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<document ID-DOI="http://doi.org/10.5281/zenodo.3794455" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962590" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794455" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. &amp; Scholtz, Clarke H." docDate="2020" docId="D474D525FFB07C49D7619954FEC5FBFA" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Neosisyphus tembyi Daniel &amp; Davis &amp; Sole &amp; Scholtz 2020, sp. n." docType="treatment" docVersion="6" lastPageId="45" lastPageNumber="46" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="43" pageNumber="44" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel, Gimo M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davis, Adrian Lv.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sole, Catherine L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Insect Systematics &amp; Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-01-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>61</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
<mods:identifier type="GBIF-Dataset">a725007b-2d51-4c92-838a-9e8fd27484c8</mods:identifier>
<mods:identifier type="ISSN">1876-312X</mods:identifier>
<mods:identifier type="Zenodo-Dep">3786629</mods:identifier>
<mods:identifier type="ZooBank">701C1742-718D-4486-A158-AEA608BA8576</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794455" ID-GBIF-Taxon="163962590" ID-Zenodo-Dep="3794455" LSID="urn:lsid:plazi:treatment:D474D525FFB07C49D7619954FEC5FBFA" httpUri="http://treatment.plazi.org/id/D474D525FFB07C49D7619954FEC5FBFA" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection box="[98,744,1295,1322]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph blockId="43.[98,1008,1295,1356]" box="[98,744,1295,1322]" pageId="43" pageNumber="44">
<heading box="[98,744,1295,1322]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="3">
<emphasis bold="true" box="[98,594,1295,1322]" pageId="43" pageNumber="44">
<taxonomicName authority="Daniel &amp; Davis" authorityName="Daniel &amp; Davis &amp; Sole &amp; Scholtz" authorityYear="2020" box="[98,522,1295,1322]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tembyi" status="sp. n.">
<emphasis bold="true" box="[98,327,1295,1322]" italics="true" pageId="43" pageNumber="44">Neosisyphus tembyi</emphasis>
Daniel &amp; Davis
</taxonomicName>
<taxonomicNameLabel box="[529,594,1296,1322]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[609,687,1296,1322]" captionStart="Fig" captionStartId="23.[98,131,990,1012]" captionTargetBox="[189,976,131,962]" captionTargetId="figure@23.[189,976,131,965]" captionTargetPageId="23" captionText="Fig. 7. Distribution map for the goryi species-group." figureDoi="http://doi.org/10.5281/zenodo.3786644" httpUri="https://zenodo.org/record/3786644/files/figure.png" pageId="43" pageNumber="44">Figs. 7</figureCitation>
,
<figureCitation box="[700,735,1296,1322]" captionStart="Fig" captionStartId="25.[98,131,1628,1650]" captionTargetBox="[362,802,1074,1603]" captionTargetId="figure@25.[362,802,1074,1603]" captionTargetPageId="25" captionText="Fig. 8. Images of the habitus of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786646" httpUri="https://zenodo.org/record/3786646/files/figure.png" pageId="43" pageNumber="44">8C</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[98,1008,1330,1356]" pageId="43" pageNumber="44" type="description">
<paragraph blockId="43.[98,1008,1295,1356]" box="[98,1008,1330,1356]" pageId="43" pageNumber="44">
ZooBank: http://zoobank.org/
<uuid box="[445,1008,1330,1356]" pageId="43" pageNumber="44">476EBD87-13C2-4DE0-A31E-883EEC69DA66</uuid>
</paragraph>
</subSubSection>
<subSubSection lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="diagnosis">
<paragraph blockId="43.[98,1066,1399,1635]" lastBlockId="44.[106,1074,741,906]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<emphasis bold="true" box="[98,217,1399,1425]" pageId="43" pageNumber="44">Diagnosis</emphasis>
:
<taxonomicName authority="Daniel &amp; Davis &amp; Sole &amp; Scholtz, 2020" authorityName="Daniel &amp; Davis &amp; Sole &amp; Scholtz" authorityYear="2020" box="[236,349,1400,1426]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tembyi" status="sp. n.">
<emphasis box="[236,349,1400,1426]" italics="true" pageId="43" pageNumber="44">N. tembyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[361,430,1400,1426]" pageId="43" pageNumber="44">
<taxonomicNameLabel box="[361,430,1400,1426]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[593,994,1400,1427]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="jossoi">
<emphasis box="[593,786,1400,1426]" italics="true" pageId="43" pageNumber="44">Neosisyphus jossoi</emphasis>
<bibRefCitation author="Montreuil, O." box="[798,994,1400,1427]" pageId="43" pageNumber="44" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
</taxonomicName>
. Both species are roughly the same size and lack sharp spines on the posterior margin of the metafemur. However,
<taxonomicName authority="Daniel &amp; Davis &amp; Sole &amp; Scholtz, 2020" authorityName="Daniel &amp; Davis &amp; Sole &amp; Scholtz" authorityYear="2020" box="[350,459,1469,1495]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tembyi" status="sp. n.">
<emphasis box="[350,459,1469,1495]" italics="true" pageId="43" pageNumber="44">N. tembyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[467,532,1469,1495]" pageId="43" pageNumber="44">
<taxonomicNameLabel box="[467,532,1469,1495]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
bears a small angled protuberance on the midposterior margin of the metafemur in males. The metatrochanter projection in major males of the new species is ½ of the total length of the metafemur and is somewhat inwardly projected at the tip. Unlike in
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[550,641,1574,1599]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="jossoi">
<emphasis box="[550,641,1574,1599]" italics="true" pageId="43" pageNumber="44">N. jossoi</emphasis>
</taxonomicName>
, in which the metatrochanter projection is ¾ of the total length of the metafemur in major males and strongly inwardly projected. Southern African specimens of
<taxonomicName authority="Roth (1851)" authorityName="Roth" authorityYear="1851" box="[586,841,741,768]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[586,690,742,767]" italics="true" pageId="44" pageNumber="45">N. setiger</emphasis>
Roth (1851)
</taxonomicName>
differ from the new species by bearing a shorter metatrochanter in males, which is ¼ of the total length of the metafemur. Furthermore, the metatrochanter in Southern African
<taxonomicName authorityName="Roth" authorityYear="1851" box="[967,1074,811,836]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="setiger">
<emphasis box="[967,1074,811,836]" italics="true" pageId="44" pageNumber="45">N. setiger</emphasis>
</taxonomicName>
projects straight and is never bent on the tip. The phallobase and parameres of all of the above species are distinctly different.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3786668" ID-Zenodo-Dep="3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="44" pageNumber="45" startId="44.[106,139,640,662]" targetBox="[413,766,132,615]" targetPageId="44">
<paragraph blockId="44.[106,1073,640,692]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[106,185,640,662]" pageId="44" pageNumber="45">Fig. 18.</emphasis>
Images of the aedeagi of the
<taxonomicName authorityName="Wiedemann" authorityYear="1823" box="[491,584,641,662]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="barbarossa">
<emphasis box="[491,584,641,662]" italics="true" pageId="44" pageNumber="45">barbarossa</emphasis>
</taxonomicName>
species-group:
<emphasis bold="true" box="[737,752,640,661]" pageId="44" pageNumber="45">A</emphasis>
.
<taxonomicName box="[769,896,641,663]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="barabrossa">
<emphasis box="[769,896,641,663]" italics="true" pageId="44" pageNumber="45">N. barabrossa</emphasis>
</taxonomicName>
;
<emphasis bold="true" box="[914,929,641,662]" pageId="44" pageNumber="45">B</emphasis>
.
<taxonomicName authorityName="Klug" authorityYear="1855" box="[946,1068,641,663]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis box="[946,1068,641,663]" italics="true" pageId="44" pageNumber="45">N. calcaratus</emphasis>
</taxonomicName>
;
<emphasis bold="true" box="[106,123,670,692]" pageId="44" pageNumber="45">C</emphasis>
.
<taxonomicName authority="Daniel &amp; Davis &amp; Sole &amp; Scholtz, 2020" authorityName="Daniel &amp; Davis &amp; Sole &amp; Scholtz" authorityYear="2020" box="[134,223,670,692]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tembyi" status="sp. n">
<emphasis box="[134,223,670,692]" italics="true" pageId="44" pageNumber="45">N. tembyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[229,276,670,691]" pageId="44" pageNumber="45">
<taxonomicNameLabel box="[229,276,670,691]" pageId="44" pageNumber="45" rank="species">sp. n</taxonomicNameLabel>
</emphasis>
;
<emphasis bold="true" box="[288,307,670,691]" pageId="44" pageNumber="45">D</emphasis>
.
<taxonomicName authorityName="Roth" authorityYear="1851" box="[319,405,671,692]" class="Insecta" family="Scarabaeidae" genus="Neosisyphus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="setiger">N. setiger</taxonomicName>
.
</paragraph>
</caption>
<subSubSection lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
<paragraph blockId="44.[106,1074,948,1634]" lastBlockId="45.[98,1066,134,404]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis bold="true" box="[106,248,949,975]" pageId="44" pageNumber="45">Description</emphasis>
: Male
<typeStatus box="[331,442,949,975]" pageId="44" pageNumber="45">Holotype</typeStatus>
:
<emphasis bold="true" box="[455,502,948,974]" pageId="44" pageNumber="45">Size</emphasis>
. Male: length:
<quantity box="[681,776,949,976]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="44" pageNumber="45" unit="mm" value="7.3">7.3 mm</quantity>
; width:
<quantity box="[872,968,949,976]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="44" pageNumber="45" unit="mm" value="4.5">4.5 mm</quantity>
.
<emphasis bold="true" box="[981,1066,949,975]" pageId="44" pageNumber="45">Colour</emphasis>
. Black body with brown setae; meso- and metasternum black; antennal club brown.
<emphasis bold="true" box="[106,170,1018,1044]" pageId="44" pageNumber="45">Head</emphasis>
. Anterior edge of clypeus with shallow sinuosity between medial teeth; lateral extreme of clypeal margin simply arcuate. Clypeo-genal suture markedly cleft; genae elongate. Frons and vertex dorsally granular, setose and minutely punctate, epicranial suture clearly visible.
<emphasis bold="true" box="[355,478,1122,1148]" pageId="44" pageNumber="45">Pronotum</emphasis>
. Convex with a sheen on dorsal surface; maximum width as long as the maximum length; setae inserted between ocellate punctures. Incomplete lateral prothoracic ridge between pronotal disc and prothoracic episternum.
<emphasis bold="true" box="[106,178,1226,1252]" pageId="44" pageNumber="45">Elytra</emphasis>
. Broad proximally and narrow posteriorly; well-spaced setae on interstriae; elytral surface shiny, distinct antero-lateral humeral callus; striae fine with double marginal lines interrupted by smooth ocellate strial punctures; granulation basally on elytral striae 47.
<emphasis bold="true" box="[197,314,1330,1356]" pageId="44" pageNumber="45">Pygidium</emphasis>
. Bearing U-shaped, ocellate punctation, a metallic sheen and setation.
<emphasis bold="true" box="[166,269,1364,1390]" pageId="44" pageNumber="45">Sternites</emphasis>
. Margins between abdominal sternites setigerous laterally, and glabrous medially with minute ocellate punctation; meso-metasternal suture visible; meso- and metasternum finely punctate and setose; metasternum with a punctate depression on the postero-medial edge. Mesepimeron and metepisternum finely punctate and setose.
<emphasis bold="true" box="[106,160,1504,1530]" pageId="44" pageNumber="45">Legs</emphasis>
. In ventral view, profemur carinate antero-laterally, punctate and internally pubescent with a lateral row of fine and well-developed setae; mesofemur finely punctate and setose, lacking any projection posteriorly; metafemur with an angled-protuberance mid-posteriorly with fine punctation and setae; metatrochanter projected backwards almost reaching top of angled protuberance; tip of metatrochanter slightly project- ed inward, but always parallel with metafemur. Metacoxa punctate and shagreened ventro-anteriorly. Protibia with three teeth and single terminal spine; meso-metatibia densely setose, serrated laterally, with two terminal spines on mesotibia, and two terminal spurs on metatibia; pro-, meso- and metatarsus five-segmented, with two claws; mostly setose.
<emphasis bold="true" box="[265,377,307,333]" pageId="45" pageNumber="46">Aedeagus</emphasis>
. Parameres separated by visible dorsal membranous portion extending nearly to the middle; arciform ridge bordering the membrane laterally, deeply notched medially; inwardly curved on the apex (
<figureCitation box="[672,776,377,403]" captionStart="Fig" captionStartId="44.[106,139,640,662]" captionTargetBox="[413,766,132,615]" captionTargetId="figure@44.[413,766,131,615]" captionTargetPageId="44" captionText="Fig. 18. Images of the aedeagi of the barbarossa species-group: A. N. barabrossa; B. N. calcaratus; C. N. tembyi sp. n; D. N. setiger." figureDoi="http://doi.org/10.5281/zenodo.3786668" httpUri="https://zenodo.org/record/3786668/files/figure.png" pageId="45" pageNumber="46">Fig. 18C</figureCitation>
).
</paragraph>
<paragraph blockId="45.[98,391,481,507]" box="[98,391,481,507]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[98,391,481,507]" pageId="45" pageNumber="46">Morphological variation</emphasis>
</paragraph>
<paragraph blockId="45.[98,1066,533,629]" pageId="45" pageNumber="46">Males bear a backwardly projected metatrochanter that is curved inwardly at the tip; this projection is absent in females where the metatrochanter is contiguous with the metafemur. The pronotum in males is larger than in females.</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="materials_examined">
<paragraph blockId="45.[98,383,671,698]" box="[98,383,671,698]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[98,383,671,698]" pageId="45" pageNumber="46">Examined type material</emphasis>
</paragraph>
<paragraph blockId="45.[98,1066,724,820]" pageId="45" pageNumber="46">
<materialsCitation ID-GBIF-Occurrence="2610400349" collectingDate="1974-12-18" collectionCode="SANC" collectorName="Davis &amp; Temby &amp; Ex &amp; Div. Entomology S. African Station" country="Angola" latitude="-14.816667" location="Paiva Coucero" longLatPrecision="537" longitude="15.55" municipality="African Station" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[98,209,724,750]" pageId="45" pageNumber="46">Holotype</typeStatus>
: (
<specimenCount box="[232,266,724,751]" pageId="45" pageNumber="46" type="male">1♂</specimenCount>
<collectionCode box="[274,352,724,750]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="45" pageNumber="46" type="Museum">SANC</collectionCode>
) /
<collectingCountry box="[375,459,724,750]" name="Angola" pageId="45" pageNumber="46">Angola</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:D474D525FFB07C49D7619954FEC5FBFA:590232E8FFB67C49D6D49E8FFDF8FD32" box="[471,634,724,750]" country="Angola" latitude="-14.816667" longLatPrecision="537" longitude="15.55" municipality="African Station" name="Paiva Coucero" pageId="45" pageNumber="46">Paiva Coucero</location>
(
<geoCoordinate box="[649,798,724,750]" degrees="14.816667" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="1" value="-14.816667">14.816667°S</geoCoordinate>
<geoCoordinate box="[806,898,724,751]" degrees="15.55" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="555" value="15.55">15.55°E</geoCoordinate>
) (50 &amp;
<quantity box="[992,1066,724,751]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="45" pageNumber="46" unit="km" value="45.0">45 km</quantity>
E),
<date box="[141,274,758,784]" pageId="45" pageNumber="46" value="1974-12-18">
<collectingDate box="[141,274,758,784]" pageId="45" pageNumber="46" value="1974-12-18">18.xii.1974</collectingDate>
</date>
, leg:
<collectorName box="[336,400,758,784]" pageId="45" pageNumber="46">Davis</collectorName>
&amp;
<collectorName box="[442,524,758,784]" pageId="45" pageNumber="46">Temby</collectorName>
/, /
<collectorName box="[554,585,759,785]" pageId="45" pageNumber="46">Ex</collectorName>
coll CSIRO,
<collectorName pageId="45" pageNumber="46">
Div. Entomology S.
<collectingMunicipality pageId="45" pageNumber="46">African Station</collectingMunicipality>
</collectorName>
/.
<typeStatus box="[203,313,794,820]" pageId="45" pageNumber="46">Paratypes</typeStatus>
: (
<specimenCount box="[335,370,793,820]" pageId="45" pageNumber="46" type="female">1♀</specimenCount>
SANC) same data as
<typeStatus box="[617,720,793,819]" pageId="45" pageNumber="46">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="distribution">
<paragraph blockId="45.[98,1066,862,923]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[98,249,862,888]" pageId="45" pageNumber="46">Distribution</emphasis>
: The species is only known from woodland vegetation in southern
<collectingCountry name="Angola" pageId="45" pageNumber="46">Angola</collectingCountry>
(Paiva Coucero) (
<figureCitation box="[349,432,897,923]" captionStart="Fig" captionStartId="45.[98,131,1593,1615]" captionTargetId="figure@45.[254,910,1138,1568]" captionTargetPageId="45" captionText="Fig. 19. Distribution map for the barbarossa species-group." figureDoi="http://doi.org/10.5281/zenodo.3786670" httpUri="https://zenodo.org/record/3786670/files/figure.png" pageId="45" pageNumber="46">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="etymology">
<paragraph blockId="45.[98,1066,966,1062]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[98,228,966,992]" pageId="45" pageNumber="46">Etymology</emphasis>
: The patronym is named after Ian Temby, a former researcher at the CSIRO, Division of Entomology, South African station; who was one of the collectors of the
<typeStatus box="[171,219,1036,1062]" pageId="45" pageNumber="46">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>