93" ID-CoL="8CJ5P" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[1062,1436,1264,1286]" class="Insecta" family="Hybotidae" genus="Leptopeza" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="sphenoptera">
<emphasis id="B9277E07BA36FFF3FC5FFB448D2CFA93" box="[1062,1302,1264,1286]" italics="true" pageId="2" pageNumber="503">Leptopeza sphenoptera</emphasis>
<bibRefCitation id="EFC2DFE4BA36FFF3FD59FB448DA6FA93" author="Loew" box="[1312,1436,1264,1286]" pageId="2" pageNumber="503" refString="Loew, H. (1873) Beschreibung europaischer Dipteren. Systematische Beschreibung der bekannten europaischen zweflugeligen Insecten, von Johann Wilhelm Meigen. Vol. 3. Zehnter Theil oder vierter Supplementband. Halle, 320 pp." type="book" year="1873">Loew, 1873</bibRefCitation>
</taxonomicName>
(original designation).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BECA215BA36FFF3F8EEFA988CCFFAD6" blockId="2.[151,1436,1264,1347]" box="[151,1269,1325,1347]" pageId="2" pageNumber="503">
<emphasis id="B9277E07BA36FFF3F8EEFA988906FAD6" box="[151,316,1325,1347]" italics="true" pageId="2" pageNumber="503">Leptometopiella</emphasis>
<bibRefCitation id="EFC2DFE4BA36FFF3F93BFA9889DEFAD6" author="Melander" box="[322,484,1325,1347]" pageId="2" pageNumber="503" refString="Melander, A. L. (1928) Diptera, Fam. Empididae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 185 (1927). Louis Desmet- Verteneuil, Bruxelles, 434 pp." type="book" year="1928">Melander, 1928</bibRefCitation>
: 70.
<typeStatus id="54E81CB7BA36FFF3FA6DFA9B8A7CFAD6" box="[532,582,1326,1347]" pageId="2" pageNumber="503">Type</typeStatus>
species:
<taxonomicName id="4C53D996BA36FFF3FADAFA9B8C37FAD6" ID-CoL="8CJ5P" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[675,1037,1325,1347]" class="Insecta" family="Hybotidae" genus="Leptopeza" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="sphenoptera">
<emphasis id="B9277E07BA36FFF3FADAFA9B8BB5FAD6" box="[675,911,1325,1347]" italics="true" pageId="2" pageNumber="503">Leptopeza sphenoptera</emphasis>
<bibRefCitation id="EFC2DFE4BA36FFF3FBECFA9B8C37FAD6" author="Loew" box="[917,1037,1325,1347]" pageId="2" pageNumber="503" refString="Loew, H. (1873) Beschreibung europaischer Dipteren. Systematische Beschreibung der bekannten europaischen zweflugeligen Insecten, von Johann Wilhelm Meigen. Vol. 3. Zehnter Theil oder vierter Supplementband. Halle, 320 pp." type="book" year="1873">Loew, 1873</bibRefCitation>
</taxonomicName>
(original designation).
</paragraph>
</subSubSection>
<subSubSection id="C349F19EBA36FFF3F8EEFADA8B0DF96C" pageId="2" pageNumber="503" type="diagnosis">
<paragraph id="8BECA215BA36FFF3F8EEFADA8B0DF96C" blockId="2.[151,1436,1391,2037]" pageId="2" pageNumber="503">
<emphasis id="B9277E07BA36FFF3F8EEFADA8930FA1D" bold="true" box="[151,266,1391,1416]" pageId="2" pageNumber="503">Diagnosis</emphasis>
. The genus is characterized by: scape as long or longer than pedicel; antennae inserted above middle of head (except
<taxonomicName id="4C53D996BA36FFF3F956FA2089FFFA38" box="[303,453,1428,1453]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="longifrons" status="sp. nov.">
<emphasis id="B9277E07BA36FFF3F956FA2089FFFA38" box="[303,453,1428,1453]" italics="true" pageId="2" pageNumber="503">O. longifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A214C37CBA36FFF3F9B7FA218A1DFA39" box="[462,551,1428,1453]" pageId="2" pageNumber="503" rank="species">
<emphasis id="B9277E07BA36FFF3F9B7FA218A18FA38" bold="true" box="[462,546,1428,1453]" pageId="2" pageNumber="503">sp. nov</emphasis>
.
</taxonomicNameLabel>
); face long and holoptic; vein Rs long, arising before middle of cell bm; two veins emitted from cell dm reaching, or nearly reaching, wing margin (in
<taxonomicName id="4C53D996BA36FFF3FBB2FA0D8C4DFA45" box="[971,1143,1463,1488]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B9277E07BA36FFF3FBB2FA0D8C4DFA45" box="[971,1143,1463,1488]" italics="true" pageId="2" pageNumber="503">O. abdominalis</emphasis>
</taxonomicName>
three veins are issue from cell dm but only one reaches wing margin); cell dm slender and with parallel sides (except
<taxonomicName id="4C53D996BA36FFF3FCE5FA688D73FA60" box="[1180,1353,1500,1525]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B9277E07BA36FFF3FCE5FA688D73FA60" box="[1180,1353,1500,1525]" italics="true" pageId="2" pageNumber="503">O. abdominalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C53D996BA36FFF3FDF8FA68893DF98D" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="B9277E07BA36FFF3FDF8FA68893DF98D" italics="true" pageId="2" pageNumber="503">O. antennata</emphasis>
</taxonomicName>
); vein R4+5 reaching Costa below the apex of the wing; anal angle of wing poorly developed; cell
<emphasis id="B9277E07BA36FFF3FD0DF9B48DA6F98D" box="[1396,1436,1537,1560]" italics="true" pageId="2" pageNumber="503">cup</emphasis>
equal or more than two-thirds the length of cell bm; CuA2 sinuous (also in
<taxonomicName id="4C53D996BA36FFF3FB93F9938C6FF9AA" box="[1002,1109,1574,1599]" class="Insecta" family="Hybotidae" genus="Chvalaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3FB93F9938C6FF9AA" box="[1002,1109,1574,1599]" italics="true" pageId="2" pageNumber="503">Chvalaea</emphasis>
</taxonomicName>
; CuA2 absent in
<taxonomicName id="4C53D996BA36FFF3FD6EF9928DB1F9AA" box="[1303,1419,1574,1599]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B9277E07BA36FFF3FD6EF9928DB1F9AA" box="[1303,1419,1574,1599]" italics="true" pageId="2" pageNumber="503">O. bicolor</emphasis>
</taxonomicName>
); when present, pterostigma confined to cell sc at the apex of R1; a pair of posterior dorsocentral bristles usually differentiated; hypopygium asymmetrical, rotated approximately 90º to right; epandrium shallowly or deeply cleft; cercus thinly sclerotized; ejaculatory apodeme fused to base of phallus; bacilliform sclerites well sclerotized; phallic shaft arched, cylindrical; articulated distal portion of phallus bifid apically (simple in
<taxonomicName id="4C53D996BA36FFF3FCB8F9088D7CF941" box="[1217,1350,1725,1748]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="tanycera">
<emphasis id="B9277E07BA36FFF3FCB8F9088D7CF941" box="[1217,1350,1725,1748]" italics="true" pageId="2" pageNumber="503">O. tanycera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C53D996BA36FFF3FDF8F908890CF96C" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="diminuloruma">
<emphasis id="B9277E07BA36FFF3FDF8F908890CF96C" italics="true" pageId="2" pageNumber="503">O. diminuloruma</emphasis>
</taxonomicName>
) bearing a basal process with variable length.
</paragraph>
</subSubSection>
<subSubSection id="C349F19EBA36FFF3F8BEF8B189EFF860" pageId="2" pageNumber="503" type="discussion">
<paragraph id="8BECA215BA36FFF3F8BEF8B189EFF860" blockId="2.[151,1436,1391,2037]" pageId="2" pageNumber="503">
<emphasis id="B9277E07BA36FFF3F8BEF8B1890EF888" bold="true" box="[199,308,1796,1821]" pageId="2" pageNumber="503">Remarks</emphasis>
. The generic limits of
<taxonomicName id="4C53D996BA36FFF3FA33F8B18AFFF888" box="[586,709,1796,1821]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3FA33F8B18AFFF888" box="[586,709,1796,1821]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
are not well defined. As shown in the cladistic analysis (see below),
<taxonomicName id="4C53D996BA36FFF3F880F89D894FF8D4" box="[249,373,1832,1857]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F880F89D894FF8D4" box="[249,373,1832,1857]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
is apparently a non-monophyletic group, since
<taxonomicName id="4C53D996BA36FFF3FBC0F89D8C1FF8D4" box="[953,1061,1832,1857]" class="Insecta" family="Hybotidae" genus="Chvalaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3FBC0F89D8C1FF8D4" box="[953,1061,1832,1857]" italics="true" pageId="2" pageNumber="503">Chvalaea</emphasis>
</taxonomicName>
appears as a sub-group within
<taxonomicName id="4C53D996BA36FFF3F8EEF8F98929F8F0" box="[151,275,1868,1893]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F8EEF8F98929F8F0" box="[151,275,1868,1893]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
. Moreover,
<taxonomicName id="4C53D996BA36FFF3F9DDF8F98A2AF8F0" box="[420,528,1868,1893]" class="Insecta" family="Hybotidae" genus="Chvalaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F9DDF8F98A2AF8F0" box="[420,528,1868,1893]" italics="true" pageId="2" pageNumber="503">Chvalaea</emphasis>
</taxonomicName>
forms a well-supported monophyletic group. Therefore, the species currently allocated to
<taxonomicName id="4C53D996BA36FFF3F951F8C5899EF81C" box="[296,420,1904,1929]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F951F8C5899EF81C" box="[296,420,1904,1929]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
can be promptly distinguished from
<emphasis id="B9277E07BA36FFF3FB22F8C58BEEF81C" box="[859,980,1904,1929]" italics="true" pageId="2" pageNumber="503">Chavalaea</emphasis>
species by the heavily sclerotized and punctated surface of the abdomen and cell
<emphasis id="B9277E07BA36FFF3FA0EF8208A9AF839" box="[631,672,1941,1964]" italics="true" pageId="2" pageNumber="503">cup</emphasis>
approximately half length of the second basal cell in
<taxonomicName id="4C53D996BA36FFF3FC81F8218D5EF838" box="[1272,1380,1940,1965]" class="Insecta" family="Hybotidae" genus="Chvalaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3FC81F8218D5EF838" box="[1272,1380,1940,1965]" italics="true" pageId="2" pageNumber="503">Chvalaea</emphasis>
</taxonomicName>
(one species in
<taxonomicName id="4C53D996BA36FFF3F976F80D89B1F844" box="[271,395,1976,2001]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F976F80D89B1F844" box="[271,395,1976,2001]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C53D996BA36FFF3F9E3F80C8A73F844" box="[410,585,1976,2001]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B9277E07BA36FFF3F9E3F80C8A73F844" box="[410,585,1976,2001]" italics="true" pageId="2" pageNumber="503">O. abdominalis</emphasis>
</taxonomicName>
, has the abdomen darkly sclerotized but not punctated, and the cell
<emphasis id="B9277E07BA36FFF3FD21F80C8DBAF845" box="[1368,1408,1977,2000]" italics="true" pageId="2" pageNumber="503">cup</emphasis>
is similar to other
<taxonomicName id="4C53D996BA36FFF3F930F86989FEF860" box="[329,452,2012,2037]" class="Insecta" family="Hybotidae" genus="Oropezella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B9277E07BA36FFF3F930F86989FEF860" box="[329,452,2012,2037]" italics="true" pageId="2" pageNumber="503">Oropezella</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>