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<document id="DCDAD23A38D4C89ACC5D799F8375AEF5" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D3787848917FFC421CA3C56FC4AFE7D" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus punctigerus Broun" docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="11" updateTime="1698393955637" updateUser="plazi">
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<mods:title id="7561656C2D1A8D794A70CEEFC181A1DC">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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<mods:namePart id="23058E4F13C75527F8082147BE50AB75">Sokolov, Igor M.</mods:namePart>
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<mods:date id="33A6A1E8A61E92E68E0606E07F41600F">2016</mods:date>
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<treatment id="8D3787848917FFC421CA3C56FC4AFE7D" ID-DOI="http://doi.org/10.5281/zenodo.6089674" ID-GBIF-Taxon="124814317" ID-Zenodo-Dep="6089674" LSID="urn:lsid:plazi:treatment:8D3787848917FFC421CA3C56FC4AFE7D" httpUri="http://treatment.plazi.org/id/8D3787848917FFC421CA3C56FC4AFE7D" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11">
<subSubSection id="4D8465198917FFC121CA3C56FBB6F8C4" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="052136928917FFC121CA3C56FDB5F8E7" blockId="10.[151,1034,1794,1854]" box="[151,521,1794,1821]" pageId="10" pageNumber="11">
<heading id="5E6981FE8917FFC121CA3C56FDB5F8E7" bold="true" box="[151,521,1794,1821]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="C29E4D118917FFC121CA3C56FDB5F8E7" ID-CoL="8M63K" authority="Broun" authorityName="Broun" box="[151,521,1794,1821]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808917FFC121CA3C56FDB5F8E7" bold="true" box="[151,521,1794,1821]" pageId="10" pageNumber="11">
<emphasis id="37EAEA808917FFC121CA3C56FE23F8E6" bold="true" box="[151,415,1794,1820]" italics="true" pageId="10" pageNumber="11">Zeanillus punctigerus</emphasis>
(Broun)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="052136928917FFC121CA3C71FBB6F8C4" blockId="10.[151,1034,1794,1854]" box="[151,1034,1829,1854]" pageId="10" pageNumber="11">
<figureCitation id="9DA52A178917FFC121CA3C71FF5CF8C4" box="[151,224,1829,1854]" captionStart="FIGURE 1" captionStartId="8.[151,250,1917,1939]" captionTargetBox="[172,1423,199,1885]" captionTargetId="figure@8.[164,1430,193,1891]" captionTargetPageId="8" captionText="FIGURE 1. SEM illustrations of head, dorsal aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. pallidus; D— Z. pellucidus; E— Z. lescheni; F— Z. nanus. Legend: cl—clypeus; cs—clypeal seta; fcc—fronto-clypeal carina; fs—frontal seta; lb—labrum; mp 3 — maxillary palpomere 3; mp 4 — maxillary palpomere 4; pos—postorbital seta; ssa—anterior supraorbital seta; ssp—posterior supraorbital seta. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/257540/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
B, 4B, 5B, 6B, 7B, 8A, 9B, 10BGM, 11B, 12AD, 14B, 15B, 16B, 18A
</paragraph>
</subSubSection>
<subSubSection id="4D8465198917FFC121CA3C39FB27F879" box="[151,1179,1901,1923]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="052136928917FFC121CA3C39FB27F879" blockId="10.[151,1179,1901,1923]" box="[151,1179,1901,1923]" pageId="10" pageNumber="11">
<materialsCitation id="B5F63CCF8917FFC121CA3C39FB27F879" ID-GBIF-Occurrence="1341350623" box="[151,1179,1901,1923]" location="Mount Hutt" municipality="Methven" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="258E11BC8917FFC121CA3C39FB27F879" box="[151,1179,1901,1923]" pageId="10" pageNumber="11">
<taxonomicName id="C29E4D118917FFC121CA3C39FDABF879" ID-CoL="8L2MS" authority="Broun, 1914: 153" authorityName="Broun" authorityPageNumber="153" authorityYear="1914" box="[151,535,1901,1923]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808917FFC121CA3C39FEE1F879" box="[151,349,1901,1923]" italics="true" pageId="10" pageNumber="11">Anillus punctigerus</emphasis>
<treatmentCitation id="843F10838917FFC1203F3C3AFDABF879" author="Broun" box="[354,535,1901,1923]" page="153" pageId="10" pageNumber="11" year="1914">
<bibRefCitation id="610F4B638917FFC1203F3C3AFE58F879" author="Broun" box="[354,484,1901,1923]" pageId="10" pageNumber="11" refString="Broun, T. (1914) Descriptions of new genera and species of Coleoptera, Part III. New Zealand Institute Bulletin, 1, 143 - 266." type="journal article" year="1914">Broun, 1914</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="DA2588308917FFC1237C3C3AFDEFF879" box="[545,595,1902,1923]" pageId="10" pageNumber="11">Type</typeStatus>
locality: «
<location id="004160498917FFC123E33C3AFC84F879" LSID="urn:lsid:plazi:treatment:8D3787848917FFC421CA3C56FC4AFE7D:004160498917FFC123E33C3AFC84F879" box="[702,824,1902,1923]" municipality="Methven" name="Mount Hutt" pageId="10" pageNumber="11">Mount Hutt</location>
, near
<collectingMunicipality id="E545ACE88917FFC122283C39FC6CF879" box="[885,976,1901,1923]" pageId="10" pageNumber="11">Methven</collectingMunicipality>
» (original citation).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4D8465198917FFC521CA3CFBFA28FA49" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="052136928917FFC121CA3CFBFDF1F816" blockId="10.[151,1436,1967,2029]" pageId="10" pageNumber="11">
<materialsCitation id="B5F63CCF8917FFC121CA3CFBFDF1F816" ID-GBIF-Occurrence="1341350630" location="Sharplin Falls" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Canterbury">
<emphasis id="37EAEA808917FFC121CA3CFBFE30F832" bold="true" box="[151,396,1967,1992]" pageId="10" pageNumber="11">Examined material.</emphasis>
Thirteen specimens (5 dissected) from the following localities:
<collectingRegion id="C75AF8708917FFC125FF3CFBFA98F832" box="[1186,1316,1967,1992]" country="New Zealand" name="Canterbury" pageId="10" pageNumber="11">Canterbury</collectingRegion>
: Staveley,
<location id="004160498917FFC121CA3C80FE8BF817" LSID="urn:lsid:plazi:treatment:8D3787848917FFC421CA3C56FC4AFE7D:004160498917FFC121CA3C80FE8BF817" box="[151,311,2004,2029]" name="Sharplin Falls" pageId="10" pageNumber="11" stateProvince="Canterbury">Sharplin Falls</location>
; Geraldine, Orari Gorge.
</materialsCitation>
</paragraph>
<caption id="51E1661A8916FFC021CA3C02FDE0F81D" httpUri="https://zenodo.org/record/257542/files/figure.png" pageId="11" pageNumber="12" targetBox="[154,1433,204,1852]" targetPageId="11">
<paragraph id="052136928916FFC021CA3C02FDE0F81D" blockId="11.[151,1436,1878,2023]" pageId="11" pageNumber="12">
<emphasis id="37EAEA808916FFC021CA3C02FEA8F896" bold="true" box="[151,276,1878,1900]" pageId="11" pageNumber="12">FIGURE 4.</emphasis>
SEM illustrations of labial complex, ventral aspect, of
<taxonomicName id="C29E4D118916FFC0220A3C02FC09F896" box="[855,949,1878,1900]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808916FFC0220A3C02FC09F896" box="[855,949,1878,1900]" italics="true" pageId="11" pageNumber="12">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118916FFC0251C3C0CFB7BF896" box="[1089,1223,1878,1901]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808916FFC0251C3C0CFB7BF896" box="[1089,1223,1878,1901]" italics="true" pageId="11" pageNumber="12">Z. phyllobius</emphasis>
</taxonomicName>
; B—
<taxonomicName id="C29E4D118916FFC0245C3C0CFA28F897" box="[1281,1428,1880,1901]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808916FFC0245C3C0CFA28F897" box="[1281,1428,1880,1901]" italics="true" pageId="11" pageNumber="12">Z. punctigerus</emphasis>
</taxonomicName>
; C—
<taxonomicName id="C29E4D118916FFC0219C3C22FE88F871" box="[193,308,1909,1931]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808916FFC0219C3C22FE88F871" box="[193,308,1909,1931]" italics="true" pageId="11" pageNumber="12">Z. pallidus</emphasis>
</taxonomicName>
; D—
<taxonomicName id="C29E4D118916FFC0202F3C22FE47F871" box="[370,507,1909,1931]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808916FFC0202F3C22FE47F871" box="[370,507,1909,1931]" italics="true" pageId="11" pageNumber="12">Z. pellucidus</emphasis>
</taxonomicName>
; E—
<taxonomicName id="C29E4D118916FFC0236A3C22FD17F871" box="[567,683,1909,1931]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808916FFC0236A3C22FD17F871" box="[567,683,1909,1931]" italics="true" pageId="11" pageNumber="12">Z. lescheni</emphasis>
</taxonomicName>
; F—
<taxonomicName id="C29E4D118916FFC023B83C22FCF4F871" box="[741,840,1909,1931]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808916FFC023B83C22FCF4F871" box="[741,840,1909,1931]" italics="true" pageId="11" pageNumber="12">Z. brouni</emphasis>
</taxonomicName>
. Legend: alms—anterolateral mental seta; gsc—glossal sclerite; lms—lateral mental seta; lss—lateral submental seta; m—mentum; mp3—maxillary palpomere 3; mp4—maxillary palpomere 4; mt—mental tooth; mss—mental-submental suture; pms—paramedial mental seta; prss—primary basal submental seta; sm—submentum. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="51E1661A8911FFC721CA3C14FCF2F848" httpUri="https://zenodo.org/record/257543/files/figure.png" pageId="12" pageNumber="13" targetBox="[157,1428,200,1826]" targetPageId="12">
<paragraph id="052136928911FFC721CA3C14FCF2F848" blockId="12.[151,1437,1856,1970]" pageId="12" pageNumber="13">
<emphasis id="37EAEA808911FFC721CA3C14FEA8F8AC" bold="true" box="[151,276,1856,1878]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
SEM illustrations of pronotum, dorsal aspect, of
<taxonomicName id="C29E4D118911FFC722493C14FCCEF8AC" box="[788,882,1856,1878]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808911FFC722493C14FCCEF8AC" box="[788,882,1856,1878]" italics="true" pageId="12" pageNumber="13">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118911FFC722A13C16FB3EF8AC" box="[1020,1154,1856,1879]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808911FFC722A13C16FB3EF8AC" box="[1020,1154,1856,1879]" italics="true" pageId="12" pageNumber="13">Z. phyllobius</emphasis>
</taxonomicName>
; B—
<taxonomicName id="C29E4D118911FFC725E63C16FAF2F8AC" box="[1211,1358,1857,1879]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808911FFC725E63C16FAF2F8AC" box="[1211,1358,1857,1879]" italics="true" pageId="12" pageNumber="13">Z. punctigerus</emphasis>
</taxonomicName>
; C—
<taxonomicName id="C29E4D118911FFC724DA3C16FF55F88F" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808911FFC724DA3C16FF55F88F" italics="true" pageId="12" pageNumber="13">Z. pallidus</emphasis>
</taxonomicName>
; D—
<taxonomicName id="C29E4D118911FFC720793C34FE15F88F" box="[292,425,1887,1909]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808911FFC720793C34FE15F88F" box="[292,425,1887,1909]" italics="true" pageId="12" pageNumber="13">Z. pellucidus</emphasis>
</taxonomicName>
; E—
<taxonomicName id="C29E4D118911FFC720BF3C34FDEEF88F" box="[482,594,1887,1909]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808911FFC720BF3C34FDEEF88F" box="[482,594,1887,1909]" italics="true" pageId="12" pageNumber="13">Z. lescheni</emphasis>
</taxonomicName>
; F—
<taxonomicName id="C29E4D118911FFC723D73C34FD58F88F" box="[650,740,1888,1909]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="37EAEA808911FFC723D73C34FD58F88F" box="[650,740,1888,1909]" italics="true" pageId="12" pageNumber="13">Z. nanus</emphasis>
</taxonomicName>
. Legend: aps—apicolateral pronotal seta; bdt—basilateral pronotal denticle; bs—basilateral pronotal seta; bsn—basilateral pronotal sinuation; ed2—scutellar seta; ls—midlateral pronotal seta; mg—marginal pronotal gutter; sct—scutellum. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="51E1661A8910FFC621CA3C76FAB8F88C" httpUri="https://zenodo.org/record/257544/files/figure.png" pageId="13" pageNumber="14" targetBox="[160,1428,202,1799]" targetPageId="13">
<paragraph id="052136928910FFC621CA3C76FAB8F88C" blockId="13.[151,1437,1826,1910]" pageId="13" pageNumber="14">
<emphasis id="37EAEA808910FFC621CA3C76FEAFF8CD" bold="true" box="[151,275,1826,1848]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
SEM illustrations of prothorax, ventral aspect, of
<taxonomicName id="C29E4D118910FFC6224A3C76FCC9F8C2" box="[791,885,1826,1848]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808910FFC6224A3C76FCC9F8C2" box="[791,885,1826,1848]" italics="true" pageId="13" pageNumber="14">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118910FFC622A33C77FB38F8C2" box="[1022,1156,1826,1848]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808910FFC622A33C77FB38F8C2" box="[1022,1156,1826,1848]" italics="true" pageId="13" pageNumber="14">Z. phyllobius</emphasis>
</taxonomicName>
; B—
<taxonomicName id="C29E4D118910FFC625E13C77FAF2F8C2" box="[1212,1358,1827,1848]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808910FFC625E13C77FAF2F8C2" box="[1212,1358,1827,1848]" italics="true" pageId="13" pageNumber="14">Z. punctigerus</emphasis>
</taxonomicName>
; C—
<taxonomicName id="C29E4D118910FFC624DA3C77FF55F8AD" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808910FFC624DA3C77FF55F8AD" italics="true" pageId="13" pageNumber="14">Z. pallidus</emphasis>
</taxonomicName>
; D—
<taxonomicName id="C29E4D118910FFC6207B3C16FE11F8AD" box="[294,429,1857,1879]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808910FFC6207B3C16FE11F8AD" box="[294,429,1857,1879]" italics="true" pageId="13" pageNumber="14">Z. pellucidus</emphasis>
</taxonomicName>
; E—
<taxonomicName id="C29E4D118910FFC620B53C16FDE7F8AD" box="[488,603,1857,1879]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808910FFC620B53C16FDE7F8AD" box="[488,603,1857,1879]" italics="true" pageId="13" pageNumber="14">Z. lescheni</emphasis>
</taxonomicName>
; F—
<taxonomicName id="C29E4D118910FFC623C93C16FD4AF8AD" box="[660,758,1857,1879]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808910FFC623C93C16FD4AF8AD" box="[660,758,1857,1879]" italics="true" pageId="13" pageNumber="14">Z. brouni</emphasis>
</taxonomicName>
. Legend: pas—prosternal ambulatory seta; pep—proepipleuron; pes—proepisternum; prcx—procoxa; ps—prosternum; psp—prosternal intercoxal process. Scale bars = 0.1mm.
</paragraph>
</caption>
<caption id="51E1661A8913FFC521CA3FE0FCF1FADC" httpUri="https://zenodo.org/record/167655/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1428,193,1181]" targetPageId="14">
<paragraph id="052136928913FFC521CA3FE0FCF1FADC" blockId="14.[151,1437,1204,1318]" pageId="14" pageNumber="15">
<emphasis id="37EAEA808913FFC521CA3FE0FEAEFB33" bold="true" box="[151,274,1204,1226]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
Digital images of pterothorax and first abdominal ventrites, ventral aspect, of
<taxonomicName id="C29E4D118913FFC525763FE0FB35FB30" box="[1067,1161,1204,1226]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808913FFC525763FE0FB35FB30" box="[1067,1161,1204,1226]" italics="true" pageId="14" pageNumber="15">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118913FFC5244C3FE1FA28FB30" box="[1297,1428,1204,1226]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808913FFC5244C3FE1FA28FB30" box="[1297,1428,1204,1226]" italics="true" pageId="14" pageNumber="15">Z. phyllobius</emphasis>
</taxonomicName>
; B—
<taxonomicName id="C29E4D118913FFC5219C3F87FEE9FB12" box="[193,341,1235,1256]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808913FFC5219C3F87FEE9FB12" box="[193,341,1235,1256]" italics="true" pageId="14" pageNumber="15">Z. punctigerus</emphasis>
</taxonomicName>
; C—
<taxonomicName id="C29E4D118913FFC520D33F87FE58FB12" box="[398,484,1235,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA808913FFC520D33F87FE58FB12" box="[398,484,1235,1256]" italics="true" pageId="14" pageNumber="15">Z. nunni</emphasis>
</taxonomicName>
; D—
<taxonomicName id="C29E4D118913FFC523423F87FD32FB12" box="[543,654,1234,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808913FFC523423F87FD32FB12" box="[543,654,1234,1256]" italics="true" pageId="14" pageNumber="15">Z. pallidus</emphasis>
</taxonomicName>
; E—
<taxonomicName id="C29E4D118913FFC5239B3F87FCF6FB12" box="[710,842,1234,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808913FFC5239B3F87FCF6FB12" box="[710,842,1234,1256]" italics="true" pageId="14" pageNumber="15">Z. pellucidus</emphasis>
</taxonomicName>
; F—
<taxonomicName id="C29E4D118913FFC522DC3F87FC63FB12" box="[897,991,1234,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808913FFC522DC3F87FC63FB12" box="[897,991,1234,1256]" italics="true" pageId="14" pageNumber="15">Z. brouni</emphasis>
</taxonomicName>
; G—
<taxonomicName id="C29E4D118913FFC525463F87FB36FB12" box="[1051,1162,1234,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808913FFC525463F87FB36FB12" box="[1051,1162,1234,1256]" italics="true" pageId="14" pageNumber="15">Z. lescheni</emphasis>
</taxonomicName>
; H—
<taxonomicName id="C29E4D118913FFC525983F87FAEBFB12" box="[1221,1367,1235,1256]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808913FFC525983F87FAEBFB12" box="[1221,1367,1235,1256]" italics="true" pageId="14" pageNumber="15">Z. montivagus</emphasis>
</taxonomicName>
; I—
<taxonomicName id="C29E4D118913FFC524DA3F87FF6FFAFD" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="37EAEA808913FFC524DA3F87FF6FFAFD" italics="true" pageId="14" pageNumber="15">Z. nanus</emphasis>
</taxonomicName>
. Legend: ipa—intercoxal process of abdominal ventrite 2; mes—metendosternite; mte—metanepisternum; mtvmetaventrite; v1v2—abdominal ventrites 12. Scale bars = 0.2mm.
</paragraph>
</caption>
<paragraph id="052136928913FFC5219A3E06FA28FA49" blockId="14.[151,1437,1362,2035]" pageId="14" pageNumber="15">
<emphasis id="37EAEA808913FFC5219A3E06FEE1FA91" bold="true" box="[199,349,1362,1387]" pageId="14" pageNumber="15">Recognition.</emphasis>
Adults of this species (
<figureCitation id="9DA52A178913FFC5233E3E06FD0EFA91" box="[611,690,1362,1387]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
B) are practically indistinguishable externally from the adults of
<taxonomicName id="C29E4D118913FFC524D83E07FEB0FA74" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808913FFC524D83E07FEB0FA74" italics="true" pageId="14" pageNumber="15">Z. phyllobius</emphasis>
</taxonomicName>
and
<taxonomicName id="C29E4D118913FFC5201A3E23FE1BFA74" box="[327,423,1399,1422]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA808913FFC5201A3E23FE1BFA74" box="[327,423,1399,1422]" italics="true" pageId="14" pageNumber="15">Z. nunni</emphasis>
</taxonomicName>
, and are distinguished from the males of those by the structure of male genitalia. From other congeners, adults of
<taxonomicName id="C29E4D118913FFC520E23ECFFDDCFA48" box="[447,608,1435,1458]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808913FFC520E23ECFFDDCFA48" box="[447,608,1435,1458]" italics="true" pageId="14" pageNumber="15">Z. punctigerus</emphasis>
</taxonomicName>
can be distinguished by the characters listed in the subgeneric diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="4D8465198913FFC4219A3EEAFC4AFE7D" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="052136928913FFC5219A3EEAFAF4FA2D" blockId="14.[151,1437,1362,2035]" box="[199,1352,1470,1495]" pageId="14" pageNumber="15">
<emphasis id="37EAEA808913FFC5219A3EEAFEE5FA2D" bold="true" box="[199,345,1470,1495]" pageId="14" pageNumber="15">Description.</emphasis>
Size. Medium to large for genus (SBL range
<quantity id="C2669B778913FFC5220A3EEAFC40FA2C" box="[855,1020,1470,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" metricValueMax="1.94" metricValueMin="1.64" pageId="14" pageNumber="15" unit="mm" value="1.79" valueMax="1.94" valueMin="1.64">1.641.94 mm</quantity>
, mean 1.80±
<quantity id="C2669B778913FFC525D73EEBFABCFA2C" box="[1162,1280,1471,1495]" metricMagnitude="-4" metricUnit="m" metricValue="1.07" pageId="14" pageNumber="15" unit="mm" value="0.107">0.107 mm</quantity>
, n=5).
</paragraph>
<paragraph id="052136928913FFC5219A3EB6FE4DF9B9" blockId="14.[151,1437,1362,2035]" pageId="14" pageNumber="15">Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.35±0.010), head of moderate width for genus compared to pronotum (WH/WPm 0.76±0.016), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.012).</paragraph>
<paragraph id="052136928913FFC5219A3D1AFCF7F99C" blockId="14.[151,1437,1362,2035]" box="[199,843,1614,1639]" pageId="14" pageNumber="15">Color. Body color rufotestaceous, appendages testaceous.</paragraph>
<paragraph id="052136928913FFC5219A3D26FC20F929" blockId="14.[151,1437,1362,2035]" pageId="14" pageNumber="15">Prothorax. Pronotum moderately long (LP/LE 0.44±0.029) and moderately elongate (WPm/LP 1.29±0.032), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.70±0.118). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.33±0.090).</paragraph>
<paragraph id="052136928913FFC5219A3D8AFE86F8C5" blockId="14.[151,1437,1362,2035]" pageId="14" pageNumber="15">Elytra. Slightly depressed along suture, comparatively short (LE/SBL 0.47±0.010) and moderately wide (WE/ LE 0.74±0.028). Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.</paragraph>
<paragraph id="052136928913FFC5219A3C1EFD5DF835" blockId="14.[151,1437,1362,2035]" pageId="14" pageNumber="15">
Male genitalia. Median lobe (
<figureCitation id="9DA52A178913FFC5234D3C1EFDE3F899" box="[528,607,1866,1891]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
A) with straight ventral margin and semicircular short apex. Apical orifice area with a pair of long ostial fields. Sclerites of internal sac united basally, long, slightly shorter than length between apical and basal orifices. Left paramere with apical constriction (
<figureCitation id="9DA52A178913FFC5228C3CC6FB9FF851" box="[977,1059,1938,1963]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
C). Ring sclerite with long handle of approximately equal length and width (
<figureCitation id="9DA52A178913FFC523303CE2FD03F835" box="[621,703,1974,1999]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="14" pageNumber="15">Fig. 14</figureCitation>
B).
</paragraph>
<paragraph id="052136928913FFC4219A3C8EFD92FF2F" blockId="14.[151,1437,1362,2035]" lastBlockId="15.[151,1436,151,392]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Female internal genitalia. Goncoxite 2 with short ensiform setae (
<figureCitation id="9DA52A178913FFC522FD3C8EFC4CF809" box="[928,1008,2010,2035]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="14" pageNumber="15">Fig. 15</figureCitation>
B). Spermatheca sclerotized, elongate, with three compartments (
<figureCitation id="9DA52A178912FFC420E63BC3FDB0FF4A" box="[443,524,151,176]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="15" pageNumber="16">Fig. 16</figureCitation>
B). Spermathecal duct short with few coils. Attachments of spermathecal duct and gland to spermatheca close together.
</paragraph>
<paragraph id="052136928912FFC4219A3B8BFCCEFEE7" blockId="15.[151,1436,151,392]" pageId="15" pageNumber="16">
<materialsCitation id="B5F63CCF8912FFC4219A3B8BFCCEFEE7" ID-GBIF-Occurrence="1341350625" location="Canterbury" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Canterbury">
<emphasis id="37EAEA808912FFC4219A3B8BFDB8FF02" bold="true" box="[199,516,223,248]" pageId="15" pageNumber="16">Geographical distribution.</emphasis>
This species is known from the eastern ranges of
<collectingRegion id="C75AF8708912FFC425733B8BFB10FF02" box="[1070,1196,223,248]" country="New Zealand" name="Canterbury" pageId="15" pageNumber="16">Canterbury</collectingRegion>
lying in basins of the Rakaia, Ashburton and Rangitātā rivers (
<figureCitation id="9DA52A178912FFC4233C3A50FD0DFEE6" box="[609,689,260,285]" captionStart="FIGURE 18" captionStartId="30.[151,250,1918,1940]" captionTargetBox="[198,1387,199,1885]" captionTargetId="figure@30.[189,1398,193,1897]" captionTargetPageId="30" captionText="FIGURE 18. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Zeanillus. A: white circles, Z. phyllobius; green circles, Z. punctigerus; red circles, Z. nunni; blue diamond, Z. pellucidus; green diamond, Z. pallidus. B: white triangles, Z. brouni; green triangles, Z. nanus; red triangles, Z. lescheni; yellow triangle, Z. carltoni; black triangle, Z. montivagus. Elevation scale bars are given in meters." httpUri="https://zenodo.org/record/167666/files/figure.png" pageId="15" pageNumber="16">Fig. 18</figureCitation>
A, green circles).
</materialsCitation>
</paragraph>
<paragraph id="052136928912FFC4219A3A73FCC2FEBA" blockId="15.[151,1436,151,392]" box="[199,894,295,320]" pageId="15" pageNumber="16">
<emphasis id="37EAEA808912FFC4219A3A73FEF3FEBA" bold="true" box="[199,335,295,320]" pageId="15" pageNumber="16">Way of life.</emphasis>
Specimens were collected in the broadleaf forest.
</paragraph>
<paragraph id="052136928912FFC4219A3A18FC4AFE7D" blockId="15.[151,1436,151,392]" pageId="15" pageNumber="16">
<emphasis id="37EAEA808912FFC4219A3A18FECCFE9F" bold="true" box="[199,368,332,357]" pageId="15" pageNumber="16">Relationships.</emphasis>
The structure of male genitalia and spermatheca suggests that
<taxonomicName id="C29E4D118912FFC4256D3A19FB6EFE9E" box="[1072,1234,333,356]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808912FFC4256D3A19FB6EFE9E" box="[1072,1234,333,356]" italics="true" pageId="15" pageNumber="16">Z. punctigerus</emphasis>
</taxonomicName>
and the allopatric
<taxonomicName id="C29E4D118912FFC421CA3A25FE95FE72" box="[151,297,368,392]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808912FFC421CA3A25FE95FE72" box="[151,297,368,392]" italics="true" pageId="15" pageNumber="16">Z. phyllobius</emphasis>
</taxonomicName>
from the coastal hills of the Banks Peninsula are the sister taxa.
</paragraph>
</subSubSection>
</treatment>
</document>