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<document ID-DOI="http://doi.org/10.5281/zenodo.3794437" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962606" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794437" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. &amp; Scholtz, Clarke H." docDate="2020" docId="D474D525FF957C6BD7699847FF3FFEF7" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Sisyphus manni Montreuil 2015" docType="treatment" docVersion="6" lastPageId="15" lastPageNumber="16" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="14" pageNumber="15" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel, Gimo M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davis, Adrian Lv.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sole, Catherine L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Insect Systematics &amp; Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-01-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>1</mods:start>
<mods:end>61</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
<mods:identifier type="GBIF-Dataset">a725007b-2d51-4c92-838a-9e8fd27484c8</mods:identifier>
<mods:identifier type="ISSN">1876-312X</mods:identifier>
<mods:identifier type="Zenodo-Dep">3786629</mods:identifier>
<mods:identifier type="ZooBank">701C1742-718D-4486-A158-AEA608BA8576</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3794437" ID-GBIF-Taxon="163962606" ID-Zenodo-Dep="3794437" LSID="urn:lsid:plazi:treatment:D474D525FF957C6BD7699847FF3FFEF7" httpUri="http://treatment.plazi.org/id/D474D525FF957C6BD7699847FF3FFEF7" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection pageId="14" pageNumber="15" type="nomenclature">
<paragraph blockId="14.[106,1073,1052,1219]" box="[106,613,1052,1079]" pageId="14" pageNumber="15">
<heading box="[106,613,1052,1079]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="3">
<taxonomicName authority="Montreuil, 2015" authorityName="Montreuil" authorityYear="2015" box="[106,497,1052,1079]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="manni">
<emphasis bold="true" box="[106,497,1052,1079]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[106,292,1052,1079]" italics="true" pageId="14" pageNumber="15">Sisyphus manni</emphasis>
<bibRefCitation author="Montreuil, O." box="[300,497,1053,1079]" pageId="14" pageNumber="15" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015">Montreuil, 2015</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[513,600,1053,1079]" captionStart="Fig" captionStartId="11.[98,131,957,979]" captionTargetBox="[99,1066,132,930]" captionTargetId="figure@11.[98,1066,131,931]" captionTargetPageId="11" captionText="Fig. 2. Images of the habitus of the muricatus species-group: A. S. muricatus; B. S. neobornemisszanus; C. S. fasciculatus; D. S. perissinottoi; E. S. manni; F. S. swazi sp. n. G. S. alveatus. I. Fully developed hind wing, which is typical to most of Sisyphus species in southern Africa H. Brachypterous hind wing of S. alveatus. J. Sclerite represents a reduced hind wing of S. swazi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786634" httpUri="https://zenodo.org/record/3786634/files/figure.png" pageId="14" pageNumber="15">Fig. 2E</figureCitation>
).
</heading>
</paragraph>
<paragraph blockId="14.[106,1073,1052,1219]" box="[106,358,1088,1115]" pageId="14" pageNumber="15">
<bibRefCitation author="Montreuil" box="[106,358,1088,1115]" firstAuthor="Montreuil" pageId="14" pageNumber="15" pagination="281 - 293" refId="ref30219" refString="Montreuil, O. (2015 c) Premiers cas de brachypterisme dans le genre Sisyphus Latreille (Coleoptera, Scarabaeidae, Sisyphini). Annales de la Societe entomologique de France, 51 (4): 281 - 293." type="journal article" year="2015" yearSuffix="c">Montreuil 2015c: 23</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="materials_examined">
<paragraph blockId="14.[106,1073,1052,1219]" box="[135,675,1122,1149]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[135,288,1122,1148]" pageId="14" pageNumber="15">Type locality</emphasis>
:
<collectingRegion box="[303,411,1122,1148]" country="South Africa" name="Limpopo" pageId="14" pageNumber="15">Limpopo</collectingRegion>
, Kruger National Park.
</paragraph>
<paragraph blockId="14.[106,1073,1052,1219]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[135,182,1156,1182]" pageId="14" pageNumber="15">Size</emphasis>
: Male: length: 5.54.0 mm; width: 2.52.0 mm. Female: length: 5.54.0 mm, width 2.52.0 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="diagnosis">
<paragraph blockId="14.[106,1074,1261,1392]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[106,225,1261,1287]" pageId="14" pageNumber="15">Diagnosis</emphasis>
: In
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[272,373,1261,1287]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="manni">
<emphasis box="[272,373,1261,1287]" italics="true" pageId="14" pageNumber="15">S. manni</emphasis>
</taxonomicName>
, proximal elytral setae are primarily distributed uniformly becoming arranged in sparse tufts mid-basally and posteriorly, unlike in
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[915,1073,1296,1322]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="perissinottoi">
<emphasis box="[915,1073,1296,1322]" italics="true" pageId="14" pageNumber="15">S. perissinottoi</emphasis>
</taxonomicName>
where dense tufts of setae occur across the entire elytra.
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[750,852,1331,1357]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="manni">
<emphasis box="[750,852,1331,1357]" italics="true" pageId="14" pageNumber="15">S. manni</emphasis>
</taxonomicName>
bears three depressions on the pronotal disc, which are lacking in
<emphasis box="[646,811,1365,1391]" italics="true" pageId="14" pageNumber="15">
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[646,807,1365,1391]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="perissinottoi">S. perissinottoi</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="14.[106,390,1434,1460]" box="[106,390,1434,1460]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[106,390,1434,1460]" pageId="14" pageNumber="15">Examined type material</emphasis>
</paragraph>
<paragraph blockId="14.[106,837,1486,1513]" box="[106,837,1486,1513]" pageId="14" pageNumber="15">
<typeStatus box="[106,217,1486,1512]" pageId="14" pageNumber="15">Holotype</typeStatus>
: (MHNH, through photograph without locality data).
</paragraph>
<paragraph blockId="14.[106,830,1555,1582]" box="[106,830,1555,1582]" pageId="14" pageNumber="15">
<emphasis bold="true" box="[106,447,1555,1582]" pageId="14" pageNumber="15">Examined non-type material</emphasis>
: See Supplementary information.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="distribution">
<paragraph blockId="15.[98,1066,134,299]" pageId="15" pageNumber="16">
<emphasis bold="true" box="[98,249,134,160]" pageId="15" pageNumber="16">Distribution</emphasis>
:
<taxonomicName authorityName="Montreuil" authorityYear="2015" box="[270,377,135,161]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="manni">
<emphasis box="[270,377,135,161]" italics="true" pageId="15" pageNumber="16">S. manni</emphasis>
</taxonomicName>
is only known from
<collectingCountry box="[646,799,134,160]" name="South Africa" pageId="15" pageNumber="16">South Africa</collectingCountry>
. The species is associated with upland to highland grassland and open woodland in
<collectingRegion box="[858,956,169,195]" country="South Africa" name="Gauteng" pageId="15" pageNumber="16">Gauteng</collectingRegion>
(cited as
<taxonomicName authorityName="Boucomont" authorityYear="1935" box="[98,220,204,230]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alveatus">
<emphasis box="[98,220,204,230]" italics="true" pageId="15" pageNumber="16">S. alveatus</emphasis>
</taxonomicName>
by
<bibRefCitation author="Davis" box="[276,493,204,231]" etAl="et al." firstAuthor="Davis" pageId="15" pageNumber="16" pagination="1 - 16" refId="ref29080" refString="Davis, A. L. V., Scholtz, C. H., &amp; Deschodt, C. (2005) A dung beetle survey of selected Gauteng nature reserves: implications for conservation of the provincial scarabaeine fauna. African entomology, 13 (1): 1 - 16." type="journal article" year="2005">Davis et al. 2005</bibRefCitation>
), KwaZulu Natal,
<collectingRegion box="[724,875,204,230]" country="South Africa" name="Mpumalanga" pageId="15" pageNumber="16">Mpumalanga</collectingRegion>
and Northwest Provinces. It has also been collected in unshaded riverine vegetation in
<collectingRegion box="[961,1066,238,264]" country="South Africa" name="Limpopo" pageId="15" pageNumber="16">Limpopo</collectingRegion>
(
<figureCitation box="[106,175,273,299]" captionStart="Fig" captionStartId="13.[98,131,1559,1581]" captionTargetId="figure@13.[234,930,923,1535]" captionTargetPageId="13" captionText="Fig. 4. Distribution map for the muricatus species-groups." figureDoi="http://doi.org/10.5281/zenodo.3786638" httpUri="https://zenodo.org/record/3786638/files/figure.png" pageId="15" pageNumber="16">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>