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<mods:title id="A193D9CEEEBDA192EE82F06ABF25DC3A">A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae) from Angola, with the description of three new species</mods:title>
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<mods:affiliation id="8186A4C928128BB3829D6283977AA533">Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, Quarteirão Nimi Ya Lukemi, edifício Q 11, 3 º andar, Luanda, Angola. &amp; adez. bandeira @ gmail. com; https: // orcid. org / 0000 - 0002 - 7496 - 9119</mods:affiliation>
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<emphasis id="A3ADEA885453FFEB9EB0FD5219D1FD20" bold="true" box="[151,420,684,711]" italics="true" pageId="32" pageNumber="333">Lygodactylus nyaneka</emphasis>
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(
<tableCitation id="DC5B03215453FFEB9EB8FD2F1975FD0C" box="[159,256,721,747]" captionStart="TABLE 4" captionStartId="18.[151,239,404,428]" captionText="TABLE 4. Comparison between the different species of Lygodactylus occurring in Angola. Measurements are presented in millimeters (mm). Abbreviations are those described in Materials and Methods section." pageId="32" pageNumber="333">Tables 4</tableCitation>
,
<tableCitation id="DC5B03215453FFEB9F2BFD2F1969FD0C" box="[268,284,721,747]" captionStart="TABLE 6" captionStartId="33.[151,239,332,356]" captionText="TABLE 6. Measurements and meristic data of the type series of Lygodactylus nyaneka sp. nov. Abbreviations are those described in Materials and Methods section." pageId="32" pageNumber="333">6</tableCitation>
;
<figureCitation id="09E22A1F5453FFEB9F0EFD2F19FDFD0C" box="[297,392,721,747]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="19.[151,250,1799,1823]" captionStartId-1="20.[151,250,1754,1778]" captionStartId-2="21.[151,248,1463,1487]" captionStartId-3="22.[151,250,1008,1032]" captionTargetBox-0="[203,1396,960,1759]" captionTargetBox-1="[151,1436,200,1730]" captionTargetBox-2="[161,1426,190,1426]" captionTargetBox-3="[151,1437,181,985]" captionTargetId-0="figure-386@19.[180,1408,915,1775]" captionTargetId-1="figure-22@20.[151,1436,200,1730]" captionTargetId-2="figure-19@21.[151,1436,181,1440]" captionTargetId-3="figure-22@22.[151,1437,181,985]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="FIGURE 1. Maximum likelihood phylogeny of Lygodactylus using ND2. Nodes supported by ML and BI are indicated by black dots. nodes only supported by the ML tree are indicated with half-filled black dots. nodes only supported by BI are indicated by half-filled red dots. and nodes with low support in both the ML and BI trees have red dots. Species of Lygodactylus occurring in Angola have a star next to them in the tree. Species within the Madgascan Lygodactylus clade include L. tolampyae, L. heterurus, L. arnoulti. Species in the Greater Drakensberg group include; L. nigropunctatus, L. graniticolus, L. ocellatus, L. soutpansbergensis. L. incognitus, L. montiscaeruli, L. waterbergensis, and L. methueni. Note that this tree should be used only as a guide to shallow relationships in the genus and to indicate relative divergence." captionText-1="FIGURE 2. Comparison of the number of precloacal pores in Angolan individuals of Lygodactylus angolensis, L. baptistai sp. nov., L. nyaneka sp. nov., L. lawrencei and L. tchokwe sp. nov." captionText-2="FIGURE 3. Comparison of the nostrils position inAngolan individuals of:A. L. angolensis (AMB 9216); B. Lygodactylus baptistai sp. nov. (CAS 263557); C. L. nyaneka sp. nov. (UF 187229); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312b)." captionText-3="FIGURE 4. Comparison of the tail scales arrangement in Angolan individuals of A. Lygodactylus angolensis (MHNC-UP/REP 533); B. L. nyaneka sp. nov. (CAS 263531); C. L. baptistai sp. nov. (CAS 263557); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312d)." figureDoi-0="http://doi.org/10.5281/zenodo.4410729" figureDoi-1="http://doi.org/10.5281/zenodo.4410731" figureDoi-2="http://doi.org/10.5281/zenodo.4410733" figureDoi-3="http://doi.org/10.5281/zenodo.4410735" httpUri-0="https://zenodo.org/record/4410729/files/figure.png" httpUri-1="https://zenodo.org/record/4410731/files/figure.png" httpUri-2="https://zenodo.org/record/4410733/files/figure.png" httpUri-3="https://zenodo.org/record/4410735/files/figure.png" pageId="32" pageNumber="333">Fig. 14</figureCitation>
,
<figureCitation id="09E22A1F5453FFEB9FB3FD2F19AEFD0C" box="[404,475,721,747]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStart-3="FIGURE 17" captionStartId-0="35.[151,250,1008,1032]" captionStartId-1="35.[151,250,1973,1997]" captionStartId-2="36.[151,250,1425,1449]" captionStartId-3="38.[151,250,1943,1967]" captionTargetBox-0="[151,1437,181,985]" captionTargetBox-1="[151,1436,1093,1949]" captionTargetBox-2="[173,1414,191,1395]" captionTargetBox-3="[152,1435,184,1918]" captionTargetId-0="figure-19@35.[151,1437,181,985]" captionTargetId-1="figure-52@35.[151,1436,1093,1949]" captionTargetId-2="figure-22@36.[164,1423,181,1402]" captionTargetId-3="figure-22@38.[152,1435,181,1920]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionTargetPageId-2="36" captionTargetPageId-3="38" captionText-0="FIGURE 14. Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of Lygodactylus nyaneka sp. nov. (CAS 263418)." captionText-1="FIGURE 15. Lygodactylus nyaneka sp. nov. in life (CAS 263531). Montipa [-14.66482º, 13.25763º, 670 m a.s.l.]. Namibe Province. southwestern Angola Photos by Ishan Agarwal." captionText-2="FIGURE 16. Distribution of Lygodactylus nyaneka sp. nov.in the country. Blue circle represents records from Epupa Falls. Namibia." captionText-3="FIGURE 17. Habitats of Lygodactylus nyaneka sp. nov. Top: Fazenda Mucungo. Namibe Province. Below: Bruco Pass. Huila escarpment. Namibe Province. Photos by Luis M. P. Ceríaco." figureDoi-0="http://doi.org/10.5281/zenodo.4410755" figureDoi-1="http://doi.org/10.5281/zenodo.4410757" figureDoi-2="http://doi.org/10.5281/zenodo.4410759" figureDoi-3="http://doi.org/10.5281/zenodo.4410761" httpUri-0="https://zenodo.org/record/4410755/files/figure.png" httpUri-1="https://zenodo.org/record/4410757/files/figure.png" httpUri-2="https://zenodo.org/record/4410759/files/figure.png" httpUri-3="https://zenodo.org/record/4410761/files/figure.png" pageId="32" pageNumber="333">1417</figureCitation>
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<emphasis id="A3ADEA885453FFEB9EB0FCC719F5FCB6" box="[151,384,825,849]" italics="true" pageId="32" pageNumber="333">Hemidactylus capensis</emphasis>
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[sic] (
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<bibRefCitation id="F5484B6B5453FFEB9FE6FCC71A08FCB6" author="Bocage, J. V. B." box="[449,637,825,849]" pageId="32" pageNumber="333" pagination="217 - 228" refId="ref25473" refString="Bocage, J. V. B. (1867) Segunda lista dos reptis das possessoes portuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 217 - 228." type="journal article" year="1867">Bocage 1867: 219</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9CA0FCC71A94FCB6" author="Bocage, J. V. B." box="[647,737,825,849]" pageId="32" pageNumber="333" pagination="66 - 68" refId="ref25515" refString="Bocage, J. V. B. (1870) Description d'un &quot; saurien &quot; nouveau de l'Afrique occidentale. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 3, 66 - 68, pl. 1." type="journal article" year="1870">1870: 68</bibRefCitation>
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<emphasis id="A3ADEA885453FFEB9EB0FCA61909FC97" box="[151,380,856,880]" italics="true" pageId="32" pageNumber="333">Lygodactylus capensis</emphasis>
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(
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<bibRefCitation id="F5484B6B5453FFEB9FADFCA61A49FC97" author="Bocage, J. V. B." box="[394,572,856,880]" pageId="32" pageNumber="333" refId="ref25600" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage 1895: 15</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9D2FFCA61B95FC97" author="Loveridge, A." box="[776,992,856,880]" pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge 1947: 208</bibRefCitation>
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<treatmentCitation id="1078108B5453FFEB9DCBFCA61C23FC97" author="Loveridge, A." box="[1004,1110,856,880]" page="187" pageId="32" pageNumber="333" year="1957">
<bibRefCitation id="F5484B6B5453FFEB9DCBFCA61C23FC97" author="Loveridge, A." box="[1004,1110,856,880]" pageId="32" pageNumber="333" pagination="151 - 362" refId="ref28000" refString="Loveridge, A. (1957) Checklist of the reptiles and amphibians of East Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bulletin of the Museum of Comparative Zoology, 117, 151 - 362 + i-xxxvi." type="journal article" year="1957">1957: 187</bibRefCitation>
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<treatmentCitation id="1078108B5453FFEB9A46FCA61D79FC97" author="Frade, F." box="[1121,1292,856,880]" page="253" pageId="32" pageNumber="333" year="1963">
<bibRefCitation id="F5484B6B5453FFEB9A46FCA61D79FC97" author="Frade, F." box="[1121,1292,856,880]" pageId="32" pageNumber="333" pagination="241 - 257" refId="ref27069" refString="Frade, F. (1963) Linhas gerais da distribuicao dos vertebrados em Angola. Memorias da Junta Investigacao do Ultramar, Segunda Serie, 43, 241 - 257." type="journal article" year="1963">Frade 1963: 253</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9B3FFCA61DE8FC97" author="Pasteur, G." box="[1304,1437,856,880]" pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur 1965</bibRefCitation>
[1964]: 62;
<treatmentCitation id="1078108B5453FFEB9F18FC891982FC68" author="Branch, W. R." box="[319,503,887,911]" page="246" pageId="32" pageNumber="333" year="1998">
<bibRefCitation id="F5484B6B5453FFEB9F18FC891982FC68" author="Branch, W. R." box="[319,503,887,911]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch 1998: 246</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9C25FC891B74FC68" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[514,769,887,911]" pageId="32" pageNumber="333" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885453FFEB9C46FC891AE1FC68" box="[609,660,887,911]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2018: 190
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<paragraph id="9166369A5453FFEB9EB0FC6B1BF4FC4A" blockId="32.[151,1437,825,1033]" box="[151,897,917,941]" pageId="32" pageNumber="333">
<treatmentCitationGroup id="B1C911B45453FFEB9EB0FC6B1BF4FC4A" box="[151,897,917,941]" pageId="32" pageNumber="333">
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<emphasis id="A3ADEA885453FFEB9EB0FC6B19ADFC4A" box="[151,472,917,941]" italics="true" pageId="32" pageNumber="333">Lygodactylus capensis capensis</emphasis>
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(
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<bibRefCitation id="F5484B6B5453FFEB9FC1FC6B1AE4FC4A" author="Schmidt, K. P." box="[486,657,917,941]" pageId="32" pageNumber="333" pagination="1 - 15" refId="ref30093" refString="Schmidt, K. P. (1933) The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum, 22, 1 - 15, pls. 1 - 2." type="journal article" year="1933">Schmidt 1933: 4</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9CBBFC681B0FFC4A" author="Bates, M. F. &amp; Branch, W. R. &amp; Bauer, A. M. &amp; Burger, M. &amp; Marais, J. &amp; Alexander, G. J. &amp; de Villiers, M. S." box="[668,890,917,941]" pageId="32" pageNumber="333" refId="ref25313" refString="Bates, M. F., Branch, W. R., Bauer, A. M., Burger, M., Marais, J., Alexander, G. J. &amp; de Villiers, M. S. (2014) Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, xvii + 485 pp." type="book" year="2014">
Bates
<emphasis id="A3ADEA885453FFEB9CFCFC681B7DFC4A" box="[731,776,917,941]" italics="true" pageId="32" pageNumber="333">et al</emphasis>
. 2014: 118
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<treatmentCitationGroup id="B1C911B45453FFEB9EB0FC4A1ABAFC2B" box="[151,719,948,972]" pageId="32" pageNumber="333">
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<emphasis id="A3ADEA885453FFEB9EB0FC4A1990FC2B" box="[151,485,948,972]" italics="true" pageId="32" pageNumber="333">Lygodactylus capensis bradfieldi</emphasis>
(
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<treatmentCitationGroup id="B1C911B45453FFEB9EB0FC2D1C85FC0C" box="[151,1264,979,1003]" pageId="32" pageNumber="333">
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<emphasis id="A3ADEA885453FFEB9EB0FC2D19FDFC0C" box="[151,392,979,1003]" italics="true" pageId="32" pageNumber="333">Lygodactylus bradfieldi</emphasis>
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[part] (
<treatmentCitation id="1078108B5453FFEB9FF3FC2D1AFBFC0C" author="Branch, W. R." box="[468,654,979,1003]" page="246" pageId="32" pageNumber="333" year="1998">
<bibRefCitation id="F5484B6B5453FFEB9FF3FC2D1AFBFC0C" author="Branch, W. R." box="[468,654,979,1003]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch 1998: 246</bibRefCitation>
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<bibRefCitation id="F5484B6B5453FFEB9CBFFC2D1BF8FC0C" author="Conradie, W. R. &amp; Branch, W. R." box="[664,909,979,1003]" pageId="32" pageNumber="333" pagination="6 - 36" refId="ref26770" refString="Conradie, W. R. &amp; Branch, W. R. (2016) The herpetofauna of the Cubango, Cuito and lower Cuando river catchments of southeastern Angola. Amphibian &amp; Reptile Conservation, 10 (2), 6 - 36." type="journal article" year="2016">
Conradie
<emphasis id="A3ADEA885453FFEB9CDCFC2D1B5DFC0C" box="[763,808,979,1003]" italics="true" pageId="32" pageNumber="333">et al</emphasis>
. 2016: 24
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[?];
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Marques
<emphasis id="A3ADEA885453FFEB9A3BFC2D1C25FC0C" box="[1052,1104,979,1003]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2018
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: 189,190)
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<treatmentCitationGroup id="B1C911B45453FFEB9EB0FC0F1AEBFBEE" box="[151,670,1009,1033]" pageId="32" pageNumber="333">
<treatmentCitation id="1078108B5453FFEB9EB0FC0F1AEBFBEE" author="Baptista, N. L. &amp; Antonio, T. &amp; Branch, W. R." box="[151,670,1009,1033]" httpUri="http://treatment.plazi.org/id/7E2D87A6FF9DFFCED610FB254943FB49" page="113" pageId="32" pageNumber="333" year="2019">
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<emphasis id="A3ADEA885453FFEB9EB0FC0F19E5FBEE" box="[151,400,1009,1033]" italics="true" pageId="32" pageNumber="333">Lygodactylus angolensis</emphasis>
(
<bibRefCitation id="F5484B6B5453FFEB9FBAFC0F1AE2FBEE" author="Baptista, N. L. &amp; Antonio, T. &amp; Branch, W. R." box="[413,663,1009,1033]" pageId="32" pageNumber="333" pagination="203" refId="ref25219" refString="Baptista, N. L., Antonio, T. &amp; Branch, W. R. (2019) The herpetofauna of Bicuar National Park and surroundings southwestern Angola: a preliminary checklist. Amphibian &amp; Reptile Conservation, 13 (2), e 203, 96 - 130. [Special Section]" type="journal article" year="2019">
Baptista
<emphasis id="A3ADEA885453FFEB9FDFFC0C1A59FBEE" box="[504,556,1009,1033]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2019: 113
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The majority of the authors who historically dealt with Angolan
<taxonomicName id="56D94D195453FFEB9D57FBC91C71FBB6" box="[880,1028,1079,1105]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis id="A3ADEA885453FFEB9D57FBC91C71FBB6" box="[880,1028,1079,1105]" italics="true" pageId="32" pageNumber="333">Lygodactylus</emphasis>
</taxonomicName>
other than
<taxonomicName id="56D94D195453FFEB9AA2FBC61D6EFBB6" box="[1157,1307,1079,1105]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="A3ADEA885453FFEB9AA2FBC61D6EFBB6" box="[1157,1307,1079,1105]" italics="true" pageId="32" pageNumber="333">L. angolensis</emphasis>
</taxonomicName>
considered them to represent
<taxonomicName id="56D94D195453FFEB9F7AFBA219ACFB92" box="[349,473,1116,1141]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9F7AFBA219ACFB92" box="[349,473,1116,1141]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="F5484B6B5453FFEB9FC0FBA21A08FB92" author="Bocage, J. V. B." box="[487,637,1115,1142]" pageId="32" pageNumber="333" pagination="217 - 228" refId="ref25473" refString="Bocage, J. V. B. (1867) Segunda lista dos reptis das possessoes portuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 217 - 228." type="journal article" year="1867">Bocage 1867</bibRefCitation>
,
<bibRefCitation id="F5484B6B5453FFEB9CA0FBA21AB7FB91" author="Bocage, J. V. B." box="[647,706,1116,1142]" pageId="32" pageNumber="333" pagination="66 - 68" refId="ref25515" refString="Bocage, J. V. B. (1870) Description d'un &quot; saurien &quot; nouveau de l'Afrique occidentale. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 3, 66 - 68, pl. 1." type="journal article" year="1870">1870</bibRefCitation>
,
<bibRefCitation id="F5484B6B5453FFEB9CEBFBA51B72FB91" author="Bocage, J. V. B." box="[716,775,1115,1142]" pageId="32" pageNumber="333" refId="ref25600" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">1895</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9D35FBA51BC5FB91" author="Schmidt, K. P." box="[786,944,1115,1142]" pageId="32" pageNumber="333" pagination="1 - 15" refId="ref30093" refString="Schmidt, K. P. (1933) The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum, 22, 1 - 15, pls. 1 - 2." type="journal article" year="1933">Schmidt 1933</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9D9DFBA51C21FB91" author="Monard, A." box="[954,1108,1115,1142]" pageId="32" pageNumber="333" pagination="19 - 154" refId="ref28807" refString="Monard, A. (1937) Contribution a l'herpetologie d'Angola. Arquivos do Museu Bocage, 8, 19 - 154." type="journal article" year="1937">Monard 1937</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9A79FBA51D6AFB91" author="FitzSimons, V. F." box="[1118,1311,1115,1142]" pageId="32" pageNumber="333" refId="ref26986" refString="FitzSimons, V. F. (1943) The lizards of South Africa. Transvaal Museum Memoir No. 1. Transvaal Museum, Pretoria, xv + 528 pp., 24 pls., 1 folding map." type="book" year="1943">FitzSimons 1943</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9B0EFBA518A7FB7D" author="Loveridge, A." pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge 1947</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9EF8FB811917FB7E" author="Frade, F." box="[223,354,1151,1177]" pageId="32" pageNumber="333" pagination="241 - 257" refId="ref27069" refString="Frade, F. (1963) Linhas gerais da distribuicao dos vertebrados em Angola. Memorias da Junta Investigacao do Ultramar, Segunda Serie, 43, 241 - 257." type="journal article" year="1963">Frade 1963</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9F49FB7E1A74FB7D" author="Pasteur, G." box="[366,513,1151,1178]" pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur 1965</bibRefCitation>
[1964];
<bibRefCitation id="F5484B6B5453FFEB9C44FB811A82FB7D" author="Branch, W. R." box="[611,759,1151,1178]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch 1998</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9D24FB7E1B9EFB7D" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[771,1003,1151,1178]" pageId="32" pageNumber="333" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885453FFEB9D4AFB7E1BD3FB7E" box="[877,934,1151,1177]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2018
</bibRefCitation>
).
<bibRefCitation id="F5484B6B5453FFEB9DD9FB7E1CEBFB7E" author="Bocage, J. V. B." box="[1022,1182,1151,1178]" pageId="32" pageNumber="333" pagination="217 - 228" refId="ref25473" refString="Bocage, J. V. B. (1867) Segunda lista dos reptis das possessoes portuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 217 - 228." type="journal article" year="1867">Bocage (1867</bibRefCitation>
,
<bibRefCitation id="F5484B6B5453FFEB9A8DFB811C90FB7D" author="Bocage, J. V. B." box="[1194,1253,1151,1178]" pageId="32" pageNumber="333" refId="ref25600" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">1895</bibRefCitation>
,
<bibRefCitation id="F5484B6B5453FFEB9AD6FB811D5FFB7D" author="Bocage, J. V. B." box="[1265,1322,1151,1178]" pageId="32" pageNumber="333" pagination="105 - 114" refId="ref25633" refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114." type="journal article" year="1896">1896</bibRefCitation>
) recorded them from
<collectingRegion id="531DF8785453FFEB9F31FB5D191AFB5A" box="[278,367,1187,1213]" country="Angola" name="Namibe" pageId="32" pageNumber="333">Namibe</collectingRegion>
and
<collectingRegion id="531DF8785453FFEB9F80FB5D1990FB5A" box="[423,485,1187,1213]" country="Angola" name="Huila" pageId="32" pageNumber="333">Huíla</collectingRegion>
provinces;
<bibRefCitation id="F5484B6B5453FFEB9C4DFB5D1B6FFB59" author="Schmidt, K. P." box="[618,794,1187,1214]" pageId="32" pageNumber="333" pagination="1 - 15" refId="ref30093" refString="Schmidt, K. P. (1933) The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum, 22, 1 - 15, pls. 1 - 2." type="journal article" year="1933">Schmidt (1933)</bibRefCitation>
recorded them from
<collectingRegion id="531DF8785453FFEB9A2CFB5D1C91FB5A" box="[1035,1252,1187,1213]" country="Angola" name="Benguela" pageId="32" pageNumber="333">Benguela Province</collectingRegion>
;
<bibRefCitation id="F5484B6B5453FFEB9AD6FB5D1DE9FB59" author="Monard, A." box="[1265,1436,1187,1214]" pageId="32" pageNumber="333" pagination="19 - 154" refId="ref28807" refString="Monard, A. (1937) Contribution a l'herpetologie d'Angola. Arquivos do Museu Bocage, 8, 19 - 154." type="journal article" year="1937">Monard (1937)</bibRefCitation>
recorded the species from
<collectingRegion id="531DF8785453FFEB9FE2FB391A76FB06" box="[453,515,1223,1249]" country="Angola" name="Huila" pageId="32" pageNumber="333">Huila</collectingRegion>
and
<collectingRegion id="531DF8785453FFEB9C1BFB361AE7FB05" box="[572,658,1224,1250]" country="Angola" name="Cunene" pageId="32" pageNumber="333">Cunene</collectingRegion>
provinces;
<bibRefCitation id="F5484B6B5453FFEB9D30FB391BA8FB05" author="Loveridge, A." box="[791,989,1223,1250]" pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge (1947)</bibRefCitation>
recorded them from
<collectingRegion id="531DF8785453FFEB9AF7FB391DEDFB06" box="[1232,1432,1223,1249]" country="Angola" name="Namibe" pageId="32" pageNumber="333">Namibe Province</collectingRegion>
; and
<bibRefCitation id="F5484B6B5453FFEB9EE1FB12191BFAE1" author="Laurent, R. F." box="[198,366,1259,1286]" pageId="32" pageNumber="333" pagination="1 - 165" refId="ref27865" refString="Laurent, R. F. (1964) Reptiles et amphibiens de l'Angola (Troiseme contribution). Publicacoes Culturais. Companhia de Diamantes de Angola, 67, 1 - 165." type="journal article" year="1964">Laurent (1964)</bibRefCitation>
recorded the species from
<collectingRegion id="531DF8785453FFEB9CBFFB151B7EFAE2" box="[664,779,1259,1285]" country="Angola" name="Lunda Sul" pageId="32" pageNumber="333">Lunda Sul</collectingRegion>
and
<collectingRegion id="531DF8785453FFEB9D67FB151BE2FAE2" box="[832,919,1259,1285]" country="Angola" name="Moxico" pageId="32" pageNumber="333">Moxico</collectingRegion>
provinces. Based on these records, the species was considered to occur in
<collectingCountry id="E9CE760A5453FFEB9FE3FAF11A63FACE" box="[452,534,1295,1321]" name="Angola" pageId="32" pageNumber="333">Angola</collectingCountry>
by all authors who reviewed the genus in
<collectingCountry id="E9CE760A5453FFEB9DCEFAF11C4EFACE" box="[1001,1083,1295,1321]" name="Angola" pageId="32" pageNumber="333">Angola</collectingCountry>
(
<bibRefCitation id="F5484B6B5453FFEB9A6DFAF11D75FACD" author="Loveridge, A." box="[1098,1280,1295,1322]" pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge 1947</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9B2CFAEE1DE9FACD" author="Pasteur, G." box="[1291,1436,1295,1322]" pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur 1965</bibRefCitation>
[1964];
<bibRefCitation id="F5484B6B5453FFEB9ED6FACD19F1FAA9" author="Branch, W. R." box="[241,388,1331,1358]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch 1998</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9FB7FACA1A01FAA9" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[400,628,1331,1358]" pageId="32" pageNumber="333" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885453FFEB9FDEFACA1A47FAAA" box="[505,562,1331,1357]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2018
</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9CA7FACD1B21FAA9" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[640,852,1331,1358]" pageId="32" pageNumber="333" pagination="283 - 334" refId="ref25903" refString="Branch, W. R., Pinto. P. V., Baptista, N. &amp; Conradie, W. (2019) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B., Russo, V., Lages &amp; Ferrand, N. (Eds.), Biodiversity of Angola, Science &amp; Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13" type="book chapter" year="2019">
Branch
<emphasis id="A3ADEA885453FFEB9CFFFACA1B65FAAA" box="[728,784,1331,1357]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2019
</bibRefCitation>
). However,
<bibRefCitation id="F5484B6B5453FFEB9DFDFACD1C08FAA9" author="Branch, W. R." box="[986,1149,1331,1358]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch (1998)</bibRefCitation>
stated the possibility that besides
<taxonomicName id="56D94D195453FFEB9ED5FAA61907FA96" box="[242,370,1368,1393]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9ED5FAA61907FA96" box="[242,370,1368,1393]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
, the morphologically similar
<taxonomicName id="56D94D195453FFEB9CE1FAA61B26FA96" box="[710,851,1367,1393]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9CE1FAA61B26FA96" box="[710,851,1367,1393]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
could also occur in
<collectingCountry id="E9CE760A5453FFEB9A1AFAA91CE6FA96" box="[1085,1171,1367,1393]" name="Angola" pageId="32" pageNumber="333">Angola</collectingCountry>
.
<bibRefCitation id="F5484B6B5453FFEB9A87FAA91DE9FA95" author="Conradie, W. R. &amp; Branch, W. R." box="[1184,1436,1367,1394]" pageId="32" pageNumber="333" pagination="6 - 36" refId="ref26770" refString="Conradie, W. R. &amp; Branch, W. R. (2016) The herpetofauna of the Cubango, Cuito and lower Cuando river catchments of southeastern Angola. Amphibian &amp; Reptile Conservation, 10 (2), 6 - 36." type="journal article" year="2016">
Conradie
<emphasis id="A3ADEA885453FFEB9B28FAA61D3CFA96" box="[1295,1353,1367,1393]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
(2016)
</bibRefCitation>
provided the first records of
<taxonomicName id="56D94D195453FFEB9FF8FA821A19FA72" box="[479,620,1403,1429]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9FF8FA821A19FA72" box="[479,620,1403,1429]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
in the country based on specimens collected in Cuando Cubango province, and
<bibRefCitation id="F5484B6B5453FFEB9F22FA5E198FFA5D" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[261,506,1439,1466]" pageId="32" pageNumber="333" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885453FFEB9F48FA5E19DDFA5E" box="[367,424,1439,1465]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
(2018)
</bibRefCitation>
interpreted many of the historical records assigned to
<taxonomicName id="56D94D195453FFEB9A45FA5E1C94FA5E" box="[1122,1249,1440,1465]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9A45FA5E1C94FA5E" box="[1122,1249,1440,1465]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
as
<taxonomicName id="56D94D195453FFEB9B2FFA5E1DE0FA5E" box="[1288,1429,1439,1465]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9B2FFA5E1DE0FA5E" box="[1288,1429,1439,1465]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
. Originally described by
<bibRefCitation id="F5484B6B5453FFEB9F80FA3D1A49FA39" author="Hewitt, J." box="[423,572,1475,1502]" pageId="32" pageNumber="333" pagination="105 - 128" refId="ref27454" refString="Hewitt, J. (1932) Some new species and subspecies of South African batrachians and lizards. Annals of the Natal Museum, 7 (1), 105 - 128." type="journal article" year="1932">Hewitt (1932</bibRefCitation>
a) based on specimens from near Okahandja in central
<collectingCountry id="E9CE760A5453FFEB9A85FA3D1D70FA3A" box="[1186,1285,1475,1501]" name="Namibia" pageId="32" pageNumber="333">Namibia</collectingCountry>
,
<taxonomicName id="56D94D195453FFEB9B37FA3A1DE9FA3A" box="[1296,1436,1475,1501]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9B37FA3A1DE9FA3A" box="[1296,1436,1475,1501]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
has been recorded from numerous places in
<collectingCountry id="E9CE760A5453FFEB9CB8FA191B77F9E6" box="[671,770,1511,1537]" name="Namibia" pageId="32" pageNumber="333">Namibia</collectingCountry>
,
<collectingCountry id="E9CE760A5453FFEB9D37FA191B10F9E6" box="[784,869,1511,1537]" name="Angola" pageId="32" pageNumber="333">Angola</collectingCountry>
,
<collectingCountry id="E9CE760A5453FFEB9D53FA161B90F9E5" box="[884,997,1512,1538]" name="Botswana" pageId="32" pageNumber="333">Botswana</collectingCountry>
,
<collectingCountry id="E9CE760A5453FFEB9DD3FA191CB9F9E6" box="[1012,1228,1511,1537]" name="Zambia Zimbabwe" pageId="32" pageNumber="333">Zambia Zimbabwe</collectingCountry>
and
<collectingCountry id="E9CE760A5453FFEB9B2DFA191DE8F9E6" box="[1290,1437,1511,1537]" name="South Africa" pageId="32" pageNumber="333">South Africa</collectingCountry>
(Mertens 1971;
<bibRefCitation id="F5484B6B5453FFEB9F77F9F51A61F9C1" author="FitzSimons, V. F." box="[336,532,1547,1574]" pageId="32" pageNumber="333" refId="ref26986" refString="FitzSimons, V. F. (1943) The lizards of South Africa. Transvaal Museum Memoir No. 1. Transvaal Museum, Pretoria, xv + 528 pp., 24 pls., 1 folding map." type="book" year="1943">FitzSimons 1943</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9C04F9F51AAEF9C1" author="Loveridge, A." box="[547,731,1547,1574]" pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge 1947</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9CCFF9F51BECF9C1" author="Auerbach, R. D." box="[744,921,1547,1574]" pageId="32" pageNumber="333" refId="ref25142" refString="Auerbach, R. D. (1987) The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Gaberone, 295 pp." type="book" year="1987">Auerbach 1987</bibRefCitation>
; Broadley 1991;
<bibRefCitation id="F5484B6B5453FFEB9A45F9F51D7EF9C1" author="Jacobsen, N. H. G." box="[1122,1291,1547,1574]" pageId="32" pageNumber="333" pagination="21 - 27" refId="ref27656" refString="Jacobsen, N. H. G. (2011) The distribution of Lygodactylus bradfieldi Hewitt, 1932 in Limpopo Province, South Africa. African Herp News, 53, 21 - 27." type="journal article" year="2011">Jacobsen 2011</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9B3EF9F218A6F9AD" author="Bates, M. F. &amp; Branch, W. R. &amp; Bauer, A. M. &amp; Burger, M. &amp; Marais, J. &amp; Alexander, G. J. &amp; de Villiers, M. S." pageId="32" pageNumber="333" refId="ref25313" refString="Bates, M. F., Branch, W. R., Bauer, A. M., Burger, M., Marais, J., Alexander, G. J. &amp; de Villiers, M. S. (2014) Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, xvii + 485 pp." type="book" year="2014">
Bates
<emphasis id="A3ADEA885453FFEB9B46F9F21DE8F9C2" box="[1377,1437,1547,1573]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9EF8F9CE19B3F9AD" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[223,454,1583,1610]" pageId="32" pageNumber="333" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885453FFEB9F6FF9CE19F4F9AE" box="[328,385,1583,1609]" italics="true" pageId="32" pageNumber="333">et al.</emphasis>
2018
</bibRefCitation>
). Morphologically the species is very similar to
<taxonomicName id="56D94D195453FFEB9DC1F9CE1C11F9AE" box="[998,1124,1584,1609]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9DC1F9CE1C11F9AE" box="[998,1124,1584,1609]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
, and most of the characters that are used to distinguish the species overlap considerably, making it difficult to easily diagnose the species from one another. This morphological similarity led some authors to consider the species as western race or a synonym of
<taxonomicName id="56D94D195453FFEB9EB0F9621961F952" box="[151,276,1692,1717]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9EB0F9621961F952" box="[151,276,1692,1717]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="F5484B6B5453FFEB9F04F9651993F951" author="FitzSimons, V. F." box="[291,486,1691,1718]" pageId="32" pageNumber="333" refId="ref26986" refString="FitzSimons, V. F. (1943) The lizards of South Africa. Transvaal Museum Memoir No. 1. Transvaal Museum, Pretoria, xv + 528 pp., 24 pls., 1 folding map." type="book" year="1943">FitzSimons 1943</bibRefCitation>
;
<bibRefCitation id="F5484B6B5453FFEB9FD6F9651AD3F951" author="Loveridge, A." box="[497,678,1691,1718]" pageId="32" pageNumber="333" pagination="1 - 469" refId="ref27936" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge 1947</bibRefCitation>
,
<bibRefCitation id="F5484B6B5453FFEB9C97F9651A99F951" author="Loveridge, A." box="[688,748,1691,1718]" pageId="32" pageNumber="333" pagination="151 - 362" refId="ref28000" refString="Loveridge, A. (1957) Checklist of the reptiles and amphibians of East Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bulletin of the Museum of Comparative Zoology, 117, 151 - 362 + i-xxxvi." type="journal article" year="1957">1957</bibRefCitation>
), until
<bibRefCitation id="F5484B6B5453FFEB9D1EF9621BA6F951" author="Pasteur, G." box="[825,979,1691,1718]" pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur (1965</bibRefCitation>
[1964]) resurrected it. According to
<bibRefCitation id="F5484B6B5453FFEB9B4BF962197EF93D" author="Pasteur, G." pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur (1965</bibRefCitation>
[1964]), the most useful characters to differentiate
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<emphasis id="A3ADEA885453FFEB9D6BF93E1BADF93E" box="[844,984,1727,1753]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
from
<taxonomicName id="56D94D195453FFEB9A3AF93E1CEEF93E" box="[1053,1179,1728,1753]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9A3AF93E1CEEF93E" box="[1053,1179,1728,1753]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
are the
<typeStatus id="4E6288385453FFEB9AD2F93E1D50F93D" box="[1269,1317,1728,1754]" pageId="32" pageNumber="333">type</typeStatus>
of contact between the rostral and the naso-rostrals (rostral dividing the naso-rostrals in
<taxonomicName id="56D94D195453FFEB9DCCF91A1CB6F91A" box="[1003,1219,1763,1789]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="subSpecies" species="bradfieldi" subSpecies="versus">
<emphasis id="A3ADEA885453FFEB9DCCF91A1CB6F91A" box="[1003,1219,1763,1789]" italics="true" pageId="32" pageNumber="333">L. bradfieldi versus</emphasis>
</taxonomicName>
rostral not dividing the naso-rostrals in
<taxonomicName id="56D94D195453FFEB9F53F8F61987F8C6" box="[372,498,1800,1825]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9F53F8F61987F8C6" box="[372,498,1800,1825]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
); a lower number of mid-dorsal scales (
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<taxonomicName id="56D94D195453FFEB9A1CF8F61D63F8C6" box="[1083,1302,1799,1825]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="subSpecies" species="bradfieldi" subSpecies="versus">
<emphasis id="A3ADEA885453FFEB9A1CF8F61D63F8C6" box="[1083,1302,1799,1825]" italics="true" pageId="32" pageNumber="333">L. bradfieldi versus</emphasis>
</taxonomicName>
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<taxonomicName id="56D94D195453FFEB9EB0F8D21960F8A2" box="[151,277,1836,1861]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9EB0F8D21960F8A2" box="[151,277,1836,1861]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
); and staggered rows of subcaudal plates in original tails of
<taxonomicName id="56D94D195453FFEB9D9DF8D21C30F8A2" box="[954,1093,1835,1861]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9D9DF8D21C30F8A2" box="[954,1093,1835,1861]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
,
<emphasis id="A3ADEA885453FFEB9A73F8D21CE9F8A2" box="[1108,1180,1836,1861]" italics="true" pageId="32" pageNumber="333">versus</emphasis>
a more linear distribution of the rows in
<taxonomicName id="56D94D195453FFEB9F54F8AE1986F88E" box="[371,499,1872,1897]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9F54F8AE1986F88E" box="[371,499,1872,1897]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="F5484B6B5453FFEB9C24F8B11AA0F88D" author="FitzSimons, V. F." box="[515,725,1871,1898]" pageId="32" pageNumber="333" refId="ref26986" refString="FitzSimons, V. F. (1943) The lizards of South Africa. Transvaal Museum Memoir No. 1. Transvaal Museum, Pretoria, xv + 528 pp., 24 pls., 1 folding map." type="book" year="1943">FitzSimons (1943)</bibRefCitation>
considered that the main distinguishing character between the taxa was the number of scales entering the nostril, which in
<taxonomicName id="56D94D195453FFEB9D0DF88A1BC1F86A" box="[810,948,1907,1933]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885453FFEB9D0DF88A1BC1F86A" box="[810,948,1907,1933]" italics="true" pageId="32" pageNumber="333">L. bradfieldi</emphasis>
</taxonomicName>
is four (rostral, 1
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supralabial and two nasal scales) and in
<taxonomicName id="56D94D195453FFEB9F1DF86619CCF856" box="[314,441,1944,1969]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="333" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885453FFEB9F1DF86619CCF856" box="[314,441,1944,1969]" italics="true" pageId="32" pageNumber="333">L. capensis</emphasis>
</taxonomicName>
is five (rostral, 1
<superScript id="66AC9BD25453FFEB9C59F8661AFCF841" attach="left" box="[638,649,1944,1958]" fontSize="6" pageId="32" pageNumber="333">st</superScript>
supralabial and three nasal scales). Following
<bibRefCitation id="F5484B6B5453FFEB9ABAF8661D4CF855" author="Pasteur, G." box="[1181,1337,1943,1970]" pageId="32" pageNumber="333" pagination="1 - 132" refId="ref29152" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12." type="journal article" year="1965">Pasteur (1965</bibRefCitation>
[1964]), Broadley (1991),
<bibRefCitation id="F5484B6B5453FFEB9F7CF8451989F831" author="Branch, W. R." box="[347,508,1979,2006]" pageId="32" pageNumber="333" refId="ref25855" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062" type="book" year="1998">Branch (1998)</bibRefCitation>
and
<bibRefCitation id="F5484B6B5453FFEB9C16F8451A90F831" author="Jacobsen, N. H. G." box="[561,741,1979,2006]" pageId="32" pageNumber="333" pagination="21 - 27" refId="ref27656" refString="Jacobsen, N. H. G. (2011) The distribution of Lygodactylus bradfieldi Hewitt, 1932 in Limpopo Province, South Africa. African Herp News, 53, 21 - 27." type="journal article" year="2011">Jacobsen (2011)</bibRefCitation>
considered the arrangement of the subcaudal scales of original tails as a good character to differentiate between the species. Our results show that, contrary to earlier interpretations, the central and southwestern Angolan populations of
<taxonomicName id="56D94D195452FFEA9D1FFF691BB9FF56" box="[824,972,151,177]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="genus">
<emphasis id="A3ADEA885452FFEA9D1FFF691BB9FF56" box="[824,972,151,177]" italics="true" pageId="33" pageNumber="334">Lygodactylus</emphasis>
</taxonomicName>
belong neither to typical
<taxonomicName id="56D94D195452FFEA9AD7FF691D1BFF57" box="[1264,1390,151,176]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9AD7FF691D1BFF57" box="[1264,1390,151,176]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
nor to true
<taxonomicName id="56D94D195452FFEA9EC0FF451907FF32" box="[231,370,187,213]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885452FFEA9EC0FF451907FF32" box="[231,370,187,213]" italics="true" pageId="33" pageNumber="334">L. bradfieldi</emphasis>
</taxonomicName>
but to another of several clades within the
<taxonomicName id="56D94D195452FFEA9D70FF451BA0FF33" box="[855,981,187,212]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9D70FF451BA0FF33" box="[855,981,187,212]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
species complex. The
<typeStatus id="4E6288385452FFEA9AF2FF451D70FF32" box="[1237,1285,187,213]" pageId="33" pageNumber="334">type</typeStatus>
locality of
<taxonomicName id="56D94D195452FFEA9BA1FF451882FF1F" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9BA1FF451882FF1F" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
is not very precise (“Kaffirland, and the districts to the north of
<collectingRegion id="531DF8785452FFEA9DF7FF211C7FFF1E" box="[976,1034,223,249]" country="South Africa" name="Western Cape" pageId="33" pageNumber="334">Cape</collectingRegion>
Colony”
<bibRefCitation id="F5484B6B5452FFEA9A5FFF211C8BFF1E" author="Smith, A." box="[1144,1278,223,249]" pageId="33" pageNumber="334" pagination="1836" refId="ref30127" refString="Smith, A. (1849) s. n. In: Illustrations of the Zoology of South Africa: Consisting chiefly of Figures and Descriptions of the Objects of Natural History collected during an Expedition into the Interior of South Africa in the years 1834, 1835 and 1836. Appendix. Smith, Elder &amp; Co, London, pl 75, accompanying unnumbered text pages" type="book chapter" year="1849">Smith 1849</bibRefCitation>
), and the
<typeStatus id="4E6288385452FFEA9B4BFF211DE9FF1E" box="[1388,1436,223,249]" pageId="33" pageNumber="334">type</typeStatus>
material is lost
<emphasis id="A3ADEA885452FFEA9F6EFEFD1901FEFA" box="[329,372,259,285]" italics="true" pageId="33" pageNumber="334">fide</emphasis>
<bibRefCitation id="F5484B6B5452FFEA9F5BFEFD1A22FEFA" author="FitzSimons, V. F." box="[380,599,259,285]" pageId="33" pageNumber="334" pagination="275 - 279" refId="ref26955" refString="FitzSimons, V. F. (1937) Three new lizards from South Africa. Annals of the Transvaal Museum, 17 (4), 275 - 279." type="journal article" year="1937">FitzSimons (1937)</bibRefCitation>
. Kaffirland, or Kaffraria, referred to southeastern regions of the
<collectingRegion id="531DF8785452FFEA9B18FEFD18A6F97E" country="South Africa" name="Eastern Cape" pageId="33" pageNumber="334">Eastern Cape</collectingRegion>
, where the species is presumably not native, but has been introduced (
<bibRefCitation id="F5484B6B5452FFEA9DC4F9811CC3F97E" author="Rebelo, A. D. &amp; Bates, M. F. &amp; Burger, M. &amp; Branch, W. R. &amp; Conradie, W." box="[995,1206,1663,1689]" pageId="33" pageNumber="334" pagination="643 - 650" refId="ref29808" refString="Rebelo, A. D., Bates, M. F., Burger, M., Branch, W. R. &amp; Conradie, W. (2019) Range expansion of the common dwarf gecko, Lygodactylus capensis: South Africa's most sucessfil reptile invader. Herpetology Notes, 12, 643 - 650." type="journal article" year="2019">
Rebelo
<emphasis id="A3ADEA885452FFEA9A1CF9811C06F97E" box="[1083,1139,1663,1689]" italics="true" pageId="33" pageNumber="334">et al.</emphasis>
2019
</bibRefCitation>
), while the “distrcits to the north of
<collectingRegion id="531DF8785452FFEA9F1CF95D1900F95A" box="[315,373,1699,1725]" country="South Africa" name="Western Cape" pageId="33" pageNumber="334">Cape</collectingRegion>
Colony” was used by Smith for areas currently part of the
<collectingRegion id="531DF8785452FFEA9A27F95D1D7AF95A" box="[1024,1295,1699,1725]" country="South Africa" name="Northern Cape" pageId="33" pageNumber="334">Northern Cape Province</collectingRegion>
and/or areas eastward to modern
<collectingRegion id="531DF8785452FFEA9F5AF9391995F906" box="[381,480,1735,1761]" country="South Africa" name="Gauteng" pageId="33" pageNumber="334">Gauteng</collectingRegion>
. It is, therefore, safe to assume that the nominotypical form belongs to the South African clade (
<taxonomicName id="56D94D195452FFEA9F05F9151A6BF8E3" box="[290,542,1771,1797]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9F05F9151A6BF8E3" box="[290,542,1771,1797]" italics="true" pageId="33" pageNumber="334">Lygodactylus capensis</emphasis>
</taxonomicName>
<quantity id="56219B7F5452FFEA9C01F9151A24F8E2" box="[550,593,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="33" pageNumber="334" unit="in" value="3.0">3 in</quantity>
<figureCitation id="09E22A1F5452FFEA9C7FF9151AC2F8E2" box="[600,695,1771,1797]" captionStart="FIGURE 1" captionStartId="19.[151,250,1799,1823]" captionTargetBox="[203,1396,960,1759]" captionTargetId="figure-386@19.[180,1408,915,1775]" captionTargetPageId="19" captionText="FIGURE 1. Maximum likelihood phylogeny of Lygodactylus using ND2. Nodes supported by ML and BI are indicated by black dots. nodes only supported by the ML tree are indicated with half-filled black dots. nodes only supported by BI are indicated by half-filled red dots. and nodes with low support in both the ML and BI trees have red dots. Species of Lygodactylus occurring in Angola have a star next to them in the tree. Species within the Madgascan Lygodactylus clade include L. tolampyae, L. heterurus, L. arnoulti. Species in the Greater Drakensberg group include; L. nigropunctatus, L. graniticolus, L. ocellatus, L. soutpansbergensis. L. incognitus, L. montiscaeruli, L. waterbergensis, and L. methueni. Note that this tree should be used only as a guide to shallow relationships in the genus and to indicate relative divergence." figureDoi="http://doi.org/10.5281/zenodo.4410729" httpUri="https://zenodo.org/record/4410729/files/figure.png" pageId="33" pageNumber="334">Figure 1</figureCitation>
;
<tableCitation id="DC5B03215452FFEA9CE2F9151B6CF8E2" box="[709,793,1771,1797]" captionStart="TABLE 2" captionStartId="4.[151,231,151,172]" captionText="TABLE 2. Specimens used for genetic analysis and corresponding GenBank accession numbers for ND2 and 16S gene used in the study. Locality data are reported in the form of decimal degrees and use the WGS 84 map datum. See Materials and Methods section for collection abbreviations. Field number acronyms and Institution number not cited in the Material and Methods section follow as: MCZ: Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA; MVZ: Museum of Vertebrate Zoology, Berkeley, California, USA; NMB: National Museum Bloemfontein, Bloemfontein, South Africa; NWZB: Natural History Museum, Bulawayo, Zimbabwe; PEM: Bayworld, Port Elizabeth Museum, Port Elizabeth, South Africa; SMWN: National Museum of Namibia, Windhoek, Namibia; YPM: Peabody Museum of Natural History, Yale University, Connecticut, USA; ZSM: Zoologische Staatssammlung München, Munich, Germany / AMB: Aaron M. Bauer field numbers; DGB: Donald G. Broadley field numbers; FGMV: Frank Glaw/Miguel Vences field numbers; FGZC: Frank Glaw Zoological Collection field numbers; JB: Jon Boone (captive animal); JM: Johan Marais field numbers; JS: Jay Sommers (captive); JVV: Jens V. Vindum field numbers; LMH: Vincent Egan tissue numbers; MBUR / MB: Marius Burger field numbers; PW: Philipp Wagner field numbers; SKI: Sebastian Kirchhof field numbers; SLT: Scott L. Travers field numbers; SVN: Stuart V. Nielsen field numbers; TG: Tony Gamble (captive animal); William R. Branch field numbers." pageId="33" pageNumber="334">Table 2</tableCitation>
) and cannot be applied to the Angolan clade. Besides the original
<emphasis id="A3ADEA885452FFEA9ED1F8F11934F8CF" box="[246,321,1807,1832]" italics="true" pageId="33" pageNumber="334">nomen</emphasis>
, there are a few names in the synonymy of
<taxonomicName id="56D94D195452FFEA9D17F8F11BDBF8CF" box="[816,942,1807,1832]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9D17F8F11BDBF8CF" box="[816,942,1807,1832]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
but none apply to the Angolan populations, and therefore, the Angolan clade of
<taxonomicName id="56D94D195452FFEA9C10F8CD1AC2F8AB" box="[567,695,1843,1868]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9C10F8CD1AC2F8AB" box="[567,695,1843,1868]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
species complex remains an unnamed and undescribed taxon. A further detailed taxonomic revision of the
<taxonomicName id="56D94D195452FFEA9C4AF8A91A9FF897" box="[621,746,1879,1904]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9C4AF8A91A9FF897" box="[621,746,1879,1904]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
species complex is underway (S. Travers
<emphasis id="A3ADEA885452FFEA9A99F8A91C83F896" box="[1214,1270,1879,1905]" italics="true" pageId="33" pageNumber="334">et al.</emphasis>
, unpublished). The preliminary identification of
<taxonomicName id="56D94D195452FFEA9C28F8851BF6F872" authority="in Cuando Cubango" authorityName="in Cuando Cubango" box="[527,899,1915,1941]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885452FFEA9C28F8851AEEF872" box="[527,667,1915,1941]" italics="true" pageId="33" pageNumber="334">L. bradfieldi</emphasis>
in Cuando Cubango
</taxonomicName>
by
<bibRefCitation id="F5484B6B5452FFEA9D8AF8851CD0F871" author="Conradie, W. R. &amp; Branch, W. R." box="[941,1189,1915,1942]" pageId="33" pageNumber="334" pagination="6 - 36" refId="ref26770" refString="Conradie, W. R. &amp; Branch, W. R. (2016) The herpetofauna of the Cubango, Cuito and lower Cuando river catchments of southeastern Angola. Amphibian &amp; Reptile Conservation, 10 (2), 6 - 36." type="journal article" year="2016">
Conradie
<emphasis id="A3ADEA885452FFEA9A3CF8851C21F872" box="[1051,1108,1915,1941]" italics="true" pageId="33" pageNumber="334">et al.</emphasis>
(2016)
</bibRefCitation>
and the interpretation by
<bibRefCitation id="F5484B6B5452FFEA9E9BF86119C6F85E" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[188,435,1951,1977]" pageId="33" pageNumber="334" pagination="1 - 501" refId="ref28241" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="A3ADEA885452FFEA9F01F8611915F85E" box="[294,352,1951,1977]" italics="true" pageId="33" pageNumber="334">et al.</emphasis>
(2018)
</bibRefCitation>
who considered the records of
<taxonomicName id="56D94D195452FFEA9D3DF8611BECF85F" box="[794,921,1951,1976]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885452FFEA9D3DF8611BECF85F" box="[794,921,1951,1976]" italics="true" pageId="33" pageNumber="334">L. capensis</emphasis>
</taxonomicName>
from
<collectingRegion id="531DF8785452FFEA9DC6F8611CD3F85E" box="[993,1190,1951,1977]" country="Angola" name="Namibe" pageId="33" pageNumber="334">Namibe Province</collectingRegion>
to belong to
<taxonomicName id="56D94D195452FFEA9B18F86118A4F83A" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885452FFEA9B18F86118A4F83A" italics="true" pageId="33" pageNumber="334">L. bradfieldi</emphasis>
</taxonomicName>
, based on the latter species distribution and adaptation to arid savanna habitats, is understandable given the subtle morphological differentiation between
<taxonomicName id="56D94D195451FFE99CB1FF691B66FF57" box="[662,787,151,176]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885451FFE99CB1FF691B66FF57" box="[662,787,151,176]" italics="true" pageId="34" pageNumber="335">L. capensis</emphasis>
</taxonomicName>
species complex and
<taxonomicName id="56D94D195451FFE99A2EFF691CE1FF56" box="[1033,1172,151,177]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885451FFE99A2EFF691CE1FF56" box="[1033,1172,151,177]" italics="true" pageId="34" pageNumber="335">L. bradfieldi</emphasis>
</taxonomicName>
. A review of our newly collected specimens and those historical museum specimens recorded as
<taxonomicName id="56D94D195451FFE99DE0FF451C30FF33" box="[967,1093,187,212]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885451FFE99DE0FF451C30FF33" box="[967,1093,187,212]" italics="true" pageId="34" pageNumber="335">L. capensis</emphasis>
</taxonomicName>
for
<collectingCountry id="E9CE760A5451FFE99A55FF451CB1FF32" box="[1138,1220,187,213]" name="Angola" pageId="34" pageNumber="335">Angola</collectingCountry>
revealed that all of these specimens have the subcaudal scales arrangement typical of
<taxonomicName id="56D94D195451FFE99D5BFF211C72FF1E" box="[892,1031,223,249]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885451FFE99D5BFF211C72FF1E" box="[892,1031,223,249]" italics="true" pageId="34" pageNumber="335">L. bradfieldi</emphasis>
</taxonomicName>
and/or three nasal scales around the nostril (an
<taxonomicName id="56D94D195451FFE99F29FEFD19FFFEFB" box="[270,394,259,284]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885451FFE99F29FEFD19FFFEFB" box="[270,394,259,284]" italics="true" pageId="34" pageNumber="335">L. capensis</emphasis>
</taxonomicName>
character). However, the molecular data provide evidence of the distinctiveness of these specimens from all other taxa, and morphological data also allow us to separate them from
<emphasis id="A3ADEA885451FFE99A7AFED91D1BFEA7" box="[1117,1390,295,320]" italics="true" pageId="34" pageNumber="335">
<taxonomicName id="56D94D195451FFE99A7AFED91D6AFEA7" box="[1117,1311,295,320]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="309" phylum="Chordata" rank="subSpecies" species="capensis" subSpecies="sensu">
L. capensis
<taxonomicNameLabel id="B89E57F35451FFE99AC6FED91D6AFEA7" box="[1249,1311,295,320]" pageId="34" pageNumber="335" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
and
<emphasis id="A3ADEA885451FFE99EB0FEB519C3FE83" box="[151,438,331,357]" italics="true" pageId="34" pageNumber="335">
<taxonomicName id="56D94D195451FFE99EB0FEB51957FE82" box="[151,290,331,357]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" sensu="stricto" species="bradfiedli">L. bradfiedli</taxonomicName>
<taxonomicNameLabel id="B89E57F35451FFE99F0EFEB519C3FE83" box="[297,438,331,356]" pageId="34" pageNumber="335" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
Besides the historical data, the new taxon has been found in
<collectingRegion id="531DF8785451FFE99A7BFEB51CC0FE82" box="[1116,1205,331,357]" country="Angola" name="Namibe" pageId="34" pageNumber="335">Namibe</collectingRegion>
and
<collectingRegion id="531DF8785451FFE99ACCFEB51D5CFE82" box="[1259,1321,331,357]" country="Angola" name="Huila" pageId="34" pageNumber="335">Huíla</collectingRegion>
provinces (this paper; see Specimens examined below). Specimens cited by
<bibRefCitation id="F5484B6B5451FFE99DA2FE911C53FE6E" author="Bocage, J. V. B." box="[901,1062,367,393]" pageId="34" pageNumber="335" pagination="217 - 228" refId="ref25473" refString="Bocage, J. V. B. (1867) Segunda lista dos reptis das possessoes portuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 217 - 228." type="journal article" year="1867">Bocage (1867</bibRefCitation>
,
<bibRefCitation id="F5484B6B5451FFE99A15FE901C1EFE6E" author="Bocage, J. V. B." box="[1074,1131,366,393]" pageId="34" pageNumber="335" refId="ref25600" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">1895</bibRefCitation>
) from
<collectingRegion id="531DF8785451FFE99A9CFE911D50FE6E" box="[1211,1317,367,393]" country="Angola" name="Benguela" pageId="34" pageNumber="335">Benguela</collectingRegion>
and
<collectingRegion id="531DF8785451FFE99B79FE911DE9FE6E" box="[1374,1436,367,393]" country="Angola" name="Huila" pageId="34" pageNumber="335">Huíla</collectingRegion>
provinces were not analyzed, as they were lost in the fire that destroyed the Lisbon museum in 1978. However, the distribution of the species, as well as the limited data provided by
<bibRefCitation id="F5484B6B5451FFE99DA0FE491C53FE36" author="Bocage, J. V. B." box="[903,1062,438,465]" pageId="34" pageNumber="335" refId="ref25600" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage (1895</bibRefCitation>
,
<bibRefCitation id="F5484B6B5451FFE99A14FE491C19FE35" author="Bocage, J. V. B." box="[1075,1132,439,466]" pageId="34" pageNumber="335" pagination="105 - 114" refId="ref25633" refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114." type="journal article" year="1896">1896</bibRefCitation>
) allow us to confidentially allocate these specimens to the new taxon. The specimens from Cuando Cubango cited by
<bibRefCitation id="F5484B6B5451FFE99A86FE251DE9FE11" author="Conradie, W. R. &amp; Branch, W. R." box="[1185,1436,475,502]" pageId="34" pageNumber="335" pagination="6 - 36" refId="ref26770" refString="Conradie, W. R. &amp; Branch, W. R. (2016) The herpetofauna of the Cubango, Cuito and lower Cuando river catchments of southeastern Angola. Amphibian &amp; Reptile Conservation, 10 (2), 6 - 36." type="journal article" year="2016">
Conradie
<emphasis id="A3ADEA885451FFE99B37FE251D3FFE12" box="[1296,1354,475,501]" italics="true" pageId="34" pageNumber="335">et al.</emphasis>
(2016)
</bibRefCitation>
need molecular and morphological reevaluation.
</paragraph>
<caption id="C5A666125452FFEA9EB0FEB21A1AFE6F" pageId="33" pageNumber="334">
<paragraph id="9166369A5452FFEA9EB0FEB21A1AFE6F" blockId="33.[151,1436,332,392]" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB0FEB2197FFE83" bold="true" box="[151,266,332,356]" pageId="33" pageNumber="334">TABLE 6.</emphasis>
Measurements and meristic data of the type series of
<taxonomicName id="56D94D195452FFEA9D69FEB21C49FE83" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[846,1084,332,356]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="334" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885452FFEA9D69FEB21C49FE83" box="[846,1084,332,356]" italics="true" pageId="33" pageNumber="334">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885452FFEA9A64FEB31CECFE82" bold="true" box="[1091,1177,333,357]" pageId="33" pageNumber="334">
<taxonomicNameLabel id="B89E57F35452FFEA9A64FEB31CECFE82" box="[1091,1177,333,357]" pageId="33" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations are those described in Materials and Methods section.
</paragraph>
</caption>
<paragraph id="9166369A5452FFEA9F15FE541DE1F9D1" pageId="33" pageNumber="334">
<table id="E3D9C43A545200349EB8FE541DE0F9D1" box="[159,1429,426,1590]" gridcols="8" gridrows="29" pageId="33" pageNumber="334">
<tr id="2FE934D8545200349EB8FE541DE0FE04" box="[159,1429,426,483]" gridrow="0" pageId="33" pageNumber="334" rowspan-0="1">
<th id="6C385DA4545200349F30FE5419DFFE04" box="[279,426,426,483]" gridcol="1" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9F15FE5419EAFE05" bold="true" pageId="33" pageNumber="334">Holotype CAS 263418</emphasis>
</th>
<th id="6C385DA4545200349F9AFE541A25FE04" box="[445,592,426,483]" gridcol="2" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9FFFFE541A30FE05" bold="true" pageId="33" pageNumber="334">Paratype CAS 263420</emphasis>
</th>
<th id="6C385DA4545200349C45FE541A80FE04" box="[610,757,426,483]" gridcol="3" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9C5AFE541A9EFE05" bold="true" pageId="33" pageNumber="334">Paratype CAS 263529</emphasis>
</th>
<th id="6C385DA4545200349D2BFE541BEAFE04" box="[780,927,426,483]" gridcol="4" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9D00FE541BE1FE05" bold="true" pageId="33" pageNumber="334">Paratype CAS 263531</emphasis>
</th>
<th id="6C385DA4545200349D91FE541C3CFE04" box="[950,1097,426,483]" gridcol="5" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9DF6FE541C43FE05" bold="true" pageId="33" pageNumber="334">Paratype UF 187228</emphasis>
</th>
<th id="6C385DA4545200349A7DFE541C98FE04" box="[1114,1261,426,483]" gridcol="6" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9A52FE541CAEFE05" bold="true" pageId="33" pageNumber="334">Paratype UF 187229</emphasis>
</th>
<th id="6C385DA4545200349B25FE541DE0FE04" box="[1282,1429,426,483]" gridcol="7" gridrow="0" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9B3AFE541DF6FE05" bold="true" pageId="33" pageNumber="334">Paratype UF 187230</emphasis>
</th>
</tr>
<tr id="2FE934D8545200349EB8FE131DE0FDE4" box="[159,1429,493,515]" gridrow="1" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FE131971FDE4" box="[159,260,493,515]" gridcol="0" gridrow="1" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FE1318BBFDE4" bold="true" box="[159,206,493,515]" pageId="33" pageNumber="334">SEX</emphasis>
</th>
<td id="6C385DA4545200349F30FE1319DFFDE4" box="[279,426,493,515]" gridcol="1" gridrow="1" pageId="33" pageNumber="334">Male</td>
<td id="6C385DA4545200349F9AFE131A25FDE4" box="[445,592,493,515]" gridcol="2" gridrow="1" pageId="33" pageNumber="334">Female</td>
<td id="6C385DA4545200349C45FE131A80FDE4" box="[610,757,493,515]" gridcol="3" gridrow="1" pageId="33" pageNumber="334">Male</td>
<td id="6C385DA4545200349D2BFE131BEAFDE4" box="[780,927,493,515]" gridcol="4" gridrow="1" pageId="33" pageNumber="334">Male</td>
<td id="6C385DA4545200349D91FE131C3CFDE4" box="[950,1097,493,515]" gridcol="5" gridrow="1" pageId="33" pageNumber="334">Male</td>
<td id="6C385DA4545200349A7DFE131C98FDE4" box="[1114,1261,493,515]" gridcol="6" gridrow="1" pageId="33" pageNumber="334">Male</td>
<td id="6C385DA4545200349B25FE131DE0FDE4" box="[1282,1429,493,515]" gridcol="7" gridrow="1" pageId="33" pageNumber="334">Male</td>
</tr>
<tr id="2FE934D8545200349EB8FDF21DE0FDC4" box="[159,1429,524,547]" gridrow="2" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FDF21971FDC4" box="[159,260,524,547]" gridcol="0" gridrow="2" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FDF218BBFDC5" bold="true" box="[159,206,524,546]" pageId="33" pageNumber="334">SVL</emphasis>
</th>
<td id="6C385DA4545200349F30FDF219DFFDC4" box="[279,426,524,547]" gridcol="1" gridrow="2" pageId="33" pageNumber="334">34.1</td>
<td id="6C385DA4545200349F9AFDF21A25FDC4" box="[445,592,524,547]" gridcol="2" gridrow="2" pageId="33" pageNumber="334">32.1</td>
<td id="6C385DA4545200349C45FDF21A80FDC4" box="[610,757,524,547]" gridcol="3" gridrow="2" pageId="33" pageNumber="334">28.8</td>
<td id="6C385DA4545200349D2BFDF21BEAFDC4" box="[780,927,524,547]" gridcol="4" gridrow="2" pageId="33" pageNumber="334">32.7</td>
<td id="6C385DA4545200349D91FDF21C3CFDC4" box="[950,1097,524,547]" gridcol="5" gridrow="2" pageId="33" pageNumber="334">25.6</td>
<td id="6C385DA4545200349A7DFDF21C98FDC4" box="[1114,1261,524,547]" gridcol="6" gridrow="2" pageId="33" pageNumber="334">30.6</td>
<td id="6C385DA4545200349B25FDF21DE0FDC4" box="[1282,1429,524,547]" gridcol="7" gridrow="2" pageId="33" pageNumber="334">26.2</td>
</tr>
<tr id="2FE934D8545200349EB8FDD21DE0FDA4" box="[159,1429,556,579]" gridrow="3" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FDD21971FDA4" box="[159,260,556,579]" gridcol="0" gridrow="3" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FDD21885FDA6" bold="true" box="[159,240,556,577]" pageId="33" pageNumber="334">TrunkL</emphasis>
</th>
<td id="6C385DA4545200349F30FDD219DFFDA4" box="[279,426,556,579]" gridcol="1" gridrow="3" pageId="33" pageNumber="334">16.7</td>
<td id="6C385DA4545200349F9AFDD21A25FDA4" box="[445,592,556,579]" gridcol="2" gridrow="3" pageId="33" pageNumber="334">15.3</td>
<td id="6C385DA4545200349C45FDD21A80FDA4" box="[610,757,556,579]" gridcol="3" gridrow="3" pageId="33" pageNumber="334">12.5</td>
<td id="6C385DA4545200349D2BFDD21BEAFDA4" box="[780,927,556,579]" gridcol="4" gridrow="3" pageId="33" pageNumber="334">14.3</td>
<td id="6C385DA4545200349D91FDD21C3CFDA4" box="[950,1097,556,579]" gridcol="5" gridrow="3" pageId="33" pageNumber="334">10.7</td>
<td id="6C385DA4545200349A7DFDD21C98FDA4" box="[1114,1261,556,579]" gridcol="6" gridrow="3" pageId="33" pageNumber="334">14.4</td>
<td id="6C385DA4545200349B25FDD21DE0FDA4" box="[1282,1429,556,579]" gridcol="7" gridrow="3" pageId="33" pageNumber="334">12.1</td>
</tr>
<tr id="2FE934D8545200349EB8FDB21DE0FD85" box="[159,1429,588,610]" gridrow="4" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FDB21971FD85" box="[159,260,588,610]" gridcol="0" gridrow="4" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FDB21897FD85" bold="true" box="[159,226,588,610]" pageId="33" pageNumber="334">CrusL</emphasis>
</th>
<td id="6C385DA4545200349F30FDB219DFFD85" box="[279,426,588,610]" gridcol="1" gridrow="4" pageId="33" pageNumber="334">5.3</td>
<td id="6C385DA4545200349F9AFDB21A25FD85" box="[445,592,588,610]" gridcol="2" gridrow="4" pageId="33" pageNumber="334">4.6</td>
<td id="6C385DA4545200349C45FDB21A80FD85" box="[610,757,588,610]" gridcol="3" gridrow="4" pageId="33" pageNumber="334">4.9</td>
<td id="6C385DA4545200349D2BFDB21BEAFD85" box="[780,927,588,610]" gridcol="4" gridrow="4" pageId="33" pageNumber="334">4.8</td>
<td id="6C385DA4545200349D91FDB21C3CFD85" box="[950,1097,588,610]" gridcol="5" gridrow="4" pageId="33" pageNumber="334">4</td>
<td id="6C385DA4545200349A7DFDB21C98FD85" box="[1114,1261,588,610]" gridcol="6" gridrow="4" pageId="33" pageNumber="334">5.5</td>
<td id="6C385DA4545200349B25FDB21DE0FD85" box="[1282,1429,588,610]" gridcol="7" gridrow="4" pageId="33" pageNumber="334">4.5</td>
</tr>
<tr id="2FE934D8545200349EB8FD921DE0FD65" box="[159,1429,620,642]" gridrow="5" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FD921971FD65" box="[159,260,620,642]" gridcol="0" gridrow="5" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FD9218CAFD66" bold="true" box="[159,191,620,641]" pageId="33" pageNumber="334">TL</emphasis>
</th>
<td id="6C385DA4545200349F30FD9219DFFD65" box="[279,426,620,642]" gridcol="1" gridrow="5" pageId="33" pageNumber="334">27.2</td>
<td id="6C385DA4545200349F9AFD921A25FD65" box="[445,592,620,642]" gridcol="2" gridrow="5" pageId="33" pageNumber="334">28.2</td>
<td id="6C385DA4545200349C45FD921A80FD65" box="[610,757,620,642]" gridcol="3" gridrow="5" pageId="33" pageNumber="334">32.8</td>
<td id="6C385DA4545200349D2BFD921BEAFD65" box="[780,927,620,642]" gridcol="4" gridrow="5" pageId="33" pageNumber="334">33.3</td>
<td id="6C385DA4545200349D91FD921C3CFD65" box="[950,1097,620,642]" gridcol="5" gridrow="5" pageId="33" pageNumber="334">broken</td>
<td id="6C385DA4545200349A7DFD921C98FD65" box="[1114,1261,620,642]" gridcol="6" gridrow="5" pageId="33" pageNumber="334">broken</td>
<td id="6C385DA4545200349B25FD921DE0FD65" box="[1282,1429,620,642]" gridcol="7" gridrow="5" pageId="33" pageNumber="334">31.2</td>
</tr>
<tr id="2FE934D8545200349EB8FD721DE0FD45" box="[159,1429,652,674]" gridrow="6" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FD721971FD45" box="[159,260,652,674]" gridcol="0" gridrow="6" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FD7218B2FD46" bold="true" box="[159,199,652,673]" pageId="33" pageNumber="334">TW</emphasis>
</th>
<td id="6C385DA4545200349F30FD7219DFFD45" box="[279,426,652,674]" gridcol="1" gridrow="6" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349F9AFD721A25FD45" box="[445,592,652,674]" gridcol="2" gridrow="6" pageId="33" pageNumber="334">2.7</td>
<td id="6C385DA4545200349C45FD721A80FD45" box="[610,757,652,674]" gridcol="3" gridrow="6" pageId="33" pageNumber="334">2.4</td>
<td id="6C385DA4545200349D2BFD721BEAFD45" box="[780,927,652,674]" gridcol="4" gridrow="6" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349D91FD721C3CFD45" box="[950,1097,652,674]" gridcol="5" gridrow="6" pageId="33" pageNumber="334">broken</td>
<td id="6C385DA4545200349A7DFD721C98FD45" box="[1114,1261,652,674]" gridcol="6" gridrow="6" pageId="33" pageNumber="334">3.2</td>
<td id="6C385DA4545200349B25FD721DE0FD45" box="[1282,1429,652,674]" gridcol="7" gridrow="6" pageId="33" pageNumber="334">1.8</td>
</tr>
<tr id="2FE934D8545200349EB8FD551DE0FD25" box="[159,1429,683,706]" gridrow="7" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FD551971FD25" box="[159,260,683,706]" gridcol="0" gridrow="7" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FD5518B7FD27" bold="true" box="[159,194,683,704]" pageId="33" pageNumber="334">HL</emphasis>
</th>
<td id="6C385DA4545200349F30FD5519DFFD25" box="[279,426,683,706]" gridcol="1" gridrow="7" pageId="33" pageNumber="334">8.4</td>
<td id="6C385DA4545200349F9AFD551A25FD25" box="[445,592,683,706]" gridcol="2" gridrow="7" pageId="33" pageNumber="334">8.3</td>
<td id="6C385DA4545200349C45FD551A80FD25" box="[610,757,683,706]" gridcol="3" gridrow="7" pageId="33" pageNumber="334">8.9</td>
<td id="6C385DA4545200349D2BFD551BEAFD25" box="[780,927,683,706]" gridcol="4" gridrow="7" pageId="33" pageNumber="334">7.6</td>
<td id="6C385DA4545200349D91FD551C3CFD25" box="[950,1097,683,706]" gridcol="5" gridrow="7" pageId="33" pageNumber="334">7.9</td>
<td id="6C385DA4545200349A7DFD551C98FD25" box="[1114,1261,683,706]" gridcol="6" gridrow="7" pageId="33" pageNumber="334">8.2</td>
<td id="6C385DA4545200349B25FD551DE0FD25" box="[1282,1429,683,706]" gridcol="7" gridrow="7" pageId="33" pageNumber="334">7.4</td>
</tr>
<tr id="2FE934D8545200349EB8FD351DE0FD06" box="[159,1429,715,737]" gridrow="8" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FD351971FD06" box="[159,260,715,737]" gridcol="0" gridrow="8" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FD3518BFFD07" bold="true" box="[159,202,715,736]" pageId="33" pageNumber="334">HW</emphasis>
</th>
<td id="6C385DA4545200349F30FD3519DFFD06" box="[279,426,715,737]" gridcol="1" gridrow="8" pageId="33" pageNumber="334">5.7</td>
<td id="6C385DA4545200349F9AFD351A25FD06" box="[445,592,715,737]" gridcol="2" gridrow="8" pageId="33" pageNumber="334">5.2</td>
<td id="6C385DA4545200349C45FD351A80FD06" box="[610,757,715,737]" gridcol="3" gridrow="8" pageId="33" pageNumber="334">4.6</td>
<td id="6C385DA4545200349D2BFD351BEAFD06" box="[780,927,715,737]" gridcol="4" gridrow="8" pageId="33" pageNumber="334">5.4</td>
<td id="6C385DA4545200349D91FD351C3CFD06" box="[950,1097,715,737]" gridcol="5" gridrow="8" pageId="33" pageNumber="334">4.8</td>
<td id="6C385DA4545200349A7DFD351C98FD06" box="[1114,1261,715,737]" gridcol="6" gridrow="8" pageId="33" pageNumber="334">5.8</td>
<td id="6C385DA4545200349B25FD351DE0FD06" box="[1282,1429,715,737]" gridcol="7" gridrow="8" pageId="33" pageNumber="334">4.5</td>
</tr>
<tr id="2FE934D8545200349EB8FD151DE0FCE6" box="[159,1429,747,769]" gridrow="9" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FD151971FCE6" box="[159,260,747,769]" gridcol="0" gridrow="9" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FD1518B1FCE7" bold="true" box="[159,196,747,768]" pageId="33" pageNumber="334">HH</emphasis>
</th>
<td id="6C385DA4545200349F30FD1519DFFCE6" box="[279,426,747,769]" gridcol="1" gridrow="9" pageId="33" pageNumber="334">3.8</td>
<td id="6C385DA4545200349F9AFD151A25FCE6" box="[445,592,747,769]" gridcol="2" gridrow="9" pageId="33" pageNumber="334">3.5</td>
<td id="6C385DA4545200349C45FD151A80FCE6" box="[610,757,747,769]" gridcol="3" gridrow="9" pageId="33" pageNumber="334">3.3</td>
<td id="6C385DA4545200349D2BFD151BEAFCE6" box="[780,927,747,769]" gridcol="4" gridrow="9" pageId="33" pageNumber="334">3.3</td>
<td id="6C385DA4545200349D91FD151C3CFCE6" box="[950,1097,747,769]" gridcol="5" gridrow="9" pageId="33" pageNumber="334">2.5</td>
<td id="6C385DA4545200349A7DFD151C98FCE6" box="[1114,1261,747,769]" gridcol="6" gridrow="9" pageId="33" pageNumber="334">3.2</td>
<td id="6C385DA4545200349B25FD151DE0FCE6" box="[1282,1429,747,769]" gridcol="7" gridrow="9" pageId="33" pageNumber="334">3</td>
</tr>
<tr id="2FE934D8545200349EB8FCF51DE0FCC6" box="[159,1429,779,801]" gridrow="10" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FCF51971FCC6" box="[159,260,779,801]" gridcol="0" gridrow="10" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FCF518A3FCC7" bold="true" box="[159,214,779,800]" pageId="33" pageNumber="334">EarL</emphasis>
</th>
<td id="6C385DA4545200349F30FCF519DFFCC6" box="[279,426,779,801]" gridcol="1" gridrow="10" pageId="33" pageNumber="334">0.5</td>
<td id="6C385DA4545200349F9AFCF51A25FCC6" box="[445,592,779,801]" gridcol="2" gridrow="10" pageId="33" pageNumber="334">0.5</td>
<td id="6C385DA4545200349C45FCF51A80FCC6" box="[610,757,779,801]" gridcol="3" gridrow="10" pageId="33" pageNumber="334">0.5</td>
<td id="6C385DA4545200349D2BFCF51BEAFCC6" box="[780,927,779,801]" gridcol="4" gridrow="10" pageId="33" pageNumber="334">0.4</td>
<td id="6C385DA4545200349D91FCF51C3CFCC6" box="[950,1097,779,801]" gridcol="5" gridrow="10" pageId="33" pageNumber="334">0.3</td>
<td id="6C385DA4545200349A7DFCF51C98FCC6" box="[1114,1261,779,801]" gridcol="6" gridrow="10" pageId="33" pageNumber="334">0.5</td>
<td id="6C385DA4545200349B25FCF51DE0FCC6" box="[1282,1429,779,801]" gridcol="7" gridrow="10" pageId="33" pageNumber="334">0.4</td>
</tr>
<tr id="2FE934D8545200349EB8FCD51DE0FCA6" box="[159,1429,811,833]" gridrow="11" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FCD51971FCA6" box="[159,260,811,833]" gridcol="0" gridrow="11" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FCD5189EFCA7" bold="true" box="[159,235,811,832]" pageId="33" pageNumber="334">ForeaL</emphasis>
</th>
<td id="6C385DA4545200349F30FCD519DFFCA6" box="[279,426,811,833]" gridcol="1" gridrow="11" pageId="33" pageNumber="334">4.5</td>
<td id="6C385DA4545200349F9AFCD51A25FCA6" box="[445,592,811,833]" gridcol="2" gridrow="11" pageId="33" pageNumber="334">3.9</td>
<td id="6C385DA4545200349C45FCD51A80FCA6" box="[610,757,811,833]" gridcol="3" gridrow="11" pageId="33" pageNumber="334">3.9</td>
<td id="6C385DA4545200349D2BFCD51BEAFCA6" box="[780,927,811,833]" gridcol="4" gridrow="11" pageId="33" pageNumber="334">4.4</td>
<td id="6C385DA4545200349D91FCD51C3CFCA6" box="[950,1097,811,833]" gridcol="5" gridrow="11" pageId="33" pageNumber="334">3.8</td>
<td id="6C385DA4545200349A7DFCD51C98FCA6" box="[1114,1261,811,833]" gridcol="6" gridrow="11" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349B25FCD51DE0FCA6" box="[1282,1429,811,833]" gridcol="7" gridrow="11" pageId="33" pageNumber="334">4.3</td>
</tr>
<tr id="2FE934D8545200349EB8FCB41DE0FC87" box="[159,1429,842,864]" gridrow="12" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FCB41971FC87" box="[159,260,842,864]" gridcol="0" gridrow="12" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FCB418B5FCB8" bold="true" box="[159,192,842,863]" pageId="33" pageNumber="334">ED</emphasis>
</th>
<td id="6C385DA4545200349F30FCB419DFFC87" box="[279,426,842,864]" gridcol="1" gridrow="12" pageId="33" pageNumber="334">1.8</td>
<td id="6C385DA4545200349F9AFCB41A25FC87" box="[445,592,842,864]" gridcol="2" gridrow="12" pageId="33" pageNumber="334">1.9</td>
<td id="6C385DA4545200349C45FCB41A80FC87" box="[610,757,842,864]" gridcol="3" gridrow="12" pageId="33" pageNumber="334">1.4</td>
<td id="6C385DA4545200349D2BFCB41BEAFC87" box="[780,927,842,864]" gridcol="4" gridrow="12" pageId="33" pageNumber="334">1.7</td>
<td id="6C385DA4545200349D91FCB41C3CFC87" box="[950,1097,842,864]" gridcol="5" gridrow="12" pageId="33" pageNumber="334">1.5</td>
<td id="6C385DA4545200349A7DFCB41C98FC87" box="[1114,1261,842,864]" gridcol="6" gridrow="12" pageId="33" pageNumber="334">1.7</td>
<td id="6C385DA4545200349B25FCB41DE0FC87" box="[1282,1429,842,864]" gridcol="7" gridrow="12" pageId="33" pageNumber="334">1.5</td>
</tr>
<tr id="2FE934D8545200349EB8FC941DE0FC67" box="[159,1429,874,896]" gridrow="13" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FC941971FC67" box="[159,260,874,896]" gridcol="0" gridrow="13" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FC9418B5FC98" bold="true" box="[159,192,874,895]" pageId="33" pageNumber="334">NE</emphasis>
</th>
<td id="6C385DA4545200349F30FC9419DFFC67" box="[279,426,874,896]" gridcol="1" gridrow="13" pageId="33" pageNumber="334">2.5</td>
<td id="6C385DA4545200349F9AFC941A25FC67" box="[445,592,874,896]" gridcol="2" gridrow="13" pageId="33" pageNumber="334">2.5</td>
<td id="6C385DA4545200349C45FC941A80FC67" box="[610,757,874,896]" gridcol="3" gridrow="13" pageId="33" pageNumber="334">2.8</td>
<td id="6C385DA4545200349D2BFC941BEAFC67" box="[780,927,874,896]" gridcol="4" gridrow="13" pageId="33" pageNumber="334">2</td>
<td id="6C385DA4545200349D91FC941C3CFC67" box="[950,1097,874,896]" gridcol="5" gridrow="13" pageId="33" pageNumber="334">2</td>
<td id="6C385DA4545200349A7DFC941C98FC67" box="[1114,1261,874,896]" gridcol="6" gridrow="13" pageId="33" pageNumber="334">2.8</td>
<td id="6C385DA4545200349B25FC941DE0FC67" box="[1282,1429,874,896]" gridcol="7" gridrow="13" pageId="33" pageNumber="334">2.1</td>
</tr>
<tr id="2FE934D8545200349EB8FC771DE0FC47" box="[159,1429,905,928]" gridrow="14" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FC771971FC47" box="[159,260,905,928]" gridcol="0" gridrow="14" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FC7718C9FC78" bold="true" box="[159,188,905,927]" pageId="33" pageNumber="334">SE</emphasis>
</th>
<td id="6C385DA4545200349F30FC7719DFFC47" box="[279,426,905,928]" gridcol="1" gridrow="14" pageId="33" pageNumber="334">3.9</td>
<td id="6C385DA4545200349F9AFC771A25FC47" box="[445,592,905,928]" gridcol="2" gridrow="14" pageId="33" pageNumber="334">3.7</td>
<td id="6C385DA4545200349C45FC771A80FC47" box="[610,757,905,928]" gridcol="3" gridrow="14" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349D2BFC771BEAFC47" box="[780,927,905,928]" gridcol="4" gridrow="14" pageId="33" pageNumber="334">3.6</td>
<td id="6C385DA4545200349D91FC771C3CFC47" box="[950,1097,905,928]" gridcol="5" gridrow="14" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349A7DFC771C98FC47" box="[1114,1261,905,928]" gridcol="6" gridrow="14" pageId="33" pageNumber="334">4.1</td>
<td id="6C385DA4545200349B25FC771DE0FC47" box="[1282,1429,905,928]" gridcol="7" gridrow="14" pageId="33" pageNumber="334">2.8</td>
</tr>
<tr id="2FE934D8545200349EB8FC541DE0FC27" box="[159,1429,938,960]" gridrow="15" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FC541971FC27" box="[159,260,938,960]" gridcol="0" gridrow="15" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FC5418CAFC58" bold="true" box="[159,191,938,959]" pageId="33" pageNumber="334">EE</emphasis>
</th>
<td id="6C385DA4545200349F30FC5419DFFC27" box="[279,426,938,960]" gridcol="1" gridrow="15" pageId="33" pageNumber="334">2.6</td>
<td id="6C385DA4545200349F9AFC541A25FC27" box="[445,592,938,960]" gridcol="2" gridrow="15" pageId="33" pageNumber="334">2.5</td>
<td id="6C385DA4545200349C45FC541A80FC27" box="[610,757,938,960]" gridcol="3" gridrow="15" pageId="33" pageNumber="334">2</td>
<td id="6C385DA4545200349D2BFC541BEAFC27" box="[780,927,938,960]" gridcol="4" gridrow="15" pageId="33" pageNumber="334">2.5</td>
<td id="6C385DA4545200349D91FC541C3CFC27" box="[950,1097,938,960]" gridcol="5" gridrow="15" pageId="33" pageNumber="334">1.6</td>
<td id="6C385DA4545200349A7DFC541C98FC27" box="[1114,1261,938,960]" gridcol="6" gridrow="15" pageId="33" pageNumber="334">2.2</td>
<td id="6C385DA4545200349B25FC541DE0FC27" box="[1282,1429,938,960]" gridcol="7" gridrow="15" pageId="33" pageNumber="334">1.4</td>
</tr>
<tr id="2FE934D8545200349EB8FC371DE0FC38" box="[159,1429,969,991]" gridrow="16" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FC371971FC38" box="[159,260,969,991]" gridcol="0" gridrow="16" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FC3718CFFC39" bold="true" box="[159,186,969,990]" pageId="33" pageNumber="334">IN</emphasis>
</th>
<td id="6C385DA4545200349F30FC3719DFFC38" box="[279,426,969,991]" gridcol="1" gridrow="16" pageId="33" pageNumber="334">1.1</td>
<td id="6C385DA4545200349F9AFC371A25FC38" box="[445,592,969,991]" gridcol="2" gridrow="16" pageId="33" pageNumber="334">1.1</td>
<td id="6C385DA4545200349C45FC371A80FC38" box="[610,757,969,991]" gridcol="3" gridrow="16" pageId="33" pageNumber="334">1</td>
<td id="6C385DA4545200349D2BFC371BEAFC38" box="[780,927,969,991]" gridcol="4" gridrow="16" pageId="33" pageNumber="334">1.2</td>
<td id="6C385DA4545200349D91FC371C3CFC38" box="[950,1097,969,991]" gridcol="5" gridrow="16" pageId="33" pageNumber="334">0.8</td>
<td id="6C385DA4545200349A7DFC371C98FC38" box="[1114,1261,969,991]" gridcol="6" gridrow="16" pageId="33" pageNumber="334">0.9</td>
<td id="6C385DA4545200349B25FC371DE0FC38" box="[1282,1429,969,991]" gridcol="7" gridrow="16" pageId="33" pageNumber="334">0.7</td>
</tr>
<tr id="2FE934D8545200349EB8FC171DE0FC18" box="[159,1429,1001,1023]" gridrow="17" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FC171971FC18" box="[159,260,1001,1023]" gridcol="0" gridrow="17" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FC1718CEFC18" bold="true" box="[159,187,1001,1023]" pageId="33" pageNumber="334">IO</emphasis>
</th>
<td id="6C385DA4545200349F30FC1719DFFC18" box="[279,426,1001,1023]" gridcol="1" gridrow="17" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349F9AFC171A25FC18" box="[445,592,1001,1023]" gridcol="2" gridrow="17" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349C45FC171A80FC18" box="[610,757,1001,1023]" gridcol="3" gridrow="17" pageId="33" pageNumber="334">2.9</td>
<td id="6C385DA4545200349D2BFC171BEAFC18" box="[780,927,1001,1023]" gridcol="4" gridrow="17" pageId="33" pageNumber="334">3.1</td>
<td id="6C385DA4545200349D91FC171C3CFC18" box="[950,1097,1001,1023]" gridcol="5" gridrow="17" pageId="33" pageNumber="334">2.7</td>
<td id="6C385DA4545200349A7DFC171C98FC18" box="[1114,1261,1001,1023]" gridcol="6" gridrow="17" pageId="33" pageNumber="334">2.9</td>
<td id="6C385DA4545200349B25FC171DE0FC18" box="[1282,1429,1001,1023]" gridcol="7" gridrow="17" pageId="33" pageNumber="334">2.3</td>
</tr>
<tr id="2FE934D8545200349EB8FBF61DE0FBF8" box="[159,1429,1032,1055]" gridrow="18" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FBF61971FBF8" box="[159,260,1032,1055]" gridcol="0" gridrow="18" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FBF618BAFBF9" bold="true" box="[159,207,1032,1054]" pageId="33" pageNumber="334">PCL</emphasis>
</th>
<td id="6C385DA4545200349F30FBF619DFFBF8" box="[279,426,1032,1055]" gridcol="1" gridrow="18" pageId="33" pageNumber="334">6</td>
<td id="6C385DA4545200349F9AFBF61A25FBF8" box="[445,592,1032,1055]" gridcol="2" gridrow="18" pageId="33" pageNumber="334"></td>
<td id="6C385DA4545200349C45FBF61A80FBF8" box="[610,757,1032,1055]" gridcol="3" gridrow="18" pageId="33" pageNumber="334">6</td>
<td id="6C385DA4545200349D2BFBF61BEAFBF8" box="[780,927,1032,1055]" gridcol="4" gridrow="18" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349D91FBF61C3CFBF8" box="[950,1097,1032,1055]" gridcol="5" gridrow="18" pageId="33" pageNumber="334">6</td>
<td id="6C385DA4545200349A7DFBF61C98FBF8" box="[1114,1261,1032,1055]" gridcol="6" gridrow="18" pageId="33" pageNumber="334">4</td>
<td id="6C385DA4545200349B25FBF61DE0FBF8" box="[1282,1429,1032,1055]" gridcol="7" gridrow="18" pageId="33" pageNumber="334">6</td>
</tr>
<tr id="2FE934D8545200349EB8FBD61DE0FBD9" box="[159,1429,1064,1086]" gridrow="19" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FBD61971FBD9" box="[159,260,1064,1086]" gridcol="0" gridrow="19" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FBD618A6FBD9" bold="true" box="[159,211,1064,1086]" pageId="33" pageNumber="334">GEN</emphasis>
</th>
<td id="6C385DA4545200349F30FBD619DFFBD9" box="[279,426,1064,1086]" gridcol="1" gridrow="19" pageId="33" pageNumber="334">19</td>
<td id="6C385DA4545200349F9AFBD61A25FBD9" box="[445,592,1064,1086]" gridcol="2" gridrow="19" pageId="33" pageNumber="334"></td>
<td id="6C385DA4545200349C45FBD61A80FBD9" box="[610,757,1064,1086]" gridcol="3" gridrow="19" pageId="33" pageNumber="334">2120</td>
<td id="6C385DA4545200349D2BFBD61BEAFBD9" box="[780,927,1064,1086]" gridcol="4" gridrow="19" pageId="33" pageNumber="334"></td>
<td id="6C385DA4545200349D91FBD61C3CFBD9" box="[950,1097,1064,1086]" gridcol="5" gridrow="19" pageId="33" pageNumber="334">1920</td>
<td id="6C385DA4545200349A7DFBD61C98FBD9" box="[1114,1261,1064,1086]" gridcol="6" gridrow="19" pageId="33" pageNumber="334">20</td>
<td id="6C385DA4545200349B25FBD61DE0FBD9" box="[1282,1429,1064,1086]" gridcol="7" gridrow="19" pageId="33" pageNumber="334">1819</td>
</tr>
<tr id="2FE934D8545200349EB8FBB61DE0FBB8" box="[159,1429,1096,1119]" gridrow="20" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FBB61971FBB8" box="[159,260,1096,1119]" gridcol="0" gridrow="20" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FBB618A1FBB9" bold="true" box="[159,212,1096,1118]" pageId="33" pageNumber="334">SAM</emphasis>
</th>
<td id="6C385DA4545200349F30FBB619DFFBB8" box="[279,426,1096,1119]" gridcol="1" gridrow="20" pageId="33" pageNumber="334">16</td>
<td id="6C385DA4545200349F9AFBB61A25FBB8" box="[445,592,1096,1119]" gridcol="2" gridrow="20" pageId="33" pageNumber="334">15</td>
<td id="6C385DA4545200349C45FBB61A80FBB8" box="[610,757,1096,1119]" gridcol="3" gridrow="20" pageId="33" pageNumber="334">16</td>
<td id="6C385DA4545200349D2BFBB61BEAFBB8" box="[780,927,1096,1119]" gridcol="4" gridrow="20" pageId="33" pageNumber="334">16</td>
<td id="6C385DA4545200349D91FBB61C3CFBB8" box="[950,1097,1096,1119]" gridcol="5" gridrow="20" pageId="33" pageNumber="334">15</td>
<td id="6C385DA4545200349A7DFBB61C98FBB8" box="[1114,1261,1096,1119]" gridcol="6" gridrow="20" pageId="33" pageNumber="334">15</td>
<td id="6C385DA4545200349B25FBB61DE0FBB8" box="[1282,1429,1096,1119]" gridcol="7" gridrow="20" pageId="33" pageNumber="334">16</td>
</tr>
<tr id="2FE934D8545200349EB8FB961DE0FB99" box="[159,1429,1128,1150]" gridrow="21" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FB961971FB99" box="[159,260,1128,1150]" gridcol="0" gridrow="21" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FB961899FB99" bold="true" box="[159,236,1128,1150]" pageId="33" pageNumber="334">FFingL</emphasis>
</th>
<td id="6C385DA4545200349F30FB9619DFFB99" box="[279,426,1128,1150]" gridcol="1" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349F9AFB961A25FB99" box="[445,592,1128,1150]" gridcol="2" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349C45FB961A80FB99" box="[610,757,1128,1150]" gridcol="3" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349D2BFB961BEAFB99" box="[780,927,1128,1150]" gridcol="4" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349D91FB961C3CFB99" box="[950,1097,1128,1150]" gridcol="5" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349A7DFB961C98FB99" box="[1114,1261,1128,1150]" gridcol="6" gridrow="21" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349B25FB961DE0FB99" box="[1282,1429,1128,1150]" gridcol="7" gridrow="21" pageId="33" pageNumber="334">5</td>
</tr>
<tr id="2FE934D8545200349EB8FB761DE0FB79" box="[159,1429,1160,1182]" gridrow="22" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FB761971FB79" box="[159,260,1160,1182]" gridcol="0" gridrow="22" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FB761897FB7A" bold="true" box="[159,226,1160,1181]" pageId="33" pageNumber="334">FToeL</emphasis>
</th>
<td id="6C385DA4545200349F30FB7619DFFB79" box="[279,426,1160,1182]" gridcol="1" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349F9AFB761A25FB79" box="[445,592,1160,1182]" gridcol="2" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349C45FB761A80FB79" box="[610,757,1160,1182]" gridcol="3" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349D2BFB761BEAFB79" box="[780,927,1160,1182]" gridcol="4" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349D91FB761C3CFB79" box="[950,1097,1160,1182]" gridcol="5" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349A7DFB761C98FB79" box="[1114,1261,1160,1182]" gridcol="6" gridrow="22" pageId="33" pageNumber="334">5</td>
<td id="6C385DA4545200349B25FB761DE0FB79" box="[1282,1429,1160,1182]" gridcol="7" gridrow="22" pageId="33" pageNumber="334">5</td>
</tr>
<tr id="2FE934D8545200349EB8FB591DE0FB59" box="[159,1429,1191,1214]" gridrow="23" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FB591971FB59" box="[159,260,1191,1214]" gridcol="0" gridrow="23" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FB5918B6FB5A" bold="true" box="[159,195,1191,1213]" pageId="33" pageNumber="334">MS</emphasis>
</th>
<td id="6C385DA4545200349F30FB5919DFFB59" box="[279,426,1191,1214]" gridcol="1" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349F9AFB591A25FB59" box="[445,592,1191,1214]" gridcol="2" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349C45FB591A80FB59" box="[610,757,1191,1214]" gridcol="3" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349D2BFB591BEAFB59" box="[780,927,1191,1214]" gridcol="4" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349D91FB591C3CFB59" box="[950,1097,1191,1214]" gridcol="5" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349A7DFB591C98FB59" box="[1114,1261,1191,1214]" gridcol="6" gridrow="23" pageId="33" pageNumber="334">undivided</td>
<td id="6C385DA4545200349B25FB591DE0FB59" box="[1282,1429,1191,1214]" gridcol="7" gridrow="23" pageId="33" pageNumber="334">undivided</td>
</tr>
<tr id="2FE934D8545200349EB8FB131DE0FA88" box="[159,1429,1261,1391]" gridrow="24" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FB131971FA88" box="[159,260,1261,1391]" gridcol="0" gridrow="24" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FB131897FAAC" bold="true" pageId="33" pageNumber="334">No. scales entering nostril</emphasis>
</th>
<td id="6C385DA4545200349F30FB1319DFFA88" box="[279,426,1261,1391]" gridcol="1" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349F9AFB131A25FA88" box="[445,592,1261,1391]" gridcol="2" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349C45FB131A80FA88" box="[610,757,1261,1391]" gridcol="3" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349D2BFB131BEAFA88" box="[780,927,1261,1391]" gridcol="4" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349D91FB131C3CFA88" box="[950,1097,1261,1391]" gridcol="5" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349A7DFB131C98FA88" box="[1114,1261,1261,1391]" gridcol="6" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
<td id="6C385DA4545200349B25FB131DE0FA88" box="[1282,1429,1261,1391]" gridcol="7" gridrow="24" pageId="33" pageNumber="334">4 (two supranasal. 1st supralabial and one postnasals)</td>
</tr>
<tr id="2FE934D8545200349EB8FA871DE0FA69" box="[159,1429,1401,1422]" gridrow="25" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FA871971FA69" box="[159,260,1401,1422]" gridcol="0" gridrow="25" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FA8718B1FA69" bold="true" box="[159,196,1401,1422]" pageId="33" pageNumber="334">PM</emphasis>
</th>
<td id="6C385DA4545200349F30FA8719DFFA69" box="[279,426,1401,1422]" gridcol="1" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349F9AFA871A25FA69" box="[445,592,1401,1422]" gridcol="2" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349C45FA871A80FA69" box="[610,757,1401,1422]" gridcol="3" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349D2BFA871BEAFA69" box="[780,927,1401,1422]" gridcol="4" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349D91FA871C3CFA69" box="[950,1097,1401,1422]" gridcol="5" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349A7DFA871C98FA69" box="[1114,1261,1401,1422]" gridcol="6" gridrow="25" pageId="33" pageNumber="334">3</td>
<td id="6C385DA4545200349B25FA871DE0FA69" box="[1282,1429,1401,1422]" gridcol="7" gridrow="25" pageId="33" pageNumber="334">3</td>
</tr>
<tr id="2FE934D8545200349EB8FA661DE0FA49" box="[159,1429,1432,1454]" gridrow="26" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FA661971FA49" box="[159,260,1432,1454]" gridcol="0" gridrow="26" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FA6618C9FA49" bold="true" box="[159,188,1432,1454]" pageId="33" pageNumber="334">SL</emphasis>
</th>
<td id="6C385DA4545200349F30FA6619DFFA49" box="[279,426,1432,1454]" gridcol="1" gridrow="26" pageId="33" pageNumber="334">8</td>
<td id="6C385DA4545200349F9AFA661A25FA49" box="[445,592,1432,1454]" gridcol="2" gridrow="26" pageId="33" pageNumber="334">8</td>
<td id="6C385DA4545200349C45FA661A80FA49" box="[610,757,1432,1454]" gridcol="3" gridrow="26" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349D2BFA661BEAFA49" box="[780,927,1432,1454]" gridcol="4" gridrow="26" pageId="33" pageNumber="334">8</td>
<td id="6C385DA4545200349D91FA661C3CFA49" box="[950,1097,1432,1454]" gridcol="5" gridrow="26" pageId="33" pageNumber="334">8</td>
<td id="6C385DA4545200349A7DFA661C98FA49" box="[1114,1261,1432,1454]" gridcol="6" gridrow="26" pageId="33" pageNumber="334">8</td>
<td id="6C385DA4545200349B25FA661DE0FA49" box="[1282,1429,1432,1454]" gridcol="7" gridrow="26" pageId="33" pageNumber="334">8</td>
</tr>
<tr id="2FE934D8545200349EB8FA471DE0FA29" box="[159,1429,1465,1486]" gridrow="27" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FA471971FA29" box="[159,260,1465,1486]" gridcol="0" gridrow="27" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FA4718CDFA29" bold="true" box="[159,184,1465,1486]" pageId="33" pageNumber="334">IL</emphasis>
</th>
<td id="6C385DA4545200349F30FA4719DFFA29" box="[279,426,1465,1486]" gridcol="1" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349F9AFA471A25FA29" box="[445,592,1465,1486]" gridcol="2" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349C45FA471A80FA29" box="[610,757,1465,1486]" gridcol="3" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349D2BFA471BEAFA29" box="[780,927,1465,1486]" gridcol="4" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349D91FA471C3CFA29" box="[950,1097,1465,1486]" gridcol="5" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349A7DFA471C98FA29" box="[1114,1261,1465,1486]" gridcol="6" gridrow="27" pageId="33" pageNumber="334">7</td>
<td id="6C385DA4545200349B25FA471DE0FA29" box="[1282,1429,1465,1486]" gridcol="7" gridrow="27" pageId="33" pageNumber="334">7</td>
</tr>
<tr id="2FE934D8545200349EB8FA291DE0F9D1" box="[159,1429,1495,1590]" gridrow="28" pageId="33" pageNumber="334">
<th id="6C385DA4545200349EB8FA291971F9D1" box="[159,260,1495,1590]" gridcol="0" gridrow="28" pageId="33" pageNumber="334">
<emphasis id="A3ADEA885452FFEA9EB8FA2618DEFA0A" bold="true" box="[159,171,1496,1517]" pageId="33" pageNumber="334">1</emphasis>
<emphasis id="A3ADEA885452FFEA9E8CFA2918C9FA04" bold="true" box="[171,188,1495,1507]" pageId="33" pageNumber="334">ST</emphasis>
<emphasis id="A3ADEA885452FFEA9EE5FA26189FF9F6" bold="true" pageId="33" pageNumber="334">IL contact</emphasis>
</th>
<td id="6C385DA4545200349F30FA2919DFF9D1" box="[279,426,1495,1590]" gridcol="1" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349F9AFA291A25F9D1" box="[445,592,1495,1590]" gridcol="2" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349C45FA291A80F9D1" box="[610,757,1495,1590]" gridcol="3" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349D2BFA291BEAF9D1" box="[780,927,1495,1590]" gridcol="4" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349D91FA291C3CF9D1" box="[950,1097,1495,1590]" gridcol="5" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349A7DFA291C98F9D1" box="[1114,1261,1495,1590]" gridcol="6" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
<td id="6C385DA4545200349B25FA291DE0F9D1" box="[1282,1429,1495,1590]" gridcol="7" gridrow="28" pageId="33" pageNumber="334">2550% overlap with postpostmental</td>
</tr>
</table>
</paragraph>
<paragraph id="9166369A5451FFE99EE0FDDD1B3BFD62" blockId="34.[151,1437,151,1905]" pageId="34" pageNumber="335">
<materialsCitation id="21B13CC75451FFE99EE0FDDD1B3BFD62" ID-GBIF-Occurrence="3031141309" accessionNumber="AMB 10134" collectingDate="2017-07-21" collectionCode="AMB, T" collectorName="M. Marques &amp; Marques, L &amp; Ceriaco, B &amp; Julio &amp; M. Heinicke" country="Angola" county="Kimbo do Soba" elevation="1866" location="An" municipality="Lubango" pageId="34" pageNumber="335" specimenCode="CAS 263418" specimenCount="1" stateProvince="Huila" typeStatus="holotype">
<emphasis id="A3ADEA885451FFE99EE0FDDD194EFDDA" bold="true" box="[199,315,547,573]" pageId="34" pageNumber="335">
<typeStatus id="4E6288385451FFE99EE0FDDD1943FDDA" box="[199,310,547,573]" pageId="34" pageNumber="335" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="940660415451FFE99F64FDDD1910FDDA" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99F64FDDD1910FDDA" box="[323,357,547,573]" country="Angola" county="Kimbo do Soba" municipality="Lubango" name="An" pageId="34" pageNumber="335" stateProvince="Huila">An</location>
adult male (
<specimenCode id="C17F9EE15451FFE99FD3FDDD1AFEFDDA" box="[500,651,547,573]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="335" type="Museum">CAS 263418</specimenCode>
, field number
<collectionCode id="F7C8AE5F5451FFE99D08FDDD1B1AFDDA" box="[815,879,547,573]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10134;
<figureCitation id="09E22A1F5451FFE99DEBFDDD1C6BFDDA" box="[972,1054,547,573]" captionStart="FIGURE 14" captionStartId="35.[151,250,1008,1032]" captionTargetBox="[151,1437,181,985]" captionTargetId="figure-19@35.[151,1437,181,985]" captionTargetPageId="35" captionText="FIGURE 14. Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of Lygodactylus nyaneka sp. nov. (CAS 263418)." figureDoi="http://doi.org/10.5281/zenodo.4410755" httpUri="https://zenodo.org/record/4410755/files/figure.png" pageId="34" pageNumber="335">Fig. 14</figureCitation>
) collected at
<collectingMunicipality id="7102ACE05451FFE99A9FFDDD1D6BFDDA" box="[1208,1310,547,573]" pageId="34" pageNumber="335">Lubango</collectingMunicipality>
,
<collectingCounty id="78074E165451FFE99B0DFDDD18A7FD86" pageId="34" pageNumber="335">Kimbo do Soba</collectingCounty>
,
<collectingRegion id="531DF8785451FFE99EC7FDB919FBFD86" box="[224,398,583,609]" country="Angola" name="Huila" pageId="34" pageNumber="335">Huíla Province</collectingRegion>
, southwestern
<collectingCountry id="E9CE760A5451FFE99C1FFDB91AFFFD86" box="[568,650,583,609]" name="Angola" pageId="34" pageNumber="335">Angola</collectingCountry>
[-14.93419º, 13.46925º,
<quantity id="56219B7F5451FFE99D8CFDB91C76FD86" box="[939,1027,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="34" pageNumber="335" unit="m" value="1866.0">
<elevation id="1AF4D1A95451FFE99D8CFDB91C76FD86" box="[939,1027,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="34" pageNumber="335" unit="m" value="1866.0">1866 m</elevation>
</quantity>
a.s.l], collected by
<collectorName id="3C2C534C5451FFE99ACFFDB91D03FD86" box="[1256,1398,583,609]" pageId="34" pageNumber="335">M. Marques</collectorName>
, L.
<collectorName id="3C2C534C5451FFE99EB0FD951967FD62" box="[151,274,619,645]" pageId="34" pageNumber="335">Ceríaco, B</collectorName>
. Butler,
<collectionCode id="F7C8AE5F5451FFE99F48FD9519F4FD62" box="[367,385,619,645]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="335">T</collectionCode>
.
<collectorName id="3C2C534C5451FFE99FABFD9519B7FD62" box="[396,450,619,645]" pageId="34" pageNumber="335">Júlio</collectorName>
and
<collectorName id="3C2C534C5451FFE99FDEFD951AFDFD62" box="[505,648,619,645]" pageId="34" pageNumber="335">M. Heinicke</collectorName>
, on
<date id="E567105A5451FFE99C91FD951B3EFD62" box="[694,843,619,645]" pageId="34" pageNumber="335" value="2017-07-21">
<collectingDate id="F523E9B25451FFE99C91FD951B3EFD62" box="[694,843,619,645]" pageId="34" pageNumber="335" value="2017-07-21">21 July 2017</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="9166369A5451FFE99EE0FD711B5FFBF6" blockId="34.[151,1437,151,1905]" pageId="34" pageNumber="335">
<materialsCitation id="21B13CC75451FFE99EE0FD71199BFD2A" ID-GBIF-Occurrence="3031141328" accessionNumber="AMB 10136" collectingDate="2017-07-21" collectionCode="AMB" collectorName="M. Marques &amp; Marques, L &amp; Ceriaco, B &amp; Julio &amp; M. Heinicke" country="Angola" county="An" elevation="1866" location="All" municipality="Six" pageId="34" pageNumber="335" specimenCode="CAS 263420" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<emphasis id="A3ADEA885451FFE99EE0FD711933FD4E" bold="true" box="[199,326,655,681]" pageId="34" pageNumber="335">
<typeStatus id="4E6288385451FFE99EE0FD711934FD4E" box="[199,321,655,681]" pageId="34" pageNumber="335" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<location id="940660415451FFE99F77FD711901FD4E" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99F77FD711901FD4E" box="[336,372,655,681]" country="Angola" county="An" municipality="Six" name="All" pageId="34" pageNumber="335">All</location>
specimens from
<collectingCountry id="E9CE760A5451FFE99C1DFD711AE5FD4E" box="[570,656,655,681]" name="Angola" pageId="34" pageNumber="335">Angola</collectingCountry>
.
<collectingMunicipality id="7102ACE05451FFE99CBAFD711AB7FD4E" box="[669,706,655,681]" pageId="34" pageNumber="335">Six</collectingMunicipality>
specimens:
<collectingCounty id="78074E165451FFE99D77FD711B07FD4E" box="[848,882,655,681]" pageId="34" pageNumber="335">An</collectingCounty>
adult female (
<specimenCode id="C17F9EE15451FFE99A3BFD711CC1FD4E" box="[1052,1204,655,681]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="335" type="Museum">CAS 263420</specimenCode>
, field number
<collectionCode id="F7C8AE5F5451FFE99B7BFD711DE9FD4E" box="[1372,1436,655,681]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10136), same data as holotype
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99FD0FD4D18A6FCF2" ID-GBIF-Occurrence="3031141319" accessionNumber="AMB 10363" collectingDate="2016-11-23" collectionCode="AMB" collectorName="L. Ceriaco &amp; S. Bandeira &amp; I. Agarwal" country="Angola" elevation="864" location="An" municipality="Bibala Crossroad" pageId="34" pageNumber="335" specimenCode="CAS 263529" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<location id="940660415451FFE99FD0FD4D1A6CFD2A" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99FD0FD4D1A6CFD2A" box="[503,537,691,717]" country="Angola" municipality="Bibala Crossroad" name="An" pageId="34" pageNumber="335" stateProvince="Huila">An</location>
adult male (
<specimenCode id="C17F9EE15451FFE99C83FD4D1B4CFD2A" box="[676,825,690,717]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="335" type="Museum">CAS 263529</specimenCode>
, field number
<collectionCode id="F7C8AE5F5451FFE99DFEFD4D1C6CFD2A" box="[985,1049,691,717]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10363), collected at
<collectingMunicipality id="7102ACE05451FFE99B24FD4D18BCFD16" pageId="34" pageNumber="335">Bibala Crossroad</collectingMunicipality>
[-14.74350º, 13.33222º,
<elevation id="1AF4D1A95451FFE99FCEFD291A04FD16" box="[489,625,726,753]" metricMagnitude="2" metricUnit="m" metricValue="8.64" pageId="34" pageNumber="335" unit="m" value="864.0">
<quantity id="56219B7F5451FFE99FCEFD291A47FD16" box="[489,562,727,753]" metricMagnitude="2" metricUnit="m" metricValue="8.64" pageId="34" pageNumber="335" unit="m" value="864.0">864 m</quantity>
a.s.l.
</elevation>
], collected by
<collectorName id="3C2C534C5451FFE99D3AFD291BEDFD16" box="[797,920,727,753]" pageId="34" pageNumber="335">L. Ceríaco</collectorName>
,
<collectorName id="3C2C534C5451FFE99D82FD291C5FFD16" box="[933,1066,727,753]" pageId="34" pageNumber="335">S. Bandeira</collectorName>
&amp;
<collectorName id="3C2C534C5451FFE99A74FD291CBBFD16" box="[1107,1230,727,753]" pageId="34" pageNumber="335">I. Agarwal</collectorName>
, on
<date id="E567105A5451FFE99B26FD2918A6FCF2" pageId="34" pageNumber="335" value="2016-11-23">
<collectingDate id="F523E9B25451FFE99B26FD2918A6FCF2" pageId="34" pageNumber="335" value="2016-11-23">23 November 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99EFBFD051C56FCDE" ID-GBIF-Occurrence="3031141305" accessionNumber="AMB 10352" collectingDate="2016-11-23" collectionCode="AMB" collectorName="L. Ceriaco &amp; S. Bandeira &amp; I. Agarwal" country="Angola" elevation="669" location="An" municipality="Montipa" pageId="34" pageNumber="335" specimenCode="CAS 263531" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<location id="940660415451FFE99EFBFD05188BFCF2" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99EFBFD05188BFCF2" box="[220,254,763,789]" country="Angola" municipality="Montipa" name="An" pageId="34" pageNumber="335" stateProvince="Huila">An</location>
adult male (
<specimenCode id="C17F9EE15451FFE99FA1FD051A6EFCF2" box="[390,539,762,789]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="335" type="Museum">CAS 263531</specimenCode>
, field number
<collectionCode id="F7C8AE5F5451FFE99C9EFD051A8CFCF2" box="[697,761,763,789]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10352), collected at
<collectingMunicipality id="7102ACE05451FFE99DC6FD051C35FCF2" box="[993,1088,763,789]" pageId="34" pageNumber="335">Montipa</collectingMunicipality>
[-14.66482º, 13.25763º,
<elevation id="1AF4D1A95451FFE99B71FD0518B8FCDE" metricMagnitude="2" metricUnit="m" metricValue="6.69" pageId="34" pageNumber="335" unit="m" value="669.0">
<quantity id="56219B7F5451FFE99B71FD051DE9FCF2" box="[1366,1436,763,789]" metricMagnitude="2" metricUnit="m" metricValue="6.69" pageId="34" pageNumber="335" unit="m" value="669.0">669 m</quantity>
a.s.l.
</elevation>
], collected by
<collectorName id="3C2C534C5451FFE99F55FCE11999FCDE" box="[370,492,799,825]" pageId="34" pageNumber="335">L. Ceríaco</collectorName>
,
<collectorName id="3C2C534C5451FFE99FD0FCE11A0FFCDE" box="[503,634,799,825]" pageId="34" pageNumber="335">S. Bandeira</collectorName>
&amp;
<collectorName id="3C2C534C5451FFE99CB9FCE11B62FCDE" box="[670,791,799,825]" pageId="34" pageNumber="335">I. Agarwal</collectorName>
, on
<date id="E567105A5451FFE99D61FCE11C56FCDE" box="[838,1059,799,825]" pageId="34" pageNumber="335" value="2016-11-23">
<collectingDate id="F523E9B25451FFE99D61FCE11C56FCDE" box="[838,1059,799,825]" pageId="34" pageNumber="335" value="2016-11-23">23 November 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99A0BFCE11D5AFC66" ID-GBIF-Occurrence="3031141321" accessionNumber="UF 187228, AMB 10165" collectingDate="2017-07-27" collectionCode="UF, AMB, T" collectorName="An &amp; M. Marques &amp; L. Ceriaco &amp; B. Butler &amp; S. Bandeira &amp; M. Heinicke" country="Angola" elevation="1242" location="Bicuar National Park" municipality="Julio" pageId="34" pageNumber="335" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<collectorName id="3C2C534C5451FFE99A0BFCE11C3BFCDE" box="[1068,1102,799,825]" pageId="34" pageNumber="335">An</collectorName>
adult male (
<collectionCode id="F7C8AE5F5451FFE99AFDFCE11C8AFCDE" box="[1242,1279,799,825]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="34" pageNumber="335" type="Museum">UF</collectionCode>
187228, field number
<collectionCode id="F7C8AE5F5451FFE99ED5FCBD1947FCBA" box="[242,306,835,861]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10165), collected at
<location id="940660415451FFE99C3CFCBD1B7CFCBA" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99C3CFCBD1B7CFCBA" box="[539,777,835,861]" country="Angola" municipality="Julio" name="Bicuar National Park" pageId="34" pageNumber="335" stateProvince="Huila">Bicuar National Park</location>
, camp headquarters [-15.10161º, 14.83986º,
<elevation id="1AF4D1A95451FFE99ADAFCBD1DFBFCBA" box="[1277,1422,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.242" pageId="34" pageNumber="335" unit="m" value="1242.0">
<quantity id="56219B7F5451FFE99ADAFCBD1D27FCBA" box="[1277,1362,835,861]" metricMagnitude="3" metricUnit="m" metricValue="1.242" pageId="34" pageNumber="335" unit="m" value="1242.0">1242 m</quantity>
a.s.l.
</elevation>
], collected by
<collectorName id="3C2C534C5451FFE99F0FFC9919C0FC66" box="[296,437,871,897]" pageId="34" pageNumber="335">M. Marques</collectorName>
,
<collectorName id="3C2C534C5451FFE99FE6FC991A4EFC66" box="[449,571,871,897]" pageId="34" pageNumber="335">L. Ceríaco</collectorName>
,
<collectorName id="3C2C534C5451FFE99C61FC991AC5FC66" box="[582,688,871,897]" pageId="34" pageNumber="335">B. Butler</collectorName>
,
<collectorName id="3C2C534C5451FFE99C9BFC991B37FC66" box="[700,834,871,897]" pageId="34" pageNumber="335">S. Bandeira</collectorName>
,
<collectionCode id="F7C8AE5F5451FFE99D69FC991B15FC66" box="[846,864,871,897]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="335">T</collectionCode>
.
<collectingMunicipality id="7102ACE05451FFE99D4CFC991BD4FC66" box="[875,929,871,897]" pageId="34" pageNumber="335">Júlio</collectingMunicipality>
and
<collectorName id="3C2C534C5451FFE99DFDFC991C1FFC66" box="[986,1130,871,897]" pageId="34" pageNumber="335">M. Heinicke</collectorName>
, on
<date id="E567105A5451FFE99ABDFC991D5AFC66" box="[1178,1327,871,897]" pageId="34" pageNumber="335" value="2017-07-27">
<collectingDate id="F523E9B25451FFE99ABDFC991D5AFC66" box="[1178,1327,871,897]" pageId="34" pageNumber="335" value="2017-07-27">27 July 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99B1DFC991CB5FC2E" ID-GBIF-Occurrence="3031141307" accessionNumber="UF 187229, AMB 10790" collectingDate="2017-08-05" collectionCode="UF, AMB, T" collectorName="An &amp; Chibia &amp; M. Marques &amp; L. Ceriaco &amp; B. Butler &amp; S. Bandeira &amp; M. Heinicke" country="Angola" elevation="1467" location="Julio" pageId="34" pageNumber="335" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<collectorName id="3C2C534C5451FFE99B1DFC991D29FC66" box="[1338,1372,871,897]" pageId="34" pageNumber="335">An</collectorName>
adult male (
<collectionCode id="F7C8AE5F5451FFE99EFBFC751974FC42" box="[220,257,907,933]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="34" pageNumber="335" type="Museum">UF</collectionCode>
187229, field number
<collectionCode id="F7C8AE5F5451FFE99C27FC751A35FC42" box="[512,576,907,933]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10790), collected at
<collectorName id="3C2C534C5451FFE99D0BFC751B02FC42" box="[812,887,907,933]" pageId="34" pageNumber="335">Chibia</collectorName>
[-15.18944º, 13.69058º,
<elevation id="1AF4D1A95451FFE99AB7FC751D57FC42" box="[1168,1314,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.4669999999999999" pageId="34" pageNumber="335" unit="m" value="1467.0">
<quantity id="56219B7F5451FFE99AB7FC751C90FC42" box="[1168,1253,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.4669999999999999" pageId="34" pageNumber="335" unit="m" value="1467.0">1467 m</quantity>
a.s.l.
</elevation>
], collected by
<collectorName id="3C2C534C5451FFE99E9DFC511930FC2E" box="[186,325,943,969]" pageId="34" pageNumber="335">M. Marques</collectorName>
,
<collectorName id="3C2C534C5451FFE99F77FC5119BCFC2E" box="[336,457,943,969]" pageId="34" pageNumber="335">L. Ceríaco</collectorName>
,
<collectorName id="3C2C534C5451FFE99FF3FC511A49FC2E" box="[468,572,943,969]" pageId="34" pageNumber="335">B. Butler</collectorName>
,
<collectorName id="3C2C534C5451FFE99C61FC511AB9FC2E" box="[582,716,943,969]" pageId="34" pageNumber="335">S. Bandeira</collectorName>
,
<collectionCode id="F7C8AE5F5451FFE99CF1FC511A9DFC2E" box="[726,744,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="335">T</collectionCode>
.
<location id="940660415451FFE99CD4FC511B5CFC2E" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99CD4FC511B5CFC2E" box="[755,809,943,969]" country="Angola" name="Julio" pageId="34" pageNumber="335" stateProvince="Huila">Júlio</location>
and
<collectorName id="3C2C534C5451FFE99D78FC511B98FC2E" box="[863,1005,943,969]" pageId="34" pageNumber="335">M. Heinicke</collectorName>
, on
<date id="E567105A5451FFE99A3CFC501CB5FC2E" box="[1051,1216,942,969]" pageId="34" pageNumber="335" value="2017-08-05">
<collectingDate id="F523E9B25451FFE99A3CFC501CB5FC2E" box="[1051,1216,942,969]" pageId="34" pageNumber="335" value="2017-08-05">5 August 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99AEDFC511B53FBF6" ID-GBIF-Occurrence="3031141302" accessionNumber="UF 187230, AMB 10477" collectingDate="2016-11-28" collectionCode="UF, AMB" collectorName="L. Ceriaco &amp; S. Bandeira &amp; I. Agarwal" country="Angola" elevation="523" location="An" municipality="Virei" pageId="34" pageNumber="335" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
<location id="940660415451FFE99AEDFC511C99FC2E" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99AEDFC511C99FC2E" box="[1226,1260,943,969]" country="Angola" municipality="Virei" name="An" pageId="34" pageNumber="335" stateProvince="Huila">An</location>
adult male (
<collectionCode id="F7C8AE5F5451FFE99B50FC511DE9FC2E" box="[1399,1436,943,969]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="34" pageNumber="335" type="Museum">UF</collectionCode>
187230, field number
<collectionCode id="F7C8AE5F5451FFE99FB0FC2D19A2FC0A" box="[407,471,979,1005]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="34" pageNumber="335" type="Museum">AMB</collectionCode>
10477), collected at
<collectingMunicipality id="7102ACE05451FFE99CEBFC2D1B71FC0A" box="[716,772,979,1005]" pageId="34" pageNumber="335">Virei</collectingMunicipality>
[-16.11958º, 12.83458524º,
<elevation id="1AF4D1A95451FFE99A69FC2C1CA2FC0A" box="[1102,1239,978,1005]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="34" pageNumber="335" unit="m" value="523.0">
<quantity id="56219B7F5451FFE99A69FC2C1CEDFC0A" box="[1102,1176,978,1005]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="34" pageNumber="335" unit="m" value="523.0">523 m</quantity>
a.s.l.
</elevation>
], collected by
<collectorName id="3C2C534C5451FFE99BA3FC2D1884FBF6" pageId="34" pageNumber="335">L. Ceríaco</collectorName>
,
<collectorName id="3C2C534C5451FFE99EDBFC09190AFBF6" box="[252,383,1015,1041]" pageId="34" pageNumber="335">S. Bandeira</collectorName>
&amp;
<collectorName id="3C2C534C5451FFE99F84FC091A6EFBF6" box="[419,539,1015,1041]" pageId="34" pageNumber="335">I. Agarwal</collectorName>
, on
<date id="E567105A5451FFE99C6DFC091B53FBF6" box="[586,806,1015,1041]" pageId="34" pageNumber="335" value="2016-11-28">
<collectingDate id="F523E9B25451FFE99C6DFC091B53FBF6" box="[586,806,1015,1041]" pageId="34" pageNumber="335" value="2016-11-28">28 November 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="9166369A5451FFE99EE0FBE41CCCF9EE" blockId="34.[151,1437,151,1905]" pageId="34" pageNumber="335">
<emphasis id="A3ADEA885451FFE99EE0FBE419CFFBD3" bold="true" box="[199,442,1050,1076]" pageId="34" pageNumber="335">Additional material.</emphasis>
<materialsCitation id="21B13CC75451FFE99FE7FBE41BD5FBBE" ID-GBIF-Occurrence="3031141312" collectionCode="MHNC-UP" country="Angola" elevation="1240" location="Cassumbi village" pageId="34" pageNumber="335" specimenCode="REP 499" specimenCount="1" stateProvince="Bie">
<emphasis id="A3ADEA885451FFE99FE7FBE41A99FBD3" bold="true" box="[448,748,1050,1077]" pageId="34" pageNumber="335">
<collectingCountry id="E9CE760A5451FFE99FE7FBE41A4AFBD3" box="[448,575,1050,1076]" name="Angola" pageId="34" pageNumber="335">ANGOLA</collectingCountry>
:
<collectingRegion id="531DF8785451FFE99C6BFBE41A9DFBD3" box="[588,744,1050,1077]" country="Angola" name="Bie" pageId="34" pageNumber="335">Bié Province</collectingRegion>
:
</emphasis>
<quantity id="56219B7F5451FFE99CD3FBE51B4EFBD2" box="[756,827,1051,1077]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="34" pageNumber="335" unit="km" value="14.0">14 km</quantity>
by dirt road from
<location id="940660415451FFE99A2CFBE51CA5FBD2" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99A2CFBE51CA5FBD2" box="[1035,1232,1051,1077]" country="Angola" name="Cassumbi village" pageId="34" pageNumber="335" stateProvince="Bie">Cassumbi village</location>
, in a banana plantation [-11.08391º, 16.66367º,
<elevation id="1AF4D1A95451FFE99FD2FBC11AFFFBBE" box="[501,650,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="34" pageNumber="335" unit="m" value="1240.0">
<quantity id="56219B7F5451FFE99FD2FBC11A39FBBE" box="[501,588,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="34" pageNumber="335" unit="m" value="1240.0">1240 m</quantity>
a.s.l.
</elevation>
] (
<collectionCode id="F7C8AE5F5451FFE99C84FBC11B50FBBE" box="[675,805,1087,1113]" pageId="34" pageNumber="335">MHNC-UP</collectionCode>
/
<specimenCode id="C17F9EE15451FFE99D0AFBC11BEDFBBE" box="[813,920,1087,1113]" collectionCode="REP" country="Turkmenistan" lsid="urn:lsid:biocol.org:col:12935" name="Desert Experiment Station of the W.I.R." pageId="34" pageNumber="335" type="Herbarium">REP 499</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="21B13CC75451FFE99D8AFBC0189FFB46" ID-GBIF-Occurrence="3031141322" accessionNumber="UF 187226, AMB 10133" collectionCode="UF, INBAC, AMB" country="Angola" elevation="1866" location="Kimbo do Soba" municipality="Lubango" pageId="34" pageNumber="335" specimenCode="CAS 263417, 263419, 263421, 10137, 10134" specimenCount="1" stateProvince="Huila">
<emphasis id="A3ADEA885451FFE99D8AFBC01C19FBBF" bold="true" box="[941,1132,1086,1113]" pageId="34" pageNumber="335">
<collectingRegion id="531DF8785451FFE99D8AFBC01C12FBBF" box="[941,1127,1086,1113]" country="Angola" name="Huila" pageId="34" pageNumber="335">Huíla Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="7102ACE05451FFE99A52FBC11CAEFBBE" box="[1141,1243,1087,1113]" pageId="34" pageNumber="335">Lubango</collectingMunicipality>
,
<location id="940660415451FFE99ACFFBC11DE9FBBE" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99ACFFBC11DE9FBBE" box="[1256,1436,1087,1113]" country="Angola" municipality="Lubango" name="Kimbo do Soba" pageId="34" pageNumber="335" stateProvince="Huila">Kimbo do Soba</location>
[-14.93419º, 13.46925º,
<quantity id="56219B7F5451FFE99F8AFB9D1A71FB9A" box="[429,516,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="34" pageNumber="335" unit="m" value="1866.0">
<elevation id="1AF4D1A95451FFE99F8AFB9D1A71FB9A" box="[429,516,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="34" pageNumber="335" unit="m" value="1866.0">1866 m</elevation>
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<location id="940660415451FFE99B16FA5918A7FA02" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99B16FA5918A7FA02" country="Angola" name="Mucungu Farm" pageId="34" pageNumber="335" stateProvince="Namibe">Mucungu Farm</location>
[-14.77083º, 12.49231º,
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<quantity id="56219B7F5451FFE99FCBFA351A46FA02" box="[492,563,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="3.3299999999999996" pageId="34" pageNumber="335" unit="m" value="333.0">333 m</quantity>
a.s.l.
</elevation>
] (
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,
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)
</materialsCitation>
;
<materialsCitation id="21B13CC75451FFE99DB4FA351AEBF9EE" ID-GBIF-Occurrence="3031141306" collectionCode="MHNC-UP" country="Angola" elevation="598" location="Otchifengo" municipality="Ouipaca" pageId="34" pageNumber="335" specimenCode="REP 571" specimenCount="1" stateProvince="Namibe">
between
<collectingMunicipality id="7102ACE05451FFE99DDFFA351C20FA02" box="[1016,1109,1483,1509]" pageId="34" pageNumber="335">Ouipaca</collectingMunicipality>
and
<location id="940660415451FFE99AABFA351D7FFA02" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99AABFA351D7FFA02" box="[1164,1290,1483,1509]" country="Angola" municipality="Ouipaca" name="Otchifengo" pageId="34" pageNumber="335" stateProvince="Namibe">Otchifengo</location>
[-16.63526º, 12.9246º,
<elevation id="1AF4D1A95451FFE99F2EFA1019F9F9EE" box="[265,396,1518,1545]" metricMagnitude="2" metricUnit="m" metricValue="5.9799999999999995" pageId="34" pageNumber="335" unit="m" value="598.0">
<quantity id="56219B7F5451FFE99F2EFA101925F9EE" box="[265,336,1518,1545]" metricMagnitude="2" metricUnit="m" metricValue="5.9799999999999995" pageId="34" pageNumber="335" unit="m" value="598.0">598 m</quantity>
a.s.l.
</elevation>
] (
<collectionCode id="F7C8AE5F5451FFE99F85FA111A51F9EE" box="[418,548,1519,1545]" pageId="34" pageNumber="335">MHNC-UP</collectionCode>
/
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)
</materialsCitation>
.
<materialsCitation id="21B13CC75451FFE99C8FFA101CC0F9EE" ID-GBIF-Occurrence="3031141313" box="[680,1205,1518,1545]" collectionCode="MHNC" country="Angola" location="Angola" municipality="Undetermined" pageId="34" pageNumber="335" specimenCode="MHNC 91.0447, 48" specimenCount="1" stateProvince="Namibe">
<emphasis id="A3ADEA885451FFE99C8FFA101B2BF9EF" bold="true" box="[680,862,1518,1544]" pageId="34" pageNumber="335">
<collectingMunicipality id="7102ACE05451FFE99C8FFA101B2CF9EF" box="[680,857,1518,1544]" pageId="34" pageNumber="335">Undetermined</collectingMunicipality>
:
</emphasis>
<location id="940660415451FFE99D42FA111BC2F9EE" LSID="urn:lsid:plazi:treatment:1970878C5453FFEE9EB0FD521C06F97E:940660415451FFE99D42FA111BC2F9EE" box="[869,951,1519,1545]" country="Angola" municipality="Undetermined" name="Angola" pageId="34" pageNumber="335" stateProvince="Namibe">Angola</location>
(
<specimenCode id="C17F9EE15451FFE99DE1FA111CF7F9EE" box="[966,1154,1519,1545]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="34" pageNumber="335">MHNC 91.0447</specimenCode>
<specimenCode id="C17F9EE15451FFE99AB6FA111CD8F9EE" box="[1169,1197,1519,1545]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="34" pageNumber="335">48</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C365115451FFEF9EE0F9EC19C2F899" lastPageId="36" lastPageNumber="337" pageId="34" pageNumber="335" type="diagnosis">
<paragraph id="9166369A5451FFE99EE0F9EC19A5F896" blockId="34.[151,1437,151,1905]" pageId="34" pageNumber="335">
<emphasis id="A3ADEA885451FFE99EE0F9EC1937F9CB" bold="true" box="[199,322,1554,1580]" pageId="34" pageNumber="335">Diagnosis.</emphasis>
<taxonomicName id="56D94D195451FFE99F6EF9ED1A36F9CA" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[329,579,1555,1581]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="335" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885451FFE99F6EF9ED1A36F9CA" box="[329,579,1555,1581]" italics="true" pageId="34" pageNumber="335">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885451FFE99C6BF9ED1AD3F9CA" bold="true" box="[588,678,1555,1581]" pageId="34" pageNumber="335">
<taxonomicNameLabel id="B89E57F35451FFE99C6BF9ED1AD3F9CA" box="[588,678,1555,1581]" pageId="34" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a small dwarf gecko with a maximum SVL of
<quantity id="56219B7F5451FFE99AFCF9ED1D5BF9CA" box="[1243,1326,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="34" pageNumber="335" unit="mm" value="34.0">34 mm</quantity>
. It can be distinguished from other members of the genus occurring in
<collectingCountry id="E9CE760A5451FFE99D72F9C91BD2F9B6" box="[853,935,1591,1617]" name="Angola" pageId="34" pageNumber="335">Angola</collectingCountry>
and surroundings region by having 1) four scales surrounding the nostril (1
<superScript id="66AC9BD25451FFE99C2FF9A51A66F98E" attach="left" box="[520,531,1627,1641]" fontSize="6" pageId="34" pageNumber="335">st</superScript>
supralabial, two supranasal and one postnasals); 2) nostrils separated from the rostral and pierced above the suture between rostral and first supralabial; 3) mental followed by three postmentals; 4) first infralabial with 2550% overlap of postpostmental; 5) 16 to 17 rows of ventral scales at midbody; 6) 46 precloacal pores in males; 7) median subcaudals arranged in a pattern of one row of scales about ¼ width of the tail alternating with two rows of scales about ½ width of the tail (
<figureCitation id="09E22A1F5451FFE99D1FF9151BFAF8E2" box="[824,911,1771,1797]" captionStart="FIGURE 4" captionStartId="22.[151,250,1008,1032]" captionTargetBox="[151,1437,181,985]" captionTargetId="figure-22@22.[151,1437,181,985]" captionTargetPageId="22" captionText="FIGURE 4. Comparison of the tail scales arrangement in Angolan individuals of A. Lygodactylus angolensis (MHNC-UP/REP 533); B. L. nyaneka sp. nov. (CAS 263531); C. L. baptistai sp. nov. (CAS 263557); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312d)." figureDoi="http://doi.org/10.5281/zenodo.4410735" httpUri="https://zenodo.org/record/4410735/files/figure.png" pageId="34" pageNumber="335">Fig. 4B</figureCitation>
); 8) venter white; 9) gular region usually white, but sometimes speckled with minute black dots. Background usually grey to brown with a pair of black lines on the flanks bordering a pale stripe (not always visible) with a line of cream dots from the posterior part of the orbit to the end of the tail (
<figureCitation id="09E22A1F5451FFE99F66F8A919B1F896" box="[321,452,1878,1905]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="35.[151,250,1008,1032]" captionStartId-1="35.[151,250,1973,1997]" captionTargetBox-0="[151,1437,181,985]" captionTargetBox-1="[151,1436,1093,1949]" captionTargetId-0="figure-19@35.[151,1437,181,985]" captionTargetId-1="figure-52@35.[151,1436,1093,1949]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionText-0="FIGURE 14. Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of Lygodactylus nyaneka sp. nov. (CAS 263418)." captionText-1="FIGURE 15. Lygodactylus nyaneka sp. nov. in life (CAS 263531). Montipa [-14.66482º, 13.25763º, 670 m a.s.l.]. Namibe Province. southwestern Angola Photos by Ishan Agarwal." figureDoi-0="http://doi.org/10.5281/zenodo.4410755" figureDoi-1="http://doi.org/10.5281/zenodo.4410757" httpUri-0="https://zenodo.org/record/4410755/files/figure.png" httpUri-1="https://zenodo.org/record/4410757/files/figure.png" pageId="34" pageNumber="335">Figs 1415</figureCitation>
).
</paragraph>
<caption id="C5A666125450FFE89EB0FC0E1A6EFBCB" ID-DOI="http://doi.org/10.5281/zenodo.4410755" ID-Zenodo-Dep="4410755" httpUri="https://zenodo.org/record/4410755/files/figure.png" pageId="35" pageNumber="336" startId="35.[151,250,1008,1032]" targetBox="[151,1437,181,985]" targetPageId="35">
<paragraph id="9166369A5450FFE89EB0FC0E1A6EFBCB" blockId="35.[151,1436,1008,1069]" pageId="35" pageNumber="336">
<emphasis id="A3ADEA885450FFE89EB0FC0E1954FBEF" bold="true" box="[151,289,1008,1032]" pageId="35" pageNumber="336">FIGURE 14.</emphasis>
Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of
<taxonomicName id="56D94D195450FFE89B1AFC0F1954FBCA" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="336" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885450FFE89B1AFC0F1954FBCA" italics="true" pageId="35" pageNumber="336">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885450FFE89F0FFBEB190CFBCA" bold="true" box="[296,377,1045,1069]" pageId="35" pageNumber="336">
<taxonomicNameLabel id="B89E57F35450FFE89F0FFBEB190CFBCA" box="[296,377,1045,1069]" pageId="35" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CAS 263418).
</paragraph>
</caption>
<caption id="C5A666125450FFE89EB0F84B1AAAF816" ID-DOI="http://doi.org/10.5281/zenodo.4410757" ID-Zenodo-Dep="4410757" httpUri="https://zenodo.org/record/4410757/files/figure.png" pageId="35" pageNumber="336" startId="35.[151,250,1973,1997]" targetBox="[151,1436,1093,1949]" targetPageId="35">
<paragraph id="9166369A5450FFE89EB0F84B1AAAF816" blockId="35.[151,1436,1973,2033]" pageId="35" pageNumber="336">
<emphasis id="A3ADEA885450FFE89EB0F84B1957F82A" bold="true" box="[151,290,1973,1997]" pageId="35" pageNumber="336">FIGURE 15.</emphasis>
<taxonomicName id="56D94D195450FFE89F0CF84B1A7BF82A" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[299,526,1973,1997]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="336" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885450FFE89F0CF84B1A7BF82A" box="[299,526,1973,1997]" italics="true" pageId="35" pageNumber="336">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885450FFE89C30F84B1A1FF82A" bold="true" box="[535,618,1973,1997]" pageId="35" pageNumber="336">
<taxonomicNameLabel id="B89E57F35450FFE89C30F84B1A1FF82A" box="[535,618,1973,1997]" pageId="35" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (CAS 263531). Montipa [-14.66482º, 13.25763º, 670 m a.s.l.]. Namibe Province. southwestern Angola Photos by Ishan Agarwal.
</paragraph>
</caption>
<caption id="C5A666125457FFEF9EB0FA6F1883FA29" ID-DOI="http://doi.org/10.5281/zenodo.4410759" ID-Zenodo-Dep="4410759" httpUri="https://zenodo.org/record/4410759/files/figure.png" pageId="36" pageNumber="337" startId="36.[151,250,1425,1449]" targetBox="[173,1414,191,1395]" targetPageId="36">
<paragraph id="9166369A5457FFEF9EB0FA6F1883FA29" blockId="36.[151,1437,1425,1486]" pageId="36" pageNumber="337">
<emphasis id="A3ADEA885457FFEF9EB0FA6F1957FA4E" bold="true" box="[151,290,1425,1449]" pageId="36" pageNumber="337">FIGURE 16.</emphasis>
Distribution of
<taxonomicName id="56D94D195457FFEF9FEDFA6C1AD8FA4D" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[458,685,1426,1450]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9FEDFA6C1AD8FA4D" box="[458,685,1426,1450]" italics="true" pageId="36" pageNumber="337">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9C91FA6C1B7CFA4D" bold="true" box="[694,777,1426,1450]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9C91FA6C1B7CFA4D" box="[694,777,1426,1450]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the country. Blue circle represents records from Epupa Falls. Namibia.
</paragraph>
</caption>
<paragraph id="9166369A5457FFEF9EE0F9DE19C2F899" blockId="36.[151,1437,1567,2026]" pageId="36" pageNumber="337">
<emphasis id="A3ADEA885457FFEF9EE0F9DE1C65F9DD" bold="true" box="[199,1040,1567,1594]" pageId="36" pageNumber="337">
Comparison with other South and Southwestern African
<taxonomicName id="56D94D195457FFEF9D55F9E11C7CF9DD" box="[882,1033,1567,1594]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="A3ADEA885457FFEF9D55F9E11C7CF9DD" bold="true" box="[882,1033,1567,1594]" italics="true" pageId="36" pageNumber="337">Lygodactylus</emphasis>
</taxonomicName>
.
</emphasis>
As the molecular data provide evidence of the distinctiveness of
<taxonomicName id="56D94D195457FFEF9FDBF9BA1A83F9B9" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[508,758,1604,1630]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="nyaneka">
<emphasis id="A3ADEA885457FFEF9FDBF9BA1A83F9B9" box="[508,758,1604,1630]" italics="true" pageId="36" pageNumber="337">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9D27F9BA1B54F9B9" bold="true" box="[768,801,1604,1630]" pageId="36" pageNumber="337">sp.</emphasis>
from all other taxa, we here restrict our morphological comparisons to those named congeners occurring in
<collectingCountry id="E9CE760A5457FFEF9CCEF9961B4EF965" box="[745,827,1640,1666]" name="Angola" pageId="36" pageNumber="337">Angola</collectingCountry>
and surrounding countries.
<taxonomicName id="56D94D195457FFEF9A5EF9961D07F965" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[1145,1394,1640,1666]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9A5EF9961D07F965" box="[1145,1394,1640,1666]" italics="true" pageId="36" pageNumber="337">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9B5CF99618BDF941" bold="true" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9B5CF99618BDF941" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56D94D195457FFEF9F7CF9731984F941" box="[347,497,1676,1702]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="A3ADEA885457FFEF9F7CF9731984F941" box="[347,497,1676,1702]" italics="true" pageId="36" pageNumber="337">L. angolensis</emphasis>
</taxonomicName>
,
<emphasis id="A3ADEA885457FFEF9FD9F9731B5FF941" box="[510,810,1676,1702]" italics="true" pageId="36" pageNumber="337">
<taxonomicName id="56D94D195457FFEF9FD9F9731AE1F941" box="[510,660,1676,1702]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="chobiensis">L. chobiensis</taxonomicName>
,
<taxonomicName id="56D94D195457FFEF9CB8F9731B5FF941" box="[671,810,1676,1702]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="lawrencei">L. lawrencei</taxonomicName>
</emphasis>
and
<taxonomicName id="56D94D195457FFEF9D47F9731BACF941" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[864,985,1676,1702]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="tchokwe" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9D47F9731BACF941" box="[864,985,1676,1702]" italics="true" pageId="36" pageNumber="337">L. tchokwe</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9DC7F9721C4CF941" bold="true" box="[992,1081,1676,1702]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9DC7F9721C4CF941" box="[992,1081,1676,1702]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having 46 precloacal pores (
<emphasis id="A3ADEA885457FFEF9E86F94F189CF92D" box="[161,233,1713,1738]" italics="true" pageId="36" pageNumber="337">versus</emphasis>
<quantity id="56219B7F5457FFEF9EC8F94E1930F92D" box="[239,325,1712,1738]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.54" metricValueMin="1.778" pageId="36" pageNumber="337" unit="in" value="8.5" valueMax="10.0" valueMin="7.0">
7
<emphasis id="A3ADEA885457FFEF9EDAF94F197EF92D" box="[253,267,1713,1738]" italics="true" pageId="36" pageNumber="337"></emphasis>
10 in
</quantity>
<taxonomicName id="56D94D195457FFEF9F6BF94F19A2F92D" box="[332,471,1712,1738]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="angolensi">
<emphasis id="A3ADEA885457FFEF9F6BF94F19A2F92D" box="[332,471,1712,1738]" italics="true" pageId="36" pageNumber="337">L. angolensi</emphasis>
</taxonomicName>
s;
<quantity id="56219B7F5457FFEF9FD6F94E1A30F92D" box="[497,581,1712,1738]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.794" metricValueMin="1.778" pageId="36" pageNumber="337" unit="in" value="9.0" valueMax="11.0" valueMin="7.0">
7
<emphasis id="A3ADEA885457FFEF9FD8F94F1A78F92D" box="[511,525,1713,1738]" italics="true" pageId="36" pageNumber="337"></emphasis>
11 in
</quantity>
<taxonomicName id="56D94D195457FFEF9C6AF94F1A95F92D" box="[589,736,1712,1738]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="chobiensis">
<emphasis id="A3ADEA885457FFEF9C6AF94F1A95F92D" box="[589,736,1712,1738]" italics="true" pageId="36" pageNumber="337">L. chobiensis</emphasis>
</taxonomicName>
; two in
<taxonomicName id="56D94D195457FFEF9D19F94F1BBCF92D" box="[830,969,1712,1738]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="lawrencei">
<emphasis id="A3ADEA885457FFEF9D19F94F1BBCF92D" box="[830,969,1712,1738]" italics="true" pageId="36" pageNumber="337">L. lawrencei</emphasis>
</taxonomicName>
; two to four in
<taxonomicName id="56D94D195457FFEF9A5CF94F1C80F92D" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[1147,1269,1712,1738]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="tchokwe" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9A5CF94F1C80F92D" box="[1147,1269,1712,1738]" italics="true" pageId="36" pageNumber="337">L. tchokwe</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9ADBF94E1D23F92D" bold="true" box="[1276,1366,1712,1738]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9ADBF94E1D23F92D" box="[1276,1366,1712,1738]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="09E22A1F5457FFEF9B48F94E18DDF909" captionStart="FIGURE 2" captionStartId="20.[151,250,1754,1778]" captionTargetBox="[151,1436,200,1730]" captionTargetId="figure-22@20.[151,1436,200,1730]" captionTargetPageId="20" captionText="FIGURE 2. Comparison of the number of precloacal pores in Angolan individuals of Lygodactylus angolensis, L. baptistai sp. nov., L. nyaneka sp. nov., L. lawrencei and L. tchokwe sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4410731" httpUri="https://zenodo.org/record/4410731/files/figure.png" pageId="36" pageNumber="337">Fig. 2</figureCitation>
). The new species can be distinguished from
<taxonomicName id="56D94D195457FFEF9C80F92B1B52F909" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[679,807,1748,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="baptistai" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9C80F92B1B52F909" box="[679,807,1748,1774]" italics="true" pageId="36" pageNumber="337">L. baptistai</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9D09F92A1BFDF909" bold="true" box="[814,904,1748,1774]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9D09F92A1BFDF909" box="[814,904,1748,1774]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="56D94D195457FFEF9DB1F92B1C57F909" box="[918,1058,1748,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="bradfieldi">
<emphasis id="A3ADEA885457FFEF9DB1F92B1C57F909" box="[918,1058,1748,1774]" italics="true" pageId="36" pageNumber="337">L. bradfieldi</emphasis>
</taxonomicName>
and
<taxonomicName id="56D94D195457FFEF9A7EF92B1CA2F909" box="[1113,1239,1749,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885457FFEF9A7EF92B1CA2F909" box="[1113,1239,1749,1774]" italics="true" pageId="36" pageNumber="337">L. capensis</emphasis>
</taxonomicName>
in not having the rostral in contact with the nostrils. It can be further distinguished from nominotypical
<taxonomicName id="56D94D195457FFEF9A43F9071C96F8F5" box="[1124,1251,1785,1810]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885457FFEF9A43F9071C96F8F5" box="[1124,1251,1785,1810]" italics="true" pageId="36" pageNumber="337">L. capensis</emphasis>
</taxonomicName>
by its staggered rows of subcaudal plates in original tails (
<emphasis id="A3ADEA885457FFEF9C43F8E31AD9F8D1" box="[612,684,1821,1846]" italics="true" pageId="36" pageNumber="337">versus</emphasis>
a more linear distribution of the rows in
<taxonomicName id="56D94D195457FFEF9A4AF8E31C9FF8D1" box="[1133,1258,1821,1846]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="capensis">
<emphasis id="A3ADEA885457FFEF9A4AF8E31C9FF8D1" box="[1133,1258,1821,1846]" italics="true" pageId="36" pageNumber="337">L. capensis</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="56D94D195457FFEF9EB0F8BF195CF8BD" box="[151,297,1856,1882]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="chobiensis">
<emphasis id="A3ADEA885457FFEF9EB0F8BF195CF8BD" box="[151,297,1856,1882]" italics="true" pageId="36" pageNumber="337">L. chobiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="56D94D195457FFEF9F7AF8BF19A9F8BD" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[349,476,1856,1882]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="baptistai" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9F7AF8BF19A9F8BD" box="[349,476,1856,1882]" italics="true" pageId="36" pageNumber="337">L. baptistai</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9FC6F8BE1A4FF8BD" bold="true" box="[481,570,1856,1882]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9FC6F8BE1A4FF8BD" box="[481,570,1856,1882]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by its white ventral and gular coloration (
<emphasis id="A3ADEA885457FFEF9A20F8BF1C3AF8BD" box="[1031,1103,1857,1882]" italics="true" pageId="36" pageNumber="337">versus</emphasis>
bright yellow in
<taxonomicName id="56D94D195457FFEF9B2DF8BF1DE9F8BD" box="[1290,1436,1856,1882]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="chobiensis">
<emphasis id="A3ADEA885457FFEF9B2DF8BF1DE9F8BD" box="[1290,1436,1856,1882]" italics="true" pageId="36" pageNumber="337">L. chobiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="56D94D195457FFEF9EE0F89B1933F899" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[199,326,1892,1918]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="337" phylum="Chordata" rank="species" species="baptistai" status="sp. nov.">
<emphasis id="A3ADEA885457FFEF9EE0F89B1933F899" box="[199,326,1892,1918]" italics="true" pageId="36" pageNumber="337">L. baptistai</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885457FFEF9F6AF89A19D2F899" bold="true" box="[333,423,1892,1918]" pageId="36" pageNumber="337">
<taxonomicNameLabel id="B89E57F35457FFEF9F6AF89A19D2F899" box="[333,423,1892,1918]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="D9C365115457FFEE9EE0F8761A17FB0E" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337" type="description">
<paragraph id="9166369A5457FFEE9EE0F8761B62FCDE" blockId="36.[151,1437,1567,2026]" lastBlockId="37.[151,1437,151,1689]" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337">
<emphasis id="A3ADEA885457FFEF9EE0F8761992F845" bold="true" box="[199,487,1928,1954]" pageId="36" pageNumber="337">
Description of
<typeStatus id="4E6288385457FFEF9F5EF8761997F845" box="[377,482,1928,1954]" pageId="36" pageNumber="337" type="holotype">holotype</typeStatus>
.
</emphasis>
SVL
<quantity id="56219B7F5457FFEF9C0DF8761AE7F845" box="[554,658,1928,1954]" metricMagnitude="-2" metricUnit="m" metricValue="3.41" pageId="36" pageNumber="337" unit="mm" value="34.1">34.1 mm</quantity>
; TrunkL:
<quantity id="56219B7F5457FFEF9D25F8761B1FF845" box="[770,874,1928,1954]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="36" pageNumber="337" unit="mm" value="16.7">16.7 mm</quantity>
; CrusL
<quantity id="56219B7F5457FFEF9DE4F8761C68F845" box="[963,1053,1928,1955]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="36" pageNumber="337" unit="mm" value="5.3">5.3 mm</quantity>
; TL:
<quantity id="56219B7F5457FFEF9A7CF8761CB6F845" box="[1115,1219,1928,1954]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="36" pageNumber="337" unit="mm" value="27.2">27.2 mm</quantity>
; TW:
<quantity id="56219B7F5457FFEF9B2EF8761D16F845" box="[1289,1379,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="36" pageNumber="337" unit="mm" value="3.1">3.1 mm</quantity>
; HL:
<quantity id="56219B7F5457FFEF9EB0F8521885F821" box="[151,240,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="36" pageNumber="337" unit="mm" value="8.4">8.4 mm</quantity>
; HW:
<quantity id="56219B7F5457FFEF9F1EF85219E7F821" box="[313,402,1964,1991]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="36" pageNumber="337" unit="mm" value="5.7">5.7 mm</quantity>
;
<collectingRegion id="531DF8785457FFEF9FB9F85219BEF821" box="[414,459,1964,1990]" country="Swaziland" name="Hhohho" pageId="36" pageNumber="337">HH</collectingRegion>
:
<quantity id="56219B7F5457FFEF9FF2F8521A5BF821" box="[469,558,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="36" pageNumber="337" unit="mm" value="3.8">3.8 mm</quantity>
; EarL:
<quantity id="56219B7F5457FFEF9CA6F8521AAFF821" box="[641,730,1964,1991]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="337" unit="mm" value="0.5">0.5 mm</quantity>
; ForeaL:
<quantity id="56219B7F5457FFEF9D61F8521BEAF821" box="[838,927,1964,1991]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="36" pageNumber="337" unit="mm" value="4.5">4.5 mm</quantity>
; ED:
<quantity id="56219B7F5457FFEF9DF8F8521C4CF821" box="[991,1081,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="36" pageNumber="337" unit="mm" value="1.8">1.8 mm</quantity>
; NE:
<quantity id="56219B7F5457FFEF9A5EF8521CA7F821" box="[1145,1234,1964,1991]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="337" unit="mm" value="2.5">2.5 mm</quantity>
; SE:
<quantity id="56219B7F5457FFEF9B2AF8521D12F821" box="[1293,1383,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="36" pageNumber="337" unit="mm" value="3.9">3.9 mm</quantity>
; EE:
<quantity id="56219B7F5457FFEF9EB0F82E1885F80D" box="[151,240,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="36" pageNumber="337" unit="mm" value="2.6">2.6 mm</quantity>
; IN:
<quantity id="56219B7F5457FFEF9F01F82E190AF80D" box="[294,383,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="36" pageNumber="337" unit="mm" value="1.1">1.1 mm</quantity>
; IO:
<quantity id="56219B7F5457FFEF9F91F82E1A7BF80D" box="[438,526,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="36" pageNumber="337" unit="mm" value="3.1">3.1 mm</quantity>
. Head broad and slightly distinct from neck; snout longer than broad, the distance from its edge to the anterior border of the eye greater than the interorbital distance anteriorly, and greater than the distance between the eye and ear opening; ear opening medium and oval; snout covered with enlarged homogeneous flattened granular scales, becoming smaller laterally above the eye; scales on the snout slightly larger than those on the occiput; prominent canthus rostralis; rostral pentagonal, wider than high; four scales surrounding the nostril (1
<superScript id="66AC9BD25456FFEE9E88FED918CFFED2" attach="left" box="[175,186,295,309]" fontSize="6" pageId="37" pageNumber="338">st</superScript>
supralabial, two supranasal and one postnasals); nostrils separated from the rostral and pierced above the suture between rostral and first supralabial (
<figureCitation id="09E22A1F5456FFEE9C1FFEB51A0EFE82" box="[568,635,331,357]" captionStart="FIGURE 3" captionStartId="21.[151,248,1463,1487]" captionTargetBox="[161,1426,190,1426]" captionTargetId="figure-19@21.[151,1436,181,1440]" captionTargetPageId="21" captionText="FIGURE 3. Comparison of the nostrils position inAngolan individuals of:A. L. angolensis (AMB 9216); B. Lygodactylus baptistai sp. nov. (CAS 263557); C. L. nyaneka sp. nov. (UF 187229); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312b)." figureDoi="http://doi.org/10.5281/zenodo.4410733" httpUri="https://zenodo.org/record/4410733/files/figure.png" pageId="37" pageNumber="338">Fig. 3</figureCitation>
); mental pentagonal, longer than broad, followed by three postmentals; scales behind postmentals intermediate in size, decreasing towards base of the throat and increasing on chest and belly; supralabials 8; infralabials 7; first infralabial with 25% to 50% contact with postpostmental; gulars smooth, enlarged and forming transverse rows posteriorly, 26 between posterior extent of the infralabials/jaws; 16 rows of ventral scales at midbody; limbs short but well developed, extremities pentadactyl; 1
<superScript id="66AC9BD25456FFEE9A76FE251C29FE0E" attach="left" box="[1105,1116,475,489]" fontSize="6" pageId="37" pageNumber="338">st</superScript>
digit of both manus and pes extremely reduced and lacking claw, distal portions of remaining digits expanded, bearing claw; 5 pair of lamellae under 4
<superScript id="66AC9BD25456FFEE9ECBFDDC188CFDD7" attach="left" box="[236,249,546,560]" fontSize="6" pageId="37" pageNumber="338">th</superScript>
of digit manus and 5 pair of lamellae beneath 4
<superScript id="66AC9BD25456FFEE9D34FDDC1B55FDD7" attach="left" box="[787,800,546,560]" fontSize="6" pageId="37" pageNumber="338">th</superScript>
of digit pes; dorsum with small, homogeneous, granular and keeled scales with a similar size to those on trunk, the scales on limbs are slightly larger; venter with enlarged, smooth, homogeneous scales; scales on ventral surface of the fore- and hindlimbs are smaller to those on venter, as well as the surface scales of thighs that are slightly smaller and imbricate; precloacal pores 6 (
<figureCitation id="09E22A1F5456FFEE9AC6FD711D53FD4E" box="[1249,1318,655,681]" captionStart="FIGURE 2" captionStartId="20.[151,250,1754,1778]" captionTargetBox="[151,1436,200,1730]" captionTargetId="figure-22@20.[151,1436,200,1730]" captionTargetPageId="20" captionText="FIGURE 2. Comparison of the number of precloacal pores in Angolan individuals of Lygodactylus angolensis, L. baptistai sp. nov., L. nyaneka sp. nov., L. lawrencei and L. tchokwe sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4410731" httpUri="https://zenodo.org/record/4410731/files/figure.png" pageId="37" pageNumber="338">Fig. 2</figureCitation>
); approximately 18-19 enlarged generation glands present along the ventral surface of each thigh; tail cylindrical; posterior half regenerated; dorsal caudal scales enlarged, flattened and homogeneous; keeled, spinose and acuminate on the lateral surface of the tail; median subcaudals scales flattened and smooth, one row of scales about ¼ width of tail alternating with two rows of scales about ½ width of tail.
</paragraph>
<paragraph id="9166369A5456FFEE9EE0FCBC1981FBBE" blockId="37.[151,1437,151,1689]" pageId="37" pageNumber="338">
<emphasis id="A3ADEA885456FFEE9EE0FCBC19BAFCBA" bold="true" box="[199,463,834,861]" pageId="37" pageNumber="338">Coloration in ethanol.</emphasis>
The coloration of
<taxonomicName id="56D94D195456FFEE9C84FCBD1BE9FCBA" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[675,924,835,861]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885456FFEE9C84FCBD1BE9FCBA" box="[675,924,835,861]" italics="true" pageId="37" pageNumber="338">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885456FFEE9D83FCBD1B8AFCBA" bold="true" box="[932,1023,835,861]" pageId="37" pageNumber="338">
<taxonomicNameLabel id="B89E57F35456FFEE9D83FCBD1B8AFCBA" box="[932,1023,835,861]" pageId="37" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is brown to ashy-grey above, with a pair of black lines on the flanks from snout to the end of the tail, bordering a pale stripe with a line of cream dots that continues to the tail. Mid-dorsal region with small scattered cream dots. The head slightly darker than body and limbs, bearing a speckled pattern with some lighter irregular dots. The venter is white with no markings or speckling, although lateral margins of the belly and ventral surface of the limbs are speckled. The chin and gular regions white with no markings or speckling. Dorsal surface of the limbs with scattered cream dots. Tail brown with a pattern of two parallel cream dots, running on across the dorsal surface up to the regenerated part, ventrally white with no markings or speckling.
</paragraph>
<paragraph id="9166369A5456FFEE9EE0FB9C1A17FB0E" blockId="37.[151,1437,151,1689]" pageId="37" pageNumber="338">
<emphasis id="A3ADEA885456FFEE9EE0FB9C194AFB9B" bold="true" box="[199,319,1122,1148]" pageId="37" pageNumber="338">Variation.</emphasis>
<materialsCitation id="21B13CC75456FFEE9F6FFB9D1A2BFB0E" ID-GBIF-Occurrence="3031141301" accessionNumber="UF 187228" collectionCode="UF" pageId="37" pageNumber="338" specimenCode="CAS 263529" specimenCount="3" typeStatus="paratype">
Variation in scalation and body measurements of the type series is reported in
<tableCitation id="DC5B03215456FFEE9AECFB9D1D57FB9A" box="[1227,1314,1123,1149]" captionStart="TABLE 6" captionStartId="33.[151,239,332,356]" captionText="TABLE 6. Measurements and meristic data of the type series of Lygodactylus nyaneka sp. nov. Abbreviations are those described in Materials and Methods section." pageId="37" pageNumber="338">Table 6</tableCitation>
.
<typeStatus id="4E6288385456FFEE9B08FB9D1DE9FB9A" box="[1327,1436,1123,1149]" pageId="37" pageNumber="338" type="paratype">Paratypes</typeStatus>
agree almost entirely with the
<typeStatus id="4E6288385456FFEE9FC9FB791A26FB46" box="[494,595,1159,1185]" pageId="37" pageNumber="338" type="holotype">holotype</typeStatus>
, with the exception of two male
<typeStatus id="4E6288385456FFEE9DE3FB791C5AFB46" box="[964,1071,1159,1185]" pageId="37" pageNumber="338" type="paratype">paratypes</typeStatus>
(
<specimenCode id="C17F9EE15456FFEE9A18FB791CA2FB46" box="[1087,1239,1158,1185]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="338" type="Museum">CAS 263529</specimenCode>
;
<collectionCode id="F7C8AE5F5456FFEE9AC5FB791D73FB46" box="[1250,1286,1159,1185]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="37" pageNumber="338" type="Museum">UF</collectionCode>
187228) that differ slightly in coloration, with both specimens having a darker dorsum with a more faded pattern, and speckled black on the gular region (
<specimenCode id="C17F9EE15456FFEE9F99FB311A23FB0E" box="[446,598,1230,1257]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="338" type="Museum">CAS 263529</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C365115456FFEE9EE0FB0C1CA2F9CA" pageId="37" pageNumber="338" type="distribution">
<paragraph id="9166369A5456FFEE9EE0FB0C1962FA26" blockId="37.[151,1437,151,1689]" pageId="37" pageNumber="338">
<emphasis id="A3ADEA885456FFEE9EE0FB0C1914FAEB" bold="true" box="[199,353,1266,1292]" pageId="37" pageNumber="338">Distribution.</emphasis>
In
<collectingCountry id="E9CE760A5456FFEE9FA0FB0D19ACFAEA" box="[391,473,1267,1293]" name="Angola" pageId="37" pageNumber="338">Angola</collectingCountry>
the species is known from the central and southwestern regions of the country, from the lower areas of
<collectingRegion id="531DF8785456FFEE9F48FAE919ACFAD6" box="[367,473,1303,1329]" country="Angola" name="Benguela" pageId="37" pageNumber="338">Benguela</collectingRegion>
to
<collectingRegion id="531DF8785456FFEE9C26FAE91A28FAD6" box="[513,605,1303,1329]" country="Angola" name="Namibe" pageId="37" pageNumber="338">Namibe</collectingRegion>
,
<collectingRegion id="531DF8785456FFEE9C4DFAE91ADDFAD6" box="[618,680,1303,1329]" country="Angola" name="Huila" pageId="37" pageNumber="338">Huíla</collectingRegion>
and
<collectingRegion id="531DF8785456FFEE9CC3FAE91B7EFAD6" box="[740,779,1303,1329]" country="Angola" name="Bie" pageId="37" pageNumber="338">Bié</collectingRegion>
provinces, and potentially to Cuando Cubango Province. Based on geography, the single specimen identified by
<bibRefCitation id="F5484B6B5456FFEE9CD8FAC51B9EFAB2" author="Baptista, N. L. &amp; Antonio, T. &amp; Branch, W. R." box="[767,1003,1339,1365]" pageId="37" pageNumber="338" pagination="203" refId="ref25219" refString="Baptista, N. L., Antonio, T. &amp; Branch, W. R. (2019) The herpetofauna of Bicuar National Park and surroundings southwestern Angola: a preliminary checklist. Amphibian &amp; Reptile Conservation, 13 (2), e 203, 96 - 130. [Special Section]" type="journal article" year="2019">
Baptista
<emphasis id="A3ADEA885456FFEE9D46FAC51BECFAB2" box="[865,921,1339,1365]" italics="true" pageId="37" pageNumber="338">et al.</emphasis>
(2019)
</bibRefCitation>
from Bicuar National Park main camp as
<taxonomicName id="56D94D195456FFEE9E9FFAA119B8FA9E" box="[184,461,1375,1401]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="A3ADEA885456FFEE9E9FFAA119B8FA9E" box="[184,461,1375,1401]" italics="true" pageId="37" pageNumber="338">Lygodactylus angolensis</emphasis>
</taxonomicName>
most likely represent a misidentification of
<taxonomicName id="56D94D195456FFEE9DECFAA11CB3FA9E" authority="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer, 2020" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[971,1222,1375,1401]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="nyaneka" status="sp. nov.">
<emphasis id="A3ADEA885456FFEE9DECFAA11CB3FA9E" box="[971,1222,1375,1401]" italics="true" pageId="37" pageNumber="338">Lygodactylus nyaneka</emphasis>
</taxonomicName>
<emphasis id="A3ADEA885456FFEE9AE8FAA11D5EFA9E" bold="true" box="[1231,1323,1375,1401]" pageId="37" pageNumber="338">
<taxonomicNameLabel id="B89E57F35456FFEE9AE8FAA11D5EFA9E" box="[1231,1323,1375,1401]" pageId="37" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In neighboring
<collectingCountry id="E9CE760A5456FFEE9ECEFA7D1939FA7A" box="[233,332,1411,1437]" name="Namibia" pageId="37" pageNumber="338">Namibia</collectingCountry>
, the species is known with certainty from Epupa Falls (
<figureCitation id="09E22A1F5456FFEE9DE2FA7D1C6DFA79" box="[965,1048,1411,1438]" captionStart="FIGURE 16" captionStartId="36.[151,250,1425,1449]" captionTargetBox="[173,1414,191,1395]" captionTargetId="figure-22@36.[164,1423,181,1402]" captionTargetPageId="36" captionText="FIGURE 16. Distribution of Lygodactylus nyaneka sp. nov.in the country. Blue circle represents records from Epupa Falls. Namibia." figureDoi="http://doi.org/10.5281/zenodo.4410759" httpUri="https://zenodo.org/record/4410759/files/figure.png" pageId="37" pageNumber="338">Fig. 16</figureCitation>
) but is probably more extensively distributed.
</paragraph>
<paragraph id="9166369A5456FFEE9EE0FA341CA2F9CA" blockId="37.[151,1437,151,1689]" pageId="37" pageNumber="338">
<emphasis id="A3ADEA885456FFEE9EE0FA341A16FA02" bold="true" box="[199,611,1482,1509]" pageId="37" pageNumber="338">Habitat and natural history notes.</emphasis>
The species is usually found basking during daytime on rocky outcrops, trees, logs or human infrastructures, such as fences, houses and poles (
<figureCitation id="09E22A1F5456FFEE9D9BFA111C64F9EE" box="[956,1041,1519,1545]" captionStart="FIGURE 17" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[152,1435,184,1918]" captionTargetId="figure-22@38.[152,1435,181,1920]" captionTargetPageId="38" captionText="FIGURE 17. Habitats of Lygodactylus nyaneka sp. nov. Top: Fazenda Mucungo. Namibe Province. Below: Bruco Pass. Huila escarpment. Namibe Province. Photos by Luis M. P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.4410761" httpUri="https://zenodo.org/record/4410761/files/figure.png" pageId="37" pageNumber="338">Fig. 17</figureCitation>
). Never found on the ground. This species occurs in sympatry with
<emphasis id="A3ADEA885456FFEE9C23F9ED1B6BF9CA" box="[516,798,1555,1581]" italics="true" pageId="37" pageNumber="338">
<taxonomicName id="56D94D195456FFEE9C23F9ED1B6CF9CA" box="[516,793,1555,1581]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="angolensis">Lygodactylus angolensis</taxonomicName>
,
</emphasis>
in the northern region of
<collectingRegion id="531DF8785456FFEE9A19F9ED1CA6F9CA" box="[1086,1235,1555,1581]" country="Angola" name="Bie" pageId="37" pageNumber="338">Bié Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C365115456FFEE9EE0F9C91C06F97E" pageId="37" pageNumber="338" type="etymology">
<paragraph id="9166369A5456FFEE9EE0F9C91C06F97E" blockId="37.[151,1437,151,1689]" pageId="37" pageNumber="338">
<emphasis id="A3ADEA885456FFEE9EE0F9C9193BF9B6" bold="true" box="[199,334,1591,1617]" pageId="37" pageNumber="338">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="56D94D195456FFEE9C74F9C91AC5F9B6" authorityName="Marques &amp; Ceríaco &amp; Buehler &amp; Bandei- Ra &amp; Janota &amp; Bauer" authorityYear="2020" box="[595,688,1591,1617]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="nyaneka">
<emphasis id="A3ADEA885456FFEE9C74F9C91AC5F9B6" box="[595,688,1591,1617]" italics="true" pageId="37" pageNumber="338">nyaneka</emphasis>
</taxonomicName>
” is a noun in apposition and is given in honor of the dominant ethnolinguistic group of the region where the species exists. We suggest “Nyaneka Dwarf Gecko” and “Osga-Anã-Nhanheca” as the English and Portuguese common names, respectively, for this species.
</paragraph>
</subSubSection>
</treatment>
</document>