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<document id="42DFCC9D0656E2E4ADB19938A86B93C4" ID-CLB-Dataset="55748" ID-DOI="10.5281/zenodo.5301289" ID-GBIF-Dataset="55b2b0bc-d79f-48e3-8be1-81424185dea7" ID-ISSN="0253-116X" ID-Zenodo-Dep="5301289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630101010072" checkinUser="marcus" docAuthor="Assing, V." docDate="2013" docId="1974879EFFE4FFCBFF04EFEFFEE3FC48" docLanguage="en" docName="LinzerbiolBeitr.2013.45.2.1479-1520.pdf" docOrigin="Linzer biologische Beiträge 45 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Micrillus nigriceps Assing 2013, nov.comb., nov.stat." docType="treatment" docVersion="10" lastPageNumber="1497" masterDocId="E54DFFE6FFF5FFD9FFB6EA15FFE8FFA6" masterDocTitle="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1520" masterPageNumber="1479" pageNumber="1496" updateTime="1699152599027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4EAD1A2CD3523EB944EE7E397E653D1B">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="F9E920070F6BE74C514D4A55697DCD68">Assing, V.</mods:namePart>
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<treatment id="1974879EFFE4FFCBFF04EFEFFEE3FC48" ID-DOI="http://doi.org/10.5281/zenodo.6512328" ID-GBIF-Taxon="194955326" ID-Zenodo-Dep="6512328" LSID="urn:lsid:plazi:treatment:1974879EFFE4FFCBFF04EFEFFEE3FC48" httpUri="http://treatment.plazi.org/id/1974879EFFE4FFCBFF04EFEFFEE3FC48" lastPageId="18" lastPageNumber="1497" pageId="17" pageNumber="1496">
<subSubSection id="D9C76503FFE4FFC8FF04EFEFFCF1F9B6" box="[178,793,1530,1552]" pageId="17" pageNumber="1496" type="nomenclature">
<paragraph id="91623688FFE4FFC8FF04EFEFFCF1F9B6" blockId="17.[178,1003,1530,1552]" box="[178,793,1530,1552]" pageId="17" pageNumber="1496">
<heading id="CA2A81E4FFE4FFC8FF04EFEFFCF1F9B6" box="[178,793,1530,1552]" fontSize="8" level="2" pageId="17" pageNumber="1496" reason="2">
<emphasis id="A3A9EA9AFFE4FFC8FF04EFEFFCF1F9B6" bold="true" box="[178,793,1530,1552]" pageId="17" pageNumber="1496">
<taxonomicName id="56DD4D0BFFE4FFC8FF04EFEFFDDDF9B6" ID-CoL="9PLZ8" authority="(CAMERON 1931)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[178,565,1530,1552]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1496" phylum="Arthropoda" rank="species" species="nigriceps" status="nov.comb., nov.stat.">
<emphasis id="A3A9EA9AFFE4FFC8FF04EFEFFE9FF9B6" bold="true" box="[178,375,1530,1552]" italics="true" pageId="17" pageNumber="1496">Micrillus nigriceps</emphasis>
(
<bibRefCitation id="F54C4B79FFE4FFC8FE33EFEFFE1BF9A9" author="CAMERON M" box="[389,499,1530,1552]" pageId="17" pageNumber="1496" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON</bibRefCitation>
1931)
</taxonomicName>
,
<taxonomicNameLabel id="B89A57E1FFE4FFC8FDF7EFEFFCF1F9B6" box="[577,793,1530,1552]" pageId="17" pageNumber="1496">nov.comb., nov.stat.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE4FFC8FCA9EFEFFC03F9B6" box="[799,1003,1530,1552]" pageId="17" pageNumber="1496" type="description">
<paragraph id="91623688FFE4FFC8FCA9EFEFFC03F9B6" blockId="17.[178,1003,1530,1552]" box="[799,1003,1530,1552]" pageId="17" pageNumber="1496">
<heading id="CA2A81E4FFE4FFC8FCA9EFEFFC03F9B6" box="[799,1003,1530,1552]" fontSize="8" level="2" pageId="17" pageNumber="1496" reason="2">
(
<figureCitation id="09E62A0DFFE4FFC8FC90EFEFFC7FF9B6" box="[806,919,1530,1552]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="17" pageNumber="1496">Figs 14-16</figureCitation>
,
<figureCitation id="09E62A0DFFE4FFC8FC17EFEEFC0BF9B6" box="[929,995,1530,1552]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="17" pageNumber="1496">Map 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE4FFC8FF04EC34FD4DF993" box="[178,677,1569,1589]" pageId="17" pageNumber="1496" type="reference_group">
<paragraph id="91623688FFE4FFC8FF04EC34FD4DF993" blockId="17.[178,677,1569,1589]" box="[178,677,1569,1589]" pageId="17" pageNumber="1496">
<treatmentCitationGroup id="B1CD11A6FFE4FFC8FF04EC34FD4DF993" box="[178,677,1569,1589]" pageId="17" pageNumber="1496">
<treatmentCitation id="107C1099FFE4FFC8FF04EC34FD79F993" author="CAMERON M" box="[178,657,1569,1589]" page="214" pageId="17" pageNumber="1496" year="1931">
<taxonomicName id="56DD4D0BFFE4FFC8FF04EC34FD79F993" authority="CAMERON 1931: 214" authorityName="CAMERON" authorityPageNumber="214" authorityYear="1931" box="[178,657,1569,1589]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1496" phylum="Arthropoda" rank="subSpecies" species="indicum" subSpecies="nigriceps">
<emphasis id="A3A9EA9AFFE4FFC8FF04EC34FE23F993" box="[178,459,1569,1589]" italics="true" pageId="17" pageNumber="1496">Scymbalium indicum nigriceps</emphasis>
<bibRefCitation id="F54C4B79FFE4FFC8FE66EC34FD79F993" author="CAMERON M" box="[464,657,1569,1589]" pageId="17" pageNumber="1496" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">
CAMERON 1931:
<specimenCount id="87DBFD01FFE4FFC8FDD9EC34FD79F993" box="[623,657,1569,1589]" pageId="17" pageNumber="1496" type="generic">214</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE7FFCBFF04EAC2FD40FE35" pageId="18" pageNumber="1497" type="materials_examined">
<paragraph id="91623688FFE7FFCBFF04EAC2FE04FEFA" blockId="18.[178,1086,215,403]" pageId="18" pageNumber="1497">
Type material examined
<materialsCitation id="21B53CD5FFE7FFCBFDDFEAC2FE7AFE89" ID-GBIF-Occurrence="3760604417" collectionCode="BMNH" pageId="18" pageNumber="1497" specimenCount="1" typeStatus="lectotype">
<typeStatus id="4E66882AFFE7FFCBFDDFEAC2FD2CFF4D" box="[617,708,215,235]" pageId="18" pageNumber="1497" type="lectotype">Lectotype</typeStatus>
, present designation: &quot;Calcutta. / Atkinson Coll. 92-3 / Syntype / M. Cameron. Bequest. B.M. 1955-147. / Lectotypus
<taxonomicName id="56DD4D0BFFE7FFCBFF04EB11FE35FEBE" authority="Cameron" authorityName="Cameron" authorityYear="1931" box="[178,477,260,280]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE7FFCBFF04EB11FE97FEBE" box="[178,383,260,280]" italics="true" pageId="18" pageNumber="1497">Scymbalium nigriceps</emphasis>
Cameron
</taxonomicName>
, desig. V. Assing 2013 /
<taxonomicName id="56DD4D0BFFE7FFCBFD65EB11FC18FEBE" authority="(Cameron)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[723,1008,260,280]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="nigriceps">Micrillus nigriceps (Cameron)</taxonomicName>
, det. V. Assing 2013&quot; (
<collectionCode id="F7CCAE4DFFE7FFCBFE88EB0EFE6FFE89" box="[318,391,283,303]" pageId="18" pageNumber="1497">BMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="21B53CD5FFE7FFCBFE2FEB0EFE04FEFA" ID-GBIF-Occurrence="3760604394" collectionCode="BMNH" pageId="18" pageNumber="1497" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="4E66882AFFE7FFCBFE2FEB0EFDC9FE89" box="[409,545,283,303]" pageId="18" pageNumber="1497" type="paralectotype">Paralectotypes</typeStatus>
: 3, 2 sex? [all glued on the same label; forebodies of
<specimenCount id="87DBFD01FFE7FFCBFF04EB27FED2FEE0" box="[178,314,306,326]" pageId="18" pageNumber="1497" type="generic">two specimens</specimenCount>
missing]: same labels as lectotype, but with additional label &quot;
<taxonomicName id="56DD4D0BFFE7FFCBFCCAEB27FBD5FEE0" box="[892,1085,306,326]" pageId="18" pageNumber="1497">Scimbalium indicum</taxonomicName>
Epp.
<taxonomicName id="56DD4D0BFFE7FFCBFF54EB5DFE6DFEFA" authority="Cam." authorityName="CAMERON" authorityYear="1931" box="[226,389,328,348]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="variety" species="indicum" variety="nigriceps">v. nigriceps Cam.</taxonomicName>
&quot; (BMNH).
</materialsCitation>
</paragraph>
<paragraph id="91623688FFE7FFCBFF04EB72FD40FE35" blockId="18.[178,1086,215,403]" pageId="18" pageNumber="1497">
Additional material examined:
<materialsCitation id="21B53CD5FFE7FFCBFD12EB72FD4BFE35" ID-GBIF-Occurrence="3760604353" collectingDate="1976-05-16" collectionCode="NHMB" collectorName="Wittmer &amp; Baroni Urbani" country="India" elevation="1200" location="Cherrapunjee" pageId="18" pageNumber="1497" specimenCount="1" stateProvince="Meghalaya">
<collectingCountry id="E9CA7618FFE7FFCBFD12EB72FD0AFEDD" box="[676,738,359,379]" name="India" pageId="18" pageNumber="1497">India</collectingCountry>
:
<specimenCount id="87DBFD01FFE7FFCBFD50EB72FD07FEDD" box="[742,751,359,379]" pageId="18" pageNumber="1497">1</specimenCount>
,
<collectingRegion id="5319F86AFFE7FFCBFCBDEB72FC86FEDD" box="[779,878,359,379]" country="India" name="Meghalaya" pageId="18" pageNumber="1497">Meghalaya</collectingRegion>
,
<location id="94026053FFE7FFCBFCC0EB72FC05FEDD" LSID="urn:lsid:plazi:treatment:1974879EFFE4FFCBFF04EFEFFEE3FC48:94026053FFE7FFCBFCC0EB72FC05FEDD" box="[886,1005,359,379]" country="India" name="Cherrapunjee" pageId="18" pageNumber="1497" stateProvince="Meghalaya">Cherrapunjee</location>
,
<quantity id="56259B6DFFE7FFCBFC40EB72FBD2FEDD" box="[1014,1082,359,379]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="1497" unit="m" value="1200.0">
<elevation id="1AF0D1BBFFE7FFCBFC40EB72FBD2FEDD" box="[1014,1082,359,379]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="1497" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="E5631048FFE7FFCBFF69EB6AFED5FE35" box="[223,317,383,403]" pageId="18" pageNumber="1497" value="1976-05-16">
<collectingDate id="F527E9A0FFE7FFCBFF69EB6AFED5FE35" box="[223,317,383,403]" pageId="18" pageNumber="1497" value="1976-05-16">16.V.1976</collectingDate>
</date>
, leg.
<collectorName id="3C28535EFFE7FFCBFEDCEB6AFE5BFE35" box="[362,435,383,403]" pageId="18" pageNumber="1497">Wittmer</collectorName>
&amp;
<collectorName id="3C28535EFFE7FFCBFE78EB6AFDA4FE35" box="[462,588,383,403]" pageId="18" pageNumber="1497">Baroni Urbani</collectorName>
(
<collectionCode id="F7CCAE4DFFE7FFCBFDE1EB6AFD76FE35" box="[599,670,383,403]" pageId="18" pageNumber="1497">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE7FFCBFF04EBB4FE93FDEF" pageId="18" pageNumber="1497" type="discussion">
<paragraph id="91623688FFE7FFCBFF04EBB4FE93FDEF" blockId="18.[178,1086,417,586]" pageId="18" pageNumber="1497">
Comment:
<taxonomicName id="56DD4D0BFFE7FFCBFEDEEBB4FDA6FE11" authorityName="Cameron" authorityYear="1931" box="[360,590,417,439]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE7FFCBFEDEEBB4FDA6FE11" box="[360,590,417,439]" italics="true" pageId="18" pageNumber="1497">Scymbalium nigriceps</emphasis>
</taxonomicName>
was originally described as a &quot;variety&quot; of
<taxonomicName id="56DD4D0BFFE7FFCBFB9CEBB7FE2AFE72" authority="EPPELSHEIM. The" authorityName="EPPELSHEIM. The" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="A3A9EA9AFFE7FFCBFB9CEBB7FEEBFE72" italics="true" pageId="18" pageNumber="1497">S. indicum</emphasis>
EPPELSHEIM. The
</taxonomicName>
description is based on an unspecified number of
<typeStatus id="4E66882AFFE7FFCBFC52EBAAFBD5FE72" box="[996,1085,447,468]" pageId="18" pageNumber="1497" type="syntype">syntypes</typeStatus>
from &quot;Calcutta&quot; (
<bibRefCitation id="F54C4B79FFE7FFCBFED1EBC9FDE7FE54" author="CAMERON M" box="[359,527,476,498]" pageId="18" pageNumber="1497" refId="ref22842" refString="CAMERON M (1931) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Volume 2. - London, Taylor and Francis: viii + 257 pp." type="book" year="1931">CAMERON 1931</bibRefCitation>
). Six partly incomplete
<typeStatus id="4E66882AFFE7FFCBFCB9EBC9FC80FE57" box="[783,872,476,497]" pageId="18" pageNumber="1497" type="syntype">syntypes</typeStatus>
were located in the Cameron collection at the BMNH. A male is designated as the
<typeStatus id="4E66882AFFE7FFCBFC87EBECFC79FDA9" box="[817,913,505,527]" pageId="18" pageNumber="1497" type="lectotype">lectotype</typeStatus>
. An examination of the type material revealed that
<taxonomicName id="56DD4D0BFFE7FFCBFDBFE802FD69FD8A" authorityName="Cameron" authorityYear="1931" box="[521,641,535,556]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE7FFCBFDBFE802FD69FD8A" box="[521,641,535,556]" italics="true" pageId="18" pageNumber="1497">S. nigriceps</emphasis>
</taxonomicName>
represents a distinct species and belongs to the genus
<taxonomicName id="56DD4D0BFFE7FFCBFEAFE821FE9EFDEC" box="[281,374,564,586]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="genus">
<emphasis id="A3A9EA9AFFE7FFCBFEAFE821FE9EFDEC" box="[281,374,564,586]" italics="true" pageId="18" pageNumber="1497">Micrillus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE7FFCBFF04E84CFF1BFCB7" pageId="18" pageNumber="1497" type="description">
<paragraph id="91623688FFE7FFCBFF04E84CFBFBFD2A" blockId="18.[178,1086,601,652]" pageId="18" pageNumber="1497">
Redescription: Body length
<quantity id="56259B6DFFE7FFCBFDE5E84CFD3AFDC9" box="[595,722,601,623]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="18" pageNumber="1497" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5-5.5 mm</quantity>
; length of forebody
<quantity id="56259B6DFFE7FFCBFC0DE84CFBD2FDC9" box="[955,1082,601,623]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="18" pageNumber="1497" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7-2.9 mm</quantity>
. Coloration: body including appendages pale reddish, with the head somewhat darker.
</paragraph>
<paragraph id="91623688FFE7FFCBFF04E88EFD41FD17" blockId="18.[178,681,667,689]" box="[178,681,667,689]" pageId="18" pageNumber="1497">
External characters similar to those of
<taxonomicName id="56DD4D0BFFE7FFCBFD8FE889FD4BFD17" baseAuthorityName="Eppelsheim" baseAuthorityYear="1890" box="[569,675,667,689]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="A3A9EA9AFFE7FFCBFD8FE889FD4BFD17" box="[569,675,667,689]" italics="true" pageId="18" pageNumber="1497">M. indicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="91623688FFE7FFCBFF71E8D4FF1BFCB7" blockId="18.[178,1086,705,785]" pageId="18" pageNumber="1497">
: sternite VIII distinctly oblong, posterior excision narrow and rather deep (
<figureCitation id="09E62A0DFFE7FFCBFC54E8D4FBD9FD71" box="[994,1073,705,727]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="18" pageNumber="1497">Fig. 14</figureCitation>
); aedeagus (
<figureCitation id="09E62A0DFFE7FFCBFEABE8CBFE64FD52" box="[285,396,734,756]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="18" pageNumber="1497">Figs 15-16</figureCitation>
) very small,
<quantity id="56259B6DFFE7FFCBFDA5E8CBFD86FD52" box="[531,622,734,756]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="1497" unit="mm" value="0.35">0.35 mm</quantity>
long, strongly asymmetric, and of distinctive shape.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE7FFCBFF04E935FE3BFC0A" pageId="18" pageNumber="1497" type="discussion">
<paragraph id="91623688FFE7FFCBFF04E935FE3BFC0A" blockId="18.[178,1086,800,940]" pageId="18" pageNumber="1497">
Comparative notes: The similar external and male sexual characters suggest that
<taxonomicName id="56DD4D0BFFE7FFCBFEA7E92AFE7AFCF2" authorityName="Cameron" authorityYear="1931" box="[273,402,831,852]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE7FFCBFEA7E92AFE7AFCF2" box="[273,402,831,852]" italics="true" pageId="18" pageNumber="1497">M. nigriceps</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="56DD4D0BFFE7FFCBFDDDE92AFD30FCF2" baseAuthorityName="Eppelsheim" baseAuthorityYear="1890" box="[619,728,830,852]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1497" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="A3A9EA9AFFE7FFCBFDDDE92AFD30FCF2" box="[619,728,830,852]" italics="true" pageId="18" pageNumber="1497">M. indicus</emphasis>
</taxonomicName>
, from which it is distinguished by the darker head, the shape of the male sternite VIII (slightly more oblong), and the slightly different shape of the smaller aedeagus (apex of ventral process shorter, internal structure of different shape).
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE7FFCBFF04E9AEFEE3FC48" pageId="18" pageNumber="1497" type="distribution">
<paragraph id="91623688FFE7FFCBFF04E9AEFEE3FC48" blockId="18.[178,1085,955,1006]" pageId="18" pageNumber="1497">
Distribution: This species is currently known only from northeastern
<collectingCountry id="E9CA7618FFE7FFCBFBBFE9AEFBD5FC77" box="[1033,1085,955,977]" name="India" pageId="18" pageNumber="1497">India</collectingCountry>
(
<figureCitation id="09E62A0DFFE7FFCBFF0FE9CDFF15FC48" box="[185,253,984,1006]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="18" pageNumber="1497">Map 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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