treatments-xml/data/D4/74/D5/D474D525FF827C7ED7619F30FD13F9F2.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3794403" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962622" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794403" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. &amp; Scholtz, Clarke H." docDate="2020" docId="D474D525FF827C7ED7619F30FD13F9F2" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Sisyphus umbraphilus Daniel &amp; Davis 2016" docType="treatment" docVersion="6" lastPageId="26" lastPageNumber="27" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="25" pageNumber="26" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel, Gimo M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davis, Adrian Lv.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sole, Catherine L.</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Scholtz, Clarke H.</mods:namePart>
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<mods:title>Insect Systematics &amp; Evolution</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-01-02</mods:number>
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<mods:detail type="volume">
<mods:number>51</mods:number>
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<mods:number>1</mods:number>
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<mods:end>61</mods:end>
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<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
<mods:identifier type="GBIF-Dataset">a725007b-2d51-4c92-838a-9e8fd27484c8</mods:identifier>
<mods:identifier type="ISSN">1876-312X</mods:identifier>
<mods:identifier type="Zenodo-Dep">3786629</mods:identifier>
<mods:identifier type="ZooBank">701C1742-718D-4486-A158-AEA608BA8576</mods:identifier>
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<paragraph blockId="25.[98,1065,840,1006]" box="[98,777,875,902]" pageId="25" pageNumber="26">
<heading box="[98,777,875,902]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="3">
<taxonomicName authority="Daniel &amp; Davis, 2016" authorityName="Daniel &amp; Davis" authorityYear="2016" box="[98,629,875,902]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis bold="true" box="[98,629,875,902]" pageId="25" pageNumber="26">
<emphasis bold="true" box="[98,359,875,902]" italics="true" pageId="25" pageNumber="26">Sisyphus umbraphilus</emphasis>
<bibRefCitation author="Daniel &amp; Davis" box="[366,629,875,901]" firstAuthor="Daniel" pageId="25" pageNumber="26" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, &amp; Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel &amp; Davis, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[645,722,876,902]" captionStart="Fig" captionStartId="25.[98,131,1628,1650]" captionTargetBox="[362,802,1074,1603]" captionTargetId="figure@25.[362,802,1074,1603]" captionTargetPageId="25" captionText="Fig. 8. Images of the habitus of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786646" httpUri="https://zenodo.org/record/3786646/files/figure.png" pageId="25" pageNumber="26">Figs. 8</figureCitation>
, 
<figureCitation box="[735,769,876,902]" captionStart="Fig" captionStartId="26.[106,139,572,594]" captionTargetBox="[487,689,132,546]" captionTargetId="figure@26.[487,692,131,547]" captionTargetPageId="26" captionText="Fig. 9. Images of the aedeagi of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786648" httpUri="https://zenodo.org/record/3786648/files/figure.png" pageId="25" pageNumber="26">9A</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[98,390,910,937]" pageId="25" pageNumber="26" type="reference_group">
<paragraph blockId="25.[98,1065,840,1006]" box="[98,390,910,937]" pageId="25" pageNumber="26">
<treatmentCitationGroup box="[98,390,910,937]" pageId="25" pageNumber="26">
<treatmentCitation box="[98,390,910,937]" pageId="25" pageNumber="26">
<bibRefCitation author="Daniel" box="[98,390,910,937]" etAl="et al." firstAuthor="Daniel" pageId="25" pageNumber="26" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, &amp; Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel et al. 2016: 67–69</bibRefCitation>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="description">
<paragraph blockId="25.[98,1065,840,1006]" pageId="25" pageNumber="26">
<emphasis bold="true" box="[128,175,944,970]" pageId="25" pageNumber="26">Size</emphasis>
: Male: length: 
<quantity box="[346,490,945,972]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="7.1" metricValueMin="5.1" pageId="25" pageNumber="26" unit="mm" value="6.1" valueMax="7.1" valueMin="5.1">5.1–7.1 mm</quantity>
, width: 
<quantity box="[582,726,945,972]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.2" metricValueMin="2.9" pageId="25" pageNumber="26" unit="mm" value="3.55" valueMax="4.2" valueMin="2.9">2.9–4.2 mm</quantity>
; Female: length: 
<quantity box="[917,1061,945,972]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.4" metricValueMin="5.4" pageId="25" pageNumber="26" unit="mm" value="6.4" valueMax="7.4" valueMin="5.4">5.4–7.4 mm</quantity>
, width 
<quantity box="[172,317,980,1006]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="25" pageNumber="26" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.2–4.2 mm</quantity>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3786646" ID-Zenodo-Dep="3786646" box="[98,859,1628,1650]" httpUri="https://zenodo.org/record/3786646/files/figure.png" pageId="25" pageNumber="26" startId="25.[98,131,1628,1650]" targetBox="[362,802,1074,1603]" targetPageId="25">
<paragraph blockId="25.[98,859,1628,1650]" box="[98,859,1628,1650]" pageId="25" pageNumber="26">
<emphasis bold="true" box="[98,160,1628,1650]" pageId="25" pageNumber="26">Fig. 8.</emphasis>
Images of the habitus of the 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[436,548,1628,1649]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[436,548,1628,1649]" italics="true" pageId="25" pageNumber="26">umbraphilus</emphasis>
</taxonomicName>
species-group: 
<emphasis bold="true" box="[690,705,1628,1649]" pageId="25" pageNumber="26">A</emphasis>
. 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[717,855,1628,1650]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="umbraphilus">S. umbraphilus</taxonomicName>
.
</paragraph>
</caption>
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<paragraph blockId="26.[106,867,572,594]" box="[106,867,572,594]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[106,167,572,594]" pageId="26" pageNumber="27">Fig. 9.</emphasis>
Images of the aedeagi of the 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[443,555,572,593]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[443,555,572,593]" italics="true" pageId="26" pageNumber="27">umbraphilus</emphasis>
</taxonomicName>
species-group: 
<emphasis bold="true" box="[697,712,572,593]" pageId="26" pageNumber="27">A</emphasis>
. 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[724,863,572,594]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">S. umbraphilus</taxonomicName>
.
</paragraph>
</caption>
<subSubSection box="[135,768,668,695]" pageId="26" pageNumber="27" type="distribution">
<paragraph blockId="26.[135,768,668,695]" box="[135,768,668,695]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[135,288,668,694]" pageId="26" pageNumber="27">Type locality</emphasis>
: Umfolozi, KwaZulu Natal (
<collectingCountry box="[615,761,668,695]" name="South Africa" pageId="26" pageNumber="27">South Africa</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="diagnosis">
<paragraph blockId="26.[106,1074,741,941]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[106,225,741,767]" pageId="26" pageNumber="27">Diagnosis</emphasis>
: 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[240,405,741,768]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[240,405,741,768]" italics="true" pageId="26" pageNumber="27">S. umbraphilus</emphasis>
</taxonomicName>
belongs to its own species-group. However, it is somewhat similar to 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[224,344,776,802]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oralensis">
<emphasis box="[224,344,776,802]" italics="true" pageId="26" pageNumber="27">S. oralensis</emphasis>
</taxonomicName>
. In this case, 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[507,673,776,802]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[507,673,776,802]" italics="true" pageId="26" pageNumber="27">S. umbraphilus</emphasis>
</taxonomicName>
is distinguished by having tufts of epipleural setae. It bears relatively small ocellate punctures on the pronotum. Furthemore, In 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[246,410,845,872]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[246,410,845,872]" italics="true" pageId="26" pageNumber="27">S. umbraphilus</emphasis>
</taxonomicName>
, the parameres are simple and attenuated towards the apex (
<figureCitation box="[114,200,880,906]" captionStart="Fig" captionStartId="26.[106,139,572,594]" captionTargetBox="[487,689,132,546]" captionTargetId="figure@26.[487,692,131,547]" captionTargetPageId="26" captionText="Fig. 9. Images of the aedeagi of the umbraphilus species-group: A. S. umbraphilus." figureDoi="http://doi.org/10.5281/zenodo.3786648" httpUri="https://zenodo.org/record/3786648/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
) whereas, in 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[350,469,880,906]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oralensis">
<emphasis box="[350,469,880,906]" italics="true" pageId="26" pageNumber="27">S. oralensis</emphasis>
</taxonomicName>
they are somewhat obtuse and truncated towards the apex (
<figureCitation box="[172,275,914,941]" captionStart="Fig" captionStartId="36.[106,139,735,757]" captionTargetBox="[247,932,131,708]" captionTargetId="figure@36.[247,932,131,710]" captionTargetPageId="36" captionText="Fig. 14. Images of the habitus of the seminulum species-group: A. S. splendidus; B. S. oralensis; C. S. nanniscus; D. S. auricomus sp. n. Pronotal setation and punctation patterns of S. oralensis (E), and S. auricomus sp. n (F)." figureDoi="http://doi.org/10.5281/zenodo.3786660" httpUri="https://zenodo.org/record/3786660/files/figure.png" pageId="26" pageNumber="27">Fig. 14A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="materials_examined">
<paragraph blockId="26.[106,390,983,1010]" box="[106,390,983,1010]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[106,390,983,1010]" pageId="26" pageNumber="27">Examined type material</emphasis>
</paragraph>
<paragraph blockId="26.[106,1074,1036,1340]" pageId="26" pageNumber="27">
<materialsCitation ID-GBIF-Occurrence="2610400336" collectingDate="1970-10-01" collectingDateMax="1970-10-07" collectingDateMin="1970-10-01" collectionCode="SANC" collectorName="Bornemissza &amp; Aschenborn" country="South Africa" latitude="-28.25" location="Umfolozi" longLatPrecision="1233" longitude="31.916666" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[106,217,1036,1062]" pageId="26" pageNumber="27">Holotype</typeStatus>
: (
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SANC): 
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, KZN, 
<location LSID="urn:lsid:plazi:treatment:D474D525FF827C7ED7619F30FD13F9F2:590232E8FF817C7ED5AB9857FC91FBFA" box="[680,787,1036,1062]" country="South Africa" latitude="-28.25" longLatPrecision="1233" longitude="31.916666" name="Umfolozi" pageId="26" pageNumber="27">Umfolozi</location>
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<geoCoordinate box="[894,988,1036,1063]" degrees="31" direction="east" minutes="55" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="31.916666">31°55’E</geoCoordinate>
, 
<date pageId="26" pageNumber="27" value="1970-10-01" valueMax="1970-10-07" valueMin="1970-10-01">
<collectingDate pageId="26" pageNumber="27" value="1970-10-01" valueMax="1970-10-07" valueMin="1970-10-01">1–7.x. 1970</collectingDate>
</date>
, leg: 
<collectorName box="[233,374,1070,1096]" pageId="26" pageNumber="27">Bornemissza</collectorName>
&amp; 
<collectorName box="[421,561,1070,1096]" pageId="26" pageNumber="27">Aschenborn</collectorName>
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="2610400340" collectingDate="1978-10-15" collectingDateMax="2010-11-11" collectingDateMin="1978-10-15" collectorName="McGeogh &amp; Davis. &amp; Bornemissza &amp; Aschenborn. &amp; Weke-Weke Stream &amp; Shongweni Farm &amp; AJ Armstrong &amp; G Van Bassouw" country="South Africa" latitude="-29.8" location="Pretoria" longLatPrecision="1225" longitude="30.716667" municipality="Ithala Game Reserve" pageId="26" pageNumber="27" specimenCount="39" specimenCount-female="25" specimenCount-male="14" stateProvince="Gauteng" typeStatus="paratype">
<typeStatus box="[575,685,1071,1097]" pageId="26" pageNumber="27">Paratypes</typeStatus>
: (
<specimenCount box="[709,746,1071,1097]" pageId="26" pageNumber="27" type="male">3♂</specimenCount>
, 
<specimenCount box="[764,801,1071,1097]" pageId="26" pageNumber="27" type="female">9♀</specimenCount>
SANC) with the same data as holotype. (
<specimenCount box="[324,359,1105,1132]" pageId="26" pageNumber="27" type="male">1♂</specimenCount>
, 
<specimenCount box="[377,414,1105,1132]" pageId="26" pageNumber="27" type="female">2♀</specimenCount>
UPSA): 
<collectingCountry box="[522,740,1105,1131]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
, KZN, 
<collectingMunicipality box="[834,1074,1105,1132]" pageId="26" pageNumber="27">Ithala Game Reserve</collectingMunicipality>
<geoCoordinate box="[106,257,1140,1167]" degrees="27" direction="south" minutes="27" orientation="latitude" pageId="26" pageNumber="27" precision="1" seconds="53.6" value="-27.46489">27°27’53.6”S</geoCoordinate>
<geoCoordinate box="[264,422,1140,1167]" degrees="31" direction="east" minutes="15" orientation="longitude" pageId="26" pageNumber="27" precision="1" seconds="36.1" value="31.260027">31°15’36.1”E</geoCoordinate>
, 
<date box="[433,545,1140,1166]" pageId="26" pageNumber="27" value="1999-01-13">
<collectingDate box="[433,545,1140,1166]" pageId="26" pageNumber="27" value="1999-01-13">13.i.1999</collectingDate>
</date>
, leg: Chown, 
<collectorName box="[700,814,1140,1166]" pageId="26" pageNumber="27">McGeogh</collectorName>
&amp; 
<collectorName box="[850,922,1140,1166]" pageId="26" pageNumber="27">Davis.</collectorName>
(
<specimenCount box="[937,974,1140,1167]" pageId="26" pageNumber="27" type="male">2♂</specimenCount>
SANC): 
<collectingCountry box="[106,325,1175,1201]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
, KZN, Jozini 
<geoCoordinate box="[498,613,1174,1200]" degrees="27" direction="south" minutes="26" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="0" value="-27.433332">27°26’0”S</geoCoordinate>
<geoCoordinate box="[625,731,1175,1201]" degrees="32" direction="east" minutes="4" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="0" value="32.066666">32°4’0”E</geoCoordinate>
, 
<date box="[747,865,1174,1201]" pageId="26" pageNumber="27" value="1978-10-15">
<collectingDate box="[747,865,1174,1201]" pageId="26" pageNumber="27" value="1978-10-15">15.x.1978</collectingDate>
</date>
, leg: 
<collectorName box="[932,1073,1174,1200]" pageId="26" pageNumber="27">Bornemissza</collectorName>
&amp; 
<collectorName box="[142,286,1209,1235]" pageId="26" pageNumber="27">Aschenborn.</collectorName>
(
<specimenCount box="[307,344,1209,1236]" pageId="26" pageNumber="27" type="male">7♂</specimenCount>
, 
<specimenCount box="[363,415,1209,1236]" pageId="26" pageNumber="27" type="female">14♀</specimenCount>
SANC): 
<collectingCountry box="[532,752,1209,1235]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
, 
<collectingRegion box="[769,870,1209,1235]" country="South Africa" name="Gauteng" pageId="26" pageNumber="27">Gauteng</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:D474D525FF827C7ED7619F30FD13F9F2:590232E8FF817C7ED47598E2FC50FB0F" box="[886,978,1209,1235]" country="South Africa" latitude="-29.8" longLatPrecision="1225" longitude="30.716667" municipality="Ithala Game Reserve" name="Pretoria" pageId="26" pageNumber="27" stateProvince="Gauteng">Pretoria</location>
, 
<locationDeviation location="Pretoria Farm" pageId="26" pageNumber="27">
<quantity box="[994,1073,1209,1236]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="km" value="35.0">35 km</quantity>
NE of Pretoria Farm
</locationDeviation>
<geoCoordinate box="[350,466,1244,1271]" degrees="25" direction="south" minutes="42" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="0" value="-25.7">25°42’0”S</geoCoordinate>
<geoCoordinate box="[474,595,1244,1270]" degrees="28" direction="east" minutes="13" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="0" value="28.216667">28°13’0”E</geoCoordinate>
, 
<date box="[608,785,1244,1270]" pageId="26" pageNumber="27" value="1983-12-12" valueMax="1983-12-13" valueMin="1983-12-12">
<collectingDate box="[608,785,1244,1270]" pageId="26" pageNumber="27" value="1983-12-12" valueMax="1983-12-13" valueMin="1983-12-12">12–13.xii.1983</collectingDate>
</date>
, leg: Davis. (
<specimenCount box="[938,972,1244,1271]" pageId="26" pageNumber="27" type="male">1♂</specimenCount>
SANC): 
<collectingCountry box="[106,325,1279,1305]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
, KZN, 
<collectorName box="[422,649,1278,1305]" pageId="26" pageNumber="27">Weke-Weke Stream</collectorName>
, 
<collectorName box="[665,862,1278,1305]" pageId="26" pageNumber="27">Shongweni Farm</collectorName>
<geoCoordinate box="[874,963,1279,1305]" degrees="29" direction="south" minutes="48" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-29.8">29°48’S</geoCoordinate>
<geoCoordinate box="[974,1069,1279,1305]" degrees="30" direction="east" minutes="43" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="30.716667">30°43’E</geoCoordinate>
, 
<date box="[106,231,1313,1339]" pageId="26" pageNumber="27" value="2010-11-11">
<collectingDate box="[106,231,1313,1339]" pageId="26" pageNumber="27" value="2010-11-11">11.xi.2010</collectingDate>
</date>
, leg: 
<collectorName box="[290,448,1313,1339]" pageId="26" pageNumber="27">AJ Armstrong</collectorName>
&amp; 
<collectorName box="[486,666,1313,1340]" pageId="26" pageNumber="27">G Van Bassouw</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="26.[106,830,1382,1409]" box="[106,830,1382,1409]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[106,447,1382,1408]" pageId="26" pageNumber="27">Examined non-type material</emphasis>
: See Supplementary information.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="distribution">
<paragraph blockId="26.[106,1074,1451,1583]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[106,257,1451,1477]" pageId="26" pageNumber="27">Distribution</emphasis>
: 
<taxonomicName authorityName="Daniel &amp; Davis" authorityYear="2016" box="[275,443,1452,1478]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="umbraphilus">
<emphasis box="[275,443,1452,1478]" italics="true" pageId="26" pageNumber="27">S. umbraphilus</emphasis>
</taxonomicName>
has been recorded in patches of dense woodland and thicket in KwaZulu Natal (
<bibRefCitation author="Daniel" box="[418,630,1486,1513]" etAl="et al." firstAuthor="Daniel" pageId="26" pageNumber="27" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, &amp; Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel et al. 2016</bibRefCitation>
) as well as in dense riverine woodland in 
<collectingRegion box="[136,374,1521,1548]" country="South Africa" name="North West" pageId="26" pageNumber="27">North West Province</collectingRegion>
(
<collectingCountry box="[388,538,1521,1548]" name="South Africa" pageId="26" pageNumber="27">South Africa</collectingCountry>
). In 
<collectingRegion box="[587,688,1521,1547]" country="South Africa" name="Gauteng" pageId="26" pageNumber="27">Gauteng</collectingRegion>
, it has been collected from shaded vegetation on deep sands (
<bibRefCitation author="Davis" box="[402,536,1556,1583]" firstAuthor="Davis" pageId="26" pageNumber="27" pagination="260 - 280" refId="ref28946" refString="Davis, A. L. V. (1996) Habitat associations in a South African, summer rainfall, dung beetle community (Coleoptera: Scarabaeidae, Aphodiidae, Staphylinidae, Histeridae, Hydrophilidae). Pedobiologia, 40 (3): 260 - 280." type="journal article" year="1996">Davis 1996</bibRefCitation>
) (
<figureCitation box="[559,643,1556,1582]" captionStart="Fig" captionStartId="27.[98,131,462,484]" captionTargetId="figure@27.[323,841,131,437]" captionTargetPageId="27" captionText="Fig. 10. Distribution maps for the umbraphilus species-groups." figureDoi="http://doi.org/10.5281/zenodo.3786650" httpUri="https://zenodo.org/record/3786650/files/figure.png" pageId="26" pageNumber="27">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>