122 lines
15 KiB
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122 lines
15 KiB
XML
<document id="AC5381FD5708ACCE678BFF7D44492118" ID-CLB-Dataset="29761" ID-DOI="10.11646/zootaxa.4442.2.8" ID-GBIF-Dataset="d66ed56d-eec7-424a-a7c9-9f2c884269e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303179" ID-ZooBank="A822F33A-0E10-46A4-8630-6944D4A00307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533711971" checkinUser="plazi" docAuthor="Kopylov, D. S., Spasojevic, T. & Klopfstein, S." docDate="2018" docId="03D9905AFFD43B5E19EC7A45FB2CF870" docLanguage="en" docName="zootaxa.4442.2.8.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dolichomitus Smith 1877" docType="treatment" docVersion="7" lastPageNumber="326" masterDocId="FFE0E822FFD33B59197B7901FFFFFFAB" masterDocTitle="New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East" masterLastPageNumber="330" masterPageNumber="319" pageNumber="326" updateTime="1698552165635" updateUser="ExternalLinkService">
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<mods:titleInfo id="E1D37CB1BDE837B13CA6347593276BD4">
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<mods:title id="4556F71BB8D9055716752F89A48D6DA1">New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East</mods:title>
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<mods:name id="8A2FF72EF2AD39026995103B73ACB119" type="personal">
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<mods:namePart id="779CFE982412B01C2D2FA0015520DC5E">Kopylov, D. S.</mods:namePart>
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<mods:roleTerm id="42BE218D850EC569A68DE29F308EDE82">Author</mods:roleTerm>
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<mods:namePart id="70BCD48F4EFF0A8DED503A4A244FA92B">Spasojevic, T.</mods:namePart>
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<mods:roleTerm id="B9F48F0B5CC58155B224531E0ED5B869">Author</mods:roleTerm>
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<mods:namePart id="03923361D6131D93A85FD771450352AD">Klopfstein, S.</mods:namePart>
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<mods:title id="ADA5B43F8B361B4DAD16D6684197E760">Zootaxa</mods:title>
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<mods:part id="7375715028984F0F89E43FE70CCBF6AE">
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<mods:date id="A67008D70B59E50FAC5CBF7F23DBBD58">2018</mods:date>
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<mods:number id="FB1195D294C02B83BFEB6AD1D06AE727">2018-07-02</mods:number>
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<mods:number id="59D745F0726C2F994CC1F91230E27C2C">4442</mods:number>
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<mods:classification id="7BD0A7D0416620741CB8615C29187EF8">journal article</mods:classification>
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<treatment id="03D9905AFFD43B5E19EC7A45FB2CF870" ID-DOI="http://doi.org/10.5281/zenodo.5973714" ID-GBIF-Taxon="146044114" ID-Zenodo-Dep="5973714" LSID="urn:lsid:plazi:treatment:03D9905AFFD43B5E19EC7A45FB2CF870" httpUri="http://treatment.plazi.org/id/03D9905AFFD43B5E19EC7A45FB2CF870" lastPageNumber="326" pageId="7" pageNumber="326">
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<subSubSection id="C36A72C7FFD43B5E19EC7A45FF0AFCD4" pageId="7" pageNumber="326" type="nomenclature">
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<paragraph id="8BCF214CFFD43B5E19EC7A45FC4EFCF5" blockId="7.[151,945,836,895]" box="[151,945,836,862]" pageId="7" pageNumber="326">
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<heading id="D0879620FFD43B5E19EC7A45FC4EFCF5" bold="true" box="[151,945,836,862]" fontSize="11" level="1" pageId="7" pageNumber="326" reason="1">
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<emphasis id="B904FD5EFFD43B5E19EC7A45FC4EFCF5" bold="true" box="[151,945,836,862]" pageId="7" pageNumber="326">
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<emphasis id="B904FD5EFFD43B5E19EC7A45FE5EFCF5" bold="true" box="[151,417,836,862]" italics="true" pageId="7" pageNumber="326">
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<taxonomicName id="4C705ACFFFD43B5E19EC7A45FEC2FCF5" ID-CoL="47HK" ID-ENA="65330" authorityName="Smith" authorityYear="1877" box="[151,317,836,862]" class="Insecta" family="Ichneumonidae" genus="Dolichomitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="genus">Dolichomitus</taxonomicName>
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? saxeus
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</emphasis>
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Kopylov, Spasojevic & Klopfstein sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCF214CFFD43B5E19EC7A67FF0AFCD4" blockId="7.[151,945,836,895]" box="[151,245,870,895]" pageId="7" pageNumber="326">
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<figureCitation id="134B3DC9FFD43B5E19EC7A67FF0AFCD4" box="[151,245,870,895]" captionStart="FIGURE 5" captionStartId="8.[151,250,1744,1766]" captionTargetBox="[151,1436,195,1711]" captionTargetId="figure@8.[151,1436,195,1722]" captionTargetPageId="8" captionText="FIGURE 5. Dolichomitus? saxeus sp. nov. holotype PIN, 3364/31: a—part, b—our interpretation." httpUri="https://zenodo.org/record/1303193/files/figure.png" pageId="7" pageNumber="326">Figure 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36A72C7FFD43B5E19EC7AAFFACDFB98" pageId="7" pageNumber="326" type="materials_examined">
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<paragraph id="8BCF214CFFD43B5E19EC7AAFFACDFB98" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">
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<emphasis id="B904FD5EFFD43B5E19EC7AAFFEFBFC6C" bold="true" box="[151,260,942,967]" pageId="7" pageNumber="326">Holotype</emphasis>
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. PIN, 3364/31, part with counterpart. Imago, sex unknown, mainly dorsal aspect (but see following and description below). Head absent; mesosoma rather well preserved, but with apical part of propodeum and most of T1 obscured by coxae; legs missing excluding hind coxae and one detached femur; antennae and ovipositor missing, fore wings and one hind wing preserved. Ugolny, Russian Far East, Eocene, Tadushi Formation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36A72C7FFD43B5E19BC7D3FFF22FBD0" pageId="7" pageNumber="326" type="etymology">
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<paragraph id="8BCF214CFFD43B5E19BC7D3FFF22FBD0" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">
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<emphasis id="B904FD5EFFD43B5E19BC7D3FFEB9FBFC" bold="true" box="[199,326,1086,1111]" pageId="7" pageNumber="326">Etymology</emphasis>
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. The species name is derived from the Latin word “saxeus”, which means “rocky” or “made of rock”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36A72C7FFD43B5E19BC7D87FB2CF870" pageId="7" pageNumber="326" type="description">
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<paragraph id="8BCF214CFFD43B5E19BC7D87FEF8FB68" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">
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<emphasis id="B904FD5EFFD43B5E19BC7D87FEAEFB34" bold="true" box="[199,337,1158,1183]" pageId="7" pageNumber="326">Description</emphasis>
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. Coloration. Mesosoma, coxae and wing veins very dark, metasoma and legs somewhat lighter, brownish.
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</paragraph>
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<paragraph id="8BCF214CFFD43B5E19BC7DCEFDB2FA84" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">Mesosoma. Mesonotum coarsely punctured, as long as wide and 2.7x as long as scutellum. Notauli strong, converging, reach the middle of mesonotum. Propodeum carination unclear, with break between part and counterpart potentially inside the body.</paragraph>
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<paragraph id="8BCF214CFFD43B5E19BC7C3BFEE8F9AC" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">Wings. Fore wing 3.3× as long as wide; pterostigma and cell 3R elongate, both about 3.9× as long as wide; areolet large, 2.3× as long as wide, almost triangular, with 4-M very short (2× as long as vein width), 2+3M 1.6× 2- Rs, 1.6x r-m, petiolus of areolet (3-Rs) 0.24× as long as areolet width; 1-Rs&1-M almost straight; 1m-cu&2-Rs+M gently curved, with no sharp bend or traces of 1-Rs+M; r-rs issues at mid-length of pterostigma, s-shaped, 2.8× pterostigma width. Hind wing with 1-Rs 1.1× as long as r-m; 1-Cu 0.9× as long as cu-a, the two veins meeting at a 120° angle.</paragraph>
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<paragraph id="8BCF214CFFD43B5E19BC7F12FB51F9D8" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">Metasoma. Metasoma a little elongate, fusiform, 3.0× as long as wide and 2.1× as long as mesosoma. T1 elongate, 1.7× as long as wide, comprises 0.22 of metasoma length. T1 carinae, if present, not visible on the specimen. T2 and T3 wider than long, but still longer than posterior tergites. T5 is the widest.</paragraph>
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<paragraph id="8BCF214CFFD43B5E19BC7F7EFA62F93D" blockId="7.[151,1437,942,2011]" box="[199,1437,1662,1687]" pageId="7" pageNumber="326">
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Measurements. Mesosoma length
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<quantity id="4C888CA9FFD43B5E1B3E7F7EFD62F93D" box="[581,669,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="326" unit="mm" value="3.3">3.3 mm</quantity>
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, metasoma length
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<quantity id="4C888CA9FFD43B5E1A167F7FFC39F93D" box="[877,966,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="7" pageNumber="326" unit="mm" value="6.9">6.9 mm</quantity>
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. Fore wing length
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<quantity id="4C888CA9FFD43B5E1DED7F7EFB11F93D" box="[1174,1262,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="7" pageNumber="326" unit="mm" value="9.9">9.9 mm</quantity>
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, width 3.0 mm.
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</paragraph>
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<paragraph id="8BCF214CFFD43B5E19BC7FA3FB2CF870" blockId="7.[151,1437,942,2011]" pageId="7" pageNumber="326">
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<emphasis id="B904FD5EFFD43B5E19BC7FA3FE2AF910" bold="true" box="[199,469,1698,1723]" pageId="7" pageNumber="326">Taxonomic placement.</emphasis>
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The placement of this fossil is not straightforward, given its incompleteness and, in particular, lack of visible ovipositor. The venation of the fore wing, especially the almost triangular areolet receiving vein 2m-cu close to its outer corner, 2m-cu bowed outwards and probably with two bullae, and s-shaped vein r-rs point to the subfamily
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<taxonomicName id="4C705ACFFFD43B5E1B7A7E0FFD89F88C" box="[513,630,1806,1831]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
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. The hind wing vein 1-Cu being a bit shorter than cu-a indicates either
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<taxonomicName id="4C705ACFFFD43B5E19EC7E33FEFAF8E0" box="[151,261,1842,1867]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="tribe" tribe="Ephialtini">Ephialtini</taxonomicName>
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or
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<taxonomicName id="4C705ACFFFD43B5E18567E33FE38F8E0" box="[301,455,1842,1867]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="tribe" tribe="Delomeristini">Delomeristini</taxonomicName>
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(in
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<taxonomicName id="4C705ACFFFD43B5E188C7E33FDA5F8E0" box="[503,602,1842,1867]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="tribe" tribe="Pimplini">Pimplini</taxonomicName>
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, 1-Cu is invariably much shorter than cu-a, see
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<figureCitation id="134B3DC9FFD43B5E1D077E33FB42F8E0" box="[1148,1213,1842,1867]" captionStart="FIGURE 1" captionStartId="2.[151,250,1688,1710]" captionTargetBox="[161,1435,198,1667]" captionTargetId="figure@2.[151,1436,195,1667]" captionTargetPageId="2" captionText="FIGURE 1. Phylogeny of Pimplinae and outgroups, redrawn according to Gauld et al. (2002), with examples of propodea, first metasomal tergites and hind wings shown to the left of the genus names. The drawings are modified from Townes (1969), and the species representatives are, from top to bottom: Xorides irrigator Fabricius, Odontocolon ochropus Townes, Poemenia hectica Gravenhorst, Rhyssa persuasoria Linnaeus, Theronia atalantae Poda, Lissopimpla excelsa Costa, Xanthopimpla hova Saussure, Echthromorpha agrestoria Swederus, Itoplectis maculator Fabricius, Pimpla madecassa Saussure, Zatypota percontatoria Müller, Dreisbachia mira Tosquinet, Camptotypus pulchripennis Saussure, Scambus tecumseh Viereck, Flavopimpla nava Tosquinet, Delomerista novita Cresson, Atractogaster semisculptus Kriechbaumer, and Perithous albicinctus Gravenhorst." httpUri="https://zenodo.org/record/1303181/files/figure.png" pageId="7" pageNumber="326">Fig. 1</figureCitation>
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). Several genera in
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<taxonomicName id="4C705ACFFFD43B5E19EC7E57FEFAF8C4" box="[151,261,1878,1903]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="tribe" tribe="Ephialtini">Ephialtini</taxonomicName>
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have similarly elongate fore wings with a large triangular areolet; most of them also have elongate metasomal tergites, but with quite some variation. The incompleteness of the fossil does not allow a clear attribution to one of these genera, and we thus place it tentatively in
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<taxonomicName id="4C705ACFFFD43B5E1AD77E9FFBBCF81C" authorityName="Smith" authorityYear="1877" box="[940,1091,1950,1975]" class="Insecta" family="Ichneumonidae" genus="Dolichomitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="genus">
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<emphasis id="B904FD5EFFD43B5E1AD77E9FFBBCF81C" box="[940,1091,1950,1975]" italics="true" pageId="7" pageNumber="326">Dolichomitus</emphasis>
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</taxonomicName>
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as its wing venation, size and metasomal tergite lengths fall within the variation found in the extant species of this large genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |