727 lines
88 KiB
XML
727 lines
88 KiB
XML
<document id="EFA347BEF9D7B720DD7E34DBF0617E76" ID-CLB-Dataset="293753" ID-DOI="10.15298/rusentj.28.1.02" ID-GBIF-Dataset="54d157eb-451a-4cbf-993b-96d305f9f239" ID-ISSN="0132-8069" ID-Zenodo-Dep="10919914" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712198186010" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2019" docId="CD4787B3FFC54162FEB2FD6CFCB0FD58" docLanguage="en" docName="RussEntomolJ.28.1.5-16.pdf" docOrigin="Russian Entomological Journal 28 (1)" docSource="http://dx.doi.org/10.15298/rusentj.28.1.02" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="Eustra Schmidt-Gobel 1846" docType="treatment" docVersion="5" lastPageNumber="9" masterDocId="317EFFCBFFC44166FF93FFA7FFB3FFF4" masterDocTitle="New or little-known species of the genus Eustra Schmidt-Göbel, 1846 (Coleoptera: Carabidae: Ozaenini)" masterLastPageNumber="16" masterPageNumber="5" pageNumber="6" updateTime="1713211868065" updateUser="carolina">
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<mods:title id="5EBF5058CEFD006F6AA615A3326FFB82">New or little-known species of the genus Eustra Schmidt-Göbel, 1846 (Coleoptera: Carabidae: Ozaenini)</mods:title>
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<mods:name id="F2DF920DD18CAF595E9274A20FD6E30A" type="personal">
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<mods:namePart id="36BAE8D90BE33F1A01DCA00EE49D8A6F">Fedorenko, D. N.</mods:namePart>
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<mods:title id="1AC8B3F63865AEEC2A7E8DBF3BBE4025">Russian Entomological Journal</mods:title>
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<mods:date id="3CAE5375B0FC1C77BEFFFA0AD05B6289">2019</mods:date>
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<treatment id="CD4787B3FFC54162FEB2FD6CFCB0FD58" ID-DOI="http://doi.org/10.5281/zenodo.10970642" ID-GBIF-Taxon="225684252" ID-Zenodo-Dep="10970642" LSID="urn:lsid:plazi:treatment:CD4787B3FFC54162FEB2FD6CFCB0FD58" httpUri="http://treatment.plazi.org/id/CD4787B3FFC54162FEB2FD6CFCB0FD58" lastPageId="4" lastPageNumber="9" pageId="1" pageNumber="6">
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<subSubSection id="0DF4652EFFC54167FEB2FD6CFD11FD17" box="[289,674,715,739]" pageId="1" pageNumber="6" type="nomenclature">
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<paragraph id="455136A5FFC54167FEB2FD6CFD11FD17" blockId="1.[189,776,715,2065]" box="[289,674,715,739]" pageId="1" pageNumber="6">
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<heading id="1E1981C9FFC54167FEB2FD6CFD11FD17" box="[289,674,715,739]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="6" reason="2">
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Genus
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<taxonomicName id="82EE4D26FFC54167FEFEFD6BFD11FD17" ID-CoL="8KVTD" authority="Schmidt-Gobel, 1846" authorityName="Schmidt-Gobel" authorityYear="1846" box="[365,674,715,739]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="779AEAB7FFC54167FEFEFD6BFE07FD17" box="[365,436,716,739]" italics="true" pageId="1" pageNumber="6">Eustra</emphasis>
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<bibRefCitation id="217F4B54FFC54167FE28FD6CFD11FD17" author="Schmidt-Gobel H. M." box="[443,674,715,739]" pageId="1" pageNumber="6" pagination="1 - 94" refId="ref11582" refString="Schmidt-Gobel H. M. 1846. Faunula Coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. P. I - VIII, 1 - 94, Pl. I - III." type="journal article" year="1846">Schmidt-Göbel, 1846</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4652EFFC54167FF71FD57FDF6FCBF" pageId="1" pageNumber="6" type="reference_group">
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<paragraph id="455136A5FFC54167FF71FD57FDF6FCBF" blockId="1.[189,776,715,2065]" pageId="1" pageNumber="6">
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<bibRefCitation id="217F4B54FFC54167FF71FD57FE7DFCF7" author="Schmidt-Gobel H. M." box="[226,462,752,771]" pageId="1" pageNumber="6" pagination="1 - 94" refId="ref11582" refString="Schmidt-Gobel H. M. 1846. Faunula Coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. P. I - VIII, 1 - 94, Pl. I - III." type="journal article" year="1846">Schmidt-Göbel, 1846: 65</bibRefCitation>
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;
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<bibRefCitation id="217F4B54FFC54167FE4EFD57FD2CFCF7" author="Chaudoir M." box="[477,671,752,771]" pageId="1" pageNumber="6" pagination="279 - 352" refId="ref11002" refString="Chaudoir M. 1854. Memoire sur la Famille des Carabiques. 4 e partie // Bull. Soc. Nat. Mosc. T. 27. P. 279 - 352." type="journal article" year="1854">Chaudoir, 1854: 309</bibRefCitation>
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;
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<bibRefCitation id="217F4B54FFC54167FD3DFD57FCB0FCF7" author="Chaudoir M." box="[686,771,752,771]" pageId="1" pageNumber="6" pagination="43 - 74" refId="ref11036" refString="Chaudoir M. 1868. Revision du Groupe de Ozaenides // Ann. Soc. Ent. Belg. T. 11. P. 43 - 74." type="journal article" year="1868">1868: 71</bibRefCitation>
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;
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<bibRefCitation id="217F4B54FFC54167FF2EFCAFFEC9FCEF" author="Andrewes H. E." box="[189,378,776,795]" pageId="1" pageNumber="6" pagination="290 - 300" refId="ref10755" refString="Andrewes H. E. 1919. Papers on Oriental Carabidae. - III // Ann. Mag. Nat. Hist. Ser. 9. Vol. 4. P. 290 - 300." type="journal article" year="1919">Andrewes, 1919: 295</bibRefCitation>
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, 299; 1929: 163, 167;
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<bibRefCitation id="217F4B54FFC54167FDD0FCAFFCB0FCEF" author="Banninger M." box="[579,771,776,795]" pageId="1" pageNumber="6" pagination="177 - 216" refId="ref10861" refString="Banninger M. 1927. Monographie der Subtribus Scaritina (Col. Carab.) I, II. // D. Entomol. Zeitschr. 1927. H. 3. S. 177 - 216." type="journal article" year="1927">Bänninger, 1927: 189</bibRefCitation>
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, 206;
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<bibRefCitation id="217F4B54FFC54167FF7EFC87FE23FCC7" author="Jeannel R." box="[237,400,800,819]" pageId="1" pageNumber="6" pagination="1 - 372" refId="ref11375" refString="Jeannel R. 1946. Faune de l'Empire Francais VI. Coleopteres Carabiques de la region Malgache (premiere rartie). Paris. P. 1 - 372." type="book chapter" year="1946">Jeannel, 1946: 48</bibRefCitation>
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;
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<bibRefCitation id="217F4B54FFC54167FE0DFC87FDF6FCC7" author="Stork N. E." box="[414,581,800,819]" pageId="1" pageNumber="6" pagination="1113 - 1138" refId="ref11669" refString="Stork N. E. 1985. Dhanya, a South-east Asian genus of ozaenine ground beetles // J. Nat. Hist. Vol. 19. P. 1113 - 1138." type="journal article" year="1985">Stork, 1985: 1121</bibRefCitation>
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, 1123;
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<bibRefCitation id="217F4B54FFC54167FD1EFC87FEB9FCBF" author="Deuve Th." pageId="1" pageNumber="6" pagination="547 - 578" refId="ref11147" refString="Deuve Th. 2001. Le genre Eustra Schmidt-Goebel, 1846, insectes (Coleoptera, Paussidae, Ozaeninae) a genitalia femelles orthotopiques // Zoosystema. V. 23. P. 547 - 578." type="journal article" year="2001">Deuve, 2001: 547–578</bibRefCitation>
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;
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<bibRefCitation id="217F4B54FFC54167FE86FC9FFE4EFCBF" author="Moore W. & Song X. & Di Giulio A." box="[277,509,824,843]" pageId="1" pageNumber="6" pagination="63 - 82" refId="ref11481" refString="Moore W., Song X., Di Giulio A. 2011. The larva of Eustra (Coleoptera, Paussinae, Ozaenini): a facultative associate of ants // Zookeys. Vol. 90. P. 63 - 82." type="book chapter" year="2011">
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Moore
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<emphasis id="779AEAB7FFC54167FEC6FC9EFECEFCBF" box="[341,381,824,843]" italics="true" pageId="1" pageNumber="6">et al</emphasis>
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., 2011: 63–82
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</bibRefCitation>
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(larva).
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4652EFFC54167FF71FCF7FF42FC8F" pageId="1" pageNumber="6" type="type_taxon">
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<paragraph id="455136A5FFC54167FF71FCF7FF42FC8F" blockId="1.[189,776,715,2065]" pageId="1" pageNumber="6">
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<typeStatus id="9A558807FFC54167FF71FCF7FEBDFC97" box="[226,270,848,867]" pageId="1" pageNumber="6">Type</typeStatus>
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species:
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<taxonomicName id="82EE4D26FFC54167FECCFCF6FD15FC97" authority="Schmidt-Gobel, 1846" authorityName="Schmidt-Gobel" authorityYear="1846" box="[351,678,848,867]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="6" phylum="Arthropoda" rank="species" species="plagiata">
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<emphasis id="779AEAB7FFC54167FECCFCF6FE55FC97" box="[351,486,848,867]" italics="true" pageId="1" pageNumber="6">Eustra plagiata</emphasis>
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<bibRefCitation id="217F4B54FFC54167FE7FFCF7FD15FC97" author="Schmidt-Gobel H. M." box="[492,678,848,867]" pageId="1" pageNumber="6" pagination="1 - 94" refId="ref11582" refString="Schmidt-Gobel H. M. 1846. Faunula Coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. P. I - VIII, 1 - 94, Pl. I - III." type="journal article" year="1846">Schmidt-Göbel, 1846</bibRefCitation>
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</taxonomicName>
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(by monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4652EFFC54167FF71FC27FBA4F7E3" pageId="1" pageNumber="6" type="description">
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<paragraph id="455136A5FFC54167FF71FC27FD6CFB72" blockId="1.[189,776,715,2065]" pageId="1" pageNumber="6">
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REDESCRIPTION.Body (
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<figureCitation id="DDD52A20FFC54167FE77FC27FD85FC62" box="[484,566,896,918]" captionStart="Figs 1–6" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[200,1389,657,1951]" captionTargetId="figure-306@2.[188,1398,652,1954]" captionTargetPageId="2" captionText="Figs 1–6. Eustra spp., dorsal habitus: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii. Рис. 1–6. Eustra spp., габитус дорЗально: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii." figureDoi="http://doi.org/10.5281/zenodo.10919916" httpUri="https://zenodo.org/record/10919916/files/figure.png" pageId="1" pageNumber="6">Figs 1–6</figureCitation>
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) macropterous, small,
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<collectionCode id="23FFAE60FFC54167FF2EFC3CFF6FFC44" box="[189,220,923,944]" pageId="1" pageNumber="6">BL</collectionCode>
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ca
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<quantity id="82169B40FFC54167FF6EFC3DFEEDFC44" box="[253,350,922,944]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="6" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
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in general, shiny or slightly dull, and rather pale, yellow or pale brown; elytra bimaculate,
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<emphasis id="779AEAB7FFC54167FDECFC11FD2EFC3F" box="[639,669,950,971]" italics="true" pageId="1" pageNumber="6">i.e.</emphasis>
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, each with a darker, subquadrate, submedian macula. Dorsal microsculpture meshed, isodiametric, very superficial to obliterate; elytra laterally with aciculate microsculpture consisting of dense microscopic tubercles. Body appendages and underside rather densely pubescent; dorsal pubescence sparser, its density depends on head, pronotum or elytra, leaving elytral submarginal area glabrous; propleura generally smooth.
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</paragraph>
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<paragraph id="455136A5FFC54167FF71FB2CFE5AF8D5" blockId="1.[189,776,715,2065]" pageId="1" pageNumber="6">Head medium-sized, more or less constricted behind eyes, without fixed supra-ocular setae. Eyes round in lateral view, small and convex; genae distinct, oblique, neither toothed nor embracing eyes from behind. A well-developed ocular ridge running around and at a distance from eye; the ridge conspicuously edged anteroventrally, separating gena from maxillary fissure; dorsally it consisting of supra-antennal carina anteriorly and supra-ocular carina behind, the former running just inside the latter, with a strigose area mostly substituted for both at anterior margin of eye. Frontal foveae missing. Clypeus bisetose on each side, slightly transverse, subquadrate, gently sinuate apically; frontoclypeal suture mostly subsinuate. Labrum long trapezoidal, almost triangular, with sides straight to subsinuate and converging much apicad; apex rounded to truncate, tuberculate or crenulate, with eight close setae along apical margin and one preapical seta on each side of inner pair of setae; without or with a subtle emargination on each side of two median setae. Antennae rather short, moniliform, entirely pubescent; pilosity on antennomeres 1–4 similar to that on the rest. Scape globular, short, with or without dorsal seta, pedicel eccentrically articulated. Antennomeres 5–10 subglobose and increasingly, slightly to moderately, transverse, antennomere 11 parallel-sided, twice as long as wide and as long as antennomeres 8–10 combined.</paragraph>
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<paragraph id="455136A5FFC54167FF71F882FBCAFE9F" blockId="1.[189,776,715,2065]" lastBlockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">
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Mandibles moderately long, scrobe with 0–2 setae along ventral ridge; terebral tooth small, retinacular ridge large, bifid, basal retinacular tooth triangular, blunt, without narrow basal extension.
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<collectingRegion id="872AF847FFC54167FE1CF8D1FE52F87F" box="[399,481,1910,1931]" country="Myanmar" name="Chin" pageId="1" pageNumber="6">Mentum</collectingRegion>
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anteriorly bisetose, with vestigial median tooth; ligula subquadrate, quadrisetose, paraglossae adnate vestigial. Submentum quadrisetose, gula well-developed, moderately wide basally, almost indistinct at submentum. Maxillary cardo short, stipes bisetose. Terminal labial and maxillary palpomere fusiform, maxillary one twice as wide as penultimate palpomere. Penultimate labial palpomere with two very close setae.
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</paragraph>
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<paragraph id="455136A5FFC54167FCC1FED6FB79FD5F" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">Pronotum cordate, more or less transverse, broadest 1/3– 1/5 from apex. Base slightly narrower than apex, truncate, strongly and abruptly oblique on sides; basal angles sharp, obtuse to right. Apex deeply sinuate or subtruncate medially, apical angles acute and porrect. Sides rounded, subsinuate just behind apical angles and straight to sinuate before basal ones. Disc convex, lateral margin rather widely explanate and finely beaded, more or less reflexed basally; lateral edge setulose and tuberculate or crenulate to smooth. Basal foveae round and deep, occupying basal third, median line fine yet deep, basal transverse impression quite transverse and fairly deep, apical transverse impression less so.</paragraph>
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<paragraph id="455136A5FFC54167FCC1FD16FBFDFADF" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">
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Elytra estriate, wide, convex, broadest behind middle, with sides more or less rounded and diverging slightly apicad, more rounded behind humeri, obliquely truncate posteriorly; apices rounded separately each; lateral half of each elytron flattened and longitudinally impressed in third fourth to middle third. Base distinctly wider than that of pronotum, basal margin slightly oblique from humeri to mesothoracic peduncle; humeri prominent, with blunt or rounded apices. Basal ridge fine at humerus, vestigial inside; lateral reflexed margin narrow, interrupted before preapical plica; lateral edge setulose and almost indistinctly serrulate, sometimes (
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<taxonomicName id="82EE4D26FFC54167FAF2FC1BFC67FC1F" authorityName="Fedorenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
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<emphasis id="779AEAB7FFC54167FAF2FC1BFC67FC1F" italics="true" pageId="1" pageNumber="6">E. nageli subglabra</emphasis>
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</taxonomicName>
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<emphasis id="779AEAB7FFC54167FC4EFC71FBA4FC1F" bold="true" box="[989,1047,982,1003]" pageId="1" pageNumber="6">ssp.n.</emphasis>
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) smooth and nearly glabrous. Epipleura totally reduced behind preapical plica. Dorsum densely tuberculate and pilose, except for a smooth and glabrous submarginal area in middle third; pilosity dense to sparse, moderately long to very short. Fixed dorsal setae, basal or discal, missing; submarginal umbilical seta series (USS) continuous, consisting of larger setigerous pores, about ten (9–13) in basal 2/3 and five on a level with and behind preapical plica, the anterior and posterior pore groups being connected by an oblique row of 3–7, mostly 4–5, smaller setigerous pores. — Anterior setigerous pores increased to
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<quantity id="82169B40FFC54167FCBFFB5CFC36FAE5" box="[812,901,1275,1297]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="1" pageNumber="6" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
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number are due to the presence of a few minute pores additional to larger ten.
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</paragraph>
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<paragraph id="455136A5FFC54167FCC1FA96FCE8F955" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">
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Underside: mesocoxae separate and slightly projecting, with ventral intercoxal bridge, chiefly posterior process of mesoventrite, totally reduced, leaving mesocoxal cavities shallow, more so mesally, and apodemal pits of mesofurca (mesendosternite) exposed; anterior metaventral process either somewhat pointed, tuberculate and projecting (
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<taxonomicName id="82EE4D26FFC54167FAB3FA10FCEEFA12" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="6" phylum="Arthropoda" rank="species" species="cariniceps" status="sp.n.">
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<emphasis id="779AEAB7FFC54167FAB3FA10FCEEFA12" italics="true" pageId="1" pageNumber="6">E. cariniceps</emphasis>
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</taxonomicName>
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<emphasis id="779AEAB7FFC54167FCF5FA76FC25FA12" bold="true" box="[870,918,1489,1510]" pageId="1" pageNumber="6">
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||
<taxonomicNameLabel id="6CA957CCFFC54167FCF5FA76FC25FA12" box="[870,918,1489,1510]" pageId="1" pageNumber="6" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or apically blunt to truncate, not or barely projecting (the other congeners reviewed). Metacoxae separate. Abdominal sternites transversely sulcate at bases (which character is hardly traceable because of the sternites telescoped), with obligatory setae hardly separable from dense ventral pubescence if at all (four or more longer setae present closer to apical margin), sternite VII mostly bisetose on each side.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC54167FCC1F901FBF8F8B5" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">Legs. Chetome obscured by dense, longer and shorter, pilosity. Meso- and metatarsomere 1 long, as long as respective tarsomeres 2–4 combined, usually slightly to conspicuously longer than tarsomere 5. Protarsomeres 1–3 ventroapically rounded and setose; protarsomeres 1–2 barely dilated, with ventral pad, in male.</paragraph>
|
||
<paragraph id="455136A5FFC54167FCC1F8E1FAC4F858" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">
|
||
Aedeagus as in
|
||
<figureCitation id="DDD52A20FFC54167FC7AF8E1FBFFF8A8" box="[1001,1100,1862,1884]" captionStart-0="Figs 7–18" captionStart-1="Figs 19–25" captionStart-2="Figs 26–32" captionStart-3="Figs 33–46" captionStartId-0="5.[226,262,1969,1988]" captionStartId-1="6.[226,263,1966,1985]" captionStartId-2="8.[226,263,1917,1936]" captionStartId-3="9.[226,263,1918,1937]" captionTargetBox-0="[204,1393,463,1941]" captionTargetBox-1="[202,1396,1030,1921]" captionTargetBox-2="[192,1395,999,1881]" captionTargetBox-3="[197,1392,1006,1883]" captionTargetId-0="figure-121@5.[188,1398,450,1949]" captionTargetId-1="figure-10@6.[189,1399,1012,1947]" captionTargetId-2="figure-10@8.[189,1399,998,1876]" captionTargetId-3="figure-565@9.[188,1398,996,1901]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Figs 7–18. Median lobe of aedeagus: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — right aspect; 8, 17 — ventral aspect; 9, 12, 15, 18 — left aspect; 11, 14 — dorsal aspect;. Scale bars: 0.5 mm. Рис. 7–18. СреднЯЯ долЯ Эдеагуса: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — справа; 8, 17 — сниЗу; 9, 12, 15, 18 — слева; 11, 14 — сверху. Масштаб: 0.5 мм." captionText-1="Figs 19–25. Median lobe of aedeagus: 19–21 — Eustra nageli subglabra ssp.n.; 22–23 — E. cariniceps sp.n.; 24–25 — E. dalatensis sp.n.; 19, 22, 24 — right aspect; 20, 23 — ventral aspect; 21 — left aspect; 25 — dorsal aspect. Scale bars: 0.5 mm. Рис. 19–25. СреднЯЯ долЯ Эдеагуса: 19–21 — Eustra nageli subglabra ssp.n.; 22–23 — E. cariniceps sp.n.; 24–25 — E. dalatensis sp.n.; 19, 22, 24 — справа; 20, 23 — сниЗу; 21 — слева; 25 — сверху. Масштаб: 0.5 мм." captionText-2="Figs 26–32. Median lobe of aedeagus in plane of apex (~left ventrobasal aspect): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Scale bar 0.5 mm. Рис. 26–32. СреднЯЯ долЯ Эдеагуса в плоскости вершины (~вентробаЗально слева): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Масштаб 0.5 мм." captionText-3="Figs 33–46. Parameres: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — right paramere, left aspect; 34, 36, 38, 40, 42, 44, 46 — left paramere, right aspect. Scale bar 0.5 mm. Рис. 33–46. Парамеры: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — праваЯ парамера, вид слева; 34, 36, 38, 40, 42, 44, 46 — леваЯ парамера, вид справа. Масштаб 0.5 мм." figureDoi-0="http://doi.org/10.5281/zenodo.10919920" figureDoi-1="http://doi.org/10.5281/zenodo.10919922" figureDoi-2="http://doi.org/10.5281/zenodo.10919924" figureDoi-3="http://doi.org/10.5281/zenodo.10919928" httpUri-0="https://zenodo.org/record/10919920/files/figure.png" httpUri-1="https://zenodo.org/record/10919922/files/figure.png" httpUri-2="https://zenodo.org/record/10919924/files/figure.png" httpUri-3="https://zenodo.org/record/10919928/files/figure.png" pageId="1" pageNumber="6">Figs 7–46</figureCitation>
|
||
; right paramere narrow, longer to shorter than left one; right paramere more or less densely setose, left paramere glabrous. Internal sac mostly with a cluster of
|
||
<emphasis id="779AEAB7FFC54167FC18F830FC11F858" box="[907,930,1943,1964]" italics="true" pageId="1" pageNumber="6">ca</emphasis>
|
||
5–7, strong, spiniform, radiating apical sclerites.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC54167FCC1F816FBA4F7E3" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="6">
|
||
Female genitalia and reproductive tract. Gonocoxae entire, membranous and densely pubescent, widely separated by a round sclerotized part of
|
||
<emphasis id="779AEAB7FFC54167FBC0F841FB47F80F" box="[1107,1268,2022,2043]" italics="true" pageId="1" pageNumber="6">bursa copulatrix</emphasis>
|
||
; helminthoid sclerite well-developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFC64165FF71FE9CFD5FFD83" lastPageId="3" lastPageNumber="8" pageId="2" pageNumber="7" type="diagnosis">
|
||
<paragraph id="455136A5FFC64165FF71FE9CFD5FFD83" blockId="2.[189,775,315,631]" lastBlockId="3.[189,776,315,2071]" lastPageId="3" lastPageNumber="8" pageId="2" pageNumber="7">
|
||
DIAGNOSIS. The genus is very distinctive within Ozaenini in many points (see also ‘Comments’ to the genus). Very small body size and pale brown ground-colour, combined with fusiform terminal palpomeres, separate
|
||
<taxonomicName id="82EE4D26FFC64164FD55FE2BFCB5FE55" authorityName="Schmidt-Gobel" authorityYear="1846" box="[710,774,396,417]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FD55FE2BFCB5FE55" box="[710,774,396,417]" italics="true" pageId="2" pageNumber="7">Eustra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC64164FF74FE01FE6AFE4F" authority="Andrewes, 1919" authorityName="Andrewes" authorityYear="1919" box="[231,473,422,444]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FF74FE01FE80FE4F" box="[231,307,422,443]" italics="true" pageId="2" pageNumber="7">Dhanya</emphasis>
|
||
<bibRefCitation id="217F4B54FFC64164FEA9FE01FE6AFE4F" author="Andrewes H. E." box="[314,473,422,444]" pageId="2" pageNumber="7" pagination="290 - 300" refId="ref10755" refString="Andrewes H. E. 1919. Papers on Oriental Carabidae. - III // Ann. Mag. Nat. Hist. Ser. 9. Vol. 4. P. 290 - 300." type="journal article" year="1919">Andrewes, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
from the other ozaenines. The two genera are otherwise very distinctive. Specifically,
|
||
<taxonomicName id="82EE4D26FFC64164FD49FE66FF6AFE05" authorityName="Schmidt-Gobel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FD49FE66FF6AFE05" italics="true" pageId="2" pageNumber="7">Eustra</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguishable from
|
||
<taxonomicName id="82EE4D26FFC64164FD87FE7CFDD3FE04" authorityName="Andrewes" authorityYear="1919" box="[532,608,475,496]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FD87FE7CFDD3FE04" box="[532,608,475,496]" italics="true" pageId="2" pageNumber="7">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
in having mesocoxae conspicuously separated (
|
||
<emphasis id="779AEAB7FFC64164FE67FE50FDBCFDF8" box="[500,527,503,524]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
contiguous) and propleural pits missing (well-developed propleural pits are autapomorphy of
|
||
<taxonomicName id="82EE4D26FFC64164FEB6FD8CFEC2FDB4" authorityName="Andrewes" authorityYear="1919" box="[293,369,555,576]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FEB6FD8CFEC2FDB4" box="[293,369,555,576]" italics="true" pageId="2" pageNumber="7">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
within Ozaenini). The other differences of
|
||
<taxonomicName id="82EE4D26FFC64164FF2EFDE0FF4EFDA8" authorityName="Schmidt-Gobel" authorityYear="1846" box="[189,253,583,604]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FF2EFDE0FF4EFDA8" box="[189,253,583,604]" italics="true" pageId="2" pageNumber="7">Eustra</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="82EE4D26FFC64164FEA9FDE1FE35FDAF" authorityName="Andrewes" authorityYear="1919" box="[314,390,582,603]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC64164FEA9FDE1FE35FDAF" box="[314,390,582,603]" italics="true" pageId="2" pageNumber="7">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
are many: ocular ridge well-developed throughout or nearly so (
|
||
<emphasis id="779AEAB7FFC64164FE24FDC6FE61FD82" box="[439,466,609,630]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
indistinct posteriorly), clypeus transversely subquadrate and bisetose on each side (
|
||
<emphasis id="779AEAB7FFC64164FAB9FE9BFAF6FEA5" box="[1322,1349,316,337]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
long hexagonal and unisetose on each side), antennal scape globulose (
|
||
<emphasis id="779AEAB7FFC64164FCFDFED6FC3AFE72" box="[878,905,369,390]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
short subcylindric), antennomere 11 much longer, labrum trapezoidal (
|
||
<emphasis id="779AEAB7FFC64164FC6FFE2BFBA4FE55" box="[1020,1047,396,417]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
subquadrate), mandibles medium-sized and incurved (
|
||
<emphasis id="779AEAB7FFC64164FC7FFE00FBB4FE48" box="[1004,1031,423,444]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
large and almost straight), mandibular scrobe with several setae in general (
|
||
<emphasis id="779AEAB7FFC64164FB04FE66FB01FE22" box="[1175,1202,449,470]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
asetose), gula well-developed (
|
||
<emphasis id="779AEAB7FFC64164FC37FE7BFC0CFE05" box="[932,959,476,497]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
totally reduced), submentum bisetose (
|
||
<emphasis id="779AEAB7FFC64164FACFFE7BFAC4FE05" box="[1372,1399,476,497]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
unisetose), mentum tooth indistinct (
|
||
<emphasis id="779AEAB7FFC64164FB38FE50FB75FDF8" box="[1195,1222,503,524]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
well-developed), maxillary stipes bisetose (
|
||
<emphasis id="779AEAB7FFC64164FBB6FDB6FBF3FDD2" box="[1061,1088,529,550]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
unisetose), pronotal lateral edge setulose and crenulate or tuberculate (
|
||
<emphasis id="779AEAB7FFC64164FB34FD8BFB71FDB5" box="[1191,1218,556,577]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
smooth, with 3–4 subequally spaced setae inside), elytral epipleura missing apically (
|
||
<emphasis id="779AEAB7FFC64164FC14FDC6FC11FD82" box="[903,930,609,630]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
entire), elytral discal setae missing (
|
||
<emphasis id="779AEAB7FFC64164FA95FDC6FA92FD82" box="[1286,1313,609,630]" italics="true" pageId="2" pageNumber="7">vs.</emphasis>
|
||
multiple in sites of intervals 3 and 5), protarsomeres 1–3 ventroapically rounded (
|
||
<emphasis id="779AEAB7FFC74165FEE1FEF0FE3EFE98" box="[370,397,343,364]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
toothed at each apical angle), protarsomeres 1 and 2 slightly dilated and with ventral pad in male (
|
||
<emphasis id="779AEAB7FFC74165FF56FE2BFF6CFE55" box="[197,223,396,417]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
not dilated and without ventral pad), mesotarsomere 1 longer, as long as mesotarsomeres 2–4 combined (
|
||
<emphasis id="779AEAB7FFC74165FD31FE00FD0EFE48" box="[674,701,423,444]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
as long as those 2 and 3 combined), metatarsomere 1 much longer than (
|
||
<emphasis id="779AEAB7FFC74165FF67FE7BFEBCFE05" box="[244,271,476,497]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
as long as) metatarsomere 5, abdominal sternite VII membranous yet entire apically (
|
||
<emphasis id="779AEAB7FFC74165FD97FE50FDACFDF8" box="[516,543,503,524]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
well sclerotized, being sinuate in female in addition), with 2 apical setae on each side in general (
|
||
<emphasis id="779AEAB7FFC74165FEB9FD8BFEF6FDB5" box="[298,325,556,577]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
3–4 setae on each side), female gonocoxite IX membranous (
|
||
<emphasis id="779AEAB7FFC74165FED8FDE0FED5FDA8" box="[331,358,583,604]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
well sclerotized). Finally, the genus has ventral structure of pterothorax unique within Adephaga.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1191662DFFC64164FF71F816FD06F7E4" ID-DOI="http://doi.org/10.5281/zenodo.10919916" ID-Zenodo-Dep="10919916" httpUri="https://zenodo.org/record/10919916/files/figure.png" pageId="2" pageNumber="7" startId="2.[226,263,1969,1988]" targetBox="[200,1389,657,1951]" targetPageId="2" targetType="figure">
|
||
<paragraph id="455136A5FFC64164FF71F816FD06F828" blockId="2.[189,1398,1969,2064]" pageId="2" pageNumber="7">
|
||
Figs 1–6.
|
||
<taxonomicName id="82EE4D26FFC64164FED1F815FE1AF831" box="[322,425,1970,1989]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC64164FED1F815FEC8F830" box="[322,379,1970,1988]" italics="true" pageId="2" pageNumber="7">Eustra</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, dorsal habitus: 1 —
|
||
<taxonomicName id="82EE4D26FFC64164FDFAF815FD61F830" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[617,722,1969,1989]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="parallela" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FDFAF815FDC5F830" box="[617,630,1970,1988]" italics="true" pageId="2" pageNumber="7">E</emphasis>
|
||
.
|
||
<emphasis id="779AEAB7FFC64164FD10F816FD61F830" box="[643,722,1969,1988]" italics="true" pageId="2" pageNumber="7">parallela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FD4AF816FCB7F830" bold="true" box="[729,772,1969,1988]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FD4AF816FCB7F830" box="[729,772,1969,1988]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 2 —
|
||
<taxonomicName id="82EE4D26FFC64164FCD3F815FC00F830" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[832,947,1970,1988]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="cariniceps" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FCD3F815FC00F830" box="[832,947,1970,1988]" italics="true" pageId="2" pageNumber="7">E. cariniceps</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FC29F816FC56F830" bold="true" box="[954,997,1969,1988]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FC29F816FC56F830" box="[954,997,1969,1988]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 3 —
|
||
<taxonomicName id="82EE4D26FFC64164FBB2F815FB0AF830" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1057,1209,1970,1988]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="posteroguttata" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FBB2F815FB0AF830" box="[1057,1209,1970,1988]" italics="true" pageId="2" pageNumber="7">E. posteroguttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FB53F816FB58F830" bold="true" box="[1216,1259,1969,1988]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FB53F816FB58F830" box="[1216,1259,1969,1988]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 4 —
|
||
<taxonomicName id="82EE4D26FFC64164FAB4F815FEA7F828" authorityName="Fedorenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
|
||
<emphasis id="779AEAB7FFC64164FAB4F815FEA7F828" italics="true" pageId="2" pageNumber="7">E. nageli subglabra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FE88F86EFEFDF828" bold="true" box="[283,334,1993,2012]" pageId="2" pageNumber="7">ssp.n.</emphasis>
|
||
; 5 —
|
||
<taxonomicName id="82EE4D26FFC64164FE1AF86DFE49F828" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[393,506,1993,2012]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="dalatensis" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FE1AF86DFE49F828" box="[393,506,1993,2012]" italics="true" pageId="2" pageNumber="7">E. dalatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FD92F86EFD9FF828" bold="true" box="[513,556,1993,2012]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FD92F86EFD9FF828" box="[513,556,1993,2012]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 6 —
|
||
<emphasis id="779AEAB7FFC64164FDF4F86DFD06F828" box="[615,693,1993,2012]" italics="true" pageId="2" pageNumber="7">
|
||
<taxonomicName id="82EE4D26FFC64164FDF4F86DFD03F828" authorityName="Jedlicka" authorityYear="1968" box="[615,688,1993,2012]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="csikii">E. csikii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC64164FF71F842FD06F7E4" blockId="2.[189,1398,1969,2064]" pageId="2" pageNumber="7">
|
||
Рис. 1–6.
|
||
<taxonomicName id="82EE4D26FFC64164FEAAF841FE2DF80C" box="[313,414,2021,2040]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC64164FEAAF841FEC1F80C" box="[313,370,2022,2040]" italics="true" pageId="2" pageNumber="7">Eustra</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, габитус дорЗально: 1 —
|
||
<taxonomicName id="82EE4D26FFC64164FD15F841FD5EF80C" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[646,749,2021,2040]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="parallela" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FD15F841FD20F80C" box="[646,659,2022,2040]" italics="true" pageId="2" pageNumber="7">E</emphasis>
|
||
.
|
||
<emphasis id="779AEAB7FFC64164FD0DF842FD5EF80C" box="[670,749,2021,2040]" italics="true" pageId="2" pageNumber="7">parallela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FD61F842FCAEF80C" bold="true" box="[754,797,2021,2040]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FD61F842FCAEF80C" box="[754,797,2021,2040]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 2 —
|
||
<taxonomicName id="82EE4D26FFC64164FCC7F841FC76F80C" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[852,965,2022,2040]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="cariniceps" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FCC7F841FC76F80C" box="[852,965,2022,2040]" italics="true" pageId="2" pageNumber="7">E. cariniceps</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FC58F842FC45F80C" bold="true" box="[971,1014,2021,2040]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FC58F842FC45F80C" box="[971,1014,2021,2040]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 3 —
|
||
<taxonomicName id="82EE4D26FFC64164FBBFF841FB71F80C" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1068,1218,2022,2040]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="posteroguttata" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FBBFF841FB71F80C" box="[1068,1218,2022,2040]" italics="true" pageId="2" pageNumber="7">E. posteroguttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FB54F842FB41F80C" bold="true" box="[1223,1266,2021,2040]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FB54F842FB41F80C" box="[1223,1266,2021,2040]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 4 —
|
||
<taxonomicName id="82EE4D26FFC64164FABBF841FEA7F7E4" authorityName="Fedorenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
|
||
<emphasis id="779AEAB7FFC64164FABBF841FEA7F7E4" italics="true" pageId="2" pageNumber="7">E. nageli subglabra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FE88F85AFEFDF7E4" bold="true" box="[283,334,2045,2064]" pageId="2" pageNumber="7">ssp.n.</emphasis>
|
||
; 5 —
|
||
<taxonomicName id="82EE4D26FFC64164FE1AF859FE49F7E4" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[393,506,2045,2064]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="dalatensis" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC64164FE1AF859FE49F7E4" box="[393,506,2045,2064]" italics="true" pageId="2" pageNumber="7">E. dalatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC64164FD92F85AFD9FF7E4" bold="true" box="[513,556,2045,2064]" pageId="2" pageNumber="7">
|
||
<taxonomicNameLabel id="6CA957CCFFC64164FD92F85AFD9FF7E4" box="[513,556,2045,2064]" pageId="2" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 6 —
|
||
<taxonomicName id="82EE4D26FFC64164FDF4F859FD03F7E4" authorityName="Jedlicka" authorityYear="1968" box="[615,688,2045,2064]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="7" phylum="Arthropoda" rank="species" species="csikii">E. csikii</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0DF4652EFFC74165FF71FDDCFD5AFD0F" pageId="3" pageNumber="8" type="distribution">
|
||
<paragraph id="455136A5FFC74165FF71FDDCFD5AFD0F" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="8">
|
||
DISTRIBUTION. The range extends throughout the Oriental region from
|
||
<collectingCountry id="3DF97635FFC74165FEF1FD31FE72FD58" box="[354,449,662,684]" name="Sri Lanka" pageId="3" pageNumber="8">Sri Lanka</collectingCountry>
|
||
, southern
|
||
<collectingCountry id="3DF97635FFC74165FD8DFD31FDFCFD58" box="[542,591,662,684]" name="India" pageId="3" pageNumber="8">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="3DF97635FFC74165FDE9FD31FD06FD58" box="[634,693,662,684]" name="Nepal" pageId="3" pageNumber="8">Nepal</collectingCountry>
|
||
, through Indochina to
|
||
<collectingCountry id="3DF97635FFC74165FED6FD16FE3CFD33" box="[325,399,689,711]" name="Taiwan" pageId="3" pageNumber="8">Taiwan</collectingCountry>
|
||
, southern
|
||
<collectingCountry id="3DF97635FFC74165FE68FD16FD85FD33" box="[507,566,689,711]" name="China" pageId="3" pageNumber="8">China</collectingCountry>
|
||
, southern
|
||
<collectingCountry id="3DF97635FFC74165FD31FD16FD69FD32" box="[674,730,689,710]" name="Japan" pageId="3" pageNumber="8">Japan</collectingCountry>
|
||
, the Sunda Isles and the
|
||
<collectingCountry id="3DF97635FFC74165FEEDFD6CFE58FD15" box="[382,491,715,737]" name="Philippines" pageId="3" pageNumber="8">Philippines</collectingCountry>
|
||
, including also adjacent parts of Palearctic
|
||
<collectingCountry id="3DF97635FFC74165FEA9FD41FE25FD08" box="[314,406,742,764]" name="China" pageId="3" pageNumber="8">East Asia</collectingCountry>
|
||
in eastern
|
||
<collectingCountry id="3DF97635FFC74165FE6DFD41FD8AFD08" box="[510,569,742,764]" name="China" pageId="3" pageNumber="8">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="3DF97635FFC74165FDD0FD41FDCEFD0F" box="[579,637,742,763]" name="South Korea" pageId="3" pageNumber="8">Korea</collectingCountry>
|
||
and
|
||
<collectingCountry id="3DF97635FFC74165FD3FFD41FD56FD0F" box="[684,741,742,763]" name="Japan" pageId="3" pageNumber="8">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFC74165FF71FCA6FD47FB85" pageId="3" pageNumber="8" type="biology_ecology">
|
||
<paragraph id="455136A5FFC74165FF71FCA6FD47FB85" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="8">
|
||
HABITATS AND HABITS.The members of the genus are silvicolous or cavernicolous. Silvicoles were captured at elevations ranging between 60 and
|
||
<quantity id="82169B40FFC74165FE4AFC91FD94FCBF" box="[473,551,822,843]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="8" unit="m" value="1900.0">1,900 m</quantity>
|
||
a.s.l. Many of them are known from one to a few specimens only and rare in collections. On the other hand, most species taken by me in
|
||
<collectingCountry id="3DF97635FFC74165FD26FCCCFCB4FC75" box="[693,775,875,897]" name="Vietnam" pageId="3" pageNumber="8">Vietnam</collectingCountry>
|
||
are not uncommon lowland and piedmont forest-dwellers. Great majority of the specimens were captured by hands under bark of larger dead trees, standing or fallen, or under tree fragments on the ground (
|
||
<taxonomicName id="82EE4D26FFC74165FE7AFC70FD8AFC1F" authorityName="Jedlicka" authorityYear="1968" box="[489,569,982,1004]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="csikii">
|
||
<emphasis id="779AEAB7FFC74165FE7AFC70FD8AFC1F" box="[489,569,982,1004]" italics="true" pageId="3" pageNumber="8">E. csikii</emphasis>
|
||
</taxonomicName>
|
||
) at elevations below
|
||
<quantity id="82169B40FFC74165FF2EFC56FF4AFBF2" box="[189,249,1009,1030]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="8" unit="m" value="800.0">800 m</quantity>
|
||
a.s.l. The species were found to be associated with ants and/or termites, with two or three of them sometimes sharing their habitats with one another and often also with such carabids as
|
||
<taxonomicName id="82EE4D26FFC74165FEA7FBE6FD3CFBA2" authority="Andrewes, 1919" authorityName="Andrewes" authorityYear="1919" box="[308,655,1089,1111]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="bioculata">
|
||
<emphasis id="779AEAB7FFC74165FEA7FBE6FE56FBA2" box="[308,485,1089,1110]" italics="true" pageId="3" pageNumber="8">Dhanya bioculata</emphasis>
|
||
<bibRefCitation id="217F4B54FFC74165FE7DFBE6FD3CFBA2" author="Andrewes H. E." box="[494,655,1089,1111]" pageId="3" pageNumber="8" pagination="290 - 300" refId="ref10755" refString="Andrewes H. E. 1919. Papers on Oriental Carabidae. - III // Ann. Mag. Nat. Hist. Ser. 9. Vol. 4. P. 290 - 300." type="journal article" year="1919">Andrewes, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
(Ozaenini),
|
||
<taxonomicName id="82EE4D26FFC74165FF2EFBFBFEF4FB85" box="[189,327,1116,1137]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FF2EFBFBFEA4FB85" box="[189,279,1116,1137]" italics="true" pageId="3" pageNumber="8">Perigona</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(Perigonini), and
|
||
<taxonomicName id="82EE4D26FFC74165FE68FBFBFDC7FB85" box="[507,628,1116,1137]" class="Insecta" family="Carabidae" genus="Morion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FE68FBFBFDF0FB85" box="[507,579,1116,1137]" italics="true" pageId="3" pageNumber="8">Morion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(Morionini).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFC74165FF71FBD1FBEEFD58" pageId="3" pageNumber="8" type="discussion">
|
||
<paragraph id="455136A5FFC74165FF71FBD1FDD5FB53" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="8">COMMENTS. The above redescription only concerns non-cavernicolous species reviewed below.</paragraph>
|
||
<paragraph id="455136A5FFC74165FF71FB0CFE0DF8B5" blockId="3.[189,776,315,2071]" pageId="3" pageNumber="8">
|
||
Ventral pterothoracic structure of
|
||
<taxonomicName id="82EE4D26FFC74165FDAEFB0BFDCEFB35" authorityName="Schmidt-Gobel" authorityYear="1846" box="[573,637,1196,1217]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FDAEFB0BFDCEFB35" box="[573,637,1196,1217]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
is worthy of special mention. Even though this pattern looks very primitive, it with no doubt has emerged from much more complex structure in course of evolution. This latter groundplan is peculiar to nearly all Adephaga and includes almost internally contiguous mesocoxae, ventrally bridged with posterior process of mesoventrite and anterior process of metaventrite. These two cover mesal parts of the coxae ventrally, with apex of the latter process being superimposed on the former, to make mesocoxal cavities well separated in ventral view.
|
||
<taxonomicName id="82EE4D26FFC74165FF2EFA11FE04FA3F" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[189,439,1462,1484]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FF2EFA11FE96FA3F" box="[189,293,1462,1483]" italics="true" pageId="3" pageNumber="8">Omophron</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
has widely separated mesocoxal cavities and distinctly separated mesocoxae, which is certainly due to its nearly circular body, and the other deviations from the groundplan are only observed within Paussinae, including Ozaenini.Some of them,
|
||
<emphasis id="779AEAB7FFC74165FD94F986FD99F9C2" box="[519,554,1569,1590]" italics="true" pageId="3" pageNumber="8">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="82EE4D26FFC74165FDA0F986FF43F9A4" authority="Laporte, 1834" authorityName="Laporte" authorityYear="1834" class="Insecta" family="Carabidae" genus="Pseudozaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FDA0F986FD01F9C2" box="[563,690,1569,1590]" italics="true" pageId="3" pageNumber="8">Pseudozaena</emphasis>
|
||
Laporte, 1834
|
||
</taxonomicName>
|
||
, match well the groundplan, except only that the mesoventrite posterior to the mesothoracic peduncle is short. Many other Ozaenini demonstrate gradual reduction of the mesocoxal bridge in width and usually also in depth, as the mesocoxae become increasingly projecting and mesocoxal cavities increasingly shallow. Ventral pterothoracic structure of
|
||
<taxonomicName id="82EE4D26FFC74165FF45F97BFEA6F905" authorityName="Schmidt-Gobel" authorityYear="1846" box="[214,277,1756,1777]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FF45F97BFEA6F905" box="[214,277,1756,1777]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
has most probably evolved from separating between contiguous mesocoxae (
|
||
<taxonomicName id="82EE4D26FFC74165FE24F951FD1AF8FF" authority="Perty, 1830" authorityName="Perty" authorityYear="1830" box="[439,681,1782,1803]" class="Insecta" family="Carabidae" genus="Pachyteles" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FE24F951FD91F8FF" box="[439,546,1782,1803]" italics="true" pageId="3" pageNumber="8">Pachyteles</emphasis>
|
||
Perty, 1830
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE4D26FFC74165FD2AF951FEE8F8D3" authority="Andrewes, 1919" authorityName="Andrewes" authorityYear="1919" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FD2AF951FCB5F8FF" box="[697,774,1782,1803]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
<bibRefCitation id="217F4B54FFC74165FF2EF8B6FEE8F8D3" author="Andrewes H. E." box="[189,347,1809,1831]" pageId="3" pageNumber="8" pagination="290 - 300" refId="ref10755" refString="Andrewes H. E. 1919. Papers on Oriental Carabidae. - III // Ann. Mag. Nat. Hist. Ser. 9. Vol. 4. P. 290 - 300." type="journal article" year="1919">Andrewes, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
) following or, less likely, followed by totally reduced intercoxal bridge.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FF7DF8E1FBEEFD58" blockId="3.[189,776,315,2071]" lastBlockId="3.[812,1399,315,1244]" pageId="3" pageNumber="8">
|
||
Neotropical Ozaenini and the remaining Ozaenini + Metriini are sister-groups based on molecular evidence [
|
||
<bibRefCitation id="217F4B54FFC74165FF51F8DBFEFFF865" author="Moore W." box="[194,332,1915,1937]" pageId="3" pageNumber="8" pagination="79 - 92" refId="ref11438" refString="Moore W. 2008. Phylogeny of the Western Hemisphere Ozaenini (Coleoptera: Carabidae: Paussinae) based on DNA sequence data // Ann. Carnegi Mus. Vol. 77. No. 1. P. 79 - 92." type="journal article" year="2008">Moore, 2008</bibRefCitation>
|
||
], but
|
||
<taxonomicName id="82EE4D26FFC74165FE18F8DBFE78F865" authorityName="Schmidt-Gobel" authorityYear="1846" box="[395,459,1916,1937]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FE18F8DBFE78F865" box="[395,459,1916,1937]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC74165FD95F8DCFDE1F864" authorityName="Andrewes" authorityYear="1919" box="[518,594,1915,1936]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FD95F8DCFDE1F864" box="[518,594,1915,1936]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
were out of that analysis. While differences between
|
||
<taxonomicName id="82EE4D26FFC74165FDB7F830FDD7F858" authorityName="Schmidt-Gobel" authorityYear="1846" box="[548,612,1943,1964]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FDB7F830FDD7F858" box="[548,612,1943,1964]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC74165FD06F831FD52F85F" authorityName="Andrewes" authorityYear="1919" box="[661,737,1942,1963]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FD06F831FD52F85F" box="[661,737,1942,1963]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
are many, both share at least five derived character states: metatarsomere 1 as long as to much longer (
|
||
<emphasis id="779AEAB7FFC74165FDA0F86BFDFDF815" box="[563,590,1996,2017]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
much shorter) than metatarsomere 5; terminal maxillary and labial palpomeres acuminate (
|
||
<emphasis id="779AEAB7FFC74165FEA0F7A6FEFDF7E2" box="[307,334,2049,2070]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
subtruncate to triangular); elytra with (
|
||
<emphasis id="779AEAB7FFC74165FD7FF7A6FCB4F7E2" box="[748,775,2049,2070]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
without) aciculate microsculpture along sides; small body size; and contiguous (
|
||
<emphasis id="779AEAB7FFC74165FB94FEF0FB91FE98" box="[1031,1058,343,364]" italics="true" pageId="3" pageNumber="8">vs.</emphasis>
|
||
separated) mesocoxae/ mesocoxal cavities as primary feature. These synapomorphies combined suggest closer relationships between
|
||
<taxonomicName id="82EE4D26FFC74165FB1DFE2BFB7DFE55" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1166,1230,396,417]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB1DFE2BFB7DFE55" box="[1166,1230,396,417]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC74165FB68FE2CFAF4FE54" authorityName="Andrewes" authorityYear="1919" box="[1275,1351,395,416]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB68FE2CFAF4FE54" box="[1275,1351,395,416]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
than between either and the other Paleotropical Ozaenini, and
|
||
<taxonomicName id="82EE4D26FFC74165FACCFE00FC62FE22" authority="Stork, 1985" authorityName="Stork" authorityYear="1985" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="mulu">
|
||
<emphasis id="779AEAB7FFC74165FACCFE00FCEFFE22" italics="true" pageId="3" pageNumber="8">D. mulu</emphasis>
|
||
<bibRefCitation id="217F4B54FFC74165FCF1FE66FC62FE22" author="Stork N. E." box="[866,977,448,471]" pageId="3" pageNumber="8" pagination="1113 - 1138" refId="ref11669" refString="Stork N. E. 1985. Dhanya, a South-east Asian genus of ozaenine ground beetles // J. Nat. Hist. Vol. 19. P. 1113 - 1138." type="journal article" year="1985">Stork, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
apparently fills morphological gap between the two genera with itself. It shares more characters with
|
||
<taxonomicName id="82EE4D26FFC74165FCBFFE50FCDFFDF8" authorityName="Schmidt-Gobel" authorityYear="1846" box="[812,876,503,524]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCBFFE50FCDFFDF8" box="[812,876,503,524]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
than with
|
||
<taxonomicName id="82EE4D26FFC74165FC46FE51FB92FDFF" authorityName="Andrewes" authorityYear="1919" box="[981,1057,502,523]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FC46FE51FB92FDFF" box="[981,1057,502,523]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="779AEAB7FFC74165FBBDFE50FBFFFDF8" box="[1070,1100,503,524]" italics="true" pageId="3" pageNumber="8">i.e.</emphasis>
|
||
, clypeus bisetose on each side anteriorly, mandibles medium-sized, scape globulose, pronotal lateral edge crenulate and pilose (
|
||
<emphasis id="779AEAB7FFC74165FB1FFD8BFB13FDB5" box="[1164,1184,556,577]" italics="true" pageId="3" pageNumber="8">vs</emphasis>
|
||
. smooth, with only 3– 4 lateral setae inside lateral edge), and meso- and metatarsal formula being similar: mesotarsi with tarsomere 1 about as long as tarsomere 5 while metatarsi with tarsomere 1 considerably longer than tarsomere 5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFC74165FCC1FD16FC39FB28" pageId="3" pageNumber="8" type="reference_group">
|
||
<paragraph id="455136A5FFC74165FCC1FD16FC39FB28" blockId="3.[812,1399,315,1244]" pageId="3" pageNumber="8">
|
||
<taxonomicName id="82EE4D26FFC74165FCC1FD16FC2DFD32" ID-CoL="8KWMG" authorityName="Andrewes" authorityYear="1919" box="[850,926,689,710]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCC1FD16FC2DFD32" box="[850,926,689,710]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
is much like a very small
|
||
<taxonomicName id="82EE4D26FFC74165FB05FD16FCEFFD15" ID-CoL="8KXD4" authority="Schmidt-Gobel, 1846" authorityName="Schmidt-Gobel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Itamus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB05FD16FB64FD32" box="[1174,1239,689,710]" italics="true" pageId="3" pageNumber="8">Itamus</emphasis>
|
||
<bibRefCitation id="217F4B54FFC74165FB4EFD16FCEFFD15" author="Schmidt-Gobel H. M." pageId="3" pageNumber="8" pagination="1 - 94" refId="ref11582" refString="Schmidt-Gobel H. M. 1846. Faunula Coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. P. I - VIII, 1 - 94, Pl. I - III." type="journal article" year="1846">Schmidt-Göbel, 1846</bibRefCitation>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="82EE4D26FFC74165FCECFD6CFB14FD14" ID-CoL="8KYZB" authority="Chaudoir, 1848" authorityName="Chaudoir" authorityYear="1848" box="[895,1191,715,737]" class="Insecta" family="Carabidae" genus="Sphaerostylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCECFD6CFBB5FD14" box="[895,1030,715,736]" italics="true" pageId="3" pageNumber="8">Sphaerostylus</emphasis>
|
||
Chaudoir, 1848
|
||
</taxonomicName>
|
||
, with which it shares many characters, including more or less derived ones, such as a fairly long hexagonal clypeus, gula totally reduced behind, large mandibles, and protarsomeres 1–3 with ventro-apical angles toothed or spinose at least posteriorly (externally). Based on this evidence,
|
||
<taxonomicName id="82EE4D26FFC74165FBBBFCF6FBDBFC92" ID-CoL="8KVTD" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1064,1128,849,870]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FBBBFCF6FBDBFC92" box="[1064,1128,849,870]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
may be supposed to have evolved from
|
||
<taxonomicName id="82EE4D26FFC74165FC24FCCCFBB0FC74" ID-CoL="8KWMG" authorityName="Andrewes" authorityYear="1919" box="[951,1027,875,896]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FC24FCCCFBB0FC74" box="[951,1027,875,896]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="82EE4D26FFC74165FBAEFCCCFB3AFC74" ID-CoL="8KWMG" authorityName="Andrewes" authorityYear="1919" box="[1085,1161,875,896]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FBAEFCCCFB3AFC74" box="[1085,1161,875,896]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
from ancestors close to
|
||
<taxonomicName id="82EE4D26FFC74165FCBFFC20FCDEFC68" ID-CoL="8KXD4" authorityName="Schmidt-Gobel" authorityYear="1846" box="[812,877,903,924]" class="Insecta" family="Carabidae" genus="Itamus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCBFFC20FCDEFC68" box="[812,877,903,924]" italics="true" pageId="3" pageNumber="8">Itamus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="82EE4D26FFC74165FCE6FC21FC4FFC6F" ID-CoL="8KYZB" authorityName="Chaudoir" authorityYear="1848" box="[885,1020,902,923]" class="Insecta" family="Carabidae" genus="Sphaerostylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCE6FC21FC4FFC6F" box="[885,1020,902,923]" italics="true" pageId="3" pageNumber="8">Sphaerostylus</emphasis>
|
||
</taxonomicName>
|
||
or something. On the other hand, the gula is posteriorly distinct in
|
||
<taxonomicName id="82EE4D26FFC74165FBFAFC06FB19FC42" ID-CoL="8KVTD" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1129,1194,929,950]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FBFAFC06FB19FC42" box="[1129,1194,929,950]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
, but much reduced throughout its length in the other Ozaenini examined for comparison, ranging from nearly indistinct in width in
|
||
<taxonomicName id="82EE4D26FFC74165FAACFC70FBDBFBF3" ID-CoL="78J8P" authority="(Klug, 1831)" baseAuthorityName="Klug" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Pseudozaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="779AEAB7FFC74165FAACFC70FC52FBF2" italics="true" pageId="3" pageNumber="8">Pseudozaena orientalis</emphasis>
|
||
(Klug, 1831)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC74165FB09FC56FAF4FBF2" box="[1178,1351,1009,1030]" class="Insecta" family="Carabidae" genus="Sphaerostylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FB09FC56FA92FBF2" box="[1178,1313,1009,1030]" italics="true" pageId="3" pageNumber="8">Sphaerostylus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, and through reduced to a suture posteriorly (
|
||
<taxonomicName id="82EE4D26FFC74165FB2DFBABFB4CFBD5" ID-CoL="8KXD4" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1214,1279,1036,1057]" class="Insecta" family="Carabidae" genus="Itamus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB2DFBABFB4CFBD5" box="[1214,1279,1036,1057]" italics="true" pageId="3" pageNumber="8">Itamus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE4D26FFC74165FA9DFBACFAC5FBD4" ID-CoL="8KXS3" authorityName="Perty" authorityYear="1830" box="[1294,1398,1035,1056]" class="Insecta" family="Carabidae" genus="Pachyteles" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FA9DFBACFAC5FBD4" box="[1294,1398,1035,1056]" italics="true" pageId="3" pageNumber="8">Pachyteles</emphasis>
|
||
</taxonomicName>
|
||
(s.lato) spp.,
|
||
<taxonomicName id="82EE4D26FFC74165FC39FB81FBE3FBCF" box="[938,1104,1062,1083]" class="Insecta" family="Carabidae" genus="Goniotropis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FC39FB81FBACFBCF" box="[938,1055,1062,1083]" italics="true" pageId="3" pageNumber="8">Goniotropis</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="779AEAB7FFC74165FBCAFB81FB5CFBCF" box="[1113,1263,1062,1083]" italics="true" pageId="3" pageNumber="8">Scythropaussus</emphasis>
|
||
sp.) to totally reduced (
|
||
<taxonomicName id="82EE4D26FFC74165FC1AFBE6FBB7FBA2" box="[905,1028,1089,1110]" class="Insecta" family="Carabidae" genus="Ozaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FC1AFBE6FC61FBA2" box="[905,978,1089,1110]" italics="true" pageId="3" pageNumber="8">Ozaena</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE4D26FFC74165FB83FBE6FB78FBA2" box="[1040,1227,1089,1110]" class="Insecta" family="Carabidae" genus="Platycerozaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="779AEAB7FFC74165FB83FBE6FB15FBA2" box="[1040,1190,1089,1110]" italics="true" pageId="3" pageNumber="8">Platycerozaena</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="82EE4D26FFC74165FB44FBE6FA90FBA2" ID-CoL="8KWMG" authorityName="Andrewes" authorityYear="1919" box="[1239,1315,1089,1110]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB44FBE6FA90FBA2" box="[1239,1315,1089,1110]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
) — this is inconsistent with the above hypothesis. Another character of
|
||
<taxonomicName id="82EE4D26FFC74165FCDFFBD0FC3FFB78" ID-CoL="8KVTD" authorityName="Schmidt-Gobel" authorityYear="1846" box="[844,908,1143,1164]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCDFFBD0FC3FFB78" box="[844,908,1143,1164]" italics="true" pageId="3" pageNumber="8">Eustra</emphasis>
|
||
</taxonomicName>
|
||
, the presense of ventral protarsal pad in male accordingly means its secondary reduction in
|
||
<taxonomicName id="82EE4D26FFC74165FB6EFB36FAFAFB52" ID-CoL="8KWMG" authorityName="Andrewes" authorityYear="1919" box="[1277,1353,1169,1190]" class="Insecta" family="Carabidae" genus="Dhanya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FB6EFB36FAFAFB52" box="[1277,1353,1169,1190]" italics="true" pageId="3" pageNumber="8">Dhanya</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE4D26FFC74165FCBFFB0BFCDEFB35" ID-CoL="8KXD4" authorityName="Schmidt-Gobel" authorityYear="1846" box="[812,877,1196,1217]" class="Insecta" family="Carabidae" genus="Itamus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="779AEAB7FFC74165FCBFFB0BFCDEFB35" box="[812,877,1196,1217]" italics="true" pageId="3" pageNumber="8">Itamus</emphasis>
|
||
</taxonomicName>
|
||
, partial to complete in the former while complete in the latter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFC74162FCBFFB57FCB0FD58" lastPageId="4" lastPageNumber="9" pageId="3" pageNumber="8" type="key">
|
||
<paragraph id="455136A5FFC74165FCBFFB57FB05FAF2" blockId="3.[812,1399,1264,2069]" box="[812,1206,1264,1287]" pageId="3" pageNumber="8">
|
||
<heading id="1E1981C9FFC74165FCBFFB57FB05FAF2" box="[812,1206,1264,1287]" fontSize="8" level="2" pageId="3" pageNumber="8" reason="6">
|
||
<smallCapsWord id="43B7A079FFC74165FCBFFB57FCEBFAF1" baselines="1281,1281" box="[812,856,1264,1287]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Key" pageId="3" pageNumber="8">KEY</smallCapsWord>
|
||
<smallCapsWord id="43B7A079FFC74165FCCCFB52FCCBFAF2" baselines="1282" box="[863,888,1269,1286]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="to" pageId="3" pageNumber="8">TO</smallCapsWord>
|
||
<smallCapsWord id="43B7A079FFC74165FCECFB52FC74FAF2" baselines="1282" box="[895,967,1269,1286]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="3" pageNumber="8">SPECIES</smallCapsWord>
|
||
<smallCapsWord id="43B7A079FFC74165FC5DFB52FC55FAF2" baselines="1282" box="[974,998,1269,1286]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="of" pageId="3" pageNumber="8">OF</smallCapsWord>
|
||
<taxonomicName id="82EE4D26FFC74165FC7EFB57FB86FAF1" ID-CoL="8KVTD" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1005,1077,1264,1287]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<smallCapsWord id="43B7A079FFC74165FC7EFB57FB86FAF1" baselines="1281,1281" box="[1005,1077,1264,1287]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eustra" pageId="3" pageNumber="8">
|
||
<emphasis id="779AEAB7FFC74165FC7EFB57FB86FAF1" box="[1005,1077,1264,1287]" italics="true" pageId="3" pageNumber="8">EUSTRA</emphasis>
|
||
</smallCapsWord>
|
||
</taxonomicName>
|
||
<smallCapsWord id="43B7A079FFC74165FBAFFB52FBE7FAF2" baselines="1282" box="[1084,1108,1269,1286]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="of" pageId="3" pageNumber="8">OF</smallCapsWord>
|
||
<smallCapsWord id="43B7A079FFC74165FBC8FB57FB05FAF2" baselines="1281,1282" box="[1115,1206,1264,1287]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Vietnam" pageId="3" pageNumber="8">
|
||
<collectingCountry id="3DF97635FFC74165FBC8FB57FB05FAF2" box="[1115,1206,1264,1287]" name="Vietnam" pageId="3" pageNumber="8">VIETNAM</collectingCountry>
|
||
</smallCapsWord>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="317F6194FFC74162FCBFFAB2FCB0FD58" lastPageId="4" lastPageNumber="9" pageId="3" pageNumber="8">
|
||
<paragraph id="455136A5FFC74165FCBFFAB2FBCFFAB1" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFFAB2FBCFFAB1" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFFAB2FBCFFAB1" pageId="3" pageNumber="8">1(12) Silvicolous, macropterous and macrophthalmic species; elytra mostly bimaculate.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFFAEDFBCAFA8E" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFFAEDFBCAFA8E" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFFAEDFBCAFA8E" pageId="3" pageNumber="8">2(11) Body unicoloured yellow to pale brown, except for elytra each with a darker spot.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFFAD8FAF9FA3E" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFFAD8FAF9FA3E" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFFAD8FAF9FA3E" pageId="3" pageNumber="8">
|
||
3(10) Elytral pubescence dense and long, setae as long as or barely longer than distance between neighbouring two in longitudinal row. Body slender,
|
||
<collectionCode id="23FFAE60FFC74165FB02FA12FB1DFA3E" box="[1169,1198,1461,1482]" pageId="3" pageNumber="8">EL</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFC74165FB26FA12FB69FA3E" box="[1205,1242,1461,1482]" pageId="3" pageNumber="8">EW</collectionCode>
|
||
1.31–1.45.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFFA68FAC0F9BB" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFFA68FAC0F9BB" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFFA68FAC0F9BB" pageId="3" pageNumber="8">
|
||
4(5) Head, especially vertex, very convex, neck constriction very deep. Pronotal lateral edge nearly smooth. Elytral spots large, mostly extending from 2/5 to just before apex. — Southern and Central
|
||
<collectingCountry id="3DF97635FFC74165FB16F9B8FB6BF9C1" box="[1157,1240,1567,1589]" name="Vietnam" pageId="3" pageNumber="8">Vietnam</collectingCountry>
|
||
........................ .............................................. 3.
|
||
<taxonomicName id="82EE4D26FFC74165FB00F99CFA88F9A4" ID-CoL="8M9DX" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1171,1339,1595,1616]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="posteroguttata" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC74165FB00F99CFA88F9A4" box="[1171,1339,1595,1616]" italics="true" pageId="3" pageNumber="8">E. posteroguttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC74165FAD0F99DFAC0F9BB" bold="true" box="[1347,1395,1594,1615]" pageId="3" pageNumber="8">
|
||
<taxonomicNameLabel id="6CA957CCFFC74165FAD0F99DFAC0F9BB" box="[1347,1395,1594,1615]" pageId="3" pageNumber="8" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFF9F3FABCF96B" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFF9F3FABCF96B" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFF9F3FABCF96B" pageId="3" pageNumber="8">5(4) Head with vertex slightly convex to nearly flat, neck constriction shallow to deep. Pronotal lateral edge nearly smooth. Elytral spots either smaller or vague.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFF903FAC0F8D0" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFF903FAC0F8D0" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFF903FAC0F8D0" pageId="3" pageNumber="8">
|
||
6(7) Elytral spot median in position; ocular ridge dorsally conspicuous, entire and black. Pronotum more rectangular than in the other species and transverse,
|
||
<collectionCode id="23FFAE60FFC74165FB60F97DFAA4F91B" box="[1267,1303,1754,1775]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="8">PW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFC74165FA8DF97DFA89F91B" box="[1310,1338,1754,1775]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="8" type="Museum">PL</collectionCode>
|
||
1.47– 1.60.
|
||
<collectionCode id="23FFAE60FFC74165FC1BF952FC14F8FE" box="[904,935,1781,1802]" pageId="3" pageNumber="8">BL</collectionCode>
|
||
<quantity id="82169B40FFC74165FC3EF952FB97F8FE" box="[941,1060,1781,1802]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="3" pageNumber="8" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
|
||
. — Southern and Central
|
||
<collectingCountry id="3DF97635FFC74165FA8EF952FAC0F8FF" box="[1309,1395,1781,1803]" name="Vietnam" pageId="3" pageNumber="8">Vietnam</collectingCountry>
|
||
. ..................................................... 1.
|
||
<taxonomicName id="82EE4D26FFC74165FB28F8B7FA8FF8D1" ID-CoL="8MJP3" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1211,1340,1808,1829]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="cariniceps" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC74165FB28F8B7FA8FF8D1" box="[1211,1340,1808,1829]" italics="true" pageId="3" pageNumber="8">E. cariniceps</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC74165FAD0F8A8FAC0F8D0" bold="true" box="[1347,1395,1807,1828]" pageId="3" pageNumber="8">
|
||
<taxonomicNameLabel id="6CA957CCFFC74165FAD0F8A8FAC0F8D0" box="[1347,1395,1807,1828]" pageId="3" pageNumber="8" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFF88DFAA3F8B4" blockId="3.[812,1399,1264,2069]" box="[812,1296,1834,1856]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFF88DFAA3F8B4" box="[812,1296,1834,1856]" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFF88DFAA3F8B4" box="[812,1296,1834,1856]" pageId="3" pageNumber="8">7(6) Combination of characters other than above.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFF8E2FAC0F85E" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFF8E2FAC0F85E" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFF8E2FAC0F85E" pageId="3" pageNumber="8">
|
||
8(9) Elytra vaguely infuscated on disc, without distinct spot; pronotum cordate and narrow,
|
||
<collectionCode id="23FFAE60FFC74165FBEEF8F8FB12F880" box="[1149,1185,1887,1908]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="8">PW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFC74165FB3BF8F8FB77F880" box="[1192,1220,1887,1908]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="8" type="Museum">PL</collectionCode>
|
||
1.33. Body small,
|
||
<collectionCode id="23FFAE60FFC74165FCC1F8DDFCC2F87B" box="[850,881,1914,1935]" pageId="3" pageNumber="8">BL</collectionCode>
|
||
<quantity id="82169B40FFC74165FCEBF8DDFC62F87B" box="[888,977,1914,1936]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="3" pageNumber="8" unit="mm" value="2.95">2.95 mm</quantity>
|
||
. — Northern
|
||
<collectingCountry id="3DF97635FFC74165FBCBF8DDFB18F864" box="[1112,1195,1914,1936]" name="Vietnam" pageId="3" pageNumber="8">Vietnam</collectingCountry>
|
||
............................... ................................................ 2.
|
||
<taxonomicName id="82EE4D26FFC74165FB0AF832FAC0F85E" ID-CoL="3DBDL" authority="Deuve, 2001" authorityName="Deuve" authorityYear="2001" box="[1177,1395,1941,1962]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="gomyi">
|
||
<emphasis id="779AEAB7FFC74165FB0AF832FB43F85E" box="[1177,1264,1941,1962]" italics="true" pageId="3" pageNumber="8">E. gomyi</emphasis>
|
||
<bibRefCitation id="217F4B54FFC74165FB64F832FAC0F85E" author="Deuve Th." box="[1271,1395,1941,1962]" pageId="3" pageNumber="8" pagination="547 - 578" refId="ref11147" refString="Deuve Th. 2001. Le genre Eustra Schmidt-Goebel, 1846, insectes (Coleoptera, Paussidae, Ozaeninae) a genitalia femelles orthotopiques // Zoosystema. V. 23. P. 547 - 578." type="journal article" year="2001">Deuve, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC74165FCBFF808FAC0F7E0" blockId="3.[812,1399,1264,2069]" pageId="3" pageNumber="8">
|
||
<keyLead id="FE1F9590FFC74165FCBFF808FAC0F7E0" pageId="3" pageNumber="8">
|
||
<keyStep id="FE1A2E00FFC74165FCBFF808FAC0F7E0" pageId="3" pageNumber="8">
|
||
9(8) Elytra each with a spot extending from middle to 2/5 elytra; pronotum more transverse,
|
||
<collectionCode id="23FFAE60FFC74165FB28F86DFB6CF82B" box="[1211,1247,1994,2015]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="8">PW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFC74165FB75F86DFAB1F82B" box="[1254,1282,1994,2015]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="8" type="Museum">PL</collectionCode>
|
||
1.50–1.57. Body larger,
|
||
<collectionCode id="23FFAE60FFC74165FC40F842FC41F80E" box="[979,1010,2021,2042]" pageId="3" pageNumber="8">BL</collectionCode>
|
||
<quantity id="82169B40FFC74165FC6BF843FBC2F80E" box="[1016,1137,2020,2042]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="3" pageNumber="8" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5–3.7 mm</quantity>
|
||
. — Southern
|
||
<collectingCountry id="3DF97635FFC74165FB64F842FAF9F80F" box="[1271,1354,2021,2043]" name="Vietnam" pageId="3" pageNumber="8">Vietnam</collectingCountry>
|
||
..... ...................................................... 4.
|
||
<taxonomicName id="82EE4D26FFC74165FB2DF7A7FA8FF7E0" ID-CoL="8M5BN" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1214,1340,2047,2069]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="8" phylum="Arthropoda" rank="species" species="dalatensis" status="sp.n.">
|
||
<emphasis id="779AEAB7FFC74165FB2DF7A7FA8FF7E0" box="[1214,1340,2047,2069]" italics="true" pageId="3" pageNumber="8">E. dalatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC74165FAD0F858FAC0F7E0" bold="true" box="[1347,1395,2047,2068]" pageId="3" pageNumber="8">
|
||
<taxonomicNameLabel id="6CA957CCFFC74165FAD0F858FAC0F7E0" box="[1347,1395,2047,2068]" pageId="3" pageNumber="8" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC04162FF2EFE9CFCB0FE4F" blockId="4.[189,776,315,684]" pageId="4" pageNumber="9">
|
||
<keyLead id="FE1F9590FFC04162FF2EFE9CFCB0FE4F" pageId="4" pageNumber="9">
|
||
<keyStep id="FE1A2E00FFC04162FF2EFE9CFCB0FE4F" pageId="4" pageNumber="9">
|
||
10(3) Elytral pubescence moderately dense and very short, setae about half as long as distance between neighbouring two in longitudinal row. Body robust,
|
||
<collectionCode id="23FFAE60FFC04162FD9DFED6FD99FE72" box="[526,554,369,390]" pageId="4" pageNumber="9">EL</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFC04162FDA2FED6FDE6FE72" box="[561,597,369,390]" pageId="4" pageNumber="9">EW</collectionCode>
|
||
1.26–1.31. Elytral spots small to indistinct, between middle and 3/4 elytra. — Southern
|
||
<collectingCountry id="3DF97635FFC04162FEADFE01FE3DFE48" box="[318,398,422,444]" name="Vietnam" pageId="4" pageNumber="9">Vietnam</collectingCountry>
|
||
.............. 6.
|
||
<taxonomicName id="82EE4D26FFC04162FD9EFE00FD75FE4F" ID-CoL="8M84P" authorityName="Fedorenko" authorityYear="2019" box="[525,710,422,444]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
|
||
<emphasis id="779AEAB7FFC04162FD9EFE00FD75FE4F" box="[525,710,422,444]" italics="true" pageId="4" pageNumber="9">E. nageli subglabra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFC04162FD5FFE01FCB0FE4F" bold="true" box="[716,771,422,443]" pageId="4" pageNumber="9">ssp.n.</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC04162FF2EFE66FCB0FDA8" blockId="4.[189,776,315,684]" pageId="4" pageNumber="9">
|
||
<keyLead id="FE1F9590FFC04162FF2EFE66FCB0FDA8" pageId="4" pageNumber="9">
|
||
<keyStep id="FE1A2E00FFC04162FF2EFE66FCB0FDA8" pageId="4" pageNumber="9">
|
||
11(2) Head dark brown to black, except for pale mouthparts, labrum, and clypeus; elytra dark brown, with suture, narrow lateral margin and apex, and basal third paler, reddish-brown to brown. Pronotal lateral edge tuberculate to denticulate. Elytra densely tuberculate-pubescent. .............................................. 5.
|
||
<taxonomicName id="82EE4D26FFC04162FDB2FDE0FCB0FDA8" ID-CoL="3DBDJ" authority="Jedlicka, 1968" authorityName="Jedlicka" authorityYear="1968" box="[545,771,582,604]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="species" species="csikii">
|
||
<emphasis id="779AEAB7FFC04162FDB2FDE0FDC3FDAF" box="[545,624,582,604]" italics="true" pageId="4" pageNumber="9">E. csikii</emphasis>
|
||
<bibRefCitation id="217F4B54FFC04162FDE4FDE1FCB0FDA8" author="Jedlicka A." box="[631,771,582,604]" pageId="4" pageNumber="9" pagination="285 - 296" refId="ref11405" refString="Jedlicka A. 1968. Neue Carabiden aus der palaearktischen Region (Coleoptera, Carabidae) // Reichenbachia. Bd. 8. Nr. 34. S. 285 - 296." type="journal article" year="1968">Jedlička, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="455136A5FFC04162FF2EFDC6FCB0FD58" blockId="4.[189,776,315,684]" pageId="4" pageNumber="9">
|
||
<keyLead id="FE1F9590FFC04162FF2EFDC6FCB0FD58" pageId="4" pageNumber="9">
|
||
<keyStep id="FE1A2E00FFC04162FF2EFDC6FCB0FD58" pageId="4" pageNumber="9">
|
||
12(1) Cavernicolous, apterous and microphthalmic species of uniformly pale brown colour. — Kien Kiang Province. .....................................
|
||
<taxonomicName id="82EE4D26FFC04162FE40FD30FCB0FD58" ID-CoL="3DBDN" authority="Deuve, 1996" authorityName="Deuve" authorityYear="1996" box="[467,771,662,684]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="species" species="honchongensis">
|
||
<emphasis id="779AEAB7FFC04162FE40FD30FD33FD5F" box="[467,640,662,684]" italics="true" pageId="4" pageNumber="9">E. honchongensis</emphasis>
|
||
<bibRefCitation id="217F4B54FFC04162FD14FD31FCB0FD58" author="Deuve Th." box="[647,771,662,684]" pageId="4" pageNumber="9" pagination="23 - 26" refId="ref11102" refString="Deuve Th. 1996. Description d'un coleoptere troglobie du genre Eustra, decouvert dans un karst du Vietnam meridional (Paussidae) // Rev. Franc. d'Entomol., N. S. Vol. 18. P. 23 - 26." type="journal article" year="1996">Deuve, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |