treatments-xml/data/F9/36/97/F9369705FFE8FF9BFDB0F9C8EAF6FCDA.xml

<document id="48959B760D4EFEB7D0F7A2EC368F0AF5" ID-DOI="10.1080/00222933.2013.800610" ID-ISSN="1464-5262" ID-Zenodo-Dep="5197685" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628882431226" checkinUser="felipe" docAuthor="Guidetti, Roberto, Peluffo, Julio Ricardo, Rocha, Alejandra Mariana, Cesari, Michele &amp; Peluffo, María Cristina Moly de" docDate="2013" docId="F9369705FFE8FF9BFDB0F9C8EAF6FCDA" docLanguage="en" docName="JNatHist.47.37-38.2409-2426.pdf" docOrigin="Journal of Natural History 47 (37 - 38)" docSource="http://dx.doi.org/10.1080/00222933.2013.800610" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Macrobiotus kristenseni Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2422" masterDocId="050FEF7DFFEEFF8BFFB1FF94EB62FFA6" masterDocTitle="The morphological and molecular analyses of a new South American urban tardigrade offer new insights on the biological meaning of the Macrobiotus hufelandi group of species (Tardigrada: Macrobiotidae)" masterLastPageNumber="2426" masterPageNumber="2409" pageNumber="2413" updateTime="1699054413493" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FCE06DCF05D2C2B247326D91D58C4859">The morphological and molecular analyses of a new South American urban tardigrade offer new insights on the biological meaning of the Macrobiotus hufelandi group of species (Tardigrada: Macrobiotidae)</mods:title>
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<mods:namePart id="DA17192A006E43300B6F531DED6BA9C0">Guidetti, Roberto</mods:namePart>
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<mods:namePart id="7832EE5075A21E78646CA6A17090AFFE">Peluffo, Julio Ricardo</mods:namePart>
<mods:affiliation id="371E3633FB2FD66EB9CE24181FC96714">Faculty of Natural and Exact Sciences, National University of La Pampa, Santa Rosa, Argentina</mods:affiliation>
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<mods:namePart id="D01CB845828248A78E71E07FA95436DA">Rocha, Alejandra Mariana</mods:namePart>
<mods:affiliation id="4980618165C26D9176B2C861D3DDD657">Faculty of Natural and Exact Sciences, National University of La Pampa, Santa Rosa, Argentina</mods:affiliation>
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<mods:namePart id="BDEA0D91C4A1C88339F42778ECFE2EBA">Cesari, Michele</mods:namePart>
<mods:affiliation id="5BA4EF85305D9FCB034323224C960B5C">Department of Life Sciences, University of Modena and Reggio Emilia, Modena, Italy;</mods:affiliation>
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<mods:namePart id="3445134FD6A9B6E3282D77239BE467FD">Peluffo, María Cristina Moly de</mods:namePart>
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<mods:title id="E182C19C97738BE45FC8EC2960061C1E">Journal of Natural History</mods:title>
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<mods:number id="5DD2C3107869E10506EF5CAD93AE3660">2013-06-27</mods:number>
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<mods:number id="E6DC203A08AC0E23ACC5BA524C5739A3">47</mods:number>
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<mods:number id="183B89C561825F8CF186ECC45EE159FE">37 - 38</mods:number>
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<subSubSection id="39857598FFE8FF8DFDB0F9C8E804F9D2" box="[513,870,1628,1653]" pageId="6" pageNumber="2413" type="nomenclature">
<paragraph id="71202613FFE8FF8DFDB0F9C8E804F9D2" blockId="6.[513,870,1628,1684]" box="[513,870,1628,1653]" pageId="6" pageNumber="2413">
<heading id="2A68917FFFE8FF8DFDB0F9C8E804F9D2" box="[513,870,1628,1653]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="2413" reason="2">
<taxonomicName id="B69F5D90FFE8FF8DFDB0F9C8E868F9D3" ID-CoL="aeebfe59-de7c-4d3d-808f-84edde61f876" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[513,778,1628,1653]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="2413" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE8FF8DFDB0F9C8E868F9D3" bold="true" box="[513,778,1628,1653]" italics="true" pageId="6" pageNumber="2413">Macrobiotus kristenseni</emphasis>
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<paragraph id="71202613FFE8FF8DFDAFF9E8E859F932" blockId="6.[513,870,1628,1684]" box="[542,827,1660,1684]" pageId="6" pageNumber="2413">
(
<figureCitation id="E9A43A96FFE8FF8DFD94F9E8E9C8F932" box="[549,682,1660,1684]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="9.[143,210,1158,1180]" captionStartId-1="10.[211,278,1174,1196]" captionStartId-2="11.[143,210,948,970]" captionStartId-3="12.[211,278,1094,1116]" captionStartId-4="13.[143,210,934,956]" captionTargetBox-0="[145,1102,168,1125]" captionTargetBox-1="[213,1170,169,1142]" captionTargetBox-2="[145,1102,168,915]" captionTargetBox-3="[213,1170,168,1062]" captionTargetBox-4="[145,1102,168,901]" captionTargetId-0="figure-5@9.[145,1102,168,1125]" captionTargetId-1="figure-5@10.[213,1170,169,1142]" captionTargetId-2="figure-5@11.[145,1102,168,915]" captionTargetId-3="figure-5@12.[213,1170,168,1062]" captionTargetId-4="figure-5@13.[145,1102,168,901]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." captionText-1="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." captionText-2="Figure 3. Macrobiotus kristenseni sp. nov. (A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm." captionText-3="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." captionText-4="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi-0="http://doi.org/10.5281/zenodo.5197687" figureDoi-1="http://doi.org/10.5281/zenodo.5197689" figureDoi-2="http://doi.org/10.5281/zenodo.5197691" figureDoi-3="http://doi.org/10.5281/zenodo.5197693" figureDoi-4="http://doi.org/10.5281/zenodo.5197695" httpUri-0="https://zenodo.org/record/5197687/files/figure.png" httpUri-1="https://zenodo.org/record/5197689/files/figure.png" httpUri-2="https://zenodo.org/record/5197691/files/figure.png" httpUri-3="https://zenodo.org/record/5197693/files/figure.png" httpUri-4="https://zenodo.org/record/5197695/files/figure.png" pageId="6" pageNumber="2413">Figures 1–5</figureCitation>
and 
<tableCitation id="3C1D13A8FFE8FF8DFD53F9E8E851F932" box="[738,819,1660,1684]" captionStart="Table 2" captionStartId="7.[58,113,485,507]" captionText="Table 2. Summary of morphometric data (in µm, except pt) for Macrobiotus kristenseni sp. nov." pageId="6" pageNumber="2413">Table 2</tableCitation>
)
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<paragraph id="71202613FFE9FF8CFF8BFE71EF61FE5D" blockId="7.[58,1027,485,507]" box="[58,1027,485,507]" pageId="7" pageNumber="2414">
Table 2. Summary of morphometric data (in µm, except 
<emphasis id="43EBFA01FFE9FF8CFDCAFE71E9EDFE5D" box="[635,655,485,507]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
) for 
<taxonomicName id="B69F5D90FFE9FF8CFD71FE71E8CCFE5D" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[704,942,485,507]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="2414" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE9FF8CFD71FE71E8CCFE5D" box="[704,942,485,507]" italics="true" pageId="7" pageNumber="2414">Macrobiotus kristenseni</emphasis>
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<taxonomicNameLabel id="58D8477AFFE9FF8CFC04FE71EF61FE5D" box="[949,1027,485,507]" pageId="7" pageNumber="2414" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="71202613FFE9FF8CFF8BFD8CEEDEFBCD" pageId="7" pageNumber="2414">
<table id="039FD4B3FFE90074FF8BFD8CEEDEFBCD" box="[58,1468,536,1131]" gridcols="13" gridrows="19" pageId="7" pageNumber="2414">
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<th id="8C7E4D2DFFE90074FF8BFD8CEBFDFD88" box="[58,159,536,558]" gridcol="0" gridrow="0" pageId="7" pageNumber="2414">Character</th>
<th id="8C7E4D2DFFE90074FF7CFD8CEA02FD88" box="[205,352,536,558]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="2414">Holotype</th>
<th id="8C7E4D2DFFE90074FE28FD8CEAB9FD88" box="[409,475,536,558]" gridcol="3" gridrow="0" pageId="7" pageNumber="2414">Mean</th>
<th id="8C7E4D2DFFE90074FDBAFD8CE923FD88" box="[523,577,536,558]" gridcol="4" gridrow="0" pageId="7" pageNumber="2414">SD</th>
<th id="8C7E4D2DFFE90074FDDFFD8CE9D2FD88" box="[622,688,536,558]" gridcol="5" gridrow="0" pageId="7" pageNumber="2414">Min</th>
<th id="8C7E4D2DFFE90074FD68FD8CE879FD88" box="[729,795,536,558]" gridcol="6" gridrow="0" pageId="7" pageNumber="2414">Max</th>
<th id="8C7E4D2DFFE90074FCF8FD8CE8F8FD88" box="[841,922,536,558]" gridcol="7" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFCF8FD8CE83FFD88" box="[841,861,536,558]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
mean
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<th id="8C7E4D2DFFE90074FC61FD8CEF6EFD88" box="[976,1036,536,558]" gridcol="8" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFC61FD8CE886FD88" box="[976,996,536,558]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
s.d.
</th>
<th id="8C7E4D2DFFE90074FBF5FD8CEFE7FD88" box="[1092,1157,536,558]" gridcol="9" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFBF5FD8CEF3AFD88" box="[1092,1112,536,558]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
min
</th>
<th id="8C7E4D2DFFE90074FB0BFD8CEE62FD88" box="[1210,1280,536,558]" gridcol="10" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFB0BFD8CEFACFD88" box="[1210,1230,536,558]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
max
</th>
<th id="8C7E4D2DFFE90074FA85FD8CEE12FD88" box="[1332,1392,536,558]" gridcol="11" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFA85FD8CEE2AFD88" box="[1332,1352,536,558]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
c.v.
</th>
<th id="8C7E4D2DFFE90074FA15FD8CEEDEFD88" box="[1444,1468,536,558]" gridcol="12" gridrow="0" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFA1CFD8CEEDBFD88" box="[1453,1465,536,558]" italics="true" pageId="7" pageNumber="2414">n</emphasis>
</th>
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<td id="8C7E4D2DFFE90074FE9BFDDFEA02FDC7" box="[298,352,587,609]" gridcol="2" gridrow="1" pageId="7" pageNumber="2414">
<emphasis id="43EBFA01FFE9FF8CFEF3FDDFEA34FDC7" box="[322,342,587,609]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFD15EEDEFD3E" box="[58,1468,641,664]" gridrow="2" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFD15EBFDFD3E" box="[58,159,641,664]" gridcol="0" gridrow="2" pageId="7" pageNumber="2414">bl</th>
<td id="8C7E4D2DFFE90074FF7CFD15EA61FD3E" box="[205,259,641,664]" gridcol="1" gridrow="2" pageId="7" pageNumber="2414">408.5</td>
<td id="8C7E4D2DFFE90074FE9BFD15EA02FD3E" box="[298,352,641,664]" gridcol="2" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FE28FD15EAB9FD3E" box="[409,475,641,664]" gridcol="3" gridrow="2" pageId="7" pageNumber="2414">393.60</td>
<td id="8C7E4D2DFFE90074FDBAFD15E923FD3E" box="[523,577,641,664]" gridcol="4" gridrow="2" pageId="7" pageNumber="2414">71.24</td>
<td id="8C7E4D2DFFE90074FDDFFD15E9D2FD3E" box="[622,688,641,664]" gridcol="5" gridrow="2" pageId="7" pageNumber="2414">247.00</td>
<td id="8C7E4D2DFFE90074FD68FD15E879FD3E" box="[729,795,641,664]" gridcol="6" gridrow="2" pageId="7" pageNumber="2414">544.00</td>
<td id="8C7E4D2DFFE90074FCF8FD15E8F8FD3E" box="[841,922,641,664]" gridcol="7" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FC61FD15EF6EFD3E" box="[976,1036,641,664]" gridcol="8" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FBF5FD15EFE7FD3E" box="[1092,1157,641,664]" gridcol="9" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FB0BFD15EE62FD3E" box="[1210,1280,641,664]" gridcol="10" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FA85FD15EE12FD3E" box="[1332,1392,641,664]" gridcol="11" gridrow="2" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FA15FD15EEDEFD3E" box="[1444,1468,641,664]" gridcol="12" gridrow="2" pageId="7" pageNumber="2414">37</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFD0AEEDEFD13" box="[58,1468,670,693]" gridrow="3" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFD0AEBFDFD13" box="[58,159,670,693]" gridcol="0" gridrow="3" pageId="7" pageNumber="2414">btl</th>
<td id="8C7E4D2DFFE90074FF7CFD0AEA61FD13" box="[205,259,670,693]" gridcol="1" gridrow="3" pageId="7" pageNumber="2414">41.8</td>
<td id="8C7E4D2DFFE90074FE9BFD0AEA02FD13" box="[298,352,670,693]" gridcol="2" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FE28FD0AEAB9FD13" box="[409,475,670,693]" gridcol="3" gridrow="3" pageId="7" pageNumber="2414">41.50</td>
<td id="8C7E4D2DFFE90074FDBAFD0AE923FD13" box="[523,577,670,693]" gridcol="4" gridrow="3" pageId="7" pageNumber="2414">3.57</td>
<td id="8C7E4D2DFFE90074FDDFFD0AE9D2FD13" box="[622,688,670,693]" gridcol="5" gridrow="3" pageId="7" pageNumber="2414">33.30</td>
<td id="8C7E4D2DFFE90074FD68FD0AE879FD13" box="[729,795,670,693]" gridcol="6" gridrow="3" pageId="7" pageNumber="2414">48.40</td>
<td id="8C7E4D2DFFE90074FCF8FD0AE8F8FD13" box="[841,922,670,693]" gridcol="7" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FC61FD0AEF6EFD13" box="[976,1036,670,693]" gridcol="8" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FBF5FD0AEFE7FD13" box="[1092,1157,670,693]" gridcol="9" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FB0BFD0AEE62FD13" box="[1210,1280,670,693]" gridcol="10" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FA85FD0AEE12FD13" box="[1332,1392,670,693]" gridcol="11" gridrow="3" pageId="7" pageNumber="2414">−</td>
<td id="8C7E4D2DFFE90074FA15FD0AEEDEFD13" box="[1444,1468,670,693]" gridcol="12" gridrow="3" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFD28EEDEFD74" box="[58,1468,700,722]" gridrow="4" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFD28EBFDFD74" box="[58,159,700,722]" gridcol="0" gridrow="4" pageId="7" pageNumber="2414">btd</th>
<td id="8C7E4D2DFFE90074FF7CFD28EA61FD74" box="[205,259,700,722]" gridcol="1" gridrow="4" pageId="7" pageNumber="2414">6.5</td>
<td id="8C7E4D2DFFE90074FE9BFD28EA02FD74" box="[298,352,700,722]" gridcol="2" gridrow="4" pageId="7" pageNumber="2414">15.55</td>
<td id="8C7E4D2DFFE90074FE28FD28EAB9FD74" box="[409,475,700,722]" gridcol="3" gridrow="4" pageId="7" pageNumber="2414">4.30</td>
<td id="8C7E4D2DFFE90074FDBAFD28E923FD74" box="[523,577,700,722]" gridcol="4" gridrow="4" pageId="7" pageNumber="2414">0.96</td>
<td id="8C7E4D2DFFE90074FDDFFD28E9D2FD74" box="[622,688,700,722]" gridcol="5" gridrow="4" pageId="7" pageNumber="2414">2.60</td>
<td id="8C7E4D2DFFE90074FD68FD28E879FD74" box="[729,795,700,722]" gridcol="6" gridrow="4" pageId="7" pageNumber="2414">6.30</td>
<td id="8C7E4D2DFFE90074FCF8FD28E8F8FD74" box="[841,922,700,722]" gridcol="7" gridrow="4" pageId="7" pageNumber="2414">10.43</td>
<td id="8C7E4D2DFFE90074FC61FD28EF6EFD74" box="[976,1036,700,722]" gridcol="8" gridrow="4" pageId="7" pageNumber="2414">2.03</td>
<td id="8C7E4D2DFFE90074FBF5FD28EFE7FD74" box="[1092,1157,700,722]" gridcol="9" gridrow="4" pageId="7" pageNumber="2414">6.90</td>
<td id="8C7E4D2DFFE90074FB0BFD28EE62FD74" box="[1210,1280,700,722]" gridcol="10" gridrow="4" pageId="7" pageNumber="2414">15.40</td>
<td id="8C7E4D2DFFE90074FA85FD28EE12FD74" box="[1332,1392,700,722]" gridcol="11" gridrow="4" pageId="7" pageNumber="2414">19.43</td>
<td id="8C7E4D2DFFE90074FA15FD28EEDEFD74" box="[1444,1468,700,722]" gridcol="12" gridrow="4" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFD4DEEDEFD49" box="[58,1468,729,751]" gridrow="5" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFD4DEBFDFD49" box="[58,159,729,751]" gridcol="0" gridrow="5" pageId="7" pageNumber="2414">vl</th>
<td id="8C7E4D2DFFE90074FF7CFD4DEA61FD49" box="[205,259,729,751]" gridcol="1" gridrow="5" pageId="7" pageNumber="2414">28.2</td>
<td id="8C7E4D2DFFE90074FE9BFD4DEA02FD49" box="[298,352,729,751]" gridcol="2" gridrow="5" pageId="7" pageNumber="2414">67.46</td>
<td id="8C7E4D2DFFE90074FE28FD4DEAB9FD49" box="[409,475,729,751]" gridcol="3" gridrow="5" pageId="7" pageNumber="2414">27.20</td>
<td id="8C7E4D2DFFE90074FDBAFD4DE923FD49" box="[523,577,729,751]" gridcol="4" gridrow="5" pageId="7" pageNumber="2414">3.35</td>
<td id="8C7E4D2DFFE90074FDDFFD4DE9D2FD49" box="[622,688,729,751]" gridcol="5" gridrow="5" pageId="7" pageNumber="2414">20.60</td>
<td id="8C7E4D2DFFE90074FD68FD4DE879FD49" box="[729,795,729,751]" gridcol="6" gridrow="5" pageId="7" pageNumber="2414">33.30</td>
<td id="8C7E4D2DFFE90074FCF8FD4DE8F8FD49" box="[841,922,729,751]" gridcol="7" gridrow="5" pageId="7" pageNumber="2414">64.88</td>
<td id="8C7E4D2DFFE90074FC61FD4DEF6EFD49" box="[976,1036,729,751]" gridcol="8" gridrow="5" pageId="7" pageNumber="2414">3.01</td>
<td id="8C7E4D2DFFE90074FBF5FD4DEFE7FD49" box="[1092,1157,729,751]" gridcol="9" gridrow="5" pageId="7" pageNumber="2414">56.90</td>
<td id="8C7E4D2DFFE90074FB0BFD4DEE62FD49" box="[1210,1280,729,751]" gridcol="10" gridrow="5" pageId="7" pageNumber="2414">70.00</td>
<td id="8C7E4D2DFFE90074FA85FD4DEE12FD49" box="[1332,1392,729,751]" gridcol="11" gridrow="5" pageId="7" pageNumber="2414">4.63</td>
<td id="8C7E4D2DFFE90074FA15FD4DEEDEFD49" box="[1444,1468,729,751]" gridcol="12" gridrow="5" pageId="7" pageNumber="2414">20</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFD62EEDEFCAA" box="[58,1468,758,780]" gridrow="6" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFD62EBFDFCAA" box="[58,159,758,780]" gridcol="0" gridrow="6" pageId="7" pageNumber="2414">ssi</th>
<td id="8C7E4D2DFFE90074FF7CFD62EA61FCAA" box="[205,259,758,780]" gridcol="1" gridrow="6" pageId="7" pageNumber="2414">32.1</td>
<td id="8C7E4D2DFFE90074FE9BFD62EA02FCAA" box="[298,352,758,780]" gridcol="2" gridrow="6" pageId="7" pageNumber="2414">76.79</td>
<td id="8C7E4D2DFFE90074FE28FD62EAB9FCAA" box="[409,475,758,780]" gridcol="3" gridrow="6" pageId="7" pageNumber="2414">31.80</td>
<td id="8C7E4D2DFFE90074FDBAFD62E923FCAA" box="[523,577,758,780]" gridcol="4" gridrow="6" pageId="7" pageNumber="2414">2.78</td>
<td id="8C7E4D2DFFE90074FDDFFD62E9D2FCAA" box="[622,688,758,780]" gridcol="5" gridrow="6" pageId="7" pageNumber="2414">26.20</td>
<td id="8C7E4D2DFFE90074FD68FD62E879FCAA" box="[729,795,758,780]" gridcol="6" gridrow="6" pageId="7" pageNumber="2414">38.00</td>
<td id="8C7E4D2DFFE90074FCF8FD62E8F8FCAA" box="[841,922,758,780]" gridcol="7" gridrow="6" pageId="7" pageNumber="2414">76.72</td>
<td id="8C7E4D2DFFE90074FC61FD62EF6EFCAA" box="[976,1036,758,780]" gridcol="8" gridrow="6" pageId="7" pageNumber="2414">2.28</td>
<td id="8C7E4D2DFFE90074FBF5FD62EFE7FCAA" box="[1092,1157,758,780]" gridcol="9" gridrow="6" pageId="7" pageNumber="2414">69.60</td>
<td id="8C7E4D2DFFE90074FB0BFD62EE62FCAA" box="[1210,1280,758,780]" gridcol="10" gridrow="6" pageId="7" pageNumber="2414">80.00</td>
<td id="8C7E4D2DFFE90074FA85FD62EE12FCAA" box="[1332,1392,758,780]" gridcol="11" gridrow="6" pageId="7" pageNumber="2414">2.97</td>
<td id="8C7E4D2DFFE90074FA15FD62EEDEFCAA" box="[1444,1468,758,780]" gridcol="12" gridrow="6" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC87EEDEFC8F" box="[58,1468,787,809]" gridrow="7" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC87EBFDFC8F" box="[58,159,787,809]" gridcol="0" gridrow="7" pageId="7" pageNumber="2414">plr</th>
<td id="8C7E4D2DFFE90074FF7CFC87EA61FC8F" box="[205,259,787,809]" gridcol="1" gridrow="7" pageId="7" pageNumber="2414">17.1</td>
<td id="8C7E4D2DFFE90074FE9BFC87EA02FC8F" box="[298,352,787,809]" gridcol="2" gridrow="7" pageId="7" pageNumber="2414">40.91</td>
<td id="8C7E4D2DFFE90074FE28FC87EAB9FC8F" box="[409,475,787,809]" gridcol="3" gridrow="7" pageId="7" pageNumber="2414">15.90</td>
<td id="8C7E4D2DFFE90074FDBAFC87E923FC8F" box="[523,577,787,809]" gridcol="4" gridrow="7" pageId="7" pageNumber="2414">2.10</td>
<td id="8C7E4D2DFFE90074FDDFFC87E9D2FC8F" box="[622,688,787,809]" gridcol="5" gridrow="7" pageId="7" pageNumber="2414">10.90</td>
<td id="8C7E4D2DFFE90074FD68FC87E879FC8F" box="[729,795,787,809]" gridcol="6" gridrow="7" pageId="7" pageNumber="2414">21.40</td>
<td id="8C7E4D2DFFE90074FCF8FC87E8F8FC8F" box="[841,922,787,809]" gridcol="7" gridrow="7" pageId="7" pageNumber="2414">38.29</td>
<td id="8C7E4D2DFFE90074FC61FC87EF6EFC8F" box="[976,1036,787,809]" gridcol="8" gridrow="7" pageId="7" pageNumber="2414">3.38</td>
<td id="8C7E4D2DFFE90074FBF5FC87EFE7FC8F" box="[1092,1157,787,809]" gridcol="9" gridrow="7" pageId="7" pageNumber="2414">30.90</td>
<td id="8C7E4D2DFFE90074FB0BFC87EE62FC8F" box="[1210,1280,787,809]" gridcol="10" gridrow="7" pageId="7" pageNumber="2414">45.30</td>
<td id="8C7E4D2DFFE90074FA85FC87EE12FC8F" box="[1332,1392,787,809]" gridcol="11" gridrow="7" pageId="7" pageNumber="2414">8.84</td>
<td id="8C7E4D2DFFE90074FA15FC87EEDEFC8F" box="[1444,1468,787,809]" gridcol="12" gridrow="7" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFCBAEEDEFCE1" box="[58,1468,814,839]" gridrow="8" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFCBAEBFDFCE1" box="[58,159,814,839]" gridcol="0" gridrow="8" pageId="7" pageNumber="2414">1◦pl</th>
<td id="8C7E4D2DFFE90074FF7CFCBAEA61FCE1" box="[205,259,814,839]" gridcol="1" gridrow="8" pageId="7" pageNumber="2414">8.1</td>
<td id="8C7E4D2DFFE90074FE9BFCBAEA02FCE1" box="[298,352,814,839]" gridcol="2" gridrow="8" pageId="7" pageNumber="2414">19.38</td>
<td id="8C7E4D2DFFE90074FE28FCBAEAB9FCE1" box="[409,475,814,839]" gridcol="3" gridrow="8" pageId="7" pageNumber="2414">8.80</td>
<td id="8C7E4D2DFFE90074FDBAFCBAE923FCE1" box="[523,577,814,839]" gridcol="4" gridrow="8" pageId="7" pageNumber="2414">1.30</td>
<td id="8C7E4D2DFFE90074FDDFFCBAE9D2FCE1" box="[622,688,814,839]" gridcol="5" gridrow="8" pageId="7" pageNumber="2414">6.20</td>
<td id="8C7E4D2DFFE90074FD68FCBAE879FCE1" box="[729,795,814,839]" gridcol="6" gridrow="8" pageId="7" pageNumber="2414">11.90</td>
<td id="8C7E4D2DFFE90074FCF8FCBAE8F8FCE1" box="[841,922,814,839]" gridcol="7" gridrow="8" pageId="7" pageNumber="2414">20.70</td>
<td id="8C7E4D2DFFE90074FC61FCBAEF6EFCE1" box="[976,1036,814,839]" gridcol="8" gridrow="8" pageId="7" pageNumber="2414">2.43</td>
<td id="8C7E4D2DFFE90074FBF5FCBAEFE7FCE1" box="[1092,1157,814,839]" gridcol="9" gridrow="8" pageId="7" pageNumber="2414">14.70</td>
<td id="8C7E4D2DFFE90074FB0BFCBAEE62FCE1" box="[1210,1280,814,839]" gridcol="10" gridrow="8" pageId="7" pageNumber="2414">24.60</td>
<td id="8C7E4D2DFFE90074FA85FCBAEE12FCE1" box="[1332,1392,814,839]" gridcol="11" gridrow="8" pageId="7" pageNumber="2414">11.76</td>
<td id="8C7E4D2DFFE90074FA15FCBAEEDEFCE1" box="[1444,1468,814,839]" gridcol="12" gridrow="8" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFCDFEEDEFCC2" box="[58,1468,843,868]" gridrow="9" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFCDFEBFDFCC2" box="[58,159,843,868]" gridcol="0" gridrow="9" pageId="7" pageNumber="2414">2◦pl</th>
<td id="8C7E4D2DFFE90074FF7CFCDFEA61FCC2" box="[205,259,843,868]" gridcol="1" gridrow="9" pageId="7" pageNumber="2414">6.1</td>
<td id="8C7E4D2DFFE90074FE9BFCDFEA02FCC2" box="[298,352,843,868]" gridcol="2" gridrow="9" pageId="7" pageNumber="2414">14.59</td>
<td id="8C7E4D2DFFE90074FE28FCDFEAB9FCC2" box="[409,475,843,868]" gridcol="3" gridrow="9" pageId="7" pageNumber="2414">5.40</td>
<td id="8C7E4D2DFFE90074FDBAFCDFE923FCC2" box="[523,577,843,868]" gridcol="4" gridrow="9" pageId="7" pageNumber="2414">0.97</td>
<td id="8C7E4D2DFFE90074FDDFFCDFE9D2FCC2" box="[622,688,843,868]" gridcol="5" gridrow="9" pageId="7" pageNumber="2414">3.60</td>
<td id="8C7E4D2DFFE90074FD68FCDFE879FCC2" box="[729,795,843,868]" gridcol="6" gridrow="9" pageId="7" pageNumber="2414">7.10</td>
<td id="8C7E4D2DFFE90074FCF8FCDFE8F8FCC2" box="[841,922,843,868]" gridcol="7" gridrow="9" pageId="7" pageNumber="2414">12.91</td>
<td id="8C7E4D2DFFE90074FC61FCDFEF6EFCC2" box="[976,1036,843,868]" gridcol="8" gridrow="9" pageId="7" pageNumber="2414">1.77</td>
<td id="8C7E4D2DFFE90074FBF5FCDFEFE7FCC2" box="[1092,1157,843,868]" gridcol="9" gridrow="9" pageId="7" pageNumber="2414">9.60</td>
<td id="8C7E4D2DFFE90074FB0BFCDFEE62FCC2" box="[1210,1280,843,868]" gridcol="10" gridrow="9" pageId="7" pageNumber="2414">17.00</td>
<td id="8C7E4D2DFFE90074FA85FCDFEE12FCC2" box="[1332,1392,843,868]" gridcol="11" gridrow="9" pageId="7" pageNumber="2414">13.73</td>
<td id="8C7E4D2DFFE90074FA15FCDFEEDEFCC2" box="[1444,1468,843,868]" gridcol="12" gridrow="9" pageId="7" pageNumber="2414">35</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFCFFEEDEFC27" box="[58,1468,875,897]" gridrow="10" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFCFFEBFDFC27" box="[58,159,875,897]" gridcol="0" gridrow="10" pageId="7" pageNumber="2414">mpl</th>
<td id="8C7E4D2DFFE90074FF7CFCFFEA61FC27" box="[205,259,875,897]" gridcol="1" gridrow="10" pageId="7" pageNumber="2414">1.9</td>
<td id="8C7E4D2DFFE90074FE9BFCFFEA02FC27" box="[298,352,875,897]" gridcol="2" gridrow="10" pageId="7" pageNumber="2414">4.55</td>
<td id="8C7E4D2DFFE90074FE28FCFFEAB9FC27" box="[409,475,875,897]" gridcol="3" gridrow="10" pageId="7" pageNumber="2414">2.50</td>
<td id="8C7E4D2DFFE90074FDBAFCFFE923FC27" box="[523,577,875,897]" gridcol="4" gridrow="10" pageId="7" pageNumber="2414">0.57</td>
<td id="8C7E4D2DFFE90074FDDFFCFFE9D2FC27" box="[622,688,875,897]" gridcol="5" gridrow="10" pageId="7" pageNumber="2414">1.27</td>
<td id="8C7E4D2DFFE90074FD68FCFFE879FC27" box="[729,795,875,897]" gridcol="6" gridrow="10" pageId="7" pageNumber="2414">3.49</td>
<td id="8C7E4D2DFFE90074FCF8FCFFE8F8FC27" box="[841,922,875,897]" gridcol="7" gridrow="10" pageId="7" pageNumber="2414">6.06</td>
<td id="8C7E4D2DFFE90074FC61FCFFEF6EFC27" box="[976,1036,875,897]" gridcol="8" gridrow="10" pageId="7" pageNumber="2414">1.26</td>
<td id="8C7E4D2DFFE90074FBF5FCFFEFE7FC27" box="[1092,1157,875,897]" gridcol="9" gridrow="10" pageId="7" pageNumber="2414">3.10</td>
<td id="8C7E4D2DFFE90074FB0BFCFFEE62FC27" box="[1210,1280,875,897]" gridcol="10" gridrow="10" pageId="7" pageNumber="2414">8.30</td>
<td id="8C7E4D2DFFE90074FA85FCFFEE12FC27" box="[1332,1392,875,897]" gridcol="11" gridrow="10" pageId="7" pageNumber="2414">20.81</td>
<td id="8C7E4D2DFFE90074FA15FCFFEEDEFC27" box="[1444,1468,875,897]" gridcol="12" gridrow="10" pageId="7" pageNumber="2414">34</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC1CEEDEFC38" box="[58,1468,904,926]" gridrow="11" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC1CEBFDFC38" box="[58,159,904,926]" gridcol="0" gridrow="11" pageId="7" pageNumber="2414">pb1</th>
<td id="8C7E4D2DFFE90074FF7CFC1CEA61FC38" box="[205,259,904,926]" gridcol="1" gridrow="11" pageId="7" pageNumber="2414">11.5</td>
<td id="8C7E4D2DFFE90074FE9BFC1CEA02FC38" box="[298,352,904,926]" gridcol="2" gridrow="11" pageId="7" pageNumber="2414">27.51</td>
<td id="8C7E4D2DFFE90074FE28FC1CEAB9FC38" box="[409,475,904,926]" gridcol="3" gridrow="11" pageId="7" pageNumber="2414">13.60</td>
<td id="8C7E4D2DFFE90074FDBAFC1CE923FC38" box="[523,577,904,926]" gridcol="4" gridrow="11" pageId="7" pageNumber="2414">1.86</td>
<td id="8C7E4D2DFFE90074FDDFFC1CE9D2FC38" box="[622,688,904,926]" gridcol="5" gridrow="11" pageId="7" pageNumber="2414">9.50</td>
<td id="8C7E4D2DFFE90074FD68FC1CE879FC38" box="[729,795,904,926]" gridcol="6" gridrow="11" pageId="7" pageNumber="2414">18.70</td>
<td id="8C7E4D2DFFE90074FCF8FC1CE8F8FC38" box="[841,922,904,926]" gridcol="7" gridrow="11" pageId="7" pageNumber="2414">32.90</td>
<td id="8C7E4D2DFFE90074FC61FC1CEF6EFC38" box="[976,1036,904,926]" gridcol="8" gridrow="11" pageId="7" pageNumber="2414">4.22</td>
<td id="8C7E4D2DFFE90074FBF5FC1CEFE7FC38" box="[1092,1157,904,926]" gridcol="9" gridrow="11" pageId="7" pageNumber="2414">23.50</td>
<td id="8C7E4D2DFFE90074FB0BFC1CEE62FC38" box="[1210,1280,904,926]" gridcol="10" gridrow="11" pageId="7" pageNumber="2414">42.00</td>
<td id="8C7E4D2DFFE90074FA85FC1CEE12FC38" box="[1332,1392,904,926]" gridcol="11" gridrow="11" pageId="7" pageNumber="2414">12.86</td>
<td id="8C7E4D2DFFE90074FA15FC1CEEDEFC38" box="[1444,1468,904,926]" gridcol="12" gridrow="11" pageId="7" pageNumber="2414">27</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC31EEDEFC1D" box="[58,1468,933,955]" gridrow="12" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC31EBFDFC1D" box="[58,159,933,955]" gridcol="0" gridrow="12" pageId="7" pageNumber="2414">sb1</th>
<td id="8C7E4D2DFFE90074FF7CFC31EA61FC1D" box="[205,259,933,955]" gridcol="1" gridrow="12" pageId="7" pageNumber="2414">9.6</td>
<td id="8C7E4D2DFFE90074FE9BFC31EA02FC1D" box="[298,352,933,955]" gridcol="2" gridrow="12" pageId="7" pageNumber="2414">22.97</td>
<td id="8C7E4D2DFFE90074FE28FC31EAB9FC1D" box="[409,475,933,955]" gridcol="3" gridrow="12" pageId="7" pageNumber="2414">10.70</td>
<td id="8C7E4D2DFFE90074FDBAFC31E923FC1D" box="[523,577,933,955]" gridcol="4" gridrow="12" pageId="7" pageNumber="2414">2.45</td>
<td id="8C7E4D2DFFE90074FDDFFC31E9D2FC1D" box="[622,688,933,955]" gridcol="5" gridrow="12" pageId="7" pageNumber="2414">8.30</td>
<td id="8C7E4D2DFFE90074FD68FC31E879FC1D" box="[729,795,933,955]" gridcol="6" gridrow="12" pageId="7" pageNumber="2414">17.70</td>
<td id="8C7E4D2DFFE90074FCF8FC31E8F8FC1D" box="[841,922,933,955]" gridcol="7" gridrow="12" pageId="7" pageNumber="2414">26.33</td>
<td id="8C7E4D2DFFE90074FC61FC31EF6EFC1D" box="[976,1036,933,955]" gridcol="8" gridrow="12" pageId="7" pageNumber="2414">5.47</td>
<td id="8C7E4D2DFFE90074FBF5FC31EFE7FC1D" box="[1092,1157,933,955]" gridcol="9" gridrow="12" pageId="7" pageNumber="2414">20.00</td>
<td id="8C7E4D2DFFE90074FB0BFC31EE62FC1D" box="[1210,1280,933,955]" gridcol="10" gridrow="12" pageId="7" pageNumber="2414">38.60</td>
<td id="8C7E4D2DFFE90074FA85FC31EE12FC1D" box="[1332,1392,933,955]" gridcol="11" gridrow="12" pageId="7" pageNumber="2414">20.76</td>
<td id="8C7E4D2DFFE90074FA15FC31EEDEFC1D" box="[1444,1468,933,955]" gridcol="12" gridrow="12" pageId="7" pageNumber="2414">15</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC57EEDEFC7F" box="[58,1468,963,985]" gridrow="13" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC57EBFDFC7F" box="[58,159,963,985]" gridcol="0" gridrow="13" pageId="7" pageNumber="2414">pb2</th>
<td id="8C7E4D2DFFE90074FF7CFC57EA61FC7F" box="[205,259,963,985]" gridcol="1" gridrow="13" pageId="7" pageNumber="2414">13.4</td>
<td id="8C7E4D2DFFE90074FE9BFC57EA02FC7F" box="[298,352,963,985]" gridcol="2" gridrow="13" pageId="7" pageNumber="2414">32.06</td>
<td id="8C7E4D2DFFE90074FE28FC57EAB9FC7F" box="[409,475,963,985]" gridcol="3" gridrow="13" pageId="7" pageNumber="2414">14.50</td>
<td id="8C7E4D2DFFE90074FDBAFC57E923FC7F" box="[523,577,963,985]" gridcol="4" gridrow="13" pageId="7" pageNumber="2414">1.83</td>
<td id="8C7E4D2DFFE90074FDDFFC57E9D2FC7F" box="[622,688,963,985]" gridcol="5" gridrow="13" pageId="7" pageNumber="2414">10.80</td>
<td id="8C7E4D2DFFE90074FD68FC57E879FC7F" box="[729,795,963,985]" gridcol="6" gridrow="13" pageId="7" pageNumber="2414">18.70</td>
<td id="8C7E4D2DFFE90074FCF8FC57E8F8FC7F" box="[841,922,963,985]" gridcol="7" gridrow="13" pageId="7" pageNumber="2414">34.92</td>
<td id="8C7E4D2DFFE90074FC61FC57EF6EFC7F" box="[976,1036,963,985]" gridcol="8" gridrow="13" pageId="7" pageNumber="2414">5.02</td>
<td id="8C7E4D2DFFE90074FBF5FC57EFE7FC7F" box="[1092,1157,963,985]" gridcol="9" gridrow="13" pageId="7" pageNumber="2414">22.40</td>
<td id="8C7E4D2DFFE90074FB0BFC57EE62FC7F" box="[1210,1280,963,985]" gridcol="10" gridrow="13" pageId="7" pageNumber="2414">43.50</td>
<td id="8C7E4D2DFFE90074FA85FC57EE12FC7F" box="[1332,1392,963,985]" gridcol="11" gridrow="13" pageId="7" pageNumber="2414">14.38</td>
<td id="8C7E4D2DFFE90074FA15FC57EEDEFC7F" box="[1444,1468,963,985]" gridcol="12" gridrow="13" pageId="7" pageNumber="2414">27</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC74EEDEFC50" box="[58,1468,992,1014]" gridrow="14" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC74EBFDFC50" box="[58,159,992,1014]" gridcol="0" gridrow="14" pageId="7" pageNumber="2414">sb2</th>
<td id="8C7E4D2DFFE90074FF7CFC74EA61FC50" box="[205,259,992,1014]" gridcol="1" gridrow="14" pageId="7" pageNumber="2414">9.8</td>
<td id="8C7E4D2DFFE90074FE9BFC74EA02FC50" box="[298,352,992,1014]" gridcol="2" gridrow="14" pageId="7" pageNumber="2414">23.44</td>
<td id="8C7E4D2DFFE90074FE28FC74EAB9FC50" box="[409,475,992,1014]" gridcol="3" gridrow="14" pageId="7" pageNumber="2414">11.50</td>
<td id="8C7E4D2DFFE90074FDBAFC74E923FC50" box="[523,577,992,1014]" gridcol="4" gridrow="14" pageId="7" pageNumber="2414">1.32</td>
<td id="8C7E4D2DFFE90074FDDFFC74E9D2FC50" box="[622,688,992,1014]" gridcol="5" gridrow="14" pageId="7" pageNumber="2414">8.90</td>
<td id="8C7E4D2DFFE90074FD68FC74E879FC50" box="[729,795,992,1014]" gridcol="6" gridrow="14" pageId="7" pageNumber="2414">13.00</td>
<td id="8C7E4D2DFFE90074FCF8FC74E8F8FC50" box="[841,922,992,1014]" gridcol="7" gridrow="14" pageId="7" pageNumber="2414">28.41</td>
<td id="8C7E4D2DFFE90074FC61FC74EF6EFC50" box="[976,1036,992,1014]" gridcol="8" gridrow="14" pageId="7" pageNumber="2414">3.96</td>
<td id="8C7E4D2DFFE90074FBF5FC74EFE7FC50" box="[1092,1157,992,1014]" gridcol="9" gridrow="14" pageId="7" pageNumber="2414">22.40</td>
<td id="8C7E4D2DFFE90074FB0BFC74EE62FC50" box="[1210,1280,992,1014]" gridcol="10" gridrow="14" pageId="7" pageNumber="2414">34.80</td>
<td id="8C7E4D2DFFE90074FA85FC74EE12FC50" box="[1332,1392,992,1014]" gridcol="11" gridrow="14" pageId="7" pageNumber="2414">13.94</td>
<td id="8C7E4D2DFFE90074FA15FC74EEDEFC50" box="[1444,1468,992,1014]" gridcol="12" gridrow="14" pageId="7" pageNumber="2414">14</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFC69EEDEFBB5" box="[58,1468,1021,1043]" gridrow="15" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFC69EBFDFBB5" box="[58,159,1021,1043]" gridcol="0" gridrow="15" pageId="7" pageNumber="2414">pb3</th>
<td id="8C7E4D2DFFE90074FF7CFC69EA61FBB5" box="[205,259,1021,1043]" gridcol="1" gridrow="15" pageId="7" pageNumber="2414">13.4</td>
<td id="8C7E4D2DFFE90074FE9BFC69EA02FBB5" box="[298,352,1021,1043]" gridcol="2" gridrow="15" pageId="7" pageNumber="2414">32.06</td>
<td id="8C7E4D2DFFE90074FE28FC69EAB9FBB5" box="[409,475,1021,1043]" gridcol="3" gridrow="15" pageId="7" pageNumber="2414">14.60</td>
<td id="8C7E4D2DFFE90074FDBAFC69E923FBB5" box="[523,577,1021,1043]" gridcol="4" gridrow="15" pageId="7" pageNumber="2414">1.74</td>
<td id="8C7E4D2DFFE90074FDDFFC69E9D2FBB5" box="[622,688,1021,1043]" gridcol="5" gridrow="15" pageId="7" pageNumber="2414">10.80</td>
<td id="8C7E4D2DFFE90074FD68FC69E879FBB5" box="[729,795,1021,1043]" gridcol="6" gridrow="15" pageId="7" pageNumber="2414">17.20</td>
<td id="8C7E4D2DFFE90074FCF8FC69E8F8FBB5" box="[841,922,1021,1043]" gridcol="7" gridrow="15" pageId="7" pageNumber="2414">35.19</td>
<td id="8C7E4D2DFFE90074FC61FC69EF6EFBB5" box="[976,1036,1021,1043]" gridcol="8" gridrow="15" pageId="7" pageNumber="2414">4.35</td>
<td id="8C7E4D2DFFE90074FBF5FC69EFE7FBB5" box="[1092,1157,1021,1043]" gridcol="9" gridrow="15" pageId="7" pageNumber="2414">27.50</td>
<td id="8C7E4D2DFFE90074FB0BFC69EE62FBB5" box="[1210,1280,1021,1043]" gridcol="10" gridrow="15" pageId="7" pageNumber="2414">46.40</td>
<td id="8C7E4D2DFFE90074FA85FC69EE12FBB5" box="[1332,1392,1021,1043]" gridcol="11" gridrow="15" pageId="7" pageNumber="2414">12.35</td>
<td id="8C7E4D2DFFE90074FA15FC69EEDEFBB5" box="[1444,1468,1021,1043]" gridcol="12" gridrow="15" pageId="7" pageNumber="2414">34</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFB8EEEDEFB96" box="[58,1468,1050,1072]" gridrow="16" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFB8EEBFDFB96" box="[58,159,1050,1072]" gridcol="0" gridrow="16" pageId="7" pageNumber="2414">sb3</th>
<td id="8C7E4D2DFFE90074FF7CFB8EEA61FB96" box="[205,259,1050,1072]" gridcol="1" gridrow="16" pageId="7" pageNumber="2414">11.1</td>
<td id="8C7E4D2DFFE90074FE9BFB8EEA02FB96" box="[298,352,1050,1072]" gridcol="2" gridrow="16" pageId="7" pageNumber="2414">26.56</td>
<td id="8C7E4D2DFFE90074FE28FB8EEAB9FB96" box="[409,475,1050,1072]" gridcol="3" gridrow="16" pageId="7" pageNumber="2414">11.00</td>
<td id="8C7E4D2DFFE90074FDBAFB8EE923FB96" box="[523,577,1050,1072]" gridcol="4" gridrow="16" pageId="7" pageNumber="2414">1.61</td>
<td id="8C7E4D2DFFE90074FDDFFB8EE9D2FB96" box="[622,688,1050,1072]" gridcol="5" gridrow="16" pageId="7" pageNumber="2414">8.30</td>
<td id="8C7E4D2DFFE90074FD68FB8EE879FB96" box="[729,795,1050,1072]" gridcol="6" gridrow="16" pageId="7" pageNumber="2414">14.10</td>
<td id="8C7E4D2DFFE90074FCF8FB8EE8F8FB96" box="[841,922,1050,1072]" gridcol="7" gridrow="16" pageId="7" pageNumber="2414">27.21</td>
<td id="8C7E4D2DFFE90074FC61FB8EEF6EFB96" box="[976,1036,1050,1072]" gridcol="8" gridrow="16" pageId="7" pageNumber="2414">3.74</td>
<td id="8C7E4D2DFFE90074FBF5FB8EEFE7FB96" box="[1092,1157,1050,1072]" gridcol="9" gridrow="16" pageId="7" pageNumber="2414">20.50</td>
<td id="8C7E4D2DFFE90074FB0BFB8EEE62FB96" box="[1210,1280,1050,1072]" gridcol="10" gridrow="16" pageId="7" pageNumber="2414">33.70</td>
<td id="8C7E4D2DFFE90074FA85FB8EEE12FB96" box="[1332,1392,1050,1072]" gridcol="11" gridrow="16" pageId="7" pageNumber="2414">13.75</td>
<td id="8C7E4D2DFFE90074FA15FB8EEEDEFB96" box="[1444,1468,1050,1072]" gridcol="12" gridrow="16" pageId="7" pageNumber="2414">14</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFBACEEDEFBE8" box="[58,1468,1080,1102]" gridrow="17" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFBACEBFDFBE8" box="[58,159,1080,1102]" gridcol="0" gridrow="17" pageId="7" pageNumber="2414">pb4</th>
<td id="8C7E4D2DFFE90074FF7CFBACEA61FBE8" box="[205,259,1080,1102]" gridcol="1" gridrow="17" pageId="7" pageNumber="2414">15.8</td>
<td id="8C7E4D2DFFE90074FE9BFBACEA02FBE8" box="[298,352,1080,1102]" gridcol="2" gridrow="17" pageId="7" pageNumber="2414">37.80</td>
<td id="8C7E4D2DFFE90074FE28FBACEAB9FBE8" box="[409,475,1080,1102]" gridcol="3" gridrow="17" pageId="7" pageNumber="2414">16.90</td>
<td id="8C7E4D2DFFE90074FDBAFBACE923FBE8" box="[523,577,1080,1102]" gridcol="4" gridrow="17" pageId="7" pageNumber="2414">2.05</td>
<td id="8C7E4D2DFFE90074FDDFFBACE9D2FBE8" box="[622,688,1080,1102]" gridcol="5" gridrow="17" pageId="7" pageNumber="2414">12.50</td>
<td id="8C7E4D2DFFE90074FD68FBACE879FBE8" box="[729,795,1080,1102]" gridcol="6" gridrow="17" pageId="7" pageNumber="2414">20.80</td>
<td id="8C7E4D2DFFE90074FCF8FBACE8F8FBE8" box="[841,922,1080,1102]" gridcol="7" gridrow="17" pageId="7" pageNumber="2414">41.01</td>
<td id="8C7E4D2DFFE90074FC61FBACEF6EFBE8" box="[976,1036,1080,1102]" gridcol="8" gridrow="17" pageId="7" pageNumber="2414">5.09</td>
<td id="8C7E4D2DFFE90074FBF5FBACEFE7FBE8" box="[1092,1157,1080,1102]" gridcol="9" gridrow="17" pageId="7" pageNumber="2414">31.40</td>
<td id="8C7E4D2DFFE90074FB0BFBACEE62FBE8" box="[1210,1280,1080,1102]" gridcol="10" gridrow="17" pageId="7" pageNumber="2414">50.00</td>
<td id="8C7E4D2DFFE90074FA85FBACEE12FBE8" box="[1332,1392,1080,1102]" gridcol="11" gridrow="17" pageId="7" pageNumber="2414">12.41</td>
<td id="8C7E4D2DFFE90074FA15FBACEEDEFBE8" box="[1444,1468,1080,1102]" gridcol="12" gridrow="17" pageId="7" pageNumber="2414">33</td>
</tr>
<tr id="CFAF2451FFE90074FF8BFBC1EEDEFBCD" box="[58,1468,1109,1131]" gridrow="18" pageId="7" pageNumber="2414">
<th id="8C7E4D2DFFE90074FF8BFBC1EBFDFBCD" box="[58,159,1109,1131]" gridcol="0" gridrow="18" pageId="7" pageNumber="2414">sb4</th>
<td id="8C7E4D2DFFE90074FF7CFBC1EA61FBCD" box="[205,259,1109,1131]" gridcol="1" gridrow="18" pageId="7" pageNumber="2414">11.9</td>
<td id="8C7E4D2DFFE90074FE9BFBC1EA02FBCD" box="[298,352,1109,1131]" gridcol="2" gridrow="18" pageId="7" pageNumber="2414">28.47</td>
<td id="8C7E4D2DFFE90074FE28FBC1EAB9FBCD" box="[409,475,1109,1131]" gridcol="3" gridrow="18" pageId="7" pageNumber="2414">11.70</td>
<td id="8C7E4D2DFFE90074FDBAFBC1E923FBCD" box="[523,577,1109,1131]" gridcol="4" gridrow="18" pageId="7" pageNumber="2414">2.04</td>
<td id="8C7E4D2DFFE90074FDDFFBC1E9D2FBCD" box="[622,688,1109,1131]" gridcol="5" gridrow="18" pageId="7" pageNumber="2414">8.30</td>
<td id="8C7E4D2DFFE90074FD68FBC1E879FBCD" box="[729,795,1109,1131]" gridcol="6" gridrow="18" pageId="7" pageNumber="2414">16.70</td>
<td id="8C7E4D2DFFE90074FCF8FBC1E8F8FBCD" box="[841,922,1109,1131]" gridcol="7" gridrow="18" pageId="7" pageNumber="2414">28.51</td>
<td id="8C7E4D2DFFE90074FC61FBC1EF6EFBCD" box="[976,1036,1109,1131]" gridcol="8" gridrow="18" pageId="7" pageNumber="2414">4.23</td>
<td id="8C7E4D2DFFE90074FBF5FBC1EFE7FBCD" box="[1092,1157,1109,1131]" gridcol="9" gridrow="18" pageId="7" pageNumber="2414">22.20</td>
<td id="8C7E4D2DFFE90074FB0BFBC1EE62FBCD" box="[1210,1280,1109,1131]" gridcol="10" gridrow="18" pageId="7" pageNumber="2414">36.40</td>
<td id="8C7E4D2DFFE90074FA85FBC1EE12FBCD" box="[1332,1392,1109,1131]" gridcol="11" gridrow="18" pageId="7" pageNumber="2414">14.83</td>
<td id="8C7E4D2DFFE90074FA15FBC1EEDEFBCD" box="[1444,1468,1109,1131]" gridcol="12" gridrow="18" pageId="7" pageNumber="2414">15</td>
</tr>
</table>
</paragraph>
<paragraph id="71202613FFE9FF8CFF8BFB10E978FAA9" blockId="7.[58,1492,1156,1295]" pageId="7" pageNumber="2414">
<tableNote id="8C79279DFFE9FF8CFF8BFB10E978FAA9" pageId="7" pageNumber="2414" targetBox="[58,1468,536,1131]" targetPageId="7">
Notes: bl = body length; btl = buccal tube length; btd = diameter of the buccal tube; vl = ventral lamina length; ssi = insertion of stylet supports; plr = macroplacoid row length; 1 
<superScript id="86EA8B5BFFE9FF8CFE46FB0AEA9DFB0A" attach="right" box="[503,511,1182,1196]" fontSize="6" pageId="7" pageNumber="2414">◦</superScript>
pl = first macroplacoid length; 2 
<superScript id="86EA8B5BFFE9FF8CFCFEFB0AE835FB0A" attach="right" box="[847,855,1182,1196]" fontSize="6" pageId="7" pageNumber="2414">◦</superScript>
pl = second macroplacoid length; mpl = microplacoid length; pb1-4 = length of primary branch of the claw on first-fourth leg pair; sb1-4 length of secondary the claw on first-fourth leg pair; 
<emphasis id="43EBFA01FFE9FF8CFAFEFB2BEE01FB73" box="[1359,1379,1215,1237]" italics="true" pageId="7" pageNumber="2414">pt</emphasis>
= index pt (length of a structure 
<emphasis id="43EBFA01FFE9FF8CFEBFFB48EA7BFB54" bold="true" box="[270,281,1244,1266]" italics="true" pageId="7" pageNumber="2414">/</emphasis>
buccal tube length 
<superScript id="86EA8B5BFFE9FF8CFE62FB4DEAB9FB41" attach="left" box="[467,475,1241,1255]" fontSize="6" pageId="7" pageNumber="2414">∗</superScript>
100; 
<bibRefCitation id="150E5BE2FFE9FF8CFDBCFB48E9E6FB54" author="Pilato G" box="[525,644,1244,1266]" pageId="7" pageNumber="2414" pagination="51 - 57" refId="ref6856" refString="Pilato G. 1981. Analisi di nuovi caratteri nello studio degli Eutardigradi. Animalia. 8: 51 - 57." type="journal article" year="1981">Pilato, 1981</bibRefCitation>
); min = minimum; max = maximum; SD = standard deviation; c.v. = coefficient of variation; 
<emphasis id="43EBFA01FFE9FF8CFF10FB6DEBCFFAA9" box="[161,173,1273,1295]" italics="true" pageId="7" pageNumber="2414">n</emphasis>
= number of considered specimens.
</tableNote>
</paragraph>
<subSubSection id="39857598FFE6FF83FF62FF3CEAF7FE21" pageId="8" pageNumber="2415" type="diagnosis">
<paragraph id="71202613FFE6FF83FF62FF3CEA5DFF19" blockId="8.[211,1172,168,391]" box="[211,319,168,191]" pageId="8" pageNumber="2415">
<heading id="2A68917FFFE6FF83FF62FF3CEA5DFF19" box="[211,319,168,191]" fontSize="10" level="3" pageId="8" pageNumber="2415" reason="5">
<emphasis id="43EBFA01FFE6FF83FF62FF3CEA5DFF19" box="[211,319,168,191]" italics="true" pageId="8" pageNumber="2415">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE6FF83FF62FF5BEAF7FE21" blockId="8.[211,1172,168,391]" pageId="8" pageNumber="2415">Size medium. Cuticle with pearls (pores). Eye-spots present. Buccal tube of average width. Buccal armature with two bands of teeth and transverse ridges. Bulb with two macroplacoids and small microplacoid. Long Y-shaped claws with two short accessory points on the main branch, lunule smooth or weakly crenate in the hind legs. Free eggs with processes in shape of elongated cones with truncated and enlarged apexes. Limnoterrestrial.</paragraph>
</subSubSection>
<subSubSection id="39857598FFE6FF82FF62FE4BE81DFA4B" lastPageId="9" lastPageNumber="2416" pageId="8" pageNumber="2415" type="description">
<paragraph id="71202613FFE6FF83FF62FE4BEA30FE50" blockId="8.[211,1172,479,1691]" box="[211,338,479,502]" pageId="8" pageNumber="2415">
<heading id="2A68917FFFE6FF83FF62FE4BEA30FE50" box="[211,338,479,502]" fontSize="10" level="3" pageId="8" pageNumber="2415" reason="5">
<emphasis id="43EBFA01FFE6FF83FF62FE4BEA30FE50" box="[211,338,479,502]" italics="true" pageId="8" pageNumber="2415">Description</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE6FF83FF62FD92EF7BFB7A" blockId="8.[211,1172,479,1691]" pageId="8" pageNumber="2415">
Size range: 247–544 µm long. Colourless, yellowish, greyish or light brown, sometimes with transverse bands with brown pigment. Eye-spots present in all specimens observed (
<figureCitation id="E9A43A96FFE6FF83FEF8FDD2EACAFDF8" box="[329,424,582,606]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figure 1</figureCitation>
), eye-spot colour not detected. Cuticle smooth under LM, except small granulations on outer part of legs. Cuticular pores rounded or ovate approximately 1 µm in diameter (
<figureCitation id="E9A43A96FFE6FF83FDAEFD12E9E6FD38" box="[543,644,646,670]" captionStart="Figure 2" captionStartId="10.[211,278,1174,1196]" captionTargetBox="[213,1170,169,1142]" captionTargetId="figure-5@10.[213,1170,169,1142]" captionTargetPageId="10" captionText="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="8" pageNumber="2415">Figure 2</figureCitation>
), generally with thickened edges, irregularly distributed on the body surface (dorsally and ventrally). Near the anterior end, a ring of pores with evident thickened edges and placed in a regular manner. Ten small peribuccal lamellae surround the mouth opening, followed by six buccal sensory lobes (
<figureCitation id="E9A43A96FFE6FF83FF6DFC91EA58FCBB" box="[220,314,773,797]" captionStart="Figure 3" captionStartId="11.[143,210,948,970]" captionTargetBox="[145,1102,168,915]" captionTargetId="figure-5@11.[145,1102,168,915]" captionTargetPageId="11" captionText="Figure 3. Macrobiotus kristenseni sp. nov. (A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197691" httpUri="https://zenodo.org/record/5197691/files/figure.png" pageId="8" pageNumber="2415">Figure 3</figureCitation>
). Buccal armature, poorly distinguishable under LM, formed by an anterior band of small teeth of variable thickness at the base of the buccal lamellae, sometimes reduced to a single row of teeth (
<figureCitation id="E9A43A96FFE6FF83FDE2FCD1E9DCFCFB" box="[595,702,837,861]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figures 1</figureCitation>
and 
<figureCitation id="E9A43A96FFE6FF83FD4AFCD1E86AFCFB" box="[763,776,837,861]" captionStart="Figure 3" captionStartId="11.[143,210,948,970]" captionTargetBox="[145,1102,168,915]" captionTargetId="figure-5@11.[145,1102,168,915]" captionTargetPageId="11" captionText="Figure 3. Macrobiotus kristenseni sp. nov. (A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197691" httpUri="https://zenodo.org/record/5197691/files/figure.png" pageId="8" pageNumber="2415">3</figureCitation>
); a posterior band of larger teeth, sometimes in a single row, followed by three dorsal and three ventral fine transverse ridges (
<figureCitation id="E9A43A96FFE6FF83FE9BFC11EAF7FC3B" box="[298,405,901,925]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figures 1</figureCitation>
and 
<figureCitation id="E9A43A96FFE6FF83FE64FC11EA80FC3B" box="[469,482,901,925]" captionStart="Figure 3" captionStartId="11.[143,210,948,970]" captionTargetBox="[145,1102,168,915]" captionTargetId="figure-5@11.[145,1102,168,915]" captionTargetPageId="11" captionText="Figure 3. Macrobiotus kristenseni sp. nov. (A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197691" httpUri="https://zenodo.org/record/5197691/files/figure.png" pageId="8" pageNumber="2415">3</figureCitation>
). Ventral transversal ridges clearly smaller than the dorsal, lateral ridges larger than the middle ones (
<figureCitation id="E9A43A96FFE6FF83FD04FC31E876FC1B" box="[693,788,933,957]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figure 1</figureCitation>
). Long ventral strengthening bar, about 65% of buccal tube total length, with an anterior crest on its ventral margin (
<figureCitation id="E9A43A96FFE6FF83FF6DFC70EA5BFC5A" box="[220,313,996,1020]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figure 1</figureCitation>
). Stylet support insertion point posteriorly placed, at about 75% the distance of the buccal tube from the anterior margin of the stylet sheaths. Anterior end of the piercing stylet with external edge straight and serrate, and internal edge curved and smooth (
<figureCitation id="E9A43A96FFE6FF83FE88FBD0EAFBFBFA" box="[313,409,1092,1116]" captionStart="Figure 3" captionStartId="11.[143,210,948,970]" captionTargetBox="[145,1102,168,915]" captionTargetId="figure-5@11.[145,1102,168,915]" captionTargetPageId="11" captionText="Figure 3. Macrobiotus kristenseni sp. nov. (A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197691" httpUri="https://zenodo.org/record/5197691/files/figure.png" pageId="8" pageNumber="2415">Figure 3</figureCitation>
). Pharyngeal bulb oval to circular, with well-developed pharyngeal apophyses. Two elongate macroplacoids and a small elongate microplacoid close to the second macroplacoid. First macroplacoid – the longest – with a small medial constriction, second macroplacoid quadrangular or circular (
<figureCitation id="E9A43A96FFE6FF83FC84FB30E8F7FB1A" box="[821,917,1188,1212]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figure 1</figureCitation>
). Slanting pharyngeal bars present between the apophyses and the first macroplacoids (
<figureCitation id="E9A43A96FFE6FF83FC1BFB50EF6AFB7A" box="[938,1032,1220,1244]" captionStart="Figure 1" captionStartId="9.[143,210,1158,1180]" captionTargetBox="[145,1102,168,1125]" captionTargetId="figure-5@9.[145,1102,168,1125]" captionTargetPageId="9" captionText="Figure 1. Macrobiotus kristenseni sp. nov. (A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="8" pageNumber="2415">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="71202613FFE6FF83FF4FFB70E922FA3D" blockId="8.[211,1172,479,1691]" pageId="8" pageNumber="2415">
Long double-claws. Main branch long with two fine accessory points, not reaching the distal bend of the main branch, and not clearly visible at LM (
<figureCitation id="E9A43A96FFE6FF83FC1BFA97EF65FABD" box="[938,1031,1283,1307]" captionStart="Figure 2" captionStartId="10.[211,278,1174,1196]" captionTargetBox="[213,1170,169,1142]" captionTargetId="figure-5@10.[213,1170,169,1142]" captionTargetPageId="10" captionText="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="8" pageNumber="2415">Figure 2</figureCitation>
). Secondary branch long and evidently bended. Short and large common tract, without distal part, and with an evident stalk connecting the claw to the lunule (
<figureCitation id="E9A43A96FFE6FF83FC29FAD7E89BFAFD" box="[920,1017,1347,1371]" captionStart="Figure 2" captionStartId="10.[211,278,1174,1196]" captionTargetBox="[213,1170,169,1142]" captionTargetId="figure-5@10.[213,1170,169,1142]" captionTargetPageId="10" captionText="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="8" pageNumber="2415">Figure 2</figureCitation>
). Hind claws larger than the others. Lunules smooth in legs I–III, larger and with weakly crenate edges in the hind legs (
<figureCitation id="E9A43A96FFE6FF83FE60FA17E94DFA3D" box="[465,559,1411,1435]" captionStart="Figure 2" captionStartId="10.[211,278,1174,1196]" captionTargetBox="[213,1170,169,1142]" captionTargetId="figure-5@10.[213,1170,169,1142]" captionTargetPageId="10" captionText="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="8" pageNumber="2415">Figure 2</figureCitation>
).
</paragraph>
<paragraph id="71202613FFE6FF83FF4FFA30E9E1FA7D" blockId="8.[211,1172,479,1691]" pageId="8" pageNumber="2415">
Measurements of different animal cuticular structures, 
<emphasis id="43EBFA01FFE6FF83FCDBFA30E8E2FA1D" box="[874,896,1444,1467]" italics="true" pageId="8" pageNumber="2415">pt</emphasis>
values and a descriptive statistical analysis are given in 
<tableCitation id="3C1D13A8FFE6FF83FD9AFA57E91EFA7D" box="[555,636,1475,1499]" captionStart="Table 2" captionStartId="7.[58,113,485,507]" captionText="Table 2. Summary of morphometric data (in µm, except pt) for Macrobiotus kristenseni sp. nov." pageId="8" pageNumber="2415">Table 2</tableCitation>
.
</paragraph>
<paragraph id="71202613FFE6FF82FF4FFA77E81DFA4B" blockId="8.[211,1172,479,1691]" lastBlockId="9.[143,1104,1366,1517]" lastPageId="9" lastPageNumber="2416" pageId="8" pageNumber="2415">
Ornamented round egg laid free (
<figureCitation id="E9A43A96FFE6FF83FD37FA77E990FA5D" box="[646,754,1507,1531]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="8" pageNumber="2415">Figures 4</figureCitation>
and 
<figureCitation id="E9A43A96FFE6FF83FC83FA77E823FA5D" box="[818,833,1507,1531]" captionStart="Figure 5" captionStartId="13.[143,210,934,956]" captionTargetBox="[145,1102,168,901]" captionTargetId="figure-5@13.[145,1102,168,901]" captionTargetPageId="13" captionText="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="8" pageNumber="2415">5</figureCitation>
). Egg ornamentation formed by processes in the shape of elongated empty cones (height 6.2–12.7 µm, diameter of their base 2.0–3.3 µm) with truncated and enlarged apexes (diameter less than 2 µm) (
<figureCitation id="E9A43A96FFE6FF83FE9FF9D6EAF8F9FC" box="[302,410,1602,1626]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="8" pageNumber="2415">Figures 4</figureCitation>
, 
<figureCitation id="E9A43A96FFE6FF83FE1AF9D6EADAF9FC" box="[427,440,1602,1626]" captionStart="Figure 5" captionStartId="13.[143,210,934,956]" captionTargetBox="[145,1102,168,901]" captionTargetId="figure-5@13.[145,1102,168,901]" captionTargetPageId="13" captionText="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="8" pageNumber="2415">5</figureCitation>
). Several processes with a long thick filament developing from the distal disk (
<figureCitation id="E9A43A96FFE6FF83FE30F9F6EA88F9DC" box="[385,490,1634,1658]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="8" pageNumber="2415">Figures 4</figureCitation>
, 
<figureCitation id="E9A43A96FFE6FF83FE48F9F6E964F9DC" box="[505,518,1634,1658]" captionStart="Figure 5" captionStartId="13.[143,210,934,956]" captionTargetBox="[145,1102,168,901]" captionTargetId="figure-5@13.[145,1102,168,901]" captionTargetPageId="13" captionText="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="8" pageNumber="2415">5</figureCitation>
). Ornamentations with smooth surface frequently ringed. Diameter of eggs without processes 73.0–96.3 µm, diameter of eggs with processes 94.0–125.0 µm. Number of processes present on a diameter 10–13. Chorion smooth; in some eggs a fine granulation and very small scattered pores in between processes visible by LM (
<figureCitation id="E9A43A96FFE7FF82FE8DFA01EAF8FA0B" box="[316,410,1429,1453]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="9" pageNumber="2416">Figure 4</figureCitation>
). Three embryonated eggs were found (
<figureCitation id="E9A43A96FFE7FF82FCEBFA01E8DBFA0B" box="[858,953,1429,1453]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="9" pageNumber="2416">Figure 4</figureCitation>
). In addition to the typical eggs described above, few eggs present slightly higher and more flexible processes internally septated and with a smaller base (
<figureCitation id="E9A43A96FFE7FF82FD5AFA41E836FA4B" box="[747,852,1493,1517]" captionStart="Figure 4" captionStartId="12.[211,278,1094,1116]" captionTargetBox="[213,1170,168,1062]" captionTargetId="figure-5@12.[213,1170,168,1062]" captionTargetPageId="12" captionText="Figure 4. Macrobiotus kristenseni sp. nov. (A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="9" pageNumber="2416">Figures 4</figureCitation>
, 
<figureCitation id="E9A43A96FFE7FF82FCD3FA41E80DFA4B" box="[866,879,1493,1517]" captionStart="Figure 5" captionStartId="13.[143,210,934,956]" captionTargetBox="[145,1102,168,901]" captionTargetId="figure-5@13.[145,1102,168,901]" captionTargetPageId="13" captionText="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="9" pageNumber="2416">5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="25E0769BFFE7FF82FF3EFB12EAF8FAB7" ID-DOI="http://doi.org/10.5281/zenodo.5197687" ID-Zenodo-Dep="5197687" httpUri="https://zenodo.org/record/5197687/files/figure.png" pageId="9" pageNumber="2416" startId="9.[143,210,1158,1180]" targetBox="[145,1102,168,1125]" targetPageId="9">
<paragraph id="71202613FFE7FF82FF3EFB12EAF8FAB7" blockId="9.[143,1104,1158,1298]" pageId="9" pageNumber="2416">
Figure 1. 
<taxonomicName id="B69F5D90FFE7FF82FF46FB13EA84FB3B" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[247,486,1159,1181]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="2416" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE7FF82FF46FB13EA84FB3B" box="[247,486,1159,1181]" italics="true" pageId="9" pageNumber="2416">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE7FF82FE5CFB13E958FB3A" box="[493,570,1159,1180]" pageId="9" pageNumber="2416" rank="species">sp. nov.</taxonomicNameLabel>
(A) Habitus; (B) anterior portion of the animal; (C) bucco-pharyngeal apparatus in ventral view; (D) dorsal and (E) ventral transversal crests of the buccal armature; (F) buccal pharyngeal apparatus in lateral view. (A–C) holotype (phase contrast); (D–F) paratypes (differential interference contrast). Scale bars: A = 50 µm; B, C, F, = 20 µm; D, E = 5 µm.
</paragraph>
</caption>
<subSubSection id="39857598FFE7FF80FF3EF9AFE8EBFA88" lastPageId="11" lastPageNumber="2418" pageId="9" pageNumber="2416" type="materials_examined">
<paragraph id="71202613FFE7FF82FF3EF9AFEA7CF9F4" blockId="9.[143,1103,1594,1690]" box="[143,286,1594,1618]" pageId="9" pageNumber="2416">
<heading id="2A68917FFFE7FF82FF3EF9AFEA7CF9F4" box="[143,286,1594,1618]" fontSize="10" level="3" pageId="9" pageNumber="2416" reason="5">
<emphasis id="43EBFA01FFE7FF82FF3EF9AFEA7CF9F4" box="[143,286,1594,1618]" italics="true" pageId="9" pageNumber="2416">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE7FF82FF3EF9F6E9BDF93C" blockId="9.[143,1103,1594,1690]" pageId="9" pageNumber="2416">
<materialsCitation id="C1F72C4EFFE7FF82FF3EF9F6E9B9F93C" country="Argentina" elevation="177" latitude="-36.65" location="Santa Rosa" longLatPrecision="1186" longitude="-64.28333" municipality="South America" pageId="9" pageNumber="2416" specimenCount="1" stateProvince="La Pampa">
<collectingMunicipality id="9144BC69FFE7FF82FF3EF9F6EA27F9DC" box="[143,325,1634,1658]" pageId="9" pageNumber="2416">South America</collectingMunicipality>
, 
<collectingCountry id="09886683FFE7FF82FEE8F9F6EAACF9DC" box="[345,462,1634,1658]" name="Argentina" pageId="9" pageNumber="2416">Argentina</collectingCountry>
, 
<collectingRegion id="B35BE8F1FFE7FF82FE50F9F7E9B3F9DC" box="[481,721,1634,1658]" country="Argentina" name="La Pampa" pageId="9" pageNumber="2416">La Pampa Province</collectingRegion>
, city of 
<location id="744070C8FFE7FF82FCF5F9F6E8AEF9DC" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE7FF82FCF5F9F6E8AEF9DC" box="[836,972,1634,1658]" country="Argentina" latitude="-36.65" longLatPrecision="1186" longitude="-64.28333" municipality="South America" name="Santa Rosa" pageId="9" pageNumber="2416" stateProvince="La Pampa">Santa Rosa</location>
(
<geoCoordinate id="14AB40D4FFE7FF82FC55F9F6EF29F9DC" box="[996,1099,1630,1658]" degrees="36" direction="south" minutes="39" orientation="latitude" pageId="9" pageNumber="2416" precision="925" value="-36.65">
36 
<superScript id="86EA8B5BFFE7FF82FBB1F9CAEF6BF9C9" attach="none" box="[1024,1033,1630,1647]" fontSize="7" pageId="9" pageNumber="2416">◦</superScript>
39 
<superScript id="86EA8B5BFFE7FF82FB94F9CAEF48F9C9" attach="left" box="[1061,1066,1630,1647]" fontSize="7" pageId="9" pageNumber="2416">′</superScript>
S
</geoCoordinate>
, 
<geoCoordinate id="14AB40D4FFE7FF82FF3EF916EB9AF93C" box="[143,248,1662,1690]" degrees="64" direction="west" minutes="17" orientation="longitude" pageId="9" pageNumber="2416" precision="925" value="-64.28333">
64 
<superScript id="86EA8B5BFFE7FF82FF1BF9EAEBD1F929" attach="none" box="[170,179,1662,1679]" fontSize="7" pageId="9" pageNumber="2416">◦</superScript>
17 
<superScript id="86EA8B5BFFE7FF82FF7EF9EAEBB6F929" attach="left" box="[207,212,1662,1679]" fontSize="7" pageId="9" pageNumber="2416">′</superScript>
W
</geoCoordinate>
) at 
<elevation id="FAB2C120FFE7FF82FE93F916E971F93C" box="[290,531,1666,1690]" metricMagnitude="2" metricUnit="m" metricValue="1.77" pageId="9" pageNumber="2416" unit="m" value="177.0">
<quantity id="B6678BF6FFE7FF82FE93F916EA04F93C" box="[290,358,1666,1690]" metricMagnitude="2" metricUnit="m" metricValue="1.77" pageId="9" pageNumber="2416" unit="m" value="177.0">177 m</quantity>
above sea level
</elevation>
, epiphytic lichens
</materialsCitation>
.
</paragraph>
<caption id="25E0769BFFE4FF81FF62FB02E9B0FA87" ID-DOI="http://doi.org/10.5281/zenodo.5197689" ID-Zenodo-Dep="5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="10" pageNumber="2417" startId="10.[211,278,1174,1196]" targetBox="[213,1170,169,1142]" targetPageId="10">
<paragraph id="71202613FFE4FF81FF62FB02E9B0FA87" blockId="10.[211,1172,1174,1314]" pageId="10" pageNumber="2417">
Figure 2. 
<taxonomicName id="B69F5D90FFE4FF81FE8DFB03E94EFB0B" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[316,556,1175,1197]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="2417" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE4FF81FE8DFB03E94EFB0B" box="[316,556,1175,1197]" italics="true" pageId="10" pageNumber="2417">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE4FF81FD85FB03E9E0FB0A" box="[564,642,1175,1196]" pageId="10" pageNumber="2417" rank="species">sp. nov.</taxonomicNameLabel>
(A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm.
</paragraph>
</caption>
<paragraph id="71202613FFE4FF81FF62FAC8EA0EFAD5" blockId="10.[211,1172,1371,1690]" box="[211,364,1371,1395]" pageId="10" pageNumber="2417">
<heading id="2A68917FFFE4FF81FF62FAC8EA0EFAD5" box="[211,364,1371,1395]" fontSize="10" level="3" pageId="10" pageNumber="2417" reason="5">
<emphasis id="43EBFA01FFE4FF81FF62FAC8EA0EFAD5" box="[211,364,1371,1395]" italics="true" pageId="10" pageNumber="2417">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE4FF81FF62FA17E86AF93C" blockId="10.[211,1172,1371,1690]" pageId="10" pageNumber="2417">
<materialsCitation id="C1F72C4EFFE4FF81FF62FA17E86AF93C" collectionCode="MOD" collectorName="J. Peluffo's &amp; National University &amp; The" country="Argentina" county="Museum of Natural Sciences" location="Department of Natural Sciences" municipality="National University of La Plata" pageId="10" pageNumber="2417" specimenCount="35" specimenCount-egg="1" stateProvince="La Pampa" typeStatus="holotype">
<typeStatus id="AE2498B1FFE4FF81FF62FA17EA5CFA3D" box="[211,318,1411,1435]" pageId="10" pageNumber="2417" type="holotype">Holotype</typeStatus>
(MLP-Oi 3726), 
<specimenCount id="6799ED9AFFE4FF81FDB7FA17E9C8FA3A" box="[518,682,1411,1436]" pageId="10" pageNumber="2417" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
(MLP-Oi 3727) and 
<specimenCount id="6799ED9AFFE4FF81FC2FFA17EF6DFA3A" box="[926,1039,1411,1436]" pageId="10" pageNumber="2417" type="egg">three eggs</specimenCount>
are housed at the 
<collectingCounty id="98415E9FFFE4FF81FE93FA37E915FA1D" box="[290,631,1443,1467]" pageId="10" pageNumber="2417">Museum of Natural Sciences</collectingCounty>
, 
<collectingMunicipality id="9144BC69FFE4FF81FD37FA37E89CFA1D" box="[646,1022,1443,1467]" pageId="10" pageNumber="2417">National University of La Plata</collectingMunicipality>
(
<collectingCountry id="09886683FFE4FF81FBA0FA37EFEAFA1D" box="[1041,1160,1443,1467]" name="Argentina" pageId="10" pageNumber="2417">Argentina</collectingCountry>
), and 
<specimenCount id="6799ED9AFFE4FF81FEB8FA56EAF9FA7D" box="[265,411,1474,1499]" pageId="10" pageNumber="2417" type="generic" typeStatus="paratypes">32 paratypes</specimenCount>
are housed in 
<collectorName id="DC6A43C5FFE4FF81FDE1FA57E9AEFA7C" box="[592,716,1474,1498]" pageId="10" pageNumber="2417">J. Peluffo’s</collectorName>
personal collection in the 
<location id="744070C8FFE4FF81FBBBFA57EADCFA5C" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE4FF81FBBBFA57EADCFA5C" country="Argentina" county="Museum of Natural Sciences" municipality="National University of La Plata" name="Department of Natural Sciences" pageId="10" pageNumber="2417" stateProvince="La Pampa">Department of Natural Sciences</location>
at the 
<collectorName id="DC6A43C5FFE4FF81FD95FA76E86FFA5C" box="[548,781,1506,1530]" pageId="10" pageNumber="2417">National University</collectorName>
of 
<collectingRegion id="B35BE8F1FFE4FF81FCF3FA77E8DDFA5C" box="[834,959,1507,1530]" country="Argentina" name="La Pampa" pageId="10" pageNumber="2417">La Pampa</collectingRegion>
(
<collectingCountry id="09886683FFE4FF81FC67FA76EF2CFA5C" box="[982,1102,1506,1530]" name="Argentina" pageId="10" pageNumber="2417">Argentina</collectingCountry>
). 
<collectorName id="DC6A43C5FFE4FF81FBD6FA76EFF1FA5C" box="[1127,1171,1506,1530]" pageId="10" pageNumber="2417">The</collectorName>
voucher specimens (two pictures of the hologenophore animals, 16 cryoconserved paragenophore animals, 
<specimenCount id="6799ED9AFFE4FF81FE56F9B6E9F3F99D" box="[487,657,1570,1595]" pageId="10" pageNumber="2417" type="generic" typeStatus="paratypes">eight paratypes</specimenCount>
<emphasis id="43EBFA01FFE4FF81FD20F9B7E9FFF99D" bold="true" box="[657,669,1571,1595]" italics="true" pageId="10" pageNumber="2417">/</emphasis>
paragenophore animals and eggs) and a fragment of the lichen are deposited in the 
<collectionCode id="178EBED6FFE4FF81FDCDF9D6E9A3F9FC" box="[636,705,1602,1626]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14382" lsid="urn:lsid:biocol.org:col:14382" name="University of Modena and Reggio Emilia, Department of Biology" pageId="10" pageNumber="2417" type="University or college">MOD</collectionCode>
:BRT collection of the 
<location id="744070C8FFE4FF81FC0BF9D7EA52F9DC" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE4FF81FC0BF9D7EA52F9DC" country="Argentina" county="Museum of Natural Sciences" municipality="National University of La Plata" name="Department of Life Sciences" pageId="10" pageNumber="2417" stateProvince="La Pampa">Department of Life Sciences</location>
(
<location id="744070C8FFE4FF81FE8FF9F6E95AF9DC" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE4FF81FE8FF9F6E95AF9DC" box="[318,568,1634,1658]" country="Argentina" county="Museum of Natural Sciences" municipality="National University of La Plata" name="University of Modena" pageId="10" pageNumber="2417" stateProvince="La Pampa">University of Modena</location>
and 
<location id="744070C8FFE4FF81FDDEF9F6E870F9DC" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE4FF81FDDEF9F6E870F9DC" box="[623,786,1634,1658]" country="Argentina" county="Museum of Natural Sciences" municipality="National University of La Plata" name="Reggio Emilia" pageId="10" pageNumber="2417" stateProvince="La Pampa">Reggio Emilia</location>
, 
<collectingCountry id="09886683FFE4FF81FCADF9F6E833F9DC" box="[796,849,1634,1658]" name="Italy" pageId="10" pageNumber="2417">Italy</collectingCountry>
) registered in the 
<location id="744070C8FFE4FF81FBA9F9F6EAB5F93C" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE4FF81FBA9F9F6EAB5F93C" country="Argentina" county="Museum of Natural Sciences" municipality="National University of La Plata" name="Registry of Biological Repositories" pageId="10" pageNumber="2417" stateProvince="La Pampa">Registry of Biological Repositories</location>
(www.biorepositories.org).
</materialsCitation>
</paragraph>
<caption id="25E0769BFFE5FF80FF3EFC20E83FFBDF" ID-DOI="http://doi.org/10.5281/zenodo.5197691" ID-Zenodo-Dep="5197691" httpUri="https://zenodo.org/record/5197691/files/figure.png" pageId="11" pageNumber="2418" startId="11.[143,210,948,970]" targetBox="[145,1102,168,915]" targetPageId="11">
<paragraph id="71202613FFE5FF80FF3EFC20E83FFBDF" blockId="11.[143,1104,948,1145]" pageId="11" pageNumber="2418">
Figure 3. 
<taxonomicName id="B69F5D90FFE5FF80FF4BFC20EA8EFC6C" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[250,492,948,970]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="2418" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE5FF80FF4BFC20EA8EFC6C" box="[250,492,948,970]" italics="true" pageId="11" pageNumber="2418">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE5FF80FE47FC21E927FC6C" box="[502,581,949,970]" pageId="11" pageNumber="2418" rank="species">sp. nov.</taxonomicNameLabel>
(A) Most anterior end of the animal bearing the mouth opening, regular ring of pores (arrow), buccal sensory lobes (L), peribuccal lamellae (asterisk); (B) mouth opening with buccal lamellae (asterisk) and buccal armature: first band of teeth (arrow head), second band of teeth (arrow); (C) mouth opening with buccal lamellae (asterisk) and buccal armature: second band of teeth (arrow head), transversal crests (arrow); (D) mouth opening with buccal lamellae (asterisk) and protruding piercing stylets (ps). A–D: scanning electron micrographs. Scale bars: A, D = 5 µm; B, C = 2 µm.
</paragraph>
</caption>
<paragraph id="71202613FFE5FF80FF3EFB3BEA04FB61" blockId="11.[143,1104,1199,1326]" box="[143,358,1199,1223]" pageId="11" pageNumber="2418">
<heading id="2A68917FFFE5FF80FF3EFB3BEA04FB61" box="[143,358,1199,1223]" fontSize="10" level="3" pageId="11" pageNumber="2418" reason="5">
<emphasis id="43EBFA01FFE5FF80FF3EFB3BEA04FB61" box="[143,358,1199,1223]" italics="true" pageId="11" pageNumber="2418">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE5FF80FF3EFB43E8EBFA88" blockId="11.[143,1104,1199,1326]" pageId="11" pageNumber="2418">
<materialsCitation id="C1F72C4EFFE5FF80FF3EFB43E8E4FA88" country="Argentina" county="South America" elevation="143" location="Natural Reserve" municipality="Santa Rosa" pageId="11" pageNumber="2418" specimenCount="1" stateProvince="La Pampa">
<collectingCounty id="98415E9FFFE5FF80FF3EFB43EA5CFB49" box="[143,318,1239,1263]" pageId="11" pageNumber="2418">South America</collectingCounty>
, 
<collectingCountry id="09886683FFE5FF80FEFBFB43EADDFB49" box="[330,447,1239,1263]" name="Argentina" pageId="11" pageNumber="2418">Argentina</collectingCountry>
, 
<collectingRegion id="B35BE8F1FFE5FF80FE7AFB43E9C9FB49" box="[459,683,1239,1263]" country="Argentina" name="La Pampa" pageId="11" pageNumber="2418">La Pampa Province</collectingRegion>
, city of General Pico at 
<elevation id="FAB2C120FFE5FF80FC70FB43EB8EFAA8" metricMagnitude="2" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="2418" unit="m" value="143.0">
<quantity id="B6678BF6FFE5FF80FC70FB43EF64FB48" box="[961,1030,1239,1263]" metricMagnitude="2" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="2418" unit="m" value="143.0">143 m</quantity>
above sea level
</elevation>
(
<quantity id="B6678BF6FFE5FF80FF4AFB62EA32FAA8" box="[251,336,1270,1294]" metricMagnitude="5" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="2418" unit="km" value="125.0">125 km</quantity>
northeast of 
<collectingMunicipality id="9144BC69FFE5FF80FE59FB62E908FAA8" box="[488,618,1270,1294]" pageId="11" pageNumber="2418">Santa Rosa</collectingMunicipality>
) and 
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Parque Luro (
<quantity id="B6678BF6FFE5FF80FBB9FB62EF32FAA8" box="[1032,1104,1270,1294]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="2418" unit="km" value="35.0">35 km</quantity>
south of 
<location id="744070C8FFE5FF80FF42FA82EA14FA88" LSID="urn:lsid:plazi:treatment:F9369705FFE8FF9BFDB0F9C8EAF6FCDA:744070C8FFE5FF80FF42FA82EA14FA88" box="[243,374,1302,1326]" country="Argentina" county="South America" municipality="Santa Rosa" name="Santa Rosa" pageId="11" pageNumber="2418" stateProvince="La Pampa">Santa Rosa</location>
, caldén forest), in all cases on epiphytic lichens
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39857598FFE5FF80FF3EFAE1EA47FA52" pageId="11" pageNumber="2418" type="etymology">
<paragraph id="71202613FFE5FF80FF3EFAE1EA65FA2B" blockId="11.[143,1104,1397,1524]" box="[143,263,1397,1421]" pageId="11" pageNumber="2418">
<heading id="2A68917FFFE5FF80FF3EFAE1EA65FA2B" box="[143,263,1397,1421]" fontSize="10" level="3" pageId="11" pageNumber="2418" reason="5">
<emphasis id="43EBFA01FFE5FF80FF3EFAE1EA65FA2B" box="[143,263,1397,1421]" italics="true" pageId="11" pageNumber="2418">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE5FF80FF3EFA08EA47FA52" blockId="11.[143,1104,1397,1524]" pageId="11" pageNumber="2418">
The name is dedicated in honour of Prof. Reinhardt M. Kristensen, of the Invertebrate Department of The Natural History Museum of 
<collectingCountry id="09886683FFE5FF80FD54FA28E833FA72" box="[741,849,1468,1492]" name="Denmark" pageId="11" pageNumber="2418">Denmark</collectingCountry>
of the University of Copenhagen.
</paragraph>
</subSubSection>
<subSubSection id="39857598FFE5FF9BFF3EF9AEEAF6FCDA" lastPageId="16" lastPageNumber="2423" pageId="11" pageNumber="2418" type="discussion">
<paragraph id="71202613FFE5FF80FF3EF9AEEB93F9F4" blockId="11.[143,1103,1594,1691]" box="[143,241,1594,1618]" pageId="11" pageNumber="2418">
<heading id="2A68917FFFE5FF80FF3EF9AEEB93F9F4" box="[143,241,1594,1618]" fontSize="10" level="3" pageId="11" pageNumber="2418" reason="5">
<emphasis id="43EBFA01FFE5FF80FF3EF9AEEB93F9F4" box="[143,241,1594,1618]" italics="true" pageId="11" pageNumber="2418">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE5FF87FF3EF9F6E86EFAFD" blockId="11.[143,1103,1594,1691]" lastBlockId="12.[211,1172,1283,1563]" lastPageId="12" lastPageNumber="2419" pageId="11" pageNumber="2418">
Under SEM, the general surface of the cuticle appears slightly wrinkled (
<figureCitation id="E9A43A96FFE5FF80FC6FF9F6EF5DF9DC" box="[990,1087,1634,1658]" captionStart="Figure 2" captionStartId="10.[211,278,1174,1196]" captionTargetBox="[213,1170,169,1142]" captionTargetId="figure-5@10.[213,1170,169,1142]" captionTargetPageId="10" captionText="Figure 2. Macrobiotus kristenseni sp. nov. (A) Claws of the hind pair of legs; (B) claws of the hind legs and cuticular dots on the external surface of the leg (arrow); (C) cuticular “pores” (arrow) in the body cuticle; (D) drawing of the claws of the hind legs; (E) claws of the second pair of legs; (F) claws of a hind leg. A–C: scanning electron micrographs; E, F: holotype (phase contrast). Scale bars: A–C = 5 µm; E, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5197689" httpUri="https://zenodo.org/record/5197689/files/figure.png" pageId="11" pageNumber="2418">Figure 2</figureCitation>
). It should be verified whether this character is an effect of SEM preparation. An egg observed by SEM had processes with filamentous strips on their surface (
<figureCitation id="E9A43A96FFE2FF87FB93FA97EFE1FABD" box="[1058,1155,1283,1307]" captionStart="Figure 5" captionStartId="13.[143,210,934,956]" captionTargetBox="[145,1102,168,901]" captionTargetId="figure-5@13.[145,1102,168,901]" captionTargetPageId="13" captionText="Figure 5. Macrobiotus kristenseni sp. nov. (A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="12" pageNumber="2419">Figure 5</figureCitation>
). These strips seem to be present on the same side of all processes: it is not clear whether they are artefacts or real morphological structures.
</paragraph>
<caption id="25E0769BFFE2FF87FF62FBD2EFF1FB12" ID-DOI="http://doi.org/10.5281/zenodo.5197693" ID-Zenodo-Dep="5197693" httpUri="https://zenodo.org/record/5197693/files/figure.png" pageId="12" pageNumber="2419" startId="12.[211,278,1094,1116]" targetBox="[213,1170,168,1062]" targetPageId="12">
<paragraph id="71202613FFE2FF87FF62FBD2EFF1FB12" blockId="12.[211,1172,1094,1204]" pageId="12" pageNumber="2419">
Figure 4. 
<taxonomicName id="B69F5D90FFE2FF87FE8CFBD3E94FFBFB" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[317,557,1095,1117]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE2FF87FE8CFBD3E94FFBFB" box="[317,557,1095,1117]" italics="true" pageId="12" pageNumber="2419">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE2FF87FD87FBD3E9E7FBFA" box="[566,645,1095,1116]" pageId="12" pageNumber="2419" rank="species">sp. nov.</taxonomicNameLabel>
(A) Embryonated eggs; (B) egg surface with small rounded meshes (arrow) bearing elongated conical processes; (C) egg with embryo buccopharyngeal apparatus in development (arrow); (D) egg processes with filamentous matter developing from their tips (arrow). A–D: phase contrast. Scale bars: A–C = 20 µm; D = 5 µm.
</paragraph>
</caption>
<paragraph id="71202613FFE2FF87FF4FFAF7EAECF9BC" blockId="12.[211,1172,1283,1563]" pageId="12" pageNumber="2419">
Where 
<taxonomicName id="B69F5D90FFE2FF87FEFFFAF0EA8EFADD" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[334,492,1379,1403]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE2FF87FEFFFAF0EA8EFADD" box="[334,492,1379,1403]" italics="true" pageId="12" pageNumber="2419">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE2FF87FE42FAF0E925FADD" box="[499,583,1380,1403]" pageId="12" pageNumber="2419" rank="species">sp. nov.</taxonomicNameLabel>
appears in a sample, it is either the only or the dominant tardigrade species. In the latter case, it is accompanied by one or more of the following species: 
<taxonomicName id="B69F5D90FFE2FF87FE2DFA37E897FA1D" authority="du Bois Raymond-Marcus, 1944" authorityName="du Bois Raymond-Marcus" authorityYear="1944" box="[412,1013,1443,1467]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="rufoviridis">
<emphasis id="43EBFA01FFE2FF87FE2DFA37E9E6FA1D" box="[412,644,1443,1467]" italics="true" pageId="12" pageNumber="2419">Echiniscus rufoviridis</emphasis>
du Bois Raymond-Marcus, 1944
</taxonomicName>
; 
<taxonomicName id="B69F5D90FFE2FF87FC4EFA37EA3EFA7D" authorityName="Doyere" authorityYear="1840" class="Eutardigrada" family="Milnesiidae" genus="Milnesium" kingdom="Animalia" order="Apochela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="tardigradum">
<emphasis id="43EBFA01FFE2FF87FC4EFA37EF16FA1D" box="[1023,1140,1443,1467]" italics="true" pageId="12" pageNumber="2419">Milnesium</emphasis>
cf. 
<emphasis id="43EBFA01FFE2FF87FF62FA57EA3EFA7D" box="[211,348,1475,1499]" italics="true" pageId="12" pageNumber="2419">tardigradum</emphasis>
</taxonomicName>
<emphasis id="43EBFA01FFE2FF87FEEDFA57EA9CFA7D" box="[348,510,1475,1499]" italics="true" pageId="12" pageNumber="2419">; Ramazzottius</emphasis>
cf. 
<emphasis id="43EBFA01FFE2FF87FD95FA57E9CDFA7D" box="[548,687,1475,1499]" italics="true" pageId="12" pageNumber="2419">oberhaeuseri</emphasis>
. The phorophyte of the lichen communities in which it was recorded more frequently is the autochthonous tree species 
<taxonomicName id="B69F5D90FFE2FF87FB84FA77EAE8F9BC" authority="(Burk.)" baseAuthorityName="Burk." class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="2419" phylum="Tracheophyta" rank="species" species="caldenia">
<emphasis id="43EBFA01FFE2FF87FB84FA77EA4CF9BD" italics="true" pageId="12" pageNumber="2419">Prosopis caldenia</emphasis>
(Burk.)
</taxonomicName>
.
</paragraph>
<paragraph id="71202613FFE2FF86FF62F9D7EAAAFAE9" blockId="12.[211,1172,1602,1690]" lastBlockId="13.[143,1104,1111,1359]" lastPageId="13" lastPageNumber="2420" pageId="12" pageNumber="2419">
<emphasis id="43EBFA01FFE2FF87FF62F9D7E9BBF9FC" box="[211,729,1602,1626]" italics="true" pageId="12" pageNumber="2419">
Taxonomic remarks. 
<taxonomicName id="B69F5D90FFE2FF87FE78F9D6E9BBF9FC" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[457,729,1602,1626]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">Macrobiotus kristenseni</taxonomicName>
</emphasis>
<taxonomicNameLabel id="58D8477AFFE2FF87FD56F9D7E821F9FC" box="[743,835,1603,1626]" pageId="12" pageNumber="2419" rank="species">sp. nov.</taxonomicNameLabel>
is similar to species of the 
<taxonomicName id="B69F5D90FFE2FF87FF62F9F7EA08F9DC" authorityName="C. A. S. Schultze" authorityYear="1834" box="[211,362,1634,1658]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFE2FF87FF62F9F7EA08F9DC" box="[211,362,1634,1658]" italics="true" pageId="12" pageNumber="2419">M. hufelandi</emphasis>
</taxonomicName>
group. Among these species, 
<taxonomicName id="B69F5D90FFE2FF87FD5AF9F6EA1CF93C" authority="Bertolani and Rebecchi, 1993" authorityName="Bertolani and Rebecchi" authorityYear="1993" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="2419" phylum="Tardigrada" rank="species" species="sandrae">
<emphasis id="43EBFA01FFE2FF87FD5AF9F6E8BEF9DC" box="[747,988,1634,1658]" italics="true" pageId="12" pageNumber="2419">Macrobiotus sandrae</emphasis>
<bibRefCitation id="150E5BE2FFE2FF87FC5FF9F6EA1CF93C" author="Bertolani R &amp; Rebecchi L" pageId="12" pageNumber="2419" pagination="127 - 152" refId="ref6033" refString="Bertolani R, Rebecchi L. 1993. A revision of the Macrobiotus hufelandi group (Tardigrada, Macrobiotidae), with some observations on the taxonomic characters of eutardigrades. Zool. Scripta. 22: 127 - 152." type="journal article" year="1993">Bertolani and Rebecchi, 1993</bibRefCitation>
</taxonomicName>
is the most similar, sharing a weak buccal armature, reduced ventral transversal crests, and indented lunule only on the fourth leg pair, but it differs from 
<taxonomicName id="B69F5D90FFE3FF86FF3EFBECEA53FB36" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[143,305,1144,1168]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE3FF86FF3EFBECEA53FB36" box="[143,305,1144,1168]" italics="true" pageId="13" pageNumber="2420">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE3FF86FE8DFBECEAF7FB29" box="[316,405,1144,1167]" pageId="13" pageNumber="2420" rank="species">sp. nov.</taxonomicNameLabel>
by longer claw accessory points and larger dorsal transversal crests. 
<taxonomicName id="B69F5D90FFE3FF86FEB2FB03E96CFB09" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[259,526,1175,1199]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE3FF86FEB2FB03E96CFB09" box="[259,526,1175,1199]" italics="true" pageId="13" pageNumber="2420">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE3FF86FDA6FB0CE90CFB09" box="[535,622,1176,1199]" pageId="13" pageNumber="2420" rank="species">sp. nov.</taxonomicNameLabel>
differs from all the 
<taxonomicName id="B69F5D90FFE3FF86FCE4FB0CE881FB09" authorityName="C. A. S. Schultze" authorityYear="1834" box="[853,995,1175,1199]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFE3FF86FCE4FB0CE881FB09" box="[853,995,1175,1199]" italics="true" pageId="13" pageNumber="2420">M. hufelandi</emphasis>
</taxonomicName>
group of species in that the shape of the egg processes is different from any other egg of the species group. The most similar eggs are those of 
<taxonomicName id="B69F5D90FFE3FF86FD7CFB43EF29FB49" authority="Cuenot, 1932" authorityName="Cuenot" authorityYear="1932" box="[717,1099,1239,1263]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="recens">
<emphasis id="43EBFA01FFE3FF86FD7CFB43E8C4FB49" box="[717,934,1239,1263]" italics="true" pageId="13" pageNumber="2420">Macrobiotus recens</emphasis>
Cuénot, 1932
</taxonomicName>
, which share similar thin, long, conical processes with a truncated apex but in 
<taxonomicName id="B69F5D90FFE3FF86FC55FB6CEF2DFAA9" authorityName="Cuenot" authorityYear="1932" box="[996,1103,1272,1295]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="recens">
<emphasis id="43EBFA01FFE3FF86FC55FB6CEF2DFAA9" box="[996,1103,1272,1295]" italics="true" pageId="13" pageNumber="2420">M. recens</emphasis>
</taxonomicName>
an evident series of small indentations around the base of the egg processes is present (
<bibRefCitation id="150E5BE2FFE3FF86FF26FAA3EADAFAE9" author="Pilato G &amp; Bertolani R" box="[151,440,1335,1359]" pageId="13" pageNumber="2420" pagination="1 - 7" refId="ref6878" refString="Pilato G, Bertolani R. 2004. Macrobiotus dariae sp. n., a new species of eutardigrade (Eutardigrada, Macrobiotidae) from Cyprus. Zootaxa. 638: 1 - 7." type="journal article" year="2004">Pilato and Bertolani 2004</bibRefCitation>
).
</paragraph>
<caption id="25E0769BFFE3FF86FF3EFC32E96EFBB2" ID-DOI="http://doi.org/10.5281/zenodo.5197695" ID-Zenodo-Dep="5197695" httpUri="https://zenodo.org/record/5197695/files/figure.png" pageId="13" pageNumber="2420" startId="13.[143,210,934,956]" targetBox="[145,1102,168,901]" targetPageId="13">
<paragraph id="71202613FFE3FF86FF3EFC32E96EFBB2" blockId="13.[143,1104,934,1044]" pageId="13" pageNumber="2420">
Figure 5. 
<taxonomicName id="B69F5D90FFE3FF86FF49FC33EA8AFC1B" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[248,488,935,957]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE3FF86FF49FC33EA8AFC1B" box="[248,488,935,957]" italics="true" pageId="13" pageNumber="2420">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE3FF86FE41FC33E95FFC1A" box="[496,573,935,956]" pageId="13" pageNumber="2420" rank="species">sp. nov.</taxonomicNameLabel>
(A) Egg; (B) egg processes with filamentous matter developing from their tips (arrows); (C, D) egg processes with annulated surface (arrow) and filamentous matter developing from their tips (arrow head). A–D: scanning electron micrographs. Scale bars: A = 10 µm; B–D = 2 µm.
</paragraph>
</caption>
<paragraph id="71202613FFE3FF86FF3EFA0EEA2DFA15" blockId="13.[143,1104,1434,1690]" box="[143,335,1434,1459]" pageId="13" pageNumber="2420">
<heading id="2A68917FFFE3FF86FF3EFA0EEA2DFA15" box="[143,335,1434,1459]" fontSize="10" level="3" pageId="13" pageNumber="2420" reason="5">
<emphasis id="43EBFA01FFE3FF86FF3EFA0EEA2DFA15" bold="true" box="[143,335,1434,1459]" italics="true" pageId="13" pageNumber="2420">Molecular results</emphasis>
</heading>
</paragraph>
<paragraph id="71202613FFE3FF86FF3EFA56E8F9F93C" blockId="13.[143,1104,1434,1690]" pageId="13" pageNumber="2420">
The molecular phylogeny based on the 18S rRNA gene (897 bp) indicates that 
<taxonomicName id="B69F5D90FFE3FF86FC44FA50EBB9FA5C" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE3FF86FC44FA50EBB9FA5C" italics="true" pageId="13" pageNumber="2420">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE3FF86FF57FA77EA5FFA5C" box="[230,317,1507,1530]" pageId="13" pageNumber="2420" rank="species">sp. nov.</taxonomicNameLabel>
does not share its 18S rRNA sequence with any other 
<taxonomicName id="B69F5D90FFE3FF86FC72FA77EBBEF9BC" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="undetermined">
<emphasis id="43EBFA01FFE3FF86FC72FA77EF2DFA5D" box="[963,1103,1507,1531]" italics="true" pageId="13" pageNumber="2420">Macrobiotus</emphasis>
species
</taxonomicName>
sequenced so far (
<figureCitation id="E9A43A96FFE3FF86FE79F996E94FF9BC" box="[456,557,1538,1562]" captionStart="Figure 6" captionStartId="14.[211,278,1550,1572]" captionTargetBox="[214,1169,169,1517]" captionTargetId="figure-5@14.[213,1170,169,1517]" captionTargetPageId="14" captionText="Figure 6. Unrooted neighbour joining dendrogram computed on Macrobiotus “hufelandi group” cox1 sequences. Bootstrap percentages computed after 2000 replicates are shown above branches. Macrobiotus kristenseni sp. nov. specimens are shown in bold. Acronyms as in Table 1." figureDoi="http://doi.org/10.5281/zenodo.5197697" httpUri="https://zenodo.org/record/5197697/files/figure.png" pageId="13" pageNumber="2420">Figure 6</figureCitation>
), but it is in a sister-group relationship with 
<taxonomicName id="B69F5D90FFE3FF86FF3EF9B6E9E6F99C" authority="Binda and Pilato, 1984" authorityName="Binda and Pilato" authorityYear="1984" box="[143,644,1570,1594]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="sapiens">
<emphasis id="43EBFA01FFE3FF86FF3EF9B6EA16F99C" box="[143,372,1570,1594]" italics="true" pageId="13" pageNumber="2420">Macrobiotus sapiens</emphasis>
Binda and Pilato, 1984
</taxonomicName>
within a well-supported clade grouping all the 
<taxonomicName id="B69F5D90FFE3FF86FF52F9D7EA10F9FC" authorityName="C. A. S. Schultze" authorityYear="1834" box="[227,370,1602,1626]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFE3FF86FF52F9D7EA10F9FC" box="[227,370,1602,1626]" italics="true" pageId="13" pageNumber="2420">M. hufelandi</emphasis>
</taxonomicName>
group of species (i.e. 
<emphasis id="43EBFA01FFE3FF86FDC7F9D7E994F9FC" box="[630,758,1603,1626]" italics="true" pageId="13" pageNumber="2420">
<taxonomicName id="B69F5D90FFE3FF86FDC7F9D7E990F9FC" authorityName="Binda and Pilato" authorityYear="1984" box="[630,754,1603,1626]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="sapiens">M. sapiens</taxonomicName>
,
</emphasis>
<taxonomicName id="B69F5D90FFE3FF86FCB1F9D6EA51F9DC" authority="C. A. S. Schultze, 1834" authorityName="C. A. S. Schultze" authorityYear="1834" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFE3FF86FCB1F9D6E899F9FC" box="[768,1019,1602,1626]" italics="true" pageId="13" pageNumber="2420">Macrobiotus hufelandi</emphasis>
C.A.S. Schultze, 1834
</taxonomicName>
, 
<taxonomicName id="B69F5D90FFE3FF86FE8EF9F6EF2AF9DC" authority="Pilato, Kaczmarek, Michalczyk and Lisi, 2003" authorityName="Pilato, Kaczmarek, Michalczyk and Lisi" authorityYear="2003" box="[319,1096,1634,1658]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="polonicus">
<emphasis id="43EBFA01FFE3FF86FE8EF9F6E95BF9DC" box="[319,569,1634,1658]" italics="true" pageId="13" pageNumber="2420">Macrobiotus polonicus</emphasis>
Pilato, Kaczmarek, Michalczyk and Lisi, 2003
</taxonomicName>
, 
<emphasis id="43EBFA01FFE3FF86FF3EF916EA7EF93C" box="[143,284,1666,1690]" italics="true" pageId="13" pageNumber="2420">Macrobiorus</emphasis>
<taxonomicName id="B69F5D90FFE3FF86FE92F916E8E8F93C" authority="Bertolani, Biserov, Rebecchi and Cesari, 2011" authorityName="Bertolani, Biserov, Rebecchi and Cesari" authorityYear="2011" box="[291,906,1666,1690]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="2420" phylum="Tardigrada" rank="species" species="vladimiri">
<emphasis id="43EBFA01FFE3FF86FE92F916EAE7F93C" box="[291,389,1666,1690]" italics="true" pageId="13" pageNumber="2420">vladimiri</emphasis>
Bertolani, Biserov, Rebecchi and 
<bibRefCitation id="150E5BE2FFE3FF86FCB1F916E8E8F93C" author="Cesari M &amp; Giovannini I &amp; Bertolani R &amp; Rebecchi L" box="[768,906,1666,1690]" pageId="13" pageNumber="2420" pagination="42 - 51" refId="ref6174" refString="Cesari M, Giovannini I, Bertolani R, Rebecchi L. 2011. An example of problems associated with DNA barcoding in tardigrades: a novel method for obtaining voucher specimens. Zootaxa. 3104: 42 - 51." type="journal article" year="2011">Cesari, 2011</bibRefCitation>
</taxonomicName>
).
</paragraph>
<caption id="25E0769BFFE0FF85FF62F99AEFF1F9F9" ID-DOI="http://doi.org/10.5281/zenodo.5197697" ID-Zenodo-Dep="5197697" httpUri="https://zenodo.org/record/5197697/files/figure.png" pageId="14" pageNumber="2421" startId="14.[211,278,1550,1572]" targetBox="[214,1169,169,1517]" targetPageId="14">
<paragraph id="71202613FFE0FF85FF62F99AEFF1F9F9" blockId="14.[211,1171,1550,1631]" pageId="14" pageNumber="2421">
Figure 6. Unrooted neighbour joining dendrogram computed on 
<taxonomicName id="B69F5D90FFE0FF85FC13F99BEF42F983" authorityName="C. A. S. Schultze" authorityYear="1834" box="[930,1056,1551,1573]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="2421" phylum="Tardigrada" rank="genus">
<emphasis id="43EBFA01FFE0FF85FC13F99BEF42F983" box="[930,1056,1551,1573]" italics="true" pageId="14" pageNumber="2421">Macrobiotus</emphasis>
</taxonomicName>
“ 
<taxonomicName id="B69F5D90FFE0FF85FB88F99BEFF1F983" authorityName="C. A. S. Schultze" authorityYear="1834" box="[1081,1171,1551,1573]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="2421" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFE0FF85FB88F99BEFF1F983" box="[1081,1171,1551,1573]" italics="true" pageId="14" pageNumber="2421">hufelandi</emphasis>
</taxonomicName>
group” 
<emphasis id="43EBFA01FFE0FF85FE93F9B8EA32F9E4" box="[290,336,1580,1602]" italics="true" pageId="14" pageNumber="2421">cox1</emphasis>
sequences. Bootstrap percentages computed after 2000 replicates are shown above branches. 
<taxonomicName id="B69F5D90FFE0FF85FE89F9DDE944F9F9" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[312,550,1609,1631]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="2421" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE0FF85FE89F9DDE944F9F9" box="[312,550,1609,1631]" italics="true" pageId="14" pageNumber="2421">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE0FF85FD9AF9DEE915F9F9" box="[555,631,1610,1631]" pageId="14" pageNumber="2421" rank="species">sp. nov.</taxonomicNameLabel>
specimens are shown in bold. Acronyms as in Table 1.
</paragraph>
</caption>
<caption id="25E0769BFFE1FF84FF20FF3DE85EFF7A" pageId="15" pageNumber="2422" startId="15.[145,200,169,191]">
<paragraph id="71202613FFE1FF84FF20FF3DE85EFF7A" blockId="15.[145,1101,169,220]" pageId="15" pageNumber="2422">Table 3. Kimura two-parameter distances computed among all newly analysed specimens (below the diagonal), and their standard error (above the diagonal).</paragraph>
</caption>
<paragraph id="71202613FFE1FF84FEB8FF6DE8D9FE1D" pageId="15" pageNumber="2422">
<table id="039FD4B3FFE10074FF20FF6DEF28FE1D" box="[145,1098,249,443]" gridcols="7" gridrows="6" pageId="15" pageNumber="2422">
<tr id="CFAF2451FFE10074FF20FF6DEF28FEA9" box="[145,1098,249,271]" gridrow="0" pageId="15" pageNumber="2422" rowspan-0="1">
<th id="8C7E4D2DFFE10074FF5FFF6DEAE7FEA9" box="[238,389,249,271]" gridcol="1" gridrow="0" pageId="15" pageNumber="2422">Specimen</th>
<th id="8C7E4D2DFFE10074FE6BFF6DE972FEA9" box="[474,528,249,271]" gridcol="2" gridrow="0" pageId="15" pageNumber="2422">1</th>
<th id="8C7E4D2DFFE10074FDD9FF6DE9FCFEA9" box="[616,670,249,271]" gridcol="3" gridrow="0" pageId="15" pageNumber="2422">2</th>
<th id="8C7E4D2DFFE10074FD46FF6DE84FFEA9" box="[759,813,249,271]" gridcol="4" gridrow="0" pageId="15" pageNumber="2422">3</th>
<th id="8C7E4D2DFFE10074FC34FF6DE8D9FEA9" box="[901,955,249,271]" gridcol="5" gridrow="0" pageId="15" pageNumber="2422">4</th>
<th id="8C7E4D2DFFE10074FBA5FF6DEF28FEA9" box="[1044,1098,249,271]" gridcol="6" gridrow="0" pageId="15" pageNumber="2422">5</th>
</tr>
<tr id="CFAF2451FFE10074FF20FEA4EF28FEE0" box="[145,1098,304,326]" gridrow="1" pageId="15" pageNumber="2422" rowspan-2="1">
<th id="8C7E4D2DFFE10074FF20FEA4EBFFFEE0" box="[145,157,304,326]" gridcol="0" gridrow="1" pageId="15" pageNumber="2422">1</th>
<td id="8C7E4D2DFFE10074FF5FFEA4EAE7FEE0" box="[238,389,304,326]" gridcol="1" gridrow="1" pageId="15" pageNumber="2422">C3291 A01-V1</td>
<td id="8C7E4D2DFFE10074FDD9FEA4E9FCFEE0" box="[616,670,304,326]" gridcol="3" gridrow="1" pageId="15" pageNumber="2422">0.000</td>
<td id="8C7E4D2DFFE10074FD46FEA4E84FFEE0" box="[759,813,304,326]" gridcol="4" gridrow="1" pageId="15" pageNumber="2422">0.006</td>
<td id="8C7E4D2DFFE10074FC34FEA4E8D9FEE0" box="[901,955,304,326]" gridcol="5" gridrow="1" pageId="15" pageNumber="2422">0.005</td>
<td id="8C7E4D2DFFE10074FBA5FEA4EF28FEE0" box="[1044,1098,304,326]" gridcol="6" gridrow="1" pageId="15" pageNumber="2422">0.005</td>
</tr>
<tr id="CFAF2451FFE10074FF20FED9EF28FEC5" box="[145,1098,333,355]" gridrow="2" pageId="15" pageNumber="2422" rowspan-3="1">
<th id="8C7E4D2DFFE10074FF20FED9EBFFFEC5" box="[145,157,333,355]" gridcol="0" gridrow="2" pageId="15" pageNumber="2422">2</th>
<td id="8C7E4D2DFFE10074FF5FFED9EAE7FEC5" box="[238,389,333,355]" gridcol="1" gridrow="2" pageId="15" pageNumber="2422">C3291 A02-V2</td>
<td id="8C7E4D2DFFE10074FE6BFED9E972FEC5" box="[474,528,333,355]" gridcol="2" gridrow="2" pageId="15" pageNumber="2422">0.000</td>
<td id="8C7E4D2DFFE10074FD46FED9E84FFEC5" box="[759,813,333,355]" gridcol="4" gridrow="2" pageId="15" pageNumber="2422">0.006</td>
<td id="8C7E4D2DFFE10074FC34FED9E8D9FEC5" box="[901,955,333,355]" gridcol="5" gridrow="2" pageId="15" pageNumber="2422">0.005</td>
<td id="8C7E4D2DFFE10074FBA5FED9EF28FEC5" box="[1044,1098,333,355]" gridcol="6" gridrow="2" pageId="15" pageNumber="2422">0.005</td>
</tr>
<tr id="CFAF2451FFE10074FF20FEFEEF28FE26" box="[145,1098,362,384]" gridrow="3" pageId="15" pageNumber="2422" rowspan-4="1">
<th id="8C7E4D2DFFE10074FF20FEFEEBFFFE26" box="[145,157,362,384]" gridcol="0" gridrow="3" pageId="15" pageNumber="2422">3</th>
<td id="8C7E4D2DFFE10074FF5FFEFEEAE7FE26" box="[238,389,362,384]" gridcol="1" gridrow="3" pageId="15" pageNumber="2422">C3291 A03</td>
<td id="8C7E4D2DFFE10074FE6BFEFEE972FE26" box="[474,528,362,384]" gridcol="2" gridrow="3" pageId="15" pageNumber="2422">0.014</td>
<td id="8C7E4D2DFFE10074FDD9FEFEE9FCFE26" box="[616,670,362,384]" gridcol="3" gridrow="3" pageId="15" pageNumber="2422">0.015</td>
<td id="8C7E4D2DFFE10074FC34FEFEE8D9FE26" box="[901,955,362,384]" gridcol="5" gridrow="3" pageId="15" pageNumber="2422">0.000</td>
<td id="8C7E4D2DFFE10074FBA5FEFEEF28FE26" box="[1044,1098,362,384]" gridcol="6" gridrow="3" pageId="15" pageNumber="2422">0.000</td>
</tr>
<tr id="CFAF2451FFE10074FF20FE1CEF28FE38" box="[145,1098,392,414]" gridrow="4" pageId="15" pageNumber="2422" rowspan-5="1">
<th id="8C7E4D2DFFE10074FF20FE1CEBFFFE38" box="[145,157,392,414]" gridcol="0" gridrow="4" pageId="15" pageNumber="2422">4</th>
<td id="8C7E4D2DFFE10074FF5FFE1CEAE7FE38" box="[238,389,392,414]" gridcol="1" gridrow="4" pageId="15" pageNumber="2422">C3291 A04</td>
<td id="8C7E4D2DFFE10074FE6BFE1CE972FE38" box="[474,528,392,414]" gridcol="2" gridrow="4" pageId="15" pageNumber="2422">0.014</td>
<td id="8C7E4D2DFFE10074FDD9FE1CE9FCFE38" box="[616,670,392,414]" gridcol="3" gridrow="4" pageId="15" pageNumber="2422">0.014</td>
<td id="8C7E4D2DFFE10074FD46FE1CE84FFE38" box="[759,813,392,414]" gridcol="4" gridrow="4" pageId="15" pageNumber="2422">0.000</td>
<td id="8C7E4D2DFFE10074FBA5FE1CEF28FE38" box="[1044,1098,392,414]" gridcol="6" gridrow="4" pageId="15" pageNumber="2422">0.000</td>
</tr>
<tr id="CFAF2451FFE10074FF20FE31EF28FE1D" box="[145,1098,421,443]" gridrow="5" pageId="15" pageNumber="2422" rowspan-6="1">
<th id="8C7E4D2DFFE10074FF20FE31EBFFFE1D" box="[145,157,421,443]" gridcol="0" gridrow="5" pageId="15" pageNumber="2422">5</th>
<td id="8C7E4D2DFFE10074FF5FFE31EAE7FE1D" box="[238,389,421,443]" gridcol="1" gridrow="5" pageId="15" pageNumber="2422">C3291 A05</td>
<td id="8C7E4D2DFFE10074FE6BFE31E972FE1D" box="[474,528,421,443]" gridcol="2" gridrow="5" pageId="15" pageNumber="2422">0.014</td>
<td id="8C7E4D2DFFE10074FDD9FE31E9FCFE1D" box="[616,670,421,443]" gridcol="3" gridrow="5" pageId="15" pageNumber="2422">0.014</td>
<td id="8C7E4D2DFFE10074FD46FE31E84FFE1D" box="[759,813,421,443]" gridcol="4" gridrow="5" pageId="15" pageNumber="2422">0.000</td>
<td id="8C7E4D2DFFE10074FC34FE31E8D9FE1D" box="[901,955,421,443]" gridcol="5" gridrow="5" pageId="15" pageNumber="2422">0.000</td>
</tr>
</table>
</paragraph>
<paragraph id="71202613FFE1FF84FF20FE40EAE0FE4C" blockId="15.[145,386,468,490]" box="[145,386,468,490]" pageId="15" pageNumber="2422">
<tableNote id="8C79279DFFE1FF84FF20FE40EAE0FE4C" box="[145,386,468,490]" pageId="15" pageNumber="2422" targetBox="[145,1098,249,443]" targetPageId="15">Acronyms as in Table 1.</tableNote>
</paragraph>
<caption id="25E0769BFFE1FF84FF3EFAC7EA5FFA05" ID-DOI="http://doi.org/10.5281/zenodo.5197699" ID-Zenodo-Dep="5197699" httpUri="https://zenodo.org/record/5197699/files/figure.png" pageId="15" pageNumber="2422" startId="15.[143,210,1363,1385]" targetBox="[145,1102,533,1330]" targetPageId="15">
<paragraph id="71202613FFE1FF84FF3EFAC7EA5FFA05" blockId="15.[143,1103,1363,1443]" pageId="15" pageNumber="2422">
Figure 7. Bayesian inference dendrogram computed on 18S rRNA sequences. Numbers near nodes indicate posterior probability. 
<taxonomicName id="B69F5D90FFE1FF84FDB0FAE4E98DFA20" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[513,751,1392,1414]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="2422" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE1FF84FDB0FAE4E98DFA20" box="[513,751,1392,1414]" italics="true" pageId="15" pageNumber="2422">Macrobiotus kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE1FF84FD47FAE5E820FA20" box="[758,834,1393,1414]" pageId="15" pageNumber="2422" rank="species">sp. nov.</taxonomicNameLabel>
specimen is shown in bold (C3291 A02-V2).
</paragraph>
</caption>
<paragraph id="71202613FFE1FF9BFF0BFA76EAF6FCDA" blockId="15.[143,1104,1506,1691]" lastBlockId="16.[211,1172,677,893]" lastPageId="16" lastPageNumber="2423" pageId="15" pageNumber="2422">
Molecular analysis of the mitochondrial DNA 
<emphasis id="43EBFA01FFE1FF84FD5CFA77E843FA5D" box="[749,801,1507,1531]" italics="true" pageId="15" pageNumber="2422">cox1</emphasis>
gene was carried out on sequences of 491–624 bp. Two different sequences (i.e. two haplotypes) are present in the 
<specimenCount id="6799ED9AFFE1FF84FF06F9B6EA37F99C" box="[183,341,1570,1594]" pageId="15" pageNumber="2422" type="generic">five specimens</specimenCount>
analysed: 
<specimenCount id="6799ED9AFFE1FF84FE79F9B6E91AF99C" box="[456,632,1570,1594]" pageId="15" pageNumber="2422" type="generic">three specimens</specimenCount>
are of one haplotype, 
<specimenCount id="6799ED9AFFE1FF84FCDBF9B7EF69F99C" box="[874,1035,1570,1594]" pageId="15" pageNumber="2422" type="generic">two specimens</specimenCount>
are of the other (
<tableCitation id="3C1D13A8FFE1FF84FEB5F9D6EA36F9FC" box="[260,340,1602,1626]" captionStart="Table 3" captionStartId="15.[145,200,169,191]" captionText="Table 3. Kimura two-parameter distances computed among all newly analysed specimens (below the diagonal), and their standard error (above the diagonal)." pageId="15" pageNumber="2422">Table 3</tableCitation>
; 
<figureCitation id="E9A43A96FFE1FF84FED3F9D6EAA2F9FC" box="[354,448,1602,1626]" captionStart="Figure 7" captionStartId="15.[143,210,1363,1385]" captionTargetBox="[145,1102,533,1330]" captionTargetId="figure-76@15.[145,1102,533,1330]" captionTargetPageId="15" captionText="Figure 7. Bayesian inference dendrogram computed on 18S rRNA sequences. Numbers near nodes indicate posterior probability. Macrobiotus kristenseni sp. nov. specimen is shown in bold (C3291 A02-V2)." figureDoi="http://doi.org/10.5281/zenodo.5197699" httpUri="https://zenodo.org/record/5197699/files/figure.png" pageId="15" pageNumber="2422">Figure 7</figureCitation>
). The two haplotypes are very similar and the intraspecific genetic distance among 
<taxonomicName id="B69F5D90FFE1FF84FE10F9F7E923F9DC" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[417,577,1634,1658]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="2422" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFE1FF84FE10F9F7E923F9DC" box="[417,577,1634,1658]" italics="true" pageId="15" pageNumber="2422">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFE1FF84FDFBF9F7E9C2F9DC" box="[586,672,1635,1658]" pageId="15" pageNumber="2422" rank="species">sp. nov.</taxonomicNameLabel>
specimens is low (1.4–1.5%; 
<tableCitation id="3C1D13A8FFE1FF84FC5DF9F6EF5DF9DC" box="[1004,1087,1634,1658]" captionStart="Table 3" captionStartId="15.[145,200,169,191]" captionText="Table 3. Kimura two-parameter distances computed among all newly analysed specimens (below the diagonal), and their standard error (above the diagonal)." pageId="15" pageNumber="2422">Table 3</tableCitation>
). Mean K2P distances between specimens pertaining to different species are very high (18.2–33.4%; 
<tableCitation id="3C1D13A8FFFEFF9BFEDDFD31EADCFD1B" box="[364,446,677,701]" captionStart="Table 4" captionStartId="16.[213,268,169,191]" captionText="Table 4. Mean Kimura two-parameter distances computed among different species pertaining to the Macrobiotus hufelandi group (below the diagonal), and their standard errors (above the diagonal), and within each species (column D)." pageId="16" pageNumber="2423">Table 4</tableCitation>
). In particular, among the considered species, 
<taxonomicName id="B69F5D90FFFEFF9BFC7FFD32EF0FFD1B" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[974,1133,677,701]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="2423" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFFEFF9BFC7FFD32EF0FFD1B" box="[974,1133,677,701]" italics="true" pageId="16" pageNumber="2423">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFFEFF9BFBC4FD31EA61FD7A" pageId="16" pageNumber="2423" rank="species">sp. nov.</taxonomicNameLabel>
has the greatest genetic difference with respect to the other 
<taxonomicName id="B69F5D90FFFEFF9BFC08FD52EF25FD7B" authorityName="C. A. S. Schultze" authorityYear="1834" box="[953,1095,709,733]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="2423" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFFEFF9BFC08FD52EF25FD7B" box="[953,1095,709,733]" italics="true" pageId="16" pageNumber="2423">M. hufelandi</emphasis>
</taxonomicName>
group of species (23.1–33.4%), while the difference among the considered species excluding 
<taxonomicName id="B69F5D90FFFEFF9BFF62FC91EA17FCBA" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[211,373,772,796]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="2423" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFFEFF9BFF62FC91EA17FCBA" box="[211,373,772,796]" italics="true" pageId="16" pageNumber="2423">M. kristenseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D8477AFFFEFF9BFECEFC91EAB4FCBA" box="[383,470,773,796]" pageId="16" pageNumber="2423" rank="species">sp. nov.</taxonomicNameLabel>
is 18.2–24.4%. The neighbour-joining dendrogram based on 
<emphasis id="43EBFA01FFFEFF9BFF62FCB0EA65FC9A" box="[211,263,804,828]" italics="true" pageId="16" pageNumber="2423">cox1</emphasis>
sequences confirms these differences between the 
<specimenCount id="6799ED9AFFFEFF9BFC82FCB1E8E7FC9A" box="[819,901,805,828]" pageId="16" pageNumber="2423" type="generic">
two 
<taxonomicName id="B69F5D90FFFEFF9BFCD2FCB1E8E7FC9A" authority="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo, 2013" authorityName="Guidetti &amp; Peluffo &amp; Rocha &amp; Cesari &amp; Peluffo" authorityYear="2013" box="[867,901,805,828]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="2423" phylum="Tardigrada" rank="species" species="kristenseni" status="sp. nov.">
<emphasis id="43EBFA01FFFEFF9BFCD2FCB1E8E7FC9A" box="[867,901,805,828]" italics="true" pageId="16" pageNumber="2423">M.</emphasis>
</taxonomicName>
</specimenCount>
kristenseni 
<taxonomicNameLabel id="58D8477AFFFEFF9BFBB6FCB1EF39FC9A" box="[1031,1115,805,828]" pageId="16" pageNumber="2423" rank="species">sp. nov.</taxonomicNameLabel>
haplotypes, and the differences among this species and the other species of 
<taxonomicName id="B69F5D90FFFEFF9BFBB4FCD1EFF1FCFA" authorityName="C. A. S. Schultze" authorityYear="1834" box="[1029,1171,836,860]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="2423" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="43EBFA01FFFEFF9BFBB4FCD1EFF1FCFA" box="[1029,1171,836,860]" italics="true" pageId="16" pageNumber="2423">M. hufelandi</emphasis>
</taxonomicName>
group (
<figureCitation id="E9A43A96FFFEFF9BFE94FCF0EAE1FCDA" box="[293,387,868,892]" captionStart="Figure 7" captionStartId="15.[143,210,1363,1385]" captionTargetBox="[145,1102,533,1330]" captionTargetId="figure-76@15.[145,1102,533,1330]" captionTargetPageId="15" captionText="Figure 7. Bayesian inference dendrogram computed on 18S rRNA sequences. Numbers near nodes indicate posterior probability. Macrobiotus kristenseni sp. nov. specimen is shown in bold (C3291 A02-V2)." figureDoi="http://doi.org/10.5281/zenodo.5197699" httpUri="https://zenodo.org/record/5197699/files/figure.png" pageId="16" pageNumber="2423">Figure 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>