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<document id="48E3C142DF69C4BFDA6120890482A183" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D378784891BFFCC21CA3FA1FDEFFC3D" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus nunni Sokolov, 2016, sp.n." docType="treatment" docUuid="1FDA0B65-307E-4818-AC93-20BC5FE25E11" docUuidSource="ZooBank" docVersion="9" lastPageNumber="8" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="7" updateTime="1698393955637" updateUser="plazi">
<mods:mods id="C77F713932EECC3CC5F9E8D6FB8E688C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9AD8F43A8D337A6B2ADEFFB581F253E">
<mods:title id="B83016CB71A2FB1D7138ACFF56C285BB">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
</mods:titleInfo>
<mods:name id="F07FF3CD5187E1477140AE3771B95AF1" type="personal">
<mods:role id="737F2CC4DD70A4A6195B77AC260B8423">
<mods:roleTerm id="097DAD6EDF6C86FB6D82BBEB289B1EED">Author</mods:roleTerm>
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<mods:namePart id="AC158D9E2287423F464E0B97C610CD7C">Sokolov, Igor M.</mods:namePart>
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<mods:title id="99C103BB07FCA60E1BFC8302D20039BE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B0B0AAD2572944111CE57A0436102CE5">
<mods:date id="A1BCCACBC7B49B1EBFF2A476969B40C8">2016</mods:date>
<mods:detail id="3A88AD8600F465FA392CFDF7F7C3820D" type="volume">
<mods:number id="DB09FB2BF3C1EB6DEDC7C839DA938C24">4196</mods:number>
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<mods:start id="1E4B52B03BFCA64E8AF287D22F2C9DDC">1</mods:start>
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<mods:classification id="3D7F52937BF16928D2627AC4AB90FC0E">journal article</mods:classification>
<mods:identifier id="49E7140A4743BD74205830EB9210762F" type="DOI">10.11646/zootaxa.4196.1.1</mods:identifier>
<mods:identifier id="9F672439DDC3EA712520798B8FBB1A7B" type="GBIF-Dataset">b7651a7b-df24-4347-9585-0633b88dba51</mods:identifier>
<mods:identifier id="23CE1A259BB4260229C435D2294AD9BF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B2E59EA8211711B20D34E2BD6AD3BFDB" type="Zenodo-Dep">167648</mods:identifier>
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<treatment id="8D378784891BFFCC21CA3FA1FDEFFC3D" ID-DOI="http://doi.org/10.5281/zenodo.6089670" ID-GBIF-Taxon="124814315" ID-Zenodo-Dep="6089670" LSID="urn:lsid:zoobank.org:act:1FDA0B65-307E-4818-AC93-20BC5FE25E11" httpUri="http://treatment.plazi.org/id/8D378784891BFFCC21CA3FA1FDEFFC3D" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="4D846519891BFFCD21CA3FA1FD3FFACA" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="05213692891BFFCD21CA3FA1FE1FFAF5" blockId="6.[151,643,1269,1328]" box="[151,419,1269,1295]" pageId="6" pageNumber="7">
<heading id="5E6981FE891BFFCD21CA3FA1FE1FFAF5" bold="true" box="[151,419,1269,1295]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="37EAEA80891BFFCD21CA3FA1FE1FFAF5" bold="true" box="[151,419,1269,1295]" pageId="6" pageNumber="7">
<emphasis id="37EAEA80891BFFCD21CA3FA1FEDCFAF5" bold="true" box="[151,352,1269,1295]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="C29E4D11891BFFCD21CA3FA1FEE6FAF5" ID-CoL="8MMWQ" box="[151,346,1269,1295]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nunni" status="sp. nov.">Zeanillus nunni</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="2CD957FB891BFFCD20353FA1FE1FFAF5" box="[360,419,1269,1295]" pageId="6" pageNumber="7" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="05213692891BFFCD21CA3E43FD3FFACA" blockId="6.[151,643,1269,1328]" box="[151,643,1303,1328]" pageId="6" pageNumber="7">
<figureCitation id="9DA52A17891BFFCD21CA3E43FF5FFACA" box="[151,227,1303,1328]" captionStart="FIGURE 7" captionStartId="14.[151,250,1204,1226]" captionTargetBox="[151,1428,193,1181]" captionTargetId="figure@14.[151,1435,193,1182]" captionTargetPageId="14" captionText="FIGURE 7. Digital images of pterothorax and first abdominal ventrites, ventral aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. nunni; D— Z. pallidus; E— Z. pellucidus; F— Z. brouni; G— Z. lescheni; H— Z. montivagus; I— Z. nanus. Legend: ipa—intercoxal process of abdominal ventrite 2; mes—metendosternite; mte—metanepisternum; mtvmetaventrite; v 1 v 2 — abdominal ventrites 1 2. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/167655/files/figure.png" pageId="6" pageNumber="7">Figs. 7</figureCitation>
C, 11C, 12EH, 14C, 15C, 16C, 18A
</paragraph>
</subSubSection>
<subSubSection id="4D846519891BFFCD21CA3E0BFABCF8FE" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="05213692891BFFCD21CA3E0BFC07FA66" blockId="6.[151,1437,1375,2013]" pageId="6" pageNumber="7">
<emphasis id="37EAEA80891BFFCD21CA3E0BFEFAFA82" bold="true" box="[151,326,1375,1400]" pageId="6" pageNumber="7">Type material.</emphasis>
HOLOTYPE, male, in NZAC, board-mounted, undissected, labeled: / New Zealand DN Trotters Gorge Palmerston
<date id="71201052891BFFCD20353ED0FE22FA67" box="[360,414,1412,1437]" pageId="6" pageNumber="7">9 Jul</date>
0 6 / Soil sample kanuka/kowhai/mahoe forest /.
</paragraph>
<paragraph id="05213692891BFFCD219A3EFCFABCF8FE" blockId="6.[151,1437,1375,2013]" pageId="6" pageNumber="7">
PARATYPES (32 specimens, 6 dissected), 6 males and 6 females labeled same as holotype;
<materialsCitation id="B5F63CCF891BFFCD25823EFCFA3DFA1E" ID-GBIF-Occurrence="1341350627" collectingDate="2011-08" collectorName="DN Evansdale Glen Evansdale" country="New Zealand" elevation="40" location="Washed" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<specimenCount id="1398FD1B891BFFCD25823EFCFA97FA3A" box="[1247,1323,1447,1472]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD21CA3E99FE92FA1F" box="[151,302,1484,1509]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD206B3E99FD2EFA1F" box="[310,658,1484,1509]" pageId="6" pageNumber="7">DN Evansdale Glen Evansdale</collectorName>
<collectingDate id="6164E9BA891BFFCD23C43E98FD50FA1E" box="[665,748,1484,1508]" pageId="6" pageNumber="7" value="2011-08">
<date id="71201052891BFFCD23C43E98FD50FA1E" box="[665,748,1484,1508]" pageId="6" pageNumber="7" value="2011-08">11 Aug</date>
</collectingDate>
0 7 /
<location id="00416049891BFFCD227A3E98FCC3FA1F" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD227A3E98FCC3FA1F" box="[807,895,1484,1509]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample
<elevation id="8EB3D1A1891BFFCD25533E98FBFCFA1E" box="[1038,1088,1484,1508]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="m" value="40.0">
<quantity id="C2669B77891BFFCD25533E98FBFCFA1E" box="[1038,1088,1484,1508]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="m" value="40.0">40m</quantity>
</elevation>
broadleaf forest by stream /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD24D33E98FB84F9F2" ID-GBIF-Occurrence="1341350621" collectingDate="2010-06" collectorName="DN Evansdale Glen" country="New Zealand" location="Washed" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<specimenCount id="1398FD1B891BFFCD24D33E98FF71F9F2" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD201E3EA4FE65F9F2" box="[323,473,1519,1544]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD20BD3EA4FD7CF9F2" box="[480,704,1519,1544]" pageId="6" pageNumber="7">DN Evansdale Glen</collectorName>
<collectingDate id="6164E9BA891BFFCD239B3EA4FCACF9FD" box="[710,784,1520,1544]" pageId="6" pageNumber="7" value="2010-06">
<date id="71201052891BFFCD239B3EA4FCACF9FD" box="[710,784,1520,1544]" pageId="6" pageNumber="7" value="2010-06">10 Jun</date>
</collectingDate>
0 6 /
<location id="00416049891BFFCD22143EBBFC1DF9F2" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD22143EBBFC1DF9F2" box="[841,929,1519,1544]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD25193EA4FC46F9D6" ID-GBIF-Occurrence="1341350614" collectingDate="1928-11" collectorName="DN Woodside Glen Outram" country="New Zealand" location="Washed" pageId="6" pageNumber="7" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="1398FD1B891BFFCD25193EA4FB26F9F2" box="[1092,1178,1519,1544]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
and
<specimenCount id="1398FD1B891BFFCD258C3EA4FA81F9F2" box="[1233,1341,1519,1544]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD21F83D41FE87F9D7" box="[165,315,1556,1581]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD201F3D41FDC5F9D6" box="[322,633,1556,1581]" pageId="6" pageNumber="7">DN Woodside Glen Outram</collectorName>
<collectingDate id="6164E9BA891BFFCD23DD3D40FD6FF9D6" box="[640,723,1556,1580]" pageId="6" pageNumber="7" value="1928-11">
<date id="71201052891BFFCD23DD3D40FD6FF9D6" box="[640,723,1556,1580]" pageId="6" pageNumber="7" value="1928-11">28 Nov</date>
</collectingDate>
0 6 /
<location id="00416049891BFFCD22563D40FCDFF9D7" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD22563D40FCDFF9D7" box="[779,867,1556,1581]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD255B3D40FC65F9AA" ID-GBIF-Occurrence="1341350616" collectingDate="1916-06" collectorName="DN Woodside Glen Outram" country="New Zealand" location="Washed" pageId="6" pageNumber="7" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="1398FD1B891BFFCD255B3D40FBEDF9D7" box="[1030,1105,1556,1581]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
and
<specimenCount id="1398FD1B891BFFCD25DA3D40FB4FF9D7" box="[1159,1267,1556,1581]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD243A3D41FF4DF9AA" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD21A13D6CFDFCF9AA" box="[252,576,1591,1616]" pageId="6" pageNumber="7">DN Woodside Glen Outram</collectorName>
<collectingDate id="6164E9BA891BFFCD23163D63FD25F9B5" box="[587,665,1591,1616]" pageId="6" pageNumber="7" value="1916-06">
<date id="71201052891BFFCD23163D63FD25F9B5" box="[587,665,1591,1616]" pageId="6" pageNumber="7" value="1916-06">16 Jun</date>
</collectingDate>
0 6 /
<location id="00416049891BFFCD23833D63FC8AF9AA" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD23833D63FC8AF9AA" box="[734,822,1591,1616]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD22B43D6CFC5DF98E" ID-GBIF-Occurrence="1341350626" collectingDate="1918-11" collectorName="DN Woodside Glen Outram" country="New Zealand" location="Washed" pageId="6" pageNumber="7" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="1398FD1B891BFFCD22B43D6CFBFFF9AA" box="[1001,1091,1591,1616]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
and
<specimenCount id="1398FD1B891BFFCD25DC3D6CFB5AF9AA" box="[1153,1254,1591,1616]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD243A3D6CFF4DF98F" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD21A13D09FD83F98E" box="[252,575,1628,1653]" pageId="6" pageNumber="7">DN Woodside Glen Outram</collectorName>
<collectingDate id="6164E9BA891BFFCD23163D08FD1DF98E" box="[587,673,1628,1652]" pageId="6" pageNumber="7" value="1918-11">
<date id="71201052891BFFCD23163D08FD1DF98E" box="[587,673,1628,1652]" pageId="6" pageNumber="7" value="1918-11">18 Nov</date>
</collectingDate>
0 6 /
<location id="00416049891BFFCD23BB3D08FC82F98F" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD23BB3D08FC82F98F" box="[742,830,1628,1653]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD22AD3D08FB33F962" ID-GBIF-Occurrence="1341350611" collectingDate="2008-09-13" collectorName="Woodside Glen &amp; Outram &amp; J. Nunn" country="New Zealand" elevation="50" location="New Zealand" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<specimenCount id="1398FD1B891BFFCD22AD3D08FB83F98F" box="[1008,1087,1628,1653]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD259D3D09FAE3F98F" box="[1216,1375,1628,1653]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
: DN,
<collectorName id="A86B5344891BFFCD21CA3D2BFEFAF962" box="[151,326,1663,1688]" pageId="6" pageNumber="7">Woodside Glen</collectorName>
,
<collectorName id="A86B5344891BFFCD200C3DD4FE15F962" box="[337,425,1664,1688]" pageId="6" pageNumber="7">Outram</collectorName>
,
<elevation id="8EB3D1A1891BFFCD20E83D2BFE55F962" box="[437,489,1663,1688]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="m" value="50.0">
<quantity id="C2669B77891BFFCD20E83D2BFE55F962" box="[437,489,1663,1688]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="m" value="50.0">50m</quantity>
</elevation>
, washed soil sample, broadleaf forest
<collectorName id="A86B5344891BFFCD22C93DD4FC58F96D" box="[916,996,1664,1687]" pageId="6" pageNumber="7">J.Nunn</collectorName>
<collectingDate id="6164E9BA891BFFCD22B63DD4FBC2F962" box="[1003,1150,1664,1688]" pageId="6" pageNumber="7" value="2008-09-13">
<date id="71201052891BFFCD22B63DD4FBC2F962" box="[1003,1150,1664,1688]" pageId="6" pageNumber="7" value="2008-09-13">13 Sept 2008</date>
</collectingDate>
/
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD25C13DD4FA94F946" ID-GBIF-Occurrence="1341350622" collectingDate="1921-06" collectorName="DN State Highway &amp; Rocklands Rd" country="New Zealand" location="Washed" pageId="6" pageNumber="7" specimenCount="3" specimenCount-male="3">
<specimenCount id="1398FD1B891BFFCD25C13DD4FB4FF962" box="[1180,1267,1663,1688]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD243A3DD4FF4DF947" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD21A13DF1FE6AF947" box="[252,470,1700,1725]" pageId="6" pageNumber="7">DN State Highway</collectorName>
87 at
<collectorName id="A86B5344891BFFCD237A3DF0FD74F947" box="[551,712,1700,1725]" pageId="6" pageNumber="7">Rocklands Rd</collectorName>
<collectingDate id="6164E9BA891BFFCD238F3DF0FC9CF946" box="[722,800,1700,1724]" pageId="6" pageNumber="7" value="1921-06">
<date id="71201052891BFFCD238F3DF0FC9CF946" box="[722,800,1700,1724]" pageId="6" pageNumber="7" value="1921-06">21 Jun</date>
</collectingDate>
0 9 /
<location id="00416049891BFFCD223E3DF0FC07F947" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD223E3DF0FC07F947" box="[867,955,1700,1725]" country="New Zealand" name="Washed" pageId="6" pageNumber="7">Washed</location>
soil sample, remnant tussock /
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD24653DF0FB6FF91A" ID-GBIF-Occurrence="1341350610" collectingDate="2008-04-12" collectorName="Semple Rd &amp; Caneys Creek Tck &amp; Edendale J." country="New Zealand" location="Caneys Creek" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1">
<specimenCount id="1398FD1B891BFFCD24653DF0FA20F947" box="[1336,1436,1700,1725]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD205A3D9CFE1FF91A" box="[263,419,1735,1760]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
: DN,
<collectorName id="A86B5344891BFFCD20BB3D93FDDFF91A" box="[486,611,1735,1760]" pageId="6" pageNumber="7">Semple Rd</collectorName>
/
<collectorName id="A86B5344891BFFCD23363D93FC81F91A" box="[619,829,1735,1760]" pageId="6" pageNumber="7">
<location id="00416049891BFFCD23363D93FCB4F91A" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD23363D93FCB4F91A" box="[619,776,1735,1760]" country="New Zealand" name="Caneys Creek" pageId="6" pageNumber="7">Caneys Creek</location>
Tck
</collectorName>
, nr
<collectorName id="A86B5344891BFFCD22353D93FC5BF925" box="[872,999,1735,1760]" pageId="6" pageNumber="7">Edendale J.</collectorName>
Nunn
<collectingDate id="6164E9BA891BFFCD256D3D9CFB7CF91A" box="[1072,1216,1736,1760]" pageId="6" pageNumber="7" value="2008-04-12">
<date id="71201052891BFFCD256D3D9CFB7CF91A" box="[1072,1216,1736,1760]" pageId="6" pageNumber="7" value="2008-04-12">12 Apr 2008</date>
</collectingDate>
/
</materialsCitation>
;
<materialsCitation id="B5F63CCF891BFFCD25823D9CFABCF8FE" ID-GBIF-Occurrence="1341350624" collectingDate="2 Dec" collectorName="SL Black Gully Blue Mtns" country="New Zealand" elevation="390" location="Nothophagus forest" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<specimenCount id="1398FD1B891BFFCD25823D9CFA97F91A" box="[1247,1323,1735,1760]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
labeled: /
<collectingCountry id="7D897602891BFFCD21CA3DB9FE91F8FF" box="[151,301,1772,1797]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
<collectorName id="A86B5344891BFFCD20693DB8FDDDF8FE" box="[308,609,1772,1797]" pageId="6" pageNumber="7">SL Black Gully Blue Mtns</collectorName>
<elevation id="8EB3D1A1891BFFCD233A3DB8FD1BF8FE" box="[615,679,1772,1796]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="7" unit="m" value="390.0">
<quantity id="C2669B77891BFFCD233A3DB8FD1BF8FE" box="[615,679,1772,1796]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="7" unit="m" value="390.0">390m</quantity>
</elevation>
<collectingDate id="6164E9BA891BFFCD23F33DB8FD4CF8FE" box="[686,752,1772,1796]" pageId="6" pageNumber="7">
<date id="71201052891BFFCD23F33DB8FD4CF8FE" box="[686,752,1772,1796]" pageId="6" pageNumber="7">2 Dec</date>
</collectingDate>
0 6 / Washed soil sample
<location id="00416049891BFFCD25533DB8FB56F8FF" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD25533DB8FB56F8FF" box="[1038,1258,1772,1797]" country="New Zealand" name="Nothophagus forest" pageId="6" pageNumber="7">Nothophagus forest</location>
/.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4D846519891BFFCD219A3C5BFCE4F8B7" pageId="6" pageNumber="7" type="description">
<paragraph id="05213692891BFFCD219A3C5BFCE4F8B7" blockId="6.[151,1437,1375,2013]" pageId="6" pageNumber="7">
<emphasis id="37EAEA80891BFFCD219A3C5BFE39F8D2" bold="true" box="[199,389,1807,1832]" pageId="6" pageNumber="7">Specific epithet.</emphasis>
The specific epithet is a Latinized eponym in the genitive case, and is based on the surname of John T. Nunn, the collector of the
<typeStatus id="DA258830891BFFCD23693C61FDD8F8B7" box="[564,612,1845,1869]" pageId="6" pageNumber="7">type</typeStatus>
series of this species.
</paragraph>
</subSubSection>
<subSubSection id="4D846519891BFFCD219A3C03FA2FF827" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="05213692891BFFCD219A3C03FC15F88A" blockId="6.[151,1437,1375,2013]" box="[199,937,1879,1904]" pageId="6" pageNumber="7">
<materialsCitation id="B5F63CCF891BFFCD219A3C03FC15F88A" ID-GBIF-Occurrence="1341350619" box="[199,937,1879,1904]" country="New Zealand" location="South Island" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
<emphasis id="37EAEA80891BFFCD219A3C03FED9F88A" bold="true" box="[199,357,1879,1904]" pageId="6" pageNumber="7">
<typeStatus id="DA258830891BFFCD219A3C03FEBEF88A" box="[199,258,1879,1904]" pageId="6" pageNumber="7">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="7D897602891BFFCD20313C0CFDB9F88A" box="[364,517,1879,1904]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
,
<location id="00416049891BFFCD234D3C03FD23F88A" LSID="urn:lsid:plazi:treatment:8D378784891BFFCC21CA3FA1FDEFFC3D:00416049891BFFCD234D3C03FD23F88A" box="[528,671,1879,1904]" country="New Zealand" name="South Island" pageId="6" pageNumber="7" stateProvince="Otago">South Island</location>
,
<collectingRegion id="C75AF870891BFFCD23F63C0CFD4EF88A" box="[683,754,1880,1904]" country="New Zealand" name="Otago" pageId="6" pageNumber="7">Otago</collectingRegion>
, Trotters Gorge.
</materialsCitation>
</paragraph>
<paragraph id="05213692891BFFCD219A3C28FA2FF827" blockId="6.[151,1437,1375,2013]" pageId="6" pageNumber="7">
<emphasis id="37EAEA80891BFFCD219A3C28FEE1F86F" bold="true" box="[199,349,1916,1941]" pageId="6" pageNumber="7">Recognition.</emphasis>
Adults of this species (
<figureCitation id="9DA52A17891BFFCD233E3C28FD0DF86E" box="[611,689,1916,1941]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
C) are practically indistinguishable externally from the adults of
<taxonomicName id="C29E4D11891BFFCD24D83C29FEB0F842" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA80891BFFCD24D83C29FEB0F842" italics="true" pageId="6" pageNumber="7">Z. phyllobius</emphasis>
</taxonomicName>
and
<taxonomicName id="C29E4D11891BFFCD201A3CF5FE57F842" box="[327,491,1953,1976]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA80891BFFCD201A3CF5FE57F842" box="[327,491,1953,1976]" italics="true" pageId="6" pageNumber="7">Z. punctigerus</emphasis>
</taxonomicName>
, and are distinguished from the males of those by the structure of male genitalia. From other congeners, adults of
<taxonomicName id="C29E4D11891BFFCD235E3C91FDDCF826" box="[515,608,1989,2012]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA80891BFFCD235E3C91FDDCF826" box="[515,608,1989,2012]" italics="true" pageId="6" pageNumber="7">Z. nunni</emphasis>
</taxonomicName>
can be distinguished by the characters listed in the subgeneric diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="4D846519891AFFCC219A3BC3FDEFFC3D" pageId="7" pageNumber="8" type="description">
<paragraph id="05213692891AFFCC219A3BC3FAE1FF4A" blockId="7.[151,1437,151,968]" box="[199,1373,151,176]" pageId="7" pageNumber="8">
<emphasis id="37EAEA80891AFFCC219A3BC3FEE5FF4A" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="8">Description.</emphasis>
Size. Small to medium for genus (SBL range
<quantity id="C2669B77891AFFCC22033BC3FBBFFF55" box="[862,1027,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="8" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.501.80 mm</quantity>
, mean 1.65±
<quantity id="C2669B77891AFFCC25CC3BC3FABBFF55" box="[1169,1287,151,176]" metricMagnitude="-5" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="8" unit="mm" value="0.083">0.083 mm</quantity>
, n=20).
</paragraph>
<paragraph id="05213692891AFFCC219A3BE8FE4DFEE7" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.35±0.010), head of moderate width for genus compared to pronotum (WH/WPm 0.74±0.023), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.018).</paragraph>
<paragraph id="05213692891AFFCC219A3A73FCF7FEC5" blockId="7.[151,1437,151,968]" box="[199,843,295,320]" pageId="7" pageNumber="8">Color. Body color rufotestaceous, appendages testaceous.</paragraph>
<paragraph id="05213692891AFFCC219A3A18FC20FE57" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">Prothorax. Pronotum moderately long (LP/LE 0.43±0.023) and moderately elongate (WPm/LP 1.29±0.034), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.65±0.059). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.28±0.053).</paragraph>
<paragraph id="05213692891AFFCC219A3AE3FE86FDE2" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">Elytra. Slightly depressed along suture, comparatively short (LE/SBL 0.49±0.017) and moderately wide (WE/ LE 0.73±0.030). Lateral margins slightly divergent at basal third, subparallel at middle, and evenly rounded to apex in apical third.</paragraph>
<paragraph id="05213692891AFFCC219A3970FB58FD7E" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">
Male genitalia. Median lobe (
<figureCitation id="9DA52A17891AFFCC23493970FDD9FDC6" box="[532,613,548,573]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
E) with straight ventral margin and short apex with tapering tip. Sclerites of internal sac united basally, long, almost equal the length between apical and basal orifices. Left paramere with apical constriction (
<figureCitation id="9DA52A17891AFFCC20293938FE7AFD7E" box="[372,454,620,645]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
G). Ring sclerite with short and markedly transverse handle (
<figureCitation id="9DA52A17891AFFCC252D3938FB7EFD7E" box="[1136,1218,620,645]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
C).
</paragraph>
<paragraph id="05213692891AFFCC219A39DBFD92FD0A" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">
Female internal genitalia. Goncoxite 2 with short ensiform setae (
<figureCitation id="9DA52A17891AFFCC22FD39DBFC4CFD52" box="[928,1008,655,680]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="7" pageNumber="8">Fig. 15</figureCitation>
C). Spermatheca sclerotized, elongate, with four compartments (
<figureCitation id="9DA52A17891AFFCC20EE39E0FDB8FD37" box="[435,516,692,717]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
C). Spermathecal duct short with few coils. Attachments of spermathecal duct and gland to spermatheca close together.
</paragraph>
<paragraph id="05213692891AFFCC219A39A8FB95FCC2" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">
<emphasis id="37EAEA80891AFFCC219A39A8FDB9FCEF" bold="true" box="[199,517,764,789]" pageId="7" pageNumber="8">Geographical distribution.</emphasis>
This species is known from the eastern foothills of
<collectingRegion id="C75AF870891AFFCC251139A8FB2DFCEE" box="[1100,1169,764,788]" country="New Zealand" name="Otago" pageId="7" pageNumber="8">Otago</collectingRegion>
within Dunedin area of
<bibRefCitation id="610F4B63891AFFCC21CA384BFEC4FCC2" box="[151,376,799,824]" pageId="7" pageNumber="8" refString="Crosby, T. K., Dugdale, J. S. &amp; Watt, J. C. (1998) Area codes for recording specimens localities in the New Zealand subregion. New Zealand Journal of Zoology, 25, 175 - 183. http: // dx. doi. org / 10.1080 / 03014223.1998.9518148" type="journal article">
Crosby
<emphasis id="37EAEA80891AFFCC21B23875FE9BFCC2" box="[239,295,800,824]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(1998)
</bibRefCitation>
and the Blue Mountains of
<collectingRegion id="C75AF870891AFFCC23EC384BFC9FFCC2" box="[689,803,799,824]" country="New Zealand" name="Southland" pageId="7" pageNumber="8">Southland</collectingRegion>
(
<figureCitation id="9DA52A17891AFFCC226F384BFC3EFCC2" box="[818,898,799,824]" captionStart="FIGURE 18" captionStartId="30.[151,250,1918,1940]" captionTargetBox="[198,1387,199,1885]" captionTargetId="figure@30.[189,1398,193,1897]" captionTargetPageId="30" captionText="FIGURE 18. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Zeanillus. A: white circles, Z. phyllobius; green circles, Z. punctigerus; red circles, Z. nunni; blue diamond, Z. pellucidus; green diamond, Z. pallidus. B: white triangles, Z. brouni; green triangles, Z. nanus; red triangles, Z. lescheni; yellow triangle, Z. carltoni; black triangle, Z. montivagus. Elevation scale bars are given in meters." httpUri="https://zenodo.org/record/167666/files/figure.png" pageId="7" pageNumber="8">Fig. 18</figureCitation>
A, red circles).
</paragraph>
<paragraph id="05213692891AFFCC219A3810FDA1FC7A" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">
<emphasis id="37EAEA80891AFFCC219A3810FEECFCA7" bold="true" box="[199,336,836,861]" pageId="7" pageNumber="8">Way of life.</emphasis>
Specimens were collected from soil in a broad range of habitats: tussocks,
<taxonomicName id="C29E4D11891AFFCC25FE3810FA98FCA6" box="[1187,1316,836,860]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="37EAEA80891AFFCC25FE3810FA98FCA6" box="[1187,1316,836,860]" italics="true" pageId="7" pageNumber="8">Nothofagus</emphasis>
</taxonomicName>
, broadleaf and kanuka/kowhai/mahoe forests.
</paragraph>
<paragraph id="05213692891AFFCC219A38D8FDEFFC3D" blockId="7.[151,1437,151,968]" pageId="7" pageNumber="8">
<emphasis id="37EAEA80891AFFCC219A38D8FECCFC5F" bold="true" box="[199,368,908,933]" pageId="7" pageNumber="8">Relationships.</emphasis>
Based on the structure of male genitalia and spermatheca
<taxonomicName id="C29E4D11891AFFCC22A238D9FBE1FC5E" box="[1023,1117,909,932]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA80891AFFCC22A238D9FBE1FC5E" box="[1023,1117,909,932]" italics="true" pageId="7" pageNumber="8">Z. nunni</emphasis>
</taxonomicName>
is postulated to be the sister taxon of
<taxonomicName id="C29E4D11891AFFCC21A738E5FE31FC32" box="[250,397,944,968]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA80891AFFCC21A738E5FE31FC32" box="[250,397,944,968]" italics="true" pageId="7" pageNumber="8">Z. phyllobius</emphasis>
</taxonomicName>
+
<taxonomicName id="C29E4D11891AFFCC20F738E5FDF0FC3D" box="[426,588,944,968]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA80891AFFCC20F738E5FDF0FC3D" box="[426,588,944,968]" italics="true" pageId="7" pageNumber="8">Z. punctigerus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>