305 lines
27 KiB
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305 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.148.50042" ID-GBIF-Dataset="1224cf7b-1d2d-4657-89e9-11fee1f5ea0b" ID-PMC="PMC7266835" ID-Pensoft-Pub="1314-2003-148-21" ID-Pensoft-UUID="6E288E0A8F725B2787799DD17F2B1642" ID-PubMed="32523392" ModsDocID="1314-2003-148-21" checkinTime="1590560163243" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Peterson, Paul M., Romaschenko, Konstantin, Bravo-Pedraza, William J., Cuta-Alarcon, Lia E. & Soreng, Robert J." docDate="2020" docId="4CC1F9F32CCA5761AB4565287E2DD1AA" docLanguage="en" docName="PhytoKeys 148: 21-50" docOrigin="PhytoKeys 148" docSource="http://dx.doi.org/10.3897/phytokeys.148.50042" docTitle="Podagrostis bacillata Sylvester & Soreng 2020, comb. nov." docType="treatment" docVersion="4" id="6E288E0A8F725B2787799DD17F2B1642" lastPageNumber="21" masterDocId="6E288E0A8F725B2787799DD17F2B1642" masterDocTitle="New combinations and updated descriptions in Podagrostis (Agrostidinae, Poaceae) from the Neotropics and Mexico" masterLastPageNumber="50" masterPageNumber="21" pageNumber="21" updateTime="1668139801695" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New combinations and updated descriptions in Podagrostis (Agrostidinae, Poaceae) from the Neotropics and Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Romaschenko, Konstantin</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>148</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.148.50042</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.148.50042</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-148-21</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6E288E0A8F725B2787799DD17F2B1642</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164360813" LSID="urn:lsid:plazi:treatment:4CC1F9F32CCA5761AB4565287E2DD1AA" httpUri="http://treatment.plazi.org/id/4CC1F9F32CCA5761AB4565287E2DD1AA" lastPageNumber="21" pageId="0" pageNumber="21">
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<subSubSection pageId="0" pageNumber="21" type="nomenclature">
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName LSID="4CC1F9F3-2CCA-5761-AB45-65287E2DD1AA" authority="(Hack.) Sylvester & Soreng" authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis bacillata" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="bacillata" status="comb. nov.">Podagrostis bacillata (Hack.) Sylvester & Soreng</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="21">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Podagrostis bacillata. A Whole plant B spikelets, the center spikelet at anthesis with the palea opened out and the rachilla prolongation visible (indicated with a red arrow) C leaf blades showing the densely scabrous abaxial surface and the central blade opened out to reveal the scabrous adaxial surface. A, B Images of specimen A. M. Evans 145 (US 2589766 A) courtesy of United States National Herbarium (US) C image of specimen R. W. Pohl 11705 A (US 3096988)." figureDoi="10.3897/phytokeys.148.50042.figure1" httpUri="https://binary.pensoft.net/fig/413782" pageId="0" pageNumber="21">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="reference_group">
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis bacillata" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="bacillata">Agrostis bacillata</taxonomicName>
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Hack., Oesterr. Bot. Z. 52(2): 59. 1902.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="materials_examined">
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<paragraph pageId="0" pageNumber="21">Type.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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Costa Rica. Cerro de la Muerte, Jan. 1897, H. Pittier 10477 (
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">holotype</emphasis>
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: W (W19160027240 [image!]);
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">isotypes</emphasis>
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: BAB (BAB00000206 [image!]), BR (BR0000006595845 [image!]), G (G00192030 [image!]), US (US00131726 [not seen], US00131725 [not seen], US00131136 fragm. [not seen]).
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</paragraph>
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<caption doi="10.3897/phytokeys.148.50042.figure1" httpUri="https://binary.pensoft.net/fig/413782" pageId="0" pageNumber="21" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="21">
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<emphasis bold="true" pageId="0" pageNumber="21">Figure 1.</emphasis>
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<taxonomicName authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis bacillata" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="bacillata">
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<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis bacillata</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="21">A</emphasis>
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Whole plant
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<emphasis bold="true" pageId="0" pageNumber="21">B</emphasis>
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spikelets, the center spikelet at anthesis with the palea opened out and the rachilla prolongation visible (indicated with a red arrow)
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<emphasis bold="true" pageId="0" pageNumber="21">C</emphasis>
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leaf blades showing the densely scabrous abaxial surface and the central blade opened out to reveal the scabrous adaxial surface.
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<emphasis bold="true" pageId="0" pageNumber="21">A, B</emphasis>
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Images of specimen A.M. Evans 145 (US2589766A) courtesy of United States National Herbarium (US)
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<emphasis bold="true" pageId="0" pageNumber="21">C</emphasis>
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image of specimen R.W. Pohl 11705A (US3096988).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="description">
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<paragraph pageId="0" pageNumber="21">Description.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName genus="Tufted" lsidName="Tufted perennial" pageId="0" pageNumber="21" rank="species" species="perennial">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Tufted perennial</emphasis>
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</taxonomicName>
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forming dense tufts, with the basal mats reaching c. 6-17 cm tall and inflorescences usually well-exserted from the basal foliage.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Tillers</emphasis>
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intravaginal.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Culms</emphasis>
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10-30(-50) cm tall, erect, simple, delicate;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">nodes and internodes</emphasis>
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terete, smooth, usually with at least 1 or 2 elongated internodes visible, with (0-)1-3 nodes exposed at flowering, uppermost internode c. 4-13 cm long, longer than the sheath.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Leaves</emphasis>
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generally basal;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">sheaths</emphasis>
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terete, glabrous, smooth to lightly scabrous with very short scabers; flag sheath 4-13 cm long; basal sheaths 1-4 cm long, striate, becoming fibrous;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">ligules</emphasis>
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1.7-4.3 mm long, membranous, slightly to usually strongly decurrent with the sheath; upper culm ligules acute to acuminate with a obtuse to acuminate apex, sometimes slightly erose towards the apex; ligules of tillers similar to those of the culm, sometimes slightly shorter;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">blades</emphasis>
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2-15 cm long, 0.2-0.3(-0.4) mm wide in diameter, involute or convolute, acicular to capillaceous and filiform, straight to slightly curved, abaxial surfaces glabrous, lightly to usually densely scabrous, adaxial surfaces glabrous, lightly to usually densely scabrous on the veins with scabers varying in size from short to long and robust.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Panicles</emphasis>
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(3.5-)4-11.5
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<normalizedToken originalValue="×">x</normalizedToken>
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(1-)3-5 cm, open to slightly congested when young, usually ovoid;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">panicle branches</emphasis>
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ascendant to patent, branched above the middle, filiform, with spikelets not present near the base, glabrous, smooth, longest branches 1-4.5 cm long;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">pedicels</emphasis>
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(1-)2-6 mm long, usually longer than the length of the spikelets, divaricate, glabrous, smooth.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Spikelets</emphasis>
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1.7-2 mm long;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">glumes</emphasis>
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remaining on the inflorescence at maturity, equal or subequal, the lower usually slightly longer than the upper, almost equaling the length of the floret to 0.5 mm longer, oblong-lanceolate, slightly keeled, apex broadly acute, glabrous, keels lightly scaberulous just in the distal 1/3, surfaces smooth; lower glume 1- (or 3-)veined, lateral veins usually vestigial; upper glume 1- (or 2- or 3-)veined, lateral veins usually vestigial;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">lemmas</emphasis>
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1.4-1.6 mm long, glabrous, smooth, faintly 5-veined, apex obtuse, awn lacking or to 0.6 mm long, straight, inserted medially or in the upper 1/3;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">paleas</emphasis>
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well-developed, 0.8-1.5 mm long, usually reaching from
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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to subequaling the lemma, keels slightly obscure, smooth, apex bifid and sometimes erose;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">rachilla</emphasis>
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0.3-1.4 mm long, prolonged from the base of the floret, glabrous, rarely smooth, usually scabrous, sometimes with scabers extending into very short hairs <0.2 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Calluses</emphasis>
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0.1-0.2 mm long, slightly elongated, glabrous, smooth.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Flowers; lodicules</emphasis>
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c. 0.3-0.5 mm long, lanceolate with acute apices, not lobed;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">anthers</emphasis>
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3 in number, 0.8-1 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Caryopses</emphasis>
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not seen. 2
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<emphasis italics="true" pageId="0" pageNumber="21">n</emphasis>
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= 28.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="distribution">
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<paragraph pageId="0" pageNumber="21">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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Costa Rica and
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<normalizedToken originalValue="Panamá">Panama</normalizedToken>
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,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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grasslands, 2200-3900 m alt.
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<bibRefCitation author="Pohl, RW" editor="Burger, WC" journalOrPublisher="Fieldiana Botany" pageId="0" pageNumber="21" pagination="1 - 608" refId="B20" refString="Pohl, RW, 1980. Family 15. Gramineae. In: Burger, WC, Ed., Flora Costaricensis. Fieldiana Botany 4: 1 - 608" title="Family 15. Gramineae." volume="4" volumeTitle="Flora Costaricensis." year="1980">Pohl (1980)</bibRefCitation>
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mentions that this species apparently flowers throughout the year.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="materials_examined">
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<paragraph pageId="0" pageNumber="21">Additional specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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Costa Rica.
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<emphasis bold="true" pageId="0" pageNumber="21">Cartago</emphasis>
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: Canyon of Rio Tiribi, 3 km NW of Llano Grande, 2200 m, 7 Feb. 1969, R.W. Pohl 11705A (US3096988).
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<emphasis bold="true" pageId="0" pageNumber="21">Limon</emphasis>
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: Cordillera de Talamanca, Atlantic slope, Kamuk Massif,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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north-east of the main Kamuk peak, 9°1'-9°17'N, 83°00'-83°0'W, 3000-3300 m alt., 17-18 Sep. 1984, G. Davidse 29277 (US3041608).
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<emphasis bold="true" pageId="0" pageNumber="21">San Jose</emphasis>
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: Cerro Chirripo, treeless
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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SW of and around summit, 11000-12000 ft [3353-3658 m alt.], 24-26 Aug. 1967, A.M. Evans 145 (US2589766A); Cerro de la Vueltas,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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, 2700-3000 m alt., 29 Dec. 1925-1 Jan. 1926, P.C. Standley 43625 (US1307153); P.C. Standley 43678 (US1307155); Cordillera de Talamanca, Cerro de la Muerte, Asuncion, 3250 m alt., 13 July 1968, R.W. Pohl 10693 (US3096655).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="notes">
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<paragraph pageId="0" pageNumber="21">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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The possible affinity of
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis bacillata" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="bacillata">
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<emphasis italics="true" pageId="0" pageNumber="21">Agrostis bacillata</emphasis>
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</taxonomicName>
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to
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis</emphasis>
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</taxonomicName>
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was mentioned previously (
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<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse 1994</bibRefCitation>
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;
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<bibRefCitation author="Briceno, B" editor="Morillo, GN" journalOrPublisher="2. Centro Editorial Litoramma C. A., Merida" pageId="0" pageNumber="21" pagination="599 - 711" refId="B2" refString="Briceno, B, 2010. Capitulo XXI Familia Poaceae (R.Br.) Barnhart (Gramineae). In: Morillo, GN, Briceno, B, Silva, JF, Eds., Botanica y Ecologia de las Monocotiledoneas de los Paramos en Venezuela, vol. 2. Centro Editorial Litoramma C. A., Merida: 599 - 711" title="Capitulo XXI Familia Poaceae (R. Br.) Barnhart (Gramineae)." volumeTitle="Botanica y Ecologia de las Monocotiledoneas de los Paramos en Venezuela, vol." year="2010">
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<normalizedToken originalValue="Briceño">Briceno</normalizedToken>
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2010
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</bibRefCitation>
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), with
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<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
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recommending transfer of
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<taxonomicName lsidName="A. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
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<emphasis italics="true" pageId="0" pageNumber="21">A. bacillata</emphasis>
|
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</taxonomicName>
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to
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis</emphasis>
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</taxonomicName>
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. The species is similar to
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<taxonomicName lsidName="P. trichodes" pageId="0" pageNumber="21" rank="species" species="trichodes">
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<emphasis italics="true" pageId="0" pageNumber="21">P. trichodes</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
(see notes under these taxa on how to differentiate them). The character of adaxial leaf blade surface with conical trichomes intermixed with scabers across a scaberulous surface, mentioned by
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
|
||
to be a key character for differentiating
|
||
<taxonomicName lsidName="P. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. bacillata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
, was not found in any of the Costa Rican specimens studied at US, including specimen Pohl 10693 that was cited by
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
|
||
. All specimens studied of this species were generally densely scabrous on the veins of the adaxial blade surface, with short to long and robust scabers of similar consistency to the abaxial surface. This could be attributed to a difference in interpretation by the authors, although similar long scabers, in a similar density, were found on the adaxial leaf blade surface of specimens of
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
, meaning this character is not considered useful for species delimitation of these taxa. Instead, we found that a key character for differentiating the two species was the presence or absence of scabrocities on the abaxial leaf blade surface, with
|
||
<taxonomicName lsidName="P. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. bacillata</emphasis>
|
||
</taxonomicName>
|
||
usually being densely scabrous with short hooks while
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
is smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="21">
|
||
Although we have not studied characters of foliar anatomy,
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
|
||
mention these to be important in separation of
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="P. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. bacillata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
|
||
also mention that spikelets are amply open at maturity in
|
||
<taxonomicName lsidName="P. exserta" pageId="0" pageNumber="21" rank="species" species="exserta">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. exserta</emphasis>
|
||
</taxonomicName>
|
||
, while those of
|
||
<taxonomicName lsidName="P. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. bacillata</emphasis>
|
||
</taxonomicName>
|
||
are more closed, although this distinction is not always clear. Caryopsis shape is also mentioned to be distinct (
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse 1994</bibRefCitation>
|
||
), although no fruiting specimens were found among specimens at US.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="21">
|
||
<bibRefCitation author="Pohl, RW" editor="Burger, WC" journalOrPublisher="Fieldiana Botany" pageId="0" pageNumber="21" pagination="1 - 608" refId="B20" refString="Pohl, RW, 1980. Family 15. Gramineae. In: Burger, WC, Ed., Flora Costaricensis. Fieldiana Botany 4: 1 - 608" title="Family 15. Gramineae." volume="4" volumeTitle="Flora Costaricensis." year="1980">Pohl (1980)</bibRefCitation>
|
||
mentions the glumes of
|
||
<taxonomicName lsidName="P. bacillata" pageId="0" pageNumber="21" rank="species" species="bacillata">
|
||
<emphasis italics="true" pageId="0" pageNumber="21">P. bacillata</emphasis>
|
||
</taxonomicName>
|
||
to be 3-veined, although this is not mentioned by
|
||
<bibRefCitation author="Pohl, RW" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico, D. F." pageId="0" pageNumber="21" pagination="237 - 240" refId="B21" refString="Pohl, RW, Davidse, G, 1994. 46. Agrostis L. In: Davidse, G, Sousa, M, Sanchez, MS, Chater, AO, Eds., Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico, D. F.: 237 - 240" title="46. Agrostis L." volumeTitle="Flora Mesoamericana, vol 6 Alismataceae a Cyperaceae." year="1994">Pohl and Davidse (1994)</bibRefCitation>
|
||
or
|
||
<bibRefCitation author="Morales-Quiros, JF" editor="Hammel, BE" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden 93 (3)" pageId="0" pageNumber="21" pagination="598 - 821" refId="B17" refString="Morales-Quiros, JF, 2003. Poaceae. In: Hammel, BE, Grayum, MH, Herrera-Mora, C, Zamora-Villalobos, N, Eds., Manual of Plants of Costa Rica. Monographs in Systematic Botany from the Missouri Botanical Garden 93(3): 598 - 821" title="Poaceae." volumeTitle="Manual of Plants of Costa Rica." year="2003">
|
||
<normalizedToken originalValue="Morales-Quirós">Morales-Quiros</normalizedToken>
|
||
(2003)
|
||
</bibRefCitation>
|
||
. Most of the spikelets that we examined had 1-veined glumes, with vestigial lateral veins sometimes present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |