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<document ID-DOI="http://doi.org/10.5281/zenodo.3794401" ID-GBIF-Dataset="a725007b-2d51-4c92-838a-9e8fd27484c8" ID-GBIF-Taxon="163962623" ID-ISSN="1876-312X" ID-Zenodo-Dep="3794401" ID-ZooBank="701C1742-718D-4486-A158-AEA608BA8576" checkinTime="1588685582300" checkinUser="felipe" docAuthor="Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. &amp; Scholtz, Clarke H." docDate="2020" docId="D474D525FF807C78D7619E4FFDCFFE53" docLanguage="en" docName="InsectSystEvol.51.1.1-61.pdf.imf" docOrigin="Insect Systematics &amp; Evolution 51 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2a2c15f9" docStyleName="insectsystevol.2020.journal_article" docTitle="Sisyphus impressipennis Lansberge 1886" docType="treatment" docVersion="6" lastPageId="28" lastPageNumber="29" masterDocId="284DAD5DFF9B7C64D7039C5BFF82FFDC" masterDocTitle="Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions" masterLastPageNumber="61" masterPageNumber="1" pageId="27" pageNumber="28" updateTime="1636469099162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions</mods:title>
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<mods:namePart>Daniel, Gimo M.</mods:namePart>
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<mods:namePart>Davis, Adrian Lv.</mods:namePart>
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<mods:namePart>Sole, Catherine L.</mods:namePart>
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<mods:namePart>Scholtz, Clarke H.</mods:namePart>
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<mods:title>Insect Systematics &amp; Evolution</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-01-02</mods:number>
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<mods:number>51</mods:number>
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<mods:identifier type="DOI">10.1163/1876312X-00002195</mods:identifier>
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<taxonomicName authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[98,586,532,559]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impressipennis">
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<emphasis bold="true" box="[98,382,532,559]" italics="true" pageId="27" pageNumber="28">Sisyphus impressipennis</emphasis>
<bibRefCitation author="Lansberge" box="[389,586,533,559]" firstAuthor="Lansberge" pageId="27" pageNumber="28" pagination="69 - 120" refId="ref29861" refString="Lansberge, J. W. (1886) Scarabaeides, Buprestides et Cerambycides de l'afrique occidentale. Notes from Leyden Museum, 8 (2): 69 - 120." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[602,694,533,559]" captionStart="Fig" captionStartId="28.[106,139,1481,1503]" captionTargetBox="[107,1069,760,1456]" captionTargetId="figure@28.[106,1074,758,1456]" captionTargetPageId="28" captionText="Fig. 11. Images of the habitus of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n. F. elytra composed of alternate rows of well and less-developed setae, normally seen in S. costatus and S. australis sp. n. G. Head of S. costatus shows a concavity between the medial teeth on the anterior edge of the clypeus. H. Head of S. australis sp. n shows a straight margin between the medial teeth on the anterior edge of the clypeus; there are defined carinae on the clypeo-frontal suture, and scattered punctation on the vertex. I. Frons of S. inconspicuus sp. n without clear punctation. J. S. gazanus: bearing vertex and frons densely and fully punctate." figureDoi="http://doi.org/10.5281/zenodo.3786652" httpUri="https://zenodo.org/record/3786652/files/figure.png" pageId="27" pageNumber="28">Figs. 11</figureCitation>
,
<figureCitation box="[707,759,533,559]" captionStart="Fig" captionStartId="29.[98,131,1599,1621]" captionTargetBox="[346,816,1085,1574]" captionTargetId="figure@29.[346,817,1081,1574]" captionTargetPageId="29" captionText="Fig. 12. Images of the aedeagi of the costatus species-group: A. S. impressipennis; B. S. costatus; C. S. gazanus; D. S. inconspicuus sp. n; E. S. australis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3786654" httpUri="https://zenodo.org/record/3786654/files/figure.png" pageId="27" pageNumber="28">12A</figureCitation>
).
</heading>
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<paragraph blockId="27.[98,1066,532,907]" pageId="27" pageNumber="28">
<treatmentCitationGroup pageId="27" pageNumber="28">
<treatmentCitation author="Lansberge, J. W." box="[98,382,568,594]" page="73" pageId="27" pageNumber="28" year="1886">
<bibRefCitation author="Lansberge" box="[98,382,568,594]" firstAuthor="Lansberge" pageId="27" pageNumber="28" pagination="69 - 120" refId="ref29861" refString="Lansberge, J. W. (1886) Scarabaeides, Buprestides et Cerambycides de l'afrique occidentale. Notes from Leyden Museum, 8 (2): 69 - 120." type="journal article" year="1886">Lansberge 1886: 7374</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation author="Gillet, J. J. E." box="[399,589,568,594]" page="23" pageId="27" pageNumber="28" year="1911">
<bibRefCitation author="Gillet" box="[399,589,568,594]" firstAuthor="Gillet" pageId="27" pageNumber="28" pagination="1 - 263" refId="ref29484" refString="Gillet, J. J. E. (1911) Scarabaeidae: Coprinae I. Coleopterorum Catalogus, 38: 1 - 263." type="journal article" year="1911">Gillet 1911: 23</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Arrow, G. J." box="[607,847,568,595]" page="462" pageId="27" pageNumber="28" year="1927">
<bibRefCitation author="Arrow" box="[607,847,568,595]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="456 - 465" refId="ref28180" refString="Arrow, G. J. (1927) Notes on the Coleopterous genus Sisyphus. Annals and Magazine of natural History, 9 (19): 456 - 465." type="journal article" year="1927">Arrow 1927: 4625</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Haaf, E." box="[865,1061,568,595]" page="362" pageId="27" pageNumber="28" year="1955">
<bibRefCitation author="Haaf" box="[865,1061,568,595]" firstAuthor="Haaf" pageId="27" pageNumber="28" pagination="341 - 381" refId="ref29533" refString="Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381." type="journal article" year="1955">Haaf 1955: 362</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="" box="[98,398,602,629]" page="124" pageId="27" pageNumber="28">
<bibRefCitation author="Schafer, P. &amp; Fischer, E." box="[98,398,602,629]" pageId="27" pageNumber="28" pagination="119 - 137" refId="ref30806" refString="Schafer, P., &amp; Fischer, E. (1992) Zur Scarabaeiden-Fauna Rwandas, Zentralafrika. Sisyphini (Insecta: Coleoptera: Lamellicornia). Senckenbergiana biologica, 72 (1 - 3), 119 - 137." type="journal article" year="1992">Schefer-Fischer 1992: 124</bibRefCitation>
</treatmentCitation>
,
<bibRefCitation author="Schafer, P. &amp; Fischer, E." box="[410,704,602,629]" pageId="27" pageNumber="28" pagination="1 - 93" refId="ref30770" refString="Schafer, P. &amp; Fischer, E. (2001) Zur Scarabaeidenfauna Rwandas, Zentralafrika 2, Gymnopleurini und nachtrag Sisyphini, Mitteilungen des internationalen entomologischen Vereins supplement 9: 1 - 93." type="journal article" year="2001">Schaefer-Fischer, 2001:58</bibRefCitation>
,
<treatmentCitation author="Montreuil, O." box="[716,958,602,629]" page="96" pageId="27" pageNumber="28" year="2015">
<bibRefCitation author="Montreuil" box="[716,958,602,629]" firstAuthor="Montreuil" pageId="27" pageNumber="28" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015" yearSuffix="a">Montreuil 2015a: 96</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<treatmentCitationGroup pageId="27" pageNumber="28">
<paragraph blockId="27.[98,1066,532,907]" box="[98,532,637,663]" pageId="27" pageNumber="28">
<taxonomicName authority="Peringuey, 1908" authorityName="Peringuey" authorityYear="1908" box="[98,532,637,663]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis box="[98,342,637,663]" italics="true" pageId="27" pageNumber="28">Sisyphus transvaalensis</emphasis>
Péringuey, 1908
</taxonomicName>
</paragraph>
<paragraph blockId="27.[98,1066,532,907]" pageId="27" pageNumber="28">
<treatmentCitation author="" box="[98,347,672,699]" page="557" pageId="27" pageNumber="28">
<bibRefCitation box="[98,347,672,699]" pageId="27" pageNumber="28">Péringuey 1908: 557</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Arrow, G. J." box="[363,568,672,698]" page="462" pageId="27" pageNumber="28" year="1927">
<bibRefCitation author="Arrow" box="[363,568,672,698]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="456 - 465" refId="ref28180" refString="Arrow, G. J. (1927) Notes on the Coleopterous genus Sisyphus. Annals and Magazine of natural History, 9 (19): 456 - 465." type="journal article" year="1927">Arrow 1927: 462</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Haaf, E." box="[600,789,672,699]" page="364" pageId="27" pageNumber="28" year="1955">
<bibRefCitation author="Haaf" box="[600,789,672,699]" firstAuthor="Haaf" pageId="27" pageNumber="28" pagination="341 - 381" refId="ref29533" refString="Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381." type="journal article" year="1955">Haaf 1955: 364</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Balthasar, V." box="[821,1061,672,699]" page="954" pageId="27" pageNumber="28" year="1968">
<bibRefCitation author="Balthasar" box="[821,1061,672,699]" firstAuthor="Balthasar" pageId="27" pageNumber="28" pagination="953 - 958" refId="ref28212" refString="Balthasar, V. (1968) Neue Scarabaeiden-Arten, 129. Beitrag zur kenntnis der Scarabaeoidea (Coleoptera). Beitrage zur Entomologie, 18: 953 - 958." type="journal article" year="1968">Balthasar 1968: 954</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Ferreira, M. C." box="[98,320,706,733]" page="807" pageId="27" pageNumber="28" year="1972">
<bibRefCitation author="Ferreira" box="[98,320,706,733]" firstAuthor="Ferreira" pageId="27" pageNumber="28" pagination="5 - 1088" refId="ref29371" refString="Ferreira, M. C. (1972) Os Escarabideos de Africa (sul do Saara). Revista de Entomologia de Mocambique, 11 (1968 - 1969): 5 - 1088." type="journal article" year="1972">Ferreira 1972: 807</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Montreuil, O." box="[331,568,706,733]" page="96" pageId="27" pageNumber="28" year="2015">
<bibRefCitation author="Montreuil" box="[331,568,706,733]" firstAuthor="Montreuil" pageId="27" pageNumber="28" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015" yearSuffix="a">Montreuil 2015a: 96</bibRefCitation>
</treatmentCitation>
</paragraph>
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<treatmentCitationGroup pageId="27" pageNumber="28">
<paragraph blockId="27.[98,1066,532,907]" box="[98,441,741,767]" pageId="27" pageNumber="28">
<taxonomicName authority="Arrow, 1909" authorityName="Arrow" authorityYear="1909" box="[98,441,741,767]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callosipes">
<emphasis box="[98,291,741,767]" italics="true" pageId="27" pageNumber="28">Sisyphus callosipes</emphasis>
<bibRefCitation author="Arrow" box="[298,441,741,767]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="517 - 523" refId="ref28134" refString="Arrow, G. J. (1909) On some new species of Coleoptera from Rhodesia and adjacent territories. The Annals and Magazine of natural History, including Zoology, Botany and Geology. London 8 (3): 517 - 523." type="journal article" year="1909">Arrow, 1909</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="27.[98,1066,532,907]" box="[98,958,776,803]" pageId="27" pageNumber="28">
<treatmentCitation author="Arrow, G. J." box="[98,297,776,803]" page="517" pageId="27" pageNumber="28" year="1909">
<bibRefCitation author="Arrow" box="[98,297,776,803]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="517 - 523" refId="ref28134" refString="Arrow, G. J. (1909) On some new species of Coleoptera from Rhodesia and adjacent territories. The Annals and Magazine of natural History, including Zoology, Botany and Geology. London 8 (3): 517 - 523." type="journal article" year="1909">Arrow 1909: 517</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Arrow, G. J." box="[308,507,776,802]" page="462" pageId="27" pageNumber="28" year="1927">
<bibRefCitation author="Arrow" box="[308,507,776,802]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="456 - 465" refId="ref28180" refString="Arrow, G. J. (1927) Notes on the Coleopterous genus Sisyphus. Annals and Magazine of natural History, 9 (19): 456 - 465." type="journal article" year="1927">Arrow 1927: 462</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Haaf, E." box="[519,702,776,803]" page="365" pageId="27" pageNumber="28" year="1955">
<bibRefCitation author="Haaf" box="[519,702,776,803]" firstAuthor="Haaf" pageId="27" pageNumber="28" pagination="341 - 381" refId="ref29533" refString="Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381." type="journal article" year="1955">Haaf 1955: 365</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Montreuil, O." box="[714,955,776,803]" page="96" pageId="27" pageNumber="28" year="2015">
<bibRefCitation author="Montreuil" box="[714,955,776,803]" firstAuthor="Montreuil" pageId="27" pageNumber="28" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015" yearSuffix="a">Montreuil 2015a: 96</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection box="[128,518,810,837]" pageId="27" pageNumber="28" type="distribution">
<paragraph blockId="27.[98,1066,532,907]" box="[128,518,810,837]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[128,281,810,836]" pageId="27" pageNumber="28">
<typeStatus box="[128,186,810,836]" pageId="27" pageNumber="28">Type</typeStatus>
locality
</emphasis>
: Humpata (
<collectingCountry box="[419,506,810,837]" name="Angola" pageId="27" pageNumber="28">Angola</collectingCountry>
).
</paragraph>
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<subSubSection pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[98,1066,532,907]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[128,175,845,871]" pageId="27" pageNumber="28">Size</emphasis>
: Male: length: 10.0
<quantity box="[423,517,845,872]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="27" pageNumber="28" unit="mm" value="7.1">7.1 mm</quantity>
; width: 5.64.0 mm; Female: length: 10.1
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="27" pageNumber="28" unit="mm" value="7.5">7.5 mm</quantity>
, width 5.14.0 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="diagnosis">
<paragraph blockId="27.[98,1066,949,1044]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[98,217,949,975]" pageId="27" pageNumber="28">Diagnosis</emphasis>
:
<taxonomicName authorityName="Lansberge" authorityYear="1886" box="[234,419,950,976]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impressipennis">
<emphasis box="[234,419,950,976]" italics="true" pageId="27" pageNumber="28">S. impressipennis</emphasis>
</taxonomicName>
is easily distinguished from other species in the
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1818" box="[984,1065,950,975]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="costatus">
<emphasis box="[984,1065,950,975]" italics="true" pageId="27" pageNumber="28">costatus</emphasis>
</taxonomicName>
species-group by bearing an anvil-shaped projection on the mid-ventral edge of the metafemur, mainly in males.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="materials_examined">
<paragraph blockId="27.[98,383,1087,1114]" box="[98,383,1087,1114]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[98,383,1087,1114]" pageId="27" pageNumber="28">
Examined
<typeStatus box="[225,276,1088,1114]" pageId="27" pageNumber="28">type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph blockId="27.[98,1066,1139,1479]" pageId="27" pageNumber="28">
<taxonomicName authority="Arrow, 1909" authorityName="Arrow" authorityYear="1909" box="[98,470,1139,1166]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callosipes">
<emphasis bold="true" box="[98,470,1139,1166]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[98,317,1139,1166]" italics="true" pageId="27" pageNumber="28">Sisyphus callosipes</emphasis>
<bibRefCitation author="Arrow" box="[323,470,1139,1165]" firstAuthor="Arrow" pageId="27" pageNumber="28" pagination="517 - 523" refId="ref28134" refString="Arrow, G. J. (1909) On some new species of Coleoptera from Rhodesia and adjacent territories. The Annals and Magazine of natural History, including Zoology, Botany and Geology. London 8 (3): 517 - 523." type="journal article" year="1909">Arrow, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
:
<materialsCitation ID-GBIF-Occurrence="2610400346" collectorName="Nyassa &amp; Simons &amp; Fry" country="Mozambique" location="Bacchus" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus box="[483,598,1140,1166]" pageId="27" pageNumber="28">Lectotype</typeStatus>
: (
<specimenCount box="[619,654,1140,1167]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
NHML), /
<collectingCountry box="[782,990,1140,1166]" name="Mozambique" pageId="27" pageNumber="28">MOZAMBIQUE</collectingCountry>
,
<collectorName pageId="27" pageNumber="28">Nyassa</collectorName>
/, /coll:
<collectorName box="[214,298,1174,1200]" pageId="27" pageNumber="28">Simons</collectorName>
&amp;
<collectorName box="[342,381,1175,1201]" pageId="27" pageNumber="28">Fry</collectorName>
, 1905/, handwritten on brown label /
<taxonomicName authority=", Arrow" authorityName="Arrow" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callopsis">
<emphasis box="[825,1018,1174,1200]" italics="true" pageId="27" pageNumber="28">Sisyphus callopsis,</emphasis>
Arrow
</taxonomicName>
<emphasis box="[150,192,1210,1235]" italics="true" pageId="27" pageNumber="28">type</emphasis>
/, /
<taxonomicName box="[228,412,1209,1235]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callopsis">
<emphasis box="[228,412,1209,1235]" italics="true" pageId="27" pageNumber="28">Sisyphus callopsis</emphasis>
</taxonomicName>
det. ME.
<location LSID="urn:lsid:plazi:treatment:D474D525FF807C78D7619E4FFDCFFE53:590232E8FF807C7FD51698E2FDF7FB0F" box="[533,629,1209,1235]" country="Mozambique" name="Bacchus" pageId="27" pageNumber="28">Bacchus</location>
, 1975
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2610400377" collectorName="Mozambique &amp; Nyassa &amp; Simons &amp; Fry" country="Mozambique" location="Bacchus" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus box="[716,832,1209,1235]" pageId="27" pageNumber="28">Lectotype</typeStatus>
/.
<typeStatus box="[853,1008,1209,1235]" pageId="27" pageNumber="28">Paralectotype</typeStatus>
: (
<specimenCount box="[1031,1066,1209,1236]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHML) /
<collectorName box="[214,368,1244,1270]" pageId="27" pageNumber="28">Mozambique</collectorName>
,
<collectorName box="[380,462,1244,1270]" pageId="27" pageNumber="28">Nyassa</collectorName>
/, /coll:
<collectorName box="[549,633,1244,1270]" pageId="27" pageNumber="28">Simons</collectorName>
&amp;
<collectorName box="[673,712,1244,1270]" pageId="27" pageNumber="28">Fry</collectorName>
, 1905/, handwritten on brown label/, /
<taxonomicName box="[186,371,1278,1304]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callopsis">
<emphasis box="[186,371,1278,1304]" italics="true" pageId="27" pageNumber="28">Sisyphus callopsis</emphasis>
</taxonomicName>
det. ME.
<location LSID="urn:lsid:plazi:treatment:D474D525FF807C78D7619E4FFDCFFE53:590232E8FF807C7FD6EF98A5FDCEFAC4" box="[492,588,1278,1304]" country="Mozambique" name="Bacchus" pageId="27" pageNumber="28">Bacchus</location>
, 1975
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2610400303" collectingDate="1907-10-18" collectorName="Kambove" country="Democratic Republic of the Congo" elevation="1219" location="Neave" municipality="Bacchus" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus box="[675,831,1278,1304]" pageId="27" pageNumber="28">Paralectotype</typeStatus>
/. (
<specimenCount box="[863,898,1279,1305]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
,
<specimenCount box="[914,951,1279,1305]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHML), /
<collectingCountry box="[108,173,1313,1339]" name="Democratic Republic of the Congo" pageId="27" pageNumber="28">RDC</collectingCountry>
,
<quantity box="[186,360,1313,1340]" metricMagnitude="5" metricUnit="m" metricValue="2.8163519999999997" metricValueMax="3.218688" metricValueMin="2.414016" pageId="27" pageNumber="28" unit="mi" value="175.0" valueMax="200.0" valueMin="150.0">150200 miles</quantity>
, W. of
<collectorName box="[445,552,1313,1339]" pageId="27" pageNumber="28">Kambove</collectorName>
(
<quantity box="[569,733,1313,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" metricValueMax="1.3716" metricValueMin="1.0668" pageId="27" pageNumber="28" unit="ft" value="4000.0" valueMax="4500.0" valueMin="3500.0">
<elevation box="[569,733,1313,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" metricValueMax="1.3716" metricValueMin="1.0668" pageId="27" pageNumber="28" unit="ft" value="4000.0" valueMax="4500.0" valueMin="3500.0">35004500 ft</elevation>
</quantity>
)/, /coll:
<location LSID="urn:lsid:plazi:treatment:D474D525FF807C78D7619E4FFDCFFE53:590232E8FF807C7FD43C9979FC0AFAE0" box="[831,904,1314,1340]" country="Democratic Republic of the Congo" municipality="Bacchus" name="Neave" pageId="27" pageNumber="28">Neave</location>
,
<date box="[917,1052,1313,1339]" pageId="27" pageNumber="28" value="1907-10-18">
<collectingDate box="[917,1052,1313,1339]" pageId="27" pageNumber="28" value="1907-10-18">18.10.1907</collectingDate>
</date>
/, /
<taxonomicName box="[108,290,1348,1374]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callopsis">
<emphasis box="[108,290,1348,1374]" italics="true" pageId="27" pageNumber="28">Sisyphus callopsis</emphasis>
</taxonomicName>
det. ME.
<collectingMunicipality box="[405,501,1348,1374]" pageId="27" pageNumber="28">Bacchus</collectingMunicipality>
, 1975.
<taxonomicName authority="Peringuey, 1908" authorityName="Peringuey" authorityYear="1908" box="[584,1059,1347,1374]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis bold="true" box="[584,1059,1347,1374]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[584,860,1347,1374]" italics="true" pageId="27" pageNumber="28">Sisyphus transvaalensis</emphasis>
Péringuey, 1908
</emphasis>
</taxonomicName>
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="2610400356" collectorName="Transvaal &amp; Pietersburg" country="South Africa" location="SOUTH AFRICA" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus box="[98,209,1383,1409]" pageId="27" pageNumber="28">Lectotype</typeStatus>
(designated here): (
<specimenCount box="[438,473,1383,1409]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
SAM): /
<collectingCountry box="[574,790,1383,1409]" name="South Africa" pageId="27" pageNumber="28">SOUTH AFRICA</collectingCountry>
,
<collectorName box="[801,910,1382,1408]" pageId="27" pageNumber="28">Transvaal</collectorName>
,
<collectorName box="[922,1053,1382,1408]" pageId="27" pageNumber="28">Pietersburg</collectorName>
/, handwritten on brown label, /
<taxonomicName authority="Peringuey" authorityName="Peringuey" authorityYear="1908" box="[453,827,1417,1443]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis box="[453,700,1417,1443]" italics="true" pageId="27" pageNumber="28">Sisyphus transvaalensis</emphasis>
Péringuey
</taxonomicName>
/, /red label, /TYPE/ (
<typeStatus box="[106,219,1452,1478]" pageId="27" pageNumber="28">Lectotype</typeStatus>
det. GM.
<bibRefCitation author="Daniel" box="[340,486,1452,1479]" firstAuthor="Daniel" pageId="27" pageNumber="28" pagination="67 - 74" refId="ref28393" refString="Daniel, G. M., Davis, A. L. V, &amp; Scholtz, C. H. (2016) Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa. Zootaxa, 4147 (1): 67 - 74." type="journal article" year="2016">Daniel 2016</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="27.[98,822,1515,1542]" box="[98,822,1515,1542]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[98,440,1515,1542]" pageId="27" pageNumber="28">Examined non-type material</emphasis>
: See Supplementary information.
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="distribution">
<paragraph blockId="27.[98,1066,1585,1681]" lastBlockId="28.[106,1073,134,230]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis bold="true" box="[98,249,1585,1611]" pageId="27" pageNumber="28">Distribution</emphasis>
:
<taxonomicName box="[267,459,1586,1612]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impresssipennis">
<emphasis box="[267,459,1586,1612]" italics="true" pageId="27" pageNumber="28">S. impresssipennis</emphasis>
</taxonomicName>
was described from south-west
<collectingCountry box="[832,916,1585,1611]" name="Angola" pageId="27" pageNumber="28">Angola</collectingCountry>
. It is distributed in east-central and southern Africa (
<bibRefCitation author="Davis" box="[562,757,1620,1647]" etAl="et al." firstAuthor="Davis" pageId="27" pageNumber="28" pagination="1 - 16" refId="ref29080" refString="Davis, A. L. V., Scholtz, C. H., &amp; Deschodt, C. (2005) A dung beetle survey of selected Gauteng nature reserves: implications for conservation of the provincial scarabaeine fauna. African entomology, 13 (1): 1 - 16." type="journal article" year="2005">Davis et al. 2005</bibRefCitation>
) where it is associated with shaded areas in savanna and dense miombo woodland. It has been recorded from lowlands in northern
<collectingCountry box="[355,502,134,160]" name="South Africa" pageId="28" pageNumber="29">South Africa</collectingCountry>
, in upland Chirinda Forest (
<collectingCountry box="[831,958,134,161]" name="Zimbabwe" pageId="28" pageNumber="29">Zimbabwe</collectingCountry>
), Mt
<collectingRegion country="Malawi" name="Mulanje" pageId="28" pageNumber="29">Mulanje</collectingRegion>
(
<collectingCountry box="[177,267,169,196]" name="Malawi" pageId="28" pageNumber="29">Malawi</collectingCountry>
), Central
<collectingCountry box="[383,536,169,195]" name="Mozambique" pageId="28" pageNumber="29">Mozambique</collectingCountry>
,
<collectingRegion box="[550,650,170,196]" country="Tanzania" name="Manyara" pageId="28" pageNumber="29">Manyara</collectingRegion>
National Park (
<collectingCountry box="[835,940,169,196]" name="Tanzania" pageId="28" pageNumber="29">Tanzania</collectingCountry>
) and Kambove (Republic Democratic of
<collectingCountry box="[454,533,204,230]" name="Democratic Republic of the Congo" pageId="28" pageNumber="29">Congo</collectingCountry>
) (
<figureCitation box="[556,660,204,230]" captionStart="Fig" captionStartId="30.[106,139,1628,1650]" captionTargetId="figure@30.[212,968,1010,1602]" captionTargetPageId="30" captionText="Fig. 13. Distribution map for the costatus species-group." figureDoi="http://doi.org/10.5281/zenodo.3786656" httpUri="https://zenodo.org/record/3786656/files/figure.png" pageId="28" pageNumber="29">Fig. 13C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="discussion">
<paragraph blockId="28.[106,1073,268,399]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[106,211,268,294]" pageId="28" pageNumber="29">Remarks</emphasis>
: Examination of
<taxonomicName authorityName="Arrow" authorityYear="1909" box="[411,536,268,294]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="callosipes">
<emphasis box="[411,536,268,294]" italics="true" pageId="28" pageNumber="29">S. callosipes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Peringuey" authorityYear="1908" box="[595,771,268,294]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis box="[595,771,268,294]" italics="true" pageId="28" pageNumber="29">S. transvaalensis</emphasis>
</taxonomicName>
<typeStatus box="[779,827,268,294]" pageId="28" pageNumber="29">type</typeStatus>
material showed that the external morphology and genitalia are similar to
<taxonomicName authorityName="Lansberge" authorityYear="1886" box="[716,901,303,329]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="impressipennis">
<emphasis box="[716,901,303,329]" italics="true" pageId="28" pageNumber="29">S. impressipennis</emphasis>
</taxonomicName>
. Therefore, we regard both species as junior synonyms of
<taxonomicName authorityName="Lansberge" authorityYear="1886" box="[585,768,337,363]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="impressipennis">
<emphasis box="[585,768,337,363]" italics="true" pageId="28" pageNumber="29">S. impressipennis</emphasis>
</taxonomicName>
, as previously proposed by
<bibRefCitation author="Paschalidis, K. M." box="[106,312,372,398]" pageId="28" pageNumber="29" refId="ref30542" refString="Paschalidis, K. M. (1974 a) The genus Sisyphus Latr. (Coleoptera: Scarabaeidae) in Southern Africa. MSc. Thesis, Rhodes University, Grahamstown, South Africa, 183 pp." type="book" year="1974">Paschalidis (1974)</bibRefCitation>
and
<bibRefCitation author="Montreuil" box="[369,584,372,399]" firstAuthor="Montreuil" pageId="28" pageNumber="29" pagination="91 - 102" refId="ref30147" refString="Montreuil, O. (2015 a) Nouveaux Sisyphus Latreille d'afrique (Coleoptera, Scarabaeidae, Sisyphini). Bulletin de la Societe entomologique de France, 120 (1): 91 - 102." type="journal article" year="2015" yearSuffix="a">Montreuil (2015a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>