treatments-xml/data/F7/63/07/F7630765E88959BA861E816D0E35EB0B.xml

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<mods:title id="B3811CAEDC49F9F4279E4111C9938D5E">Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007 (Coleoptera, Tenebrionidae, Ulomini): Descriptions of the larva and pupa and new distributional records</mods:title>
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<mods:namePart id="91CC2192C32DD8BCEAEC13E8DC1DF27C">Niu, Yiping</mods:namePart>
<mods:affiliation id="3B9F17BA0D9BAA31D883EB6A52486987">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Institute of Life Science and Green Development, Hebei University, 071002, Baoding, China</mods:affiliation>
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<mods:namePart id="58682DA10E2BA1D286440DC251AABB6D">Ren, Guodong</mods:namePart>
<mods:affiliation id="C8647298637DDC1FA255733608FA222F">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Institute of Life Science and Green Development, Hebei University, 071002, Baoding, China</mods:affiliation>
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<mods:namePart id="7FDB419F0EEB173B069953D087FBDDCB">Liu, Shanshan</mods:namePart>
<mods:affiliation id="224B3F0D7B47D64E9B4AD20BCEE5D42C">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Institute of Life Science and Green Development, Hebei University, 071002, Baoding, China</mods:affiliation>
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<taxonomicName id="4B40CBD77446509F2D6F91483C7249ED" LSID="F7630765-E889-59BA-861E-816D0E35EB0B" authority="Liu, Ren &amp; Wang, 2007" authorityName="Liu, Ren &amp; Wang" authorityYear="2007" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma (Uloma) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="species" species="intricornicula" subGenus="Uloma">Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007</taxonomicName>
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<paragraph id="216C29AEF8C6327BD18275AC27A3D2A6" pageId="0" pageNumber="107036">
<taxonomicName id="0777D3A6E3E18D751B25B67ADD0F6CEB" authorityName="Liu, Ren &amp; Wang" authorityYear="2007" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="species" species="intricornicula">Uloma intricornicula</taxonomicName>
Liu, Ren &amp; Wang, 2007 - 
<bibRefCitation id="96AAE08F911FB7412B26329010B7DB7D" author="Liu, S. S." journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="107036" pagination="70 - 75" refId="B9839293" refString="Liu, S. S., Ren, G. D., Wang, J. S., 2007. Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini). Acta Zootaxonomica Sinica 32 (1): 70 - 75" title="Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini)" volume="32" year="2007">Liu et al. (2007)</bibRefCitation>
: 71 (type locality: China, Fujian, Guadun).
</paragraph>
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<paragraph id="BBBC332E52FF48109AF245BECCC37FD8" pageId="0" pageNumber="107036">Description</paragraph>
<paragraph id="510EAF3F8697AEF4D05E3920B845CBF7" pageId="0" pageNumber="107036">
<emphasis id="8284E32CA720AC0CA5ADAE5966B514E9" bold="true" pageId="0" pageNumber="107036">Larva</emphasis>
: Oligopod (Fig. 
<figureCitation id="ABE3C138FD239115FE91A61BE1DD525D" captionStart="Figure 3" captionStartId="F10364806" captionText="Figure 3. Habitus of the larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. lateral view; B. dorsal view; C. ventral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure3" httpUri="https://binary.pensoft.net/fig/888235" pageId="0" pageNumber="107036">3</figureCitation>
). 
<emphasis id="2E9AFF507625E61B1C545AF54CC0D1DE" bold="true" pageId="0" pageNumber="107036">Body.</emphasis>
Length 13.5-14.0 mm. Body subcylindrical, ventrally flat and with sharp tail-end; evenly sclerotised both dorsally and ventrally; yellowish-brown, dorsum and both ends brown. Vestiture smooth, suffused with large and round punctures both dorsally and laterally.
</paragraph>
<paragraph id="999EB3E863D74D7D3E6FB254B589EE0A" pageId="0" pageNumber="107036">
<emphasis id="DCAD52FE2B05EB21268F52755012B2DD" bold="true" pageId="0" pageNumber="107036">Head.</emphasis>
Head (Fig. 
<figureCitation id="99CB87647B1E3FA9B56F576B25585AF5" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
) about as wide as prothorax, trapezoidal, slightly convex dorsally; with 10 long setae near the anterior margin, 6 in the middle and 10 near the posterior margin. Frontoclypeal suture evident, almost straight in the middle. Frons slightly convex, frontal sutures broadly V-shaped, distinctly incurved near the centre; median suture barely visible in dorsal view. Clypeus (Fig. 
<figureCitation id="770222722876F0C3592A038D99B75E12" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
C) slightly flat, with anterior margin linear, bearing four erect clypeal setae, with the central two longer. Labrum (Fig. 
<figureCitation id="C9A3A2C02C9147CCB0F83200E8DADD3A" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
C) transverse, semi-elliptic, slightly convex, with several short setae on anterior margin and four long erect median setae. Ocelli black, divided into two parts, the upper of which smaller than the lower. Antennae (Fig. 
<figureCitation id="13ADA096A12048809D7FC430DB8D9533" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
A, Fig. 
<figureCitation id="92FFC51BC3792161F01DBD6989072449" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B) shorter than half length of head; antennomere I short; II cylindrical, more than twice as wide as long and three times longer than I, sensorium nearly C-shaped; III thin and short, about 1/6 as long as II and with one long erect median seta and three very short setae around the base of long one at apex of III. Mandibles (Fig. 
<figureCitation id="9181CF842FB060030EDA41DEEA4C5C8F" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B, Fig. 
<figureCitation id="3C7C91CB8BA8C3822DE538D663AF9010" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
C) well developed, distally black, extended anteriorly; tridentate apically, with the apical tooth markedly larger than the dorsal and ventral ones. Maxillae almost parallel-sided, with dense setae on the apical inner margin; maxillary palpi (Fig. 
<figureCitation id="88ACBCA3CBB3BFB832D0E3D52DA8409F" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
A, Fig. 
<figureCitation id="0FDAC8B813A0C403F0EF56D9B3806136" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B) distolateral, subcylindrical, gradually narrowing towards the apex, with the terminal palpomere slender and short. Labial palpi (Fig. 
<figureCitation id="E40C4973F84D20AF3182602C9380631E" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
A, Fig. 
<figureCitation id="0B867BCF8F17F652E59C1848E9A79D4C" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B) 2-segmented, short. Ligula (Fig. 
<figureCitation id="B839F1F249C5F2B5F2E304CCF32338A0" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
A, Fig. 
<figureCitation id="545ED29E16C5BF1447A67391E5FBB00B" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B) slightly convex, with two long erect setae, anterior margin distinctly protruding in the middle part. Mentum (Fig. 
<figureCitation id="2D4241AC9EDCEBC145E1754F0DAF1FB0" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
A, Fig. 
<figureCitation id="DCA72D1AC28FD5932028A968DAC41295" captionStart="Figure 4" captionStartId="F10364808" captionText="Figure 4. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, ventral view; B. head, ventral view; C. head, anterior view; D. head, dorsal view. a, h. antenna; b, g. maxillary palpus; c, l. labial palpus; d, i. ligula; e, j. mentum; k, m. mandible; f, n. labrum; o. clypeus. Scale bars: 0.5 mm." figureDoi="10.3897/BDJ.11.e107036.figure4" httpUri="https://binary.pensoft.net/fig/888239" pageId="0" pageNumber="107036">4</figureCitation>
B) subhexagonal, widest in the middle, anterior margin weakly emarginate, with two long erect setae on both sides and posterior part respectively, the latter much thicker and longer.
</paragraph>
<paragraph id="901BE870AB0279957691DEC498773A73" pageId="0" pageNumber="107036">
<emphasis id="F538B67ADAFE6BE35D1F67DBE90F81B6" bold="true" pageId="0" pageNumber="107036">Thorax.</emphasis>
Thorax 3-segmented. Each thoracic tergum with long erect setae near sides of anterior and posterior margins, distributed as follows: 4 and 3 setae on anterior and posterior margins of prothoracic tergum, respectively; 1 and 2 setae on mesothoracic tergum; and 1 and 3 setae on metathoracic tergum. Prothoracic tergum longer than wide, nearly rectangular in dorsal view, about twice as long as the meso- or metathoracic tergum; ratios between thoracic terga as follows: 1.5: 0.7: 0.9. Meso- and metathoracic terga transverse, nearly rectangular in dorsal view. Mesothoracic spiracles (Fig. 
<figureCitation id="E0BF9F9EAE5B2EB0F842E254FE8B33CB" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
A) placed on anterolateral margins of tergum, near distinctly separate coxal cavity, visible in ventral view, slenderly oval in shape, large and approximately three times larger than the abdominal spiracles. Prothorax and metathorax without spiracles.
</paragraph>
<paragraph id="BDD9B9469F10F6D4075F1CCEC2A18C2F" pageId="0" pageNumber="107036">
<emphasis id="7211512C789483790891B7CEC64DD902" bold="true" pageId="0" pageNumber="107036">Legs.</emphasis>
Pro-, meso- and metathoracic legs short, subequal in length and similar in shape (Fig. 
<figureCitation id="A9D145815B5A0EA21BF918D5DE6D81FC" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
A). Coxa of prothoracic leg thick, longer than other segments, with 5-6 long spiniform setae on anterior margin and 2 on posterior margin; trochanter subtriangular, with 2 short spiniform setae on anterior margin, one on posterior margin and one long hair in the middle; femur subequal in length and width, with sparse short hairs, with 3 short spiniform setae on anterior margin and 5 on posterior margin and one long hair in the middle; tibia much more slender and shorter, with sparse short hairs, with 2 short spiniform setae on anterior margin and 5 on posterior margin; tarsungulus falciform, with 2 thinner short spiniform setae under it. Coxa of mesothoracic leg with 6-7 long spiniform setae on anterior margin and 2 on posterior margin; trochanter with 2 short spiniform setae on anterior margin and one on posterior margin; femur with 3 short spiniform setae on anterior margin and 5 on posterior margin; tibia with 2 short spiniform setae on anterior margin and 4 on posterior margin; others similar to those of prothoracic leg. Coxa of metathoracic leg with 5-6 long spiniform setae on anterior margin and 2 on posterior margin; trochanter with 2 short spiniform setae on anterior margin and only one on posterior margin; femur with 3 short spiniform setae on anterior margin and 5 on posterior margin; tibia with 2 short spiniform setae on anterior margin and 4 on posterior margin; others similar to those of prothoracic leg.
</paragraph>
<paragraph id="705F313A139384BA7065491AC816B0FB" pageId="0" pageNumber="107036">
<emphasis id="8DB2F9F8CF7F216CCE1463780CAF0DA9" bold="true" pageId="0" pageNumber="107036">Abdomen.</emphasis>
Abdomen 9-segmented, gradually and slightly darker towards apex, slightly enlarged backwards. Segments I-IX with denser punctures on the basal dorsal surface in comparison with other parts of abdominal segments. Tergites I-II wider than long, nearly rectangular in dorsal view; III-VIII subquadrate in dorsal view. Tergites I-VII with an arcuate shallow impression respectively near posterolateral margins, with 4 long erect setae on sides of posterior margins; only tergite I with 2 long erect setae on sides of anterior margin. Sternites I-VII nearly rectangular, longer than wide and with a long erect seta near each corner; sternites I with other 4 long erect setae near the anterior margin. Segment VIII without pleural sutures, with 2 long erect setae respectively on posterolateral margins, with 2 erect setae near the anterior margin and 6 near the middle of posterior margin of ventral surface. Segment IX (Fig. 
<figureCitation id="63A6BB5BB2F84855C7E64784A2FBF9DB" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
B, Fig. 
<figureCitation id="EB5001B71EFDF071364161D5779400A4" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
C) with dense even large punctures and sparse long erect setae; dorsal surface with 2 setae; ventral surface with 6 setae at the centre, anterior margin almost straight; tip-end with 4 setae surrounding it. Segment IX parabolic, as long as wide, subcircular in cross-section, tip-end slightly round with a small papilla, but without urogomphi. Anus concealed in posterior part of abdominal tergite VIII, without anal tubes. Abdominal spiracles (Fig. 
<figureCitation id="0289C477C56F4140E26A7BFA1C98B649" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
A) round, of the same size, opening on anterolateral margins of segments I-VIII.
</paragraph>
<paragraph id="70F096A2CA46110CB439B0AB5D701B92" pageId="0" pageNumber="107036">
<emphasis id="EAF4125744DECBC84749D32558FCB31E" bold="true" pageId="0" pageNumber="107036">Remarks</emphasis>
</paragraph>
<paragraph id="E0AB9A2F6D3AEA768819D2C81194DBB0" pageId="0" pageNumber="107036">
The description above is based on the last instar larvae, whose exuvia was preserved in 75% alcohol (Fig. 
<figureCitation id="157E29D3A25A8B11DCBBE667179DAEAA" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
F).
</paragraph>
<paragraph id="D50115473FE9E951680C94D810AD962A" pageId="0" pageNumber="107036">
The larvae of 
<taxonomicName id="17ABB04B96AFE1EC1767479A35C06A89" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="187244A4EAFF70E5299102BE626C0A70" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
differ from other known tenebrionine larvae by the presence of an elongate anterior extension on the hypopharygeal sclerome (as in 
<taxonomicName id="7F8F743BDEA4DA584442317A5CB4D513" authorityName="Laporte de Castelnau" authorityYear="1840" lsidName="" pageId="0" pageNumber="107036" rank="subFamily" subFamily="Alleculinae">Alleculinae</taxonomicName>
Laporte, 1840), a paraboloid abdominal segment IX with an apical point, the lack of cerci, the lack of pleurosternal sutures on abdominal segment VIII, a reduced anal region and the lack of anal tubes (
<bibRefCitation id="8AFB6603D35CB536307BD890FF6EEA38" author="Hayashi, N." journalOrPublisher="Insecta Matsumurana" pageId="0" pageNumber="107036" pagination="1 - 41" refId="B9839222" refString="Hayashi, N., 1966. A contribution to the knowledge of the larvae of Tenebrionidae occurring in Japan (Coleoptera: Cucujoidea). Insecta Matsumurana 1 (Supplement): 1 - 41" title="A contribution to the knowledge of the larvae of Tenebrionidae occurring in Japan (Coleoptera: Cucujoidea)" volume="1" year="1966">Hayashi 1966</bibRefCitation>
, 
<bibRefCitation id="BCAE27957F8E1DE965D6CFFBB68E7DBF" author="Watt, J. C." journalOrPublisher="New Zealand Journal of Zoology" pageId="0" pageNumber="107036" pagination="381 - 452" refId="B9839440" refString="Watt, J. C., 1974. A revised subfamily classification of Tenebrionidae (Coleoptera). New Zealand Journal of Zoology 1: 381 - 452" title="A revised subfamily classification of Tenebrionidae (Coleoptera)" volume="1" year="1974">Watt 1974</bibRefCitation>
). Although there is a substantial morphological homogeneity amongst larvae of different species in the genus 
<taxonomicName id="D1590E0E232FB7C4883AE9C90E01F81E" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="2B07CCE7EB01764EEB3205FC9A8E8403" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
, we found some important differences between 
<taxonomicName id="B833422E7049B9A1E81E6F39C6A9A5FC" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
and 
<taxonomicName id="C4A797529AA7B5A7A084A6752A9D4840" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
. Namely, mentum is relatively slender and widest in the middle in 
<taxonomicName id="0FD004C299CCC6C948968B1FB1EAF6CB" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
, while it has the maximum width more anteriorly (4/5 from the base) in 
<taxonomicName id="640596D37974FE2F48AC2C8D14303B75" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
; protibia with 2 short spiniform setae on the anterior margin in 
<taxonomicName id="467BE9F00651DBB030CC4B9DAD253D20" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
, with only one seta in 
<taxonomicName id="028D75195360F125EE948B57BD95062C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
; mesotibia with 2 short spiniform setae on the anterior margin in 
<taxonomicName id="22063D69F44696B440289A8DB157EBE5" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
, with only one seta in 
<taxonomicName id="D26C72986CAC302DE41AD67FEA4B96F7" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
; abdominal segment IX with slightly round tip-end and the anterior margin of the ventral surface almost straight in 
<taxonomicName id="B7487478549383520651E67F904F2CE3" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
(Fig. 
<figureCitation id="A974C17DB1B49F2E69F8D08DE63D1A6C" captionStart="Figure 5" captionStartId="F10364810" captionText="Figure 5. Larva of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. thorax segments, ventral view; a. prothoracic leg; b. mesothoracic leg; c. metathoracic leg; d. mesothoracic spiracle; e. abdominal spiracle. B. abdominal segment IX, lateral view. C. abdominal segment IX, ventral view. Scale bars: 0.1 mm (d, e), 1 mm (others). The arrow indicates that the anterior margin of the ventral surface of abdominal segment IX is almost straight." figureDoi="10.3897/BDJ.11.e107036.figure5" httpUri="https://binary.pensoft.net/fig/888241" pageId="0" pageNumber="107036">5</figureCitation>
C), abdominal segment IX with slightly pointed tip-end and the anterior margin of the ventral surface emarginate in 
<taxonomicName id="C121266B62FA769A337CAFEABC27072C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
.
</paragraph>
<paragraph id="11BB02AB07C98BB4159097299429A9C9" pageId="0" pageNumber="107036">
However, since available descriptions of other 
<taxonomicName id="C65EC319BCCE25905F8AB8D8A6DDC0A1" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="317E2E03DB31836A19F7F2109D5F0EEC" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
species are not detailed enough to permit good morphological comparisons, it is difficult to distinguish the larvae of 
<taxonomicName id="19A5B26ED11D7777B1E56C2A1BA7496C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
and 
<taxonomicName id="415B89167750C8CCC0E036C4DEE55120" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
from other 
<taxonomicName id="F4401DA7A076D579DF5A94C8F8040C8C" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="4AAB8CB2FE78B7AAE3D35219AD14BAC1" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
larvae without direct examination of the specimens.
</paragraph>
<paragraph id="F571D9089DE8D1C7E7B84A495415F1D5" pageId="0" pageNumber="107036">
<emphasis id="303E9FC6CA02D9A4B32730780B853AAB" bold="true" pageId="0" pageNumber="107036">Pupa</emphasis>
: Exarate (Fig. 
<figureCitation id="72A8A6340D917A6FF45291620F15092C" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
). 
<emphasis id="8E82B16B095EEFB1704D96C3DAB25055" bold="true" pageId="0" pageNumber="107036">Body.</emphasis>
Length 6.0-7.0 mm, width 2.9-3.1 mm. Body small, slightly elliptic, extremely arched dorsally; grey to light brownish, with darker mouthparts, legs and body backend; most characteristics similar to that of adults of the same species.
</paragraph>
<paragraph id="1CD3BB34588A7BF49A04C972EB519025" pageId="0" pageNumber="107036">
<emphasis id="9220B408854887802985B83ECDE92C74" bold="true" pageId="0" pageNumber="107036">Head.</emphasis>
Visible in ventral view (Fig. 
<figureCitation id="CF2C2926ECD5729508CD348BFF8B9628" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
B, Fig. 
<figureCitation id="5302F4A18DA6C5FCFD7368CF2D0A8CBB" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
A). Smooth, with transverse wrinkles. Frons depressed, anterior margin with 4 tubercles bearing short erect apical setae in an arc sparsely and posterior margin with 2 far apart tubercles bearing short erect apical setae. Frontoclypeal suture almost linear. Clypeus slightly linear on anterior margin, elevated with 2 small ridges at centre and with a tubercle bearing short erect setae separately on apices of 2 ridges and anterolateral clypeus. Labrum relatively broad, semi-elliptic, anterior margin emarginate at centre, with sparse short hairs. Mandible with sparse long hairs laterally, apices darker, apical tooth distinctly larger than dorsal and ventral ones. Maxillary palpi and labial palpi clearly visible, with sparse long hairs laterally. Eye ovate, with sparse short hairs around it, one of them on anterior margin relatively longer. Antenna thick, claviform, gradually widening towards apex, antennomeres VII-X of distinctly increasing width, XI semi-spherical, V and VII slightly sharply protruding on inner border, I-XI with sparse tubercles bearing short hairs on each apical side and lateral side.
</paragraph>
<paragraph id="424D1BC07BD434EF39DFAF4CF5E6CA4A" pageId="0" pageNumber="107036">
<emphasis id="2F4F8625613373C5EBFFB9B65387C094" bold="true" pageId="0" pageNumber="107036">Thorax.</emphasis>
Thorax 3-segmented. Pronotum slightly transverse, similar in shape to that of adults, about 1.35 times as wide as long, widest near the middle. Pronotum with distinct transverse wrinkles and sparse apical setose tubercles (Fig. 
<figureCitation id="ECF8467CFF903ED8F2964C962AECA9A5" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
B, Fig. 
<figureCitation id="BEEFC3C725170D901DA28B9B5C57131B" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
C), tubercles denser on anterior and posterior margins, but sparse on lateral margins and even sparser on disc. Disc slightly convex, with a transverse deep anterior impression. Anterior margin slightly emarginate at centre. Lateral margins markedly arcuate, narrowing forward and less so from widest point to base. Posterior margin slightly protruding at centre. Anterior angles nearly rectangular, posterior angles obtuse. Meso- and metanotum (Fig. 
<figureCitation id="E7933064E3D5D58BC16B9630D9AC3734" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
A) glabrous with irregular wrinkles, distinctly narrower than pronotum, mesonotum slightly wider than abdominal tergite I and metanotum distinctly wider than abdominal tergite I. Mesonotum elevated at centre. Metanotum slightly elevated. Elytral and hind wing sheaths (Fig. 
<figureCitation id="EC3F628026CBF24FEF3CFE00E977EB1F" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
B, Fig. 
<figureCitation id="E75A4F86F023BE0759DD7347C1E3A636" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
C) glabrous, with several transverse wrinkles; elytra relatively distinctly punctatostriate, but faintly punctate.
</paragraph>
<paragraph id="E6FF22B5A557A81BC74ADD1414022A25" pageId="0" pageNumber="107036">
<emphasis id="A6422508CF779EC4F443F5D8F541A016" bold="true" pageId="0" pageNumber="107036">Legs.</emphasis>
Legs similar in shape to that of adults (Fig. 
<figureCitation id="E66A72F99EDB70E1F9A16EEFD1DE9371" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
B). Femur thick, with sparse apical setose tubercles on edges. Tibia almost glabrous, with 2 apical spurs on inner edge; protibia gradually explanate towards apex significantly, meso- and metatibia more slightly explanate. Tarsus relatively slender, with small tubercles on ventral surface. Tarsal claws with 2 small tubercles at apex of ventral surface.
</paragraph>
<paragraph id="4C91CAE352DD65A196A28FF8B8BD0ECE" pageId="0" pageNumber="107036">
<emphasis id="EF8F95005F78A7D93E8F412B5DFA440F" bold="true" pageId="0" pageNumber="107036">Abdomen.</emphasis>
Abdomen 9-segmented, dorsomeson quite distinct. Abdominal tergites glabrous, relatively broad, slightly convex and with dense longitudinal wrinkles. Tergites I-VI (Fig. 
<figureCitation id="B12AA445CBE24B166B687204A70194DC" captionStart="Figure 6" captionStartId="F10364812" captionText="Figure 6. Habitus of the pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. dorsal view; B. ventral view; C. lateral view. Scale bars: 1 mm." figureDoi="10.3897/BDJ.11.e107036.figure6" httpUri="https://binary.pensoft.net/fig/888242" pageId="0" pageNumber="107036">6</figureCitation>
C) of similar form, nearly rectangular in dorsal view, length of tergites I-VI distinctly shorter than VII. Tergite VII almost linear on anterior margin, markedly convex in circular arc on posterior margin. Tergites I-VII with well developed rake-like lateral processes. Lateral processes with darker sclerotised edges and each lateral process with several conical projections bearing a short erect apical seta (Fig. 
<figureCitation id="14D415A9CE82A0B7827CD7EB702384E5" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
D). Number of projections from tergites I to VII as follows: 3, 4, 4-5, 4-5, 4-5, 4-5, 2-4. First two projections of tergites II-VI slightly separate. Opposing single projection on lateral processes of adjacent abdominal segments forming anterior and posterior curved teeth of gin-trap structures. Gin trap between segments III and IV as in Fig. 
<figureCitation id="858F218A1BD24BAE8B2312B780E41BAE" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
D. Tergite VIII very short and narrow, about half as long as VII, with sparse and long apical setose tubercles on posterior margin; lateral processes less developed, with 2 or 3 apical setose spines. Tergite IX nearly trapezoid, deeply depressed at centre, with a pair of slender subtapered divergent urogomphi at the end and with sparse small tubercles bearing long erect apical setae on lateral margins and ventral surface. Urogomphi (Fig. 
<figureCitation id="E3B0A938B70F01A912F35F39516F7FC5" captionStart="Figure 8" captionStartId="F10364816" captionText="Figure 8. Pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. urogomphi, male, dorsal view; B. urogomphi, female, ventral view; C. urogomphi, male, ventral view. Scale bars: 0.5 mm. The arrow (B) indicates a pair of tapered styluses of sternite VIII in female. The arrow (C) indicates no tapered styluses in male." figureDoi="10.3897/BDJ.11.e107036.figure8" httpUri="https://binary.pensoft.net/fig/888244" pageId="0" pageNumber="107036">8</figureCitation>
A) with dense annular wrinkles, gradually slightly narrowing towards apex, directed posteriorly and reflexed finely. Abdominal sternites I-VIII relatively smooth, with fine longitudinal wrinkles and with sparse tubercles bearing long erect apical setae near posterior margin. Sternite VII distinctly and deeply depressed at centre. Abdominal spiracles (Fig. 
<figureCitation id="C295C8EC54CF30C5DE2B221D5901219C" captionStart="Figure 7" captionStartId="F10364814" captionText="Figure 7. Pupa and exuvia of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. head, anterior view; B. pronotum, male, dorsal view; C. pronotum, female, dorsal view; D. lateral process of abdominal tergites II-V, dorsal view; E. abdominal tergites I-VII, lateral view; F. exuvia of last instar larvae, lateral view. Scale bars: 1 mm. The arrow (D) indicates the gin trap between segments III and IV. The arrow (E) indicates one of the abdominal spiracles." figureDoi="10.3897/BDJ.11.e107036.figure7" httpUri="https://binary.pensoft.net/fig/888243" pageId="0" pageNumber="107036">7</figureCitation>
E) approximately round, slightly convex, visible on anterolateral margins of abdominal sternites I-VII. Spiracles on abdominal segment I slightly enlarged and hidden beneath wing sheaths.
</paragraph>
<paragraph id="F59ABCA988D3D1156E2CBFD51A31A084" pageId="0" pageNumber="107036">
<emphasis id="EACAF4C609B7818AFB3F686E0758B05E" bold="true" pageId="0" pageNumber="107036">Remarks</emphasis>
</paragraph>
<paragraph id="D92C4D2E4E53B01C39FEB860A690AB19" pageId="0" pageNumber="107036">
The 
<taxonomicName id="C878D6D0D56DFADE274EDBEA0EA9CFB8" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="F796F886A9E5F53B281751D3120A250B" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
pupae are comparable to adults of the same species in body form, length and most characteristics.
</paragraph>
<paragraph id="E1946B314732C789C6EA96EDAF4245F8" pageId="0" pageNumber="107036">
The pupae of 
<taxonomicName id="184F26D3591451518BDB3AC40B91C7E9" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
can be distinguished from those of 
<taxonomicName id="CD3D024FFDEB66A3B332371BCCB19576" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
by the following characters: body small, length 6.0-7.0 mm (larger, length 10.5-11.0 mm, in 
<taxonomicName id="418289B6B5F825E5EBD1ED324A26B4E8" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
); pronotum without anterior impression in female (with a shallow one in 
<taxonomicName id="B22E8A80CA1A4633214DF4368B49C3D6" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
); first two projections of lateral processes of tergites II-VI slightly separate (distinctly separate in 
<taxonomicName id="6A8888670CE54D62140871C65405D1AD" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
); sternite VII distinctly and deeply depressed at the centre (hardly depressed in 
<taxonomicName id="AD5B8DEB6DF129534D18309853D6FC25" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metogana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metogana" subGenus="U.">U. (U.) metogana</taxonomicName>
).
</paragraph>
<paragraph id="0B3F65CE423FDA464F34B01859BE2CAE" pageId="0" pageNumber="107036">
In addition, we observed some differences between male and female pupae of 
<taxonomicName id="586E648E7E83F122DDE062583F4A4DB1" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
. Sternite VIII with a pair of tapered styluses pointing to the rear in female (Fig. 
<figureCitation id="24040A544BE4E9A214CEC5A94B75CB38" captionStart="Figure 8" captionStartId="F10364816" captionText="Figure 8. Pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. urogomphi, male, dorsal view; B. urogomphi, female, ventral view; C. urogomphi, male, ventral view. Scale bars: 0.5 mm. The arrow (B) indicates a pair of tapered styluses of sternite VIII in female. The arrow (C) indicates no tapered styluses in male." figureDoi="10.3897/BDJ.11.e107036.figure8" httpUri="https://binary.pensoft.net/fig/888244" pageId="0" pageNumber="107036">8</figureCitation>
B), without in male (Fig. 
<figureCitation id="C24384164155DCAE78D294162A8A656B" captionStart="Figure 8" captionStartId="F10364816" captionText="Figure 8. Pupa of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007. A. urogomphi, male, dorsal view; B. urogomphi, female, ventral view; C. urogomphi, male, ventral view. Scale bars: 0.5 mm. The arrow (B) indicates a pair of tapered styluses of sternite VIII in female. The arrow (C) indicates no tapered styluses in male." figureDoi="10.3897/BDJ.11.e107036.figure8" httpUri="https://binary.pensoft.net/fig/888244" pageId="0" pageNumber="107036">8</figureCitation>
C). Pronotum with a transverse deep anterior impression in male, without in female. Antennomeres V and VII slightly protruding at the inner border in male, not protruding in female.
</paragraph>
<paragraph id="7274B16480C1934A653D9EA54090EF7D" pageId="0" pageNumber="107036">
Like in other known species of 
<taxonomicName id="0698629EA904A01CFC4E44BC7CAF25C7" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="E050B12838A28CF16A07A33CAC709806" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
, 
<taxonomicName id="AFAE9AC2258F70F1F81C33A4439E722F" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
has the apices of urogomphi spined. The truncated urogomphi apices observed in the examined pupae of 
<taxonomicName id="7AA26CBD896566E2674638890A2A312F" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metagana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metagana" subGenus="U.">U. (U.) metagana</taxonomicName>
might be the result of damage. The apices of urogomphi are very fragile and can break off if shaken slightly. Thus, we suppose that the truncated urogomphi apices in 
<taxonomicName id="60F8843409D98F944C6641C0F6E3C467" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) metagana" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="metagana" subGenus="U.">U. (U.) metagana</taxonomicName>
is an artifact and that urogomphi are spined also in this species.
</paragraph>
</subSubSection>
<subSubSection id="AE94D79DF7727E1CD0CF037774E5EA37" pageId="0" pageNumber="107036" type="distribution">
<paragraph id="0981CDD90D9AE7FC8ACA0F2AFB792FDA" pageId="0" pageNumber="107036">Distribution</paragraph>
<paragraph id="1DDDD521BE9BE2D49B444C02997B80A0" pageId="0" pageNumber="107036">
China: Fujian (
<bibRefCitation id="D689233A88733893C21E78A9C1E96921" author="Liu, S. S." journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="107036" pagination="70 - 75" refId="B9839293" refString="Liu, S. S., Ren, G. D., Wang, J. S., 2007. Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini). Acta Zootaxonomica Sinica 32 (1): 70 - 75" title="Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini)" volume="32" year="2007">Liu et al. 2007</bibRefCitation>
), Guangxi (
<bibRefCitation id="47A3C94C67C30F3FDE2AC603917517A1" author="Liu, S. S." journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="107036" pagination="530 - 538" refId="B9839275" refString="Liu, S. S., Ren, G. D., 2007. Taxonomic study of the genus Uloma Dejean from Guangxi in China (Coleoptera, Tenebrionidae, Ulomini). Acta Zootaxonomica Sinica 32 (3): 530 - 538" title="Taxonomic study of the genus Uloma Dejean from Guangxi in China (Coleoptera, Tenebrionidae, Ulomini)" volume="32" year="2007">Liu and Ren 2007</bibRefCitation>
), Hainan (new record), Guangdong (new record), Sichuan (new record), Chongqing (new record), Guizhou (new record).
</paragraph>
</subSubSection>
<subSubSection id="14E715884B2C480CC61C065E8F0EF9D5" pageId="0" pageNumber="107036" type="notes">
<paragraph id="35BFB95BD57154F078A2C1FDE0BB98D6" pageId="0" pageNumber="107036">Notes</paragraph>
</subSubSection>
<subSubSection id="F4DAE021D1B7A07F3A3409BCB2D54159" pageId="0" pageNumber="107036" type="remarks">
<paragraph id="4FA542AF1C842B84EC1BB33FFAB8097B" pageId="0" pageNumber="107036">Remarks</paragraph>
<paragraph id="72145D9CBDC702BCEB42E5BC660FE300" pageId="0" pageNumber="107036">
<taxonomicName id="A1D2E0A61F9646E4CA5AC12D82D0FBFF" authorityName="Liu, Ren &amp; Wang" authorityYear="2007" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma (Uloma) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="species" species="intricornicula" subGenus="Uloma">Uloma (Uloma) intricornicula</taxonomicName>
was described by 
<bibRefCitation id="4533DCE85F3AA0EC2D1B9C7F08C13821" author="Liu, S. S." journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="107036" pagination="70 - 75" refId="B9839293" refString="Liu, S. S., Ren, G. D., Wang, J. S., 2007. Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini). Acta Zootaxonomica Sinica 32 (1): 70 - 75" title="Three new species of the genus Uloma Dejean, 1821 from Wuyi Mountain in China, with a new record (Coleoptera, Tenebrionidae, Ulomini)" volume="32" year="2007">Liu et al. (2007)</bibRefCitation>
from Fujian Province of China. Later this species was mentioned to occur in Guangxi Zhuang Autonomous Region (
<bibRefCitation id="CBF88DF19242FAAE3A9D7B65A6BDD562" author="Liu, S. S." journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="107036" pagination="530 - 538" refId="B9839275" refString="Liu, S. S., Ren, G. D., 2007. Taxonomic study of the genus Uloma Dejean from Guangxi in China (Coleoptera, Tenebrionidae, Ulomini). Acta Zootaxonomica Sinica 32 (3): 530 - 538" title="Taxonomic study of the genus Uloma Dejean from Guangxi in China (Coleoptera, Tenebrionidae, Ulomini)" volume="32" year="2007">Liu and Ren 2007</bibRefCitation>
). While examining the 
<taxonomicName id="EA67D8A608AE5C843AF4B533189BB053" class="Insecta" family="Tenebrionidae" genus="Uloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uloma" order="Coleoptera" pageId="0" pageNumber="107036" phylum="Arthropoda" rank="genus">
<emphasis id="FC4EB86AB48805BFCB3EDC224BBF0339" italics="true" pageId="0" pageNumber="107036">Uloma</emphasis>
</taxonomicName>
specimens in the collections of the Museum of Hebei University, we found that 
<taxonomicName id="7D0B94C8D7CFAB588F5E5772EC9B25BB" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="U. (U.) intricornicula" order="Coleoptera" pageId="0" pageNumber="107036" rank="species" species="intricornicula" subGenus="U.">U. (U.) intricornicula</taxonomicName>
was also collected from Chongqing Municipality and from Hainan, Guangdong, Sichuan and Guizhou Provinces of China (Fig. 
<figureCitation id="859D42581C835A52CD5A582675C2F30F" captionStart="Figure 9" captionStartId="F10364818" captionText="Figure 9. Distribution map of Uloma (Uloma) intricornicula Liu, Ren &amp; Wang, 2007 (red circle). The original map from National Platform for Common Geospatial Information Services." figureDoi="10.3897/BDJ.11.e107036.figure9" httpUri="https://binary.pensoft.net/fig/888249" pageId="0" pageNumber="107036">9</figureCitation>
). Thus, the species seem to be widely distributed in south-western China, where warm-temperate broad-leaved evergreen forests and tropical rain forests occur (
<bibRefCitation id="016A1556B8FD59CC6D5F4F13157C274E" author="Ni, Jian" journalOrPublisher="Folia Geobotanica" pageId="0" pageNumber="107036" pagination="113 - 129" refId="B10364741" refString="Ni, Jian, 2001. A biome classification of China based on plant functional types and the BIOME3 model. Folia Geobotanica 36 (2): 113 - 129" title="A biome classification of China based on plant functional types and the BIOME 3 model" volume="36" year="2001">Ni 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>