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<document ID-DOI="http://doi.org/10.5281/zenodo.3794463" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962585" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794463" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. &amp; Scholtz, Clarke H." docDate="2020" docId="D474D525FFB57C4AD7699CDDFF4CFA1B" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Neosisyphus setiger Roth 1851" docType="treatment" docVersion="6" lastPageNumber="47" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="46" pageNumber="47" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel, Gimo M.</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Davis, Adrian Lv.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sole, Catherine L.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Scholtz, Clarke H.</mods:namePart>
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<mods:title>Insect Systematics &amp; Evolution</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-01-02</mods:number>
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<mods:number>51</mods:number>
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<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
<mods:identifier type="GBIF-Dataset">a725007b-2d51-4c92-838a-9e8fd27484c8</mods:identifier>
<mods:identifier type="ISSN">1876-312X</mods:identifier>
<mods:identifier type="Zenodo-Dep">3786629</mods:identifier>
<mods:identifier type="ZooBank">701C1742-718D-4486-A158-AEA608BA8576</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794463" ID-GBIF-Taxon="163962585" ID-Zenodo-Dep="3794463" LSID="urn:lsid:plazi:treatment:D474D525FFB57C4AD7699CDDFF4CFA1B" httpUri="http://treatment.plazi.org/id/D474D525FFB57C4AD7699CDDFF4CFA1B" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection box="[106,637,134,161]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph blockId="46.[106,1073,134,508]" box="[106,637,134,161]" pageId="46" pageNumber="47">
<heading box="[106,637,134,161]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="3">
<taxonomicName authority="Roth (1851)" authorityName="Roth" authorityYear="1851" box="[106,485,134,161]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis bold="true" box="[106,485,134,161]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[106,329,134,161]" italics="true" pageId="46" pageNumber="47">Neosisyphus setiger</emphasis>
Roth (1851)
</emphasis>
</taxonomicName>
(
<figureCitation box="[500,578,134,161]" captionStart="Fig" captionStartId="23.[98,131,990,1012]" captionTargetBox="[189,976,131,962]" captionTargetId="figure@23.[189,976,131,965]" captionTargetPageId="23" captionText="Fig. 7. Distribution map for the goryi species-group." figureDoi="http://doi.org/10.5281/zenodo.3786644" httpUri="https://zenodo.org/record/3786644/files/figure.png" pageId="46" pageNumber="47">Figs. 7</figureCitation>
,
<figureCitation box="[590,628,135,161]" captionStart="Fig" captionStartId="25.[98,131,1628,1650]" captionTargetBox="[362,802,1074,1603]" captionTargetId="figure@25.[362,802,1074,1603]" captionTargetPageId="25" captionText="Fig. 8. Images of the habitus of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786646" httpUri="https://zenodo.org/record/3786646/files/figure.png" pageId="46" pageNumber="47">8D</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="reference_group">
<treatmentCitationGroup pageId="46" pageNumber="47">
<paragraph blockId="46.[106,1073,134,508]" box="[106,346,169,196]" pageId="46" pageNumber="47">
<bibRefCitation author="Montreuil" box="[106,346,169,196]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil 2015b: 11</bibRefCitation>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,406,204,231]" pageId="46" pageNumber="47">
<taxonomicName authority="Roth, 1851" authorityName="Roth" authorityYear="1851" box="[106,406,204,231]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[106,269,204,230]" italics="true" pageId="46" pageNumber="47">Sisyphus setiger</emphasis>
Roth, 1851
</taxonomicName>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,542,238,265]" pageId="46" pageNumber="47">
Roth 1851: 124;
<bibRefCitation author="Montreuil" box="[302,542,238,265]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil 2015b: 11</bibRefCitation>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,501,273,299]" pageId="46" pageNumber="47">
<taxonomicName authority="(Kolbe, 1914)" baseAuthorityName="Kolbe" baseAuthorityYear="1914" box="[106,501,273,299]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[106,335,273,299]" italics="true" pageId="46" pageNumber="47">Neosisyphus confrater</emphasis>
(
<bibRefCitation author="Kolbe" box="[349,493,273,299]" firstAuthor="Kolbe" pageId="46" pageNumber="47" pagination="261 - 416" refId="ref29802" refString="Kolbe, H. J. (1914) Lieferung 3: Scarabaeiden. In: Shubotz, H. (Ed.), Wissenschaftliches Ergebnisse der Deutschen Zentral-Afrika-Expedition 1907 unter Fuhrung Adolf Friedrichs, Herzog zu Mecklenburg. Zoologie III, 5. Klinkhardt &amp; Biermann, Leipzig, Germany, pp. 261 - 416." type="book chapter" year="1914">Kolbe, 1914</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,346,308,335]" pageId="46" pageNumber="47">
<bibRefCitation author="Montreuil" box="[106,346,308,335]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil 2015b: 11</bibRefCitation>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,445,342,369]" pageId="46" pageNumber="47">
<taxonomicName authority="Kolbe, 1914" authorityName="Kolbe" authorityYear="1914" box="[106,445,342,369]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[106,298,342,368]" italics="true" pageId="46" pageNumber="47">Sisyphus confrater</emphasis>
<bibRefCitation author="Kolbe" box="[305,445,342,369]" firstAuthor="Kolbe" pageId="46" pageNumber="47" pagination="261 - 416" refId="ref29802" refString="Kolbe, H. J. (1914) Lieferung 3: Scarabaeiden. In: Shubotz, H. (Ed.), Wissenschaftliches Ergebnisse der Deutschen Zentral-Afrika-Expedition 1907 unter Fuhrung Adolf Friedrichs, Herzog zu Mecklenburg. Zoologie III, 5. Klinkhardt &amp; Biermann, Leipzig, Germany, pp. 261 - 416." type="book chapter" year="1914">Kolbe, 1914</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="46.[106,1073,134,508]" box="[106,500,377,404]" pageId="46" pageNumber="47">
<bibRefCitation author="Kolbe" box="[106,301,377,403]" firstAuthor="Kolbe" pageId="46" pageNumber="47" pagination="261 - 416" refId="ref29802" refString="Kolbe, H. J. (1914) Lieferung 3: Scarabaeiden. In: Shubotz, H. (Ed.), Wissenschaftliches Ergebnisse der Deutschen Zentral-Afrika-Expedition 1907 unter Fuhrung Adolf Friedrichs, Herzog zu Mecklenburg. Zoologie III, 5. Klinkhardt &amp; Biermann, Leipzig, Germany, pp. 261 - 416." type="book chapter" year="1914">Kolbe 1914: 317</bibRefCitation>
;
<bibRefCitation author="Haaf" box="[312,496,377,404]" firstAuthor="Haaf" pageId="46" pageNumber="47" pagination="341 - 381" refId="ref29533" refString="Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381." type="journal article" year="1955">Haaf 1955: 376</bibRefCitation>
;
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection box="[135,487,412,439]" pageId="46" pageNumber="47" type="distribution">
<paragraph blockId="46.[106,1073,134,508]" box="[135,487,412,439]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[135,257,412,438]" pageId="46" pageNumber="47">
<typeStatus box="[135,193,412,438]" pageId="46" pageNumber="47">Type</typeStatus>
local
</emphasis>
: Bukoba (
<collectingCountry box="[374,479,412,439]" name="Tanzania" pageId="46" pageNumber="47">Tanzania</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="description">
<paragraph blockId="46.[106,1073,134,508]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[135,182,446,472]" pageId="46" pageNumber="47">Size</emphasis>
: Male: length: 9.05.0 mm; width: 4.83.0 mm. Female: length: 9.25.0 mm; width: 4.53.0 mm
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="diagnosis">
<paragraph blockId="46.[106,1074,550,681]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[106,225,550,576]" pageId="46" pageNumber="47">Diagnosis</emphasis>
:
<taxonomicName authorityName="Roth" authorityYear="1851" box="[242,346,551,576]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[242,346,551,576]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
is distinguished from other members of the
<taxonomicName authorityName="Wiedemann" authorityYear="1823" box="[865,979,550,576]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="barbarossa">
<emphasis box="[865,979,550,576]" italics="true" pageId="46" pageNumber="47">barbarossa</emphasis>
</taxonomicName>
speciesgroup, by a combination of the following features: a triangular protrusion on the midposterior edge of the metafemur in males. Additionally, the projected metatrochanter is relatively short, comprising ¼ of the total length of the metafemur in males.
</paragraph>
</subSubSection>
<subSubSection box="[106,830,723,750]" pageId="46" pageNumber="47" type="materials_examined">
<paragraph blockId="46.[106,830,723,750]" box="[106,830,723,750]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[106,447,723,750]" pageId="46" pageNumber="47">Examined non-type material</emphasis>
: See Supplementary information.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="distribution">
<paragraph blockId="46.[106,1074,793,854]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[106,257,793,819]" pageId="46" pageNumber="47">Distribution</emphasis>
: As defined by
<bibRefCitation author="Montreuil" box="[431,648,793,820]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil (2015b)</bibRefCitation>
,
<taxonomicName authorityName="Roth" authorityYear="1851" box="[660,762,794,819]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[660,762,794,819]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
is distributed in moist lowland and upland vegetation from northeast to southern Africa (
<figureCitation box="[818,902,828,854]" captionStart="Fig" captionStartId="45.[98,131,1593,1615]" captionTargetId="figure@45.[254,910,1138,1568]" captionTargetPageId="45" captionText="Fig. 19. Distribution map for the barbarossa species-group." figureDoi="http://doi.org/10.5281/zenodo.3786670" httpUri="https://zenodo.org/record/3786670/files/figure.png" pageId="46" pageNumber="47">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="discussion">
<paragraph blockId="46.[106,1074,897,1479]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[106,211,897,923]" pageId="46" pageNumber="47">Remarks</emphasis>
: We have compared Kenyan specimens identified as
<taxonomicName authorityName="Roth" authorityYear="1851" box="[803,904,898,923]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[803,904,898,923]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
with southeast African specimens identified as
<taxonomicName authorityName="Kolbe" authorityYear="1914" box="[465,597,932,958]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[465,597,932,958]" italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
by
<bibRefCitation author="Paschalidis, K. M." box="[640,851,932,958]" pageId="46" pageNumber="47" refId="ref30542" refString="Paschalidis, K. M. (1974 a) The genus Sisyphus Latr. (Coleoptera: Scarabaeidae) in Southern Africa. MSc. Thesis, Rhodes University, Grahamstown, South Africa, 183 pp." type="book" year="1974">Paschalidis (1974)</bibRefCitation>
. Females of
<taxonomicName authorityName="Kolbe" authorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Roth" authorityYear="1851" box="[220,323,967,992]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[220,323,967,992]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
are close morphologically making it difficult to separate them into species. However, we have observed that males of southern Africa “
<taxonomicName authorityName="Kolbe" authorityYear="1914" box="[873,1005,1001,1027]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[873,1005,1001,1027]" italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
” bear a short blunt triangular projection on the mid-posterior edge of the metafemur and a short projecting metatrochanter. By contrast, males of east African “
<taxonomicName authorityName="Roth" authorityYear="1851" box="[900,1003,1071,1096]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[900,1003,1071,1096]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
” have a sharp projecting spine on the mid-posterior edge of the metafemur and a metatrochanter as large as in
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[346,734,1140,1167]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jossoi">
<emphasis box="[346,535,1140,1166]" italics="true" pageId="46" pageNumber="47">Neosisyphus jossoi</emphasis>
<bibRefCitation author="Montreuil, O." box="[542,734,1140,1167]" pageId="46" pageNumber="47" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="46.[106,1074,897,1479]" pageId="46" pageNumber="47">
<bibRefCitation author="Montreuil" box="[135,351,1174,1201]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil (2015b)</bibRefCitation>
synonymized
<taxonomicName authorityName="Kolbe" authorityYear="1914" box="[516,648,1174,1200]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[516,648,1174,1200]" italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
with
<taxonomicName authority="(Roth 1851)" baseAuthorityName="Roth" baseAuthorityYear="1851" box="[715,1070,1174,1201]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[715,916,1174,1200]" italics="true" pageId="46" pageNumber="47">Neosisyphus setiger</emphasis>
(Roth 1851)
</taxonomicName>
. The synonymy seems to be based on the
<typeStatus box="[578,677,1209,1235]" pageId="46" pageNumber="47">holotype</typeStatus>
female of
<taxonomicName baseAuthorityName="Roth" baseAuthorityYear="1851" box="[800,904,1210,1235]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[800,904,1210,1235]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
from
<collectingCountry box="[977,1074,1209,1235]" name="Ethiopia" pageId="46" pageNumber="47">Ethiopia</collectingCountry>
and a
<typeStatus box="[181,283,1244,1270]" pageId="46" pageNumber="47">lectotype</typeStatus>
male of
<taxonomicName authorityName="Kolbe" authorityYear="1914" box="[391,525,1244,1270]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis box="[391,525,1244,1270]" italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
from
<collectingCountry box="[600,705,1244,1270]" name="Tanzania" pageId="46" pageNumber="47">Tanzania</collectingCountry>
. This is questionable as females are often difficult to separate into species as stated above. Furthermore, although the photograph of a major male of
<taxonomicName authority="(Montreuil 2015 b)" baseAuthorityName="Montreuil" baseAuthorityYear="2015" box="[481,817,1313,1340]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[481,588,1314,1339]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
(
<bibRefCitation author="Montreuil" box="[606,809,1313,1340]" firstAuthor="Montreuil" pageId="46" pageNumber="47" pagination="1 - 36" refId="ref30184" refString="Montreuil, O. (2015 b) Le genre Neosisyphus Muller em Afrique (Coleoptera, Scarabaeidae, Sisyphini). Catharsius La Revue, 12 (1): 1 - 36." type="journal article" year="2015" yearSuffix="b">Montreuil 2015b</bibRefCitation>
)
</taxonomicName>
closely resembles “
<taxonomicName authorityName="Kolbe" authorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="confrater">
<emphasis italics="true" pageId="46" pageNumber="47">N. confrater</emphasis>
</taxonomicName>
”, there is no indication of where the individual was collected. Nevertheless, we have retained the name
<taxonomicName baseAuthorityName="Montreuil" baseAuthorityYear="2015" box="[431,537,1383,1408]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[431,537,1383,1408]" italics="true" pageId="46" pageNumber="47">N. setiger</emphasis>
</taxonomicName>
for southern African material as we have not been able to compare it with the type and non-type material used in Montreuils study (2015b).
</paragraph>
</subSubSection>
</treatment>
</document>