treatments-xml/data/97/45/E9/9745E9441F0E5C99B50D8EC76072331F.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="9745E9441F0E5C99B50D8EC76072331F" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina stoliczkai Jirapatrasilp 2022, sp. nov." docType="treatment" docUuid="80A5D354-B516-4F67-9F08-817A29FBBEFB" docUuidSource="ZooBank" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1119</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>115</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199839013" LSID="urn:lsid:zoobank.org:act:80A5D354-B516-4F67-9F08-817A29FBBEFB" httpUri="http://treatment.plazi.org/id/9745E9441F0E5C99B50D8EC76072331F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/80A5D354-B516-4F67-9F08-817A29FBBEFB" authority="Jirapatrasilp" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina stoliczkai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stoliczkai" status="sp. nov.">Pupina stoliczkai Jirapatrasilp</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">Figs 36G, H</figureCitation>
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. A, B Pupina latisulci sp. nov. A holotype CUMZ 12146 / 1 and B paratype CUMZ 12146 / 2 from Khao Ok Talu, Phatthalung C, D Pupina stoliczkai sp. nov. C holotype CUMZ 12147 / 1 and D paratype NHMUK 20210336 from Wat Ratburana School, Chumpon E Pupina lowi, syntype MNHN-IM- 2000 - 35846 from Lahat, Perak F Pupina dorri dorri, paralectotype MNHN-IM- 2000 - 35836 from Haiphong, Vietnam. Photo: P. Maestrati, MNHN (E, F)." figureDoi="10.3897/zookeys.1119.85400.figure40" httpUri="https://binary.pensoft.net/fig/735401" pageId="0" pageNumber="1">, 40C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
CUMZ 12147/1 (Figs
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36G</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. A, B Pupina latisulci sp. nov. A holotype CUMZ 12146 / 1 and B paratype CUMZ 12146 / 2 from Khao Ok Talu, Phatthalung C, D Pupina stoliczkai sp. nov. C holotype CUMZ 12147 / 1 and D paratype NHMUK 20210336 from Wat Ratburana School, Chumpon E Pupina lowi, syntype MNHN-IM- 2000 - 35846 from Lahat, Perak F Pupina dorri dorri, paralectotype MNHN-IM- 2000 - 35836 from Haiphong, Vietnam. Photo: P. Maestrati, MNHN (E, F)." figureDoi="10.3897/zookeys.1119.85400.figure40" httpUri="https://binary.pensoft.net/fig/735401" pageId="0" pageNumber="1">40C</figureCitation>
), 10 Sept. 2016, coll. R. Srisonchai, A. Pholyotha, T. Seesamut. Measurement: shell height 9.4 mm, shell width 6.3 mm and
<normalizedToken originalValue="6½">61/2</normalizedToken>
whorls.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
CUMZ 12147/2 (1 specimen in ethanol) and NHMUK 20210336 (1 shell; Figs
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36H</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. A, B Pupina latisulci sp. nov. A holotype CUMZ 12146 / 1 and B paratype CUMZ 12146 / 2 from Khao Ok Talu, Phatthalung C, D Pupina stoliczkai sp. nov. C holotype CUMZ 12147 / 1 and D paratype NHMUK 20210336 from Wat Ratburana School, Chumpon E Pupina lowi, syntype MNHN-IM- 2000 - 35846 from Lahat, Perak F Pupina dorri dorri, paralectotype MNHN-IM- 2000 - 35836 from Haiphong, Vietnam. Photo: P. Maestrati, MNHN (E, F)." figureDoi="10.3897/zookeys.1119.85400.figure40" httpUri="https://binary.pensoft.net/fig/735401" pageId="0" pageNumber="1">40D</figureCitation>
), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Wat Ratburana School, Lang Suan District, Chumpon Province, Thailand,
<geoCoordinate degrees="9" direction="north" minutes="56" orientation="latitude" precision="1" seconds="18.0" value="9.9383335">9°56'18.0&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="02" orientation="longitude" precision="1" seconds="25.5" value="99.04041">99°02'25.5&quot;E</geoCoordinate>
, 20 m amsl.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. 70% of shell height. Apertural lip highly thickened and slightly expanded; with a furrow between inner and outer peristomes; inner peristome thickened, cord-like. Both parietal and columellar teeth fin-shaped, very thickened, always located next to and not covering its respective canal. Posterior canal slightly bulging outward.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. stoliczkai" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="stoliczkai">
<emphasis italics="true" pageId="0" pageNumber="1">P. stoliczkai</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
in shell shape in having both fin-shaped and highly thickened parietal and columellar teeth located next to their respective canals, and the posterior canal slightly bulging outward. However,
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. stoliczkai" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="stoliczkai">
<emphasis italics="true" pageId="0" pageNumber="1">P. stoliczkai</emphasis>
</taxonomicName>
sp. nov. has a furrow between inner and outer peristomes, with inner peristome thickened and cord-like.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell height 6.3-6.4 mm; shell width 9.0-9.5 mm. Shell ovate-fusiform, solid, semi-transparent, reddish brown, devoid of prominent sculpture on glazed smooth surface. Apex obtuse. Growth lines on shell surface inconspicuous. Whorls 6-
<normalizedToken originalValue="6½">61/2</normalizedToken>
, last whorl large (ca. 70% of shell height). Spire angle ca. 80-90°; slightly extended. Sutures slightly impressed, but shallow. Aperture circular; lip highly thickened (ca. 0.4-0.5 mm wide and 0.6-0.7 mm thick) with paler colour, slightly expanded. Aperture with a furrow between inner and outer peristomes, with inner peristome thickened, cord-like. Parietal callus sharply defined and thickened with paler colour. Peristome interrupted by two canals; posterior canal ca. 1.4 mm long and 0.7 mm at its widest, slightly bulging outward, continuing obliquely and widening vertically upward when observed from lateral view. Anterior canal curved and continuing obliquely upward ca. 2.0 mm. Parietal tooth fin-shaped, highly thickened (ca. 1.5 mm long, 0.5 mm at its widest and 0.3 mm thick), always located next to and not covering posterior canal. Columellar tooth fin-shaped, highly thickened (ca. 1.9 mm long, 0.5 mm at its widest and 0.3 mm thick), located next to anterior canal. Umbilicus closed. Operculum round, thick, flat, multispiral, whitish to pale yellow, opaque corneous with smooth edge.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet is dedicated to F. Stoliczka, a Czech palaeontologist and zoologist, who described
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
, to which this new species is associated with.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">This new species is found only from the type locality.</paragraph>
<caption doi="10.3897/zookeys.1119.85400.figure27" httpUri="https://binary.pensoft.net/fig/735388" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
Distribution map of the
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
species group:
<taxonomicName authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina peguensis</emphasis>
</taxonomicName>
(triangle),
<taxonomicName authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
(plus sign),
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina siamensis</emphasis>
</taxonomicName>
(circle),
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina bilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina bilabiata</emphasis>
</taxonomicName>
sp. nov. (square), and
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina godwinausteni</emphasis>
</taxonomicName>
sp. nov. (star). Each red symbol indicates the type locality of its respective taxon. The occurrences of
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Pfeiffer" authorityYear="1861" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina mouhoti</emphasis>
</taxonomicName>
in northern Thailand are uncertain, thus their distributions are not mapped.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1" start="Figure 28" startId="F28">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 28.</emphasis>
Shells of
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
species group from mainland Southeast Asia
<emphasis bold="true" pageId="0" pageNumber="1">A-G</emphasis>
<taxonomicName authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina peguensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype of
<taxonomicName authorityName="Theobald" authorityYear="1864" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina blanfordi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="blanfordi">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina blanfordi</emphasis>
</taxonomicName>
NHMUK 1888.12.4.100 and specimens
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
CUMZ 12105
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
CUMZ 12050
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
CUMZ 12103
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
NHMUK ex. Cuming coll.
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
CUMZ 12094, and
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
CUMZ 12108
<emphasis bold="true" pageId="0" pageNumber="1">H-J</emphasis>
<taxonomicName authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
lectotype MNHN-IM-2000-35834
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
paralectotype RBINS MT966/10591, and
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
specimen CUMZ 12049
<emphasis bold="true" pageId="0" pageNumber="1">K, L</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina siamensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
lectotype SMF 109948 and
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
specimen CUMZ 12052
<emphasis bold="true" pageId="0" pageNumber="1">M-P</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina bilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina bilabiata</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
holotype CUMZ 12073/1 and specimens
<emphasis bold="true" pageId="0" pageNumber="1">N</emphasis>
CUMZ 12079
<emphasis bold="true" pageId="0" pageNumber="1">O</emphasis>
CUMZ 12081, and
<emphasis bold="true" pageId="0" pageNumber="1">P</emphasis>
CUMZ 12086
<emphasis bold="true" pageId="0" pageNumber="1">Q, R</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina godwinausteni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Q</emphasis>
holotype CUMZ 12090/1 and
<emphasis bold="true" pageId="0" pageNumber="1">R</emphasis>
paratype CUMZ 12090/2. Photo: P. Maestrati, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
), F. Trus, RBINS (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
Shells of
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
species group from mainland Southeast Asia
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype UMZC I.103025 and
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
specimen NHMUK 1888.12.4.109.
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
<taxonomicName authorityName="Pfeiffer" authorityYear="1861" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina mouhoti</emphasis>
</taxonomicName>
, possible syntypes NHMUK ex. Cuming coll.
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName authorityName="Morelet" authorityYear="1862" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina vescoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vescoi">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina vescoi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
syntype NHMUK 1893.2.4.767 and
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
specimen SMF 109956/1
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1891" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina perakensis</emphasis>
</taxonomicName>
, lectotype SMF 109969/1
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina excisa</emphasis>
</taxonomicName>
, lectotype SMF 110778/1
<emphasis bold="true" pageId="0" pageNumber="1">I-K</emphasis>
<taxonomicName authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina exclamationis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">I, J</emphasis>
syntypes MNHN-IM-2000-35840 and
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
specimen NHMUK 1901.12.23.205 &quot;forma minor&quot;. Photo: J. Ablett, H. Taylor, NHM (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), A. Lardeur, P. Maestrati, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">I, J</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure30" httpUri="https://binary.pensoft.net/fig/735391" pageId="0" pageNumber="1" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 30.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
<taxonomicName authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina peguensis</emphasis>
</taxonomicName>
: specimens
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
CUMZ 12105 from Thep Phithak Punnaram Temple, Nakhon Ratchasima
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
CUMZ 12108 from Tham Thep Bandan Temple, Phetchabun, and
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
lectotype MNHN-IM-2000-35834 from Cambodge and
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
paralectotype RBINS MT966/10591 from Phnom-Rohan, Cambodge. Photo: P. Maestrati, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), F. Trus, RBINS (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure31" httpUri="https://binary.pensoft.net/fig/735392" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
, specimen CUMZ 12049 from Khao Jedee Temple, Nakhon Sawan
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina siamensis</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
lectotype SMF 109948 and
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
specimen CUMZ 12052 from Sri Thong Cave, Sra Keo
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina bilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina bilabiata</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
holotype CUMZ 12073/1, and specimens
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
CUMZ 12079 from Tham Pha Ya Temple, Loei and
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
CUMZ 12081 from Pha Jor Cave, Nong Bua Lam Phu.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
Live specimens of
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina siamensis</emphasis>
</taxonomicName>
, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina bilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina bilabiata</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
paratype CUMZ 12073/2 from Banpot Pisai Temple, Chumphon and specimens
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
CUMZ 12087 from Ban Yai, Surat Thani
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina godwinausteni</emphasis>
</taxonomicName>
sp. nov.: paratypes
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
CUMZ 12090/26 and
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
CUMZ 12091 from Khao Wong Cave, Uthai Thani
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
<taxonomicName authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina aureola</emphasis>
</taxonomicName>
: specimens
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 33.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina bilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina bilabiata</emphasis>
</taxonomicName>
sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
<taxonomicName authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina godwinausteni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
holotype CUMZ 12090/1 and
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
paratype CUMZ 12090/2 from Khao Wong Cave, Uthai Thani
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
, syntype UMZC I.103025
<normalizedToken originalValue="“Ind”">&quot;Ind&quot;</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Pfeiffer" authorityYear="1861" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina mouhoti</emphasis>
</taxonomicName>
, possible syntype NHMUK ex. Cuming coll. from Camboja
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Morelet" authorityYear="1862" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina vescoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vescoi">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina vescoi</emphasis>
</taxonomicName>
, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Morelet" authorityYear="1862" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina vescoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vescoi">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina vescoi</emphasis>
</taxonomicName>
, specimen SMF 109956/1 from Cochin China
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
<taxonomicName authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina exclamationis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
syntype MNHN-IM-2000-35840 from Tonkin and
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
specimen NHMUK 1901.12.23.205 &quot;forma minor&quot; from Than-Moi, Tonkin
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1891" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina perakensis</emphasis>
</taxonomicName>
, lectotype SMF 109969/1 from Bukit Pondong, Perak
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina excisa</emphasis>
</taxonomicName>
, lectotype SMF 110778/1 from Kelantan
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina aureola</emphasis>
</taxonomicName>
, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>