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<document id="AAB9812E41E793E207F7F497EFDC3885" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D378784890AFFD121CA3E50FCEFFE0E" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus brouni Sokolov, 2016, sp.n." docType="treatment" docUuid="3C5726CA-DB0E-4EC7-BE61-8641D2FA182F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="27" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="24" updateTime="1698393955637" updateUser="plazi">
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<mods:title id="8DD1A1B8BF1C84EEAF25F1CE3EF48C09">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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<mods:namePart id="E817A3EEF0EEC00379095F859BF54005">Sokolov, Igor M.</mods:namePart>
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<mods:title id="FED3EEA07D0E3E5F7E137D6367A8A362">Zootaxa</mods:title>
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<mods:part id="FD5784DB02ED80B49EC333B1911C6B04">
<mods:date id="FFE4D83474397608EC13A45CE4B2306C">2016</mods:date>
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<treatment id="8D378784890AFFD121CA3E50FCEFFE0E" ID-DOI="http://doi.org/10.5281/zenodo.6089682" ID-GBIF-Taxon="124814320" ID-Zenodo-Dep="6089682" LSID="urn:lsid:zoobank.org:act:3C5726CA-DB0E-4EC7-BE61-8641D2FA182F" httpUri="http://treatment.plazi.org/id/8D378784890AFFD121CA3E50FCEFFE0E" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
<subSubSection id="4D846519890AFFDC21CA3E50FC9EFAC5" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="05213692890AFFDC21CA3E50FE17FAE4" blockId="23.[151,802,1284,1343]" box="[151,427,1284,1310]" pageId="23" pageNumber="24">
<heading id="5E6981FE890AFFDC21CA3E50FE17FAE4" bold="true" box="[151,427,1284,1310]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="37EAEA80890AFFDC21CA3E50FE17FAE4" bold="true" box="[151,427,1284,1310]" pageId="23" pageNumber="24">
<emphasis id="37EAEA80890AFFDC21CA3E50FED5FAE4" bold="true" box="[151,361,1284,1310]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="C29E4D11890AFFDC21CA3E50FEDFFAE4" box="[151,355,1284,1310]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="brouni" status="sp. nov.">Zeanillus brouni</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="2CD957FB890AFFDC202D3E50FE17FAE4" box="[368,427,1284,1310]" pageId="23" pageNumber="24" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="05213692890AFFDC21CA3E72FC9EFAC5" blockId="23.[151,802,1284,1343]" box="[151,802,1318,1343]" pageId="23" pageNumber="24">
<figureCitation id="9DA52A17890AFFDC21CA3E72FF5FFAC5" box="[151,227,1318,1343]" captionStart="FIGURE 4" captionStartId="11.[151,250,1878,1900]" captionTargetBox="[154,1433,204,1852]" captionTargetId="figure@11.[151,1435,193,1857]" captionTargetPageId="11" captionText="FIGURE 4. SEM illustrations of labial complex, ventral aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. pallidus; D— Z. pellucidus; E— Z. lescheni; F— Z. brouni. Legend: alms—anterolateral mental seta; gsc—glossal sclerite; lms—lateral mental seta; lss—lateral submental seta; m—mentum; mp 3 — maxillary palpomere 3; mp 4 — maxillary palpomere 4; mt—mental tooth; mss—mental-submental suture; pms—paramedial mental seta; prss—primary basal submental seta; sm—submentum. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/257542/files/figure.png" pageId="23" pageNumber="24">Figs. 4</figureCitation>
F, 6F, 7F, 10EKQ, 11F, 13QT, 14H, 15F, 16F, 18B
</paragraph>
</subSubSection>
<subSubSection id="4D846519890AFFDC21CA3E3AFA21F95D" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="05213692890AFFDC21CA3E3AFD54FA51" blockId="23.[151,1437,1390,2027]" pageId="23" pageNumber="24">
<materialsCitation id="B5F63CCF890AFFDC21CA3E3AFD54FA51" ID-GBIF-Occurrence="1341350588" collectingDate="1919-01" collectionCode="NZAC" collectorName="DN Glen Warren Res Oamaru" country="New Zealand" location="New Zealand" pageId="23" pageNumber="24" specimenCount="2" typeStatus="holotype">
<emphasis id="37EAEA80890AFFDC21CA3E3AFEF5FA7D" bold="true" box="[151,329,1390,1415]" pageId="23" pageNumber="24">
<typeStatus id="DA258830890AFFDC21CA3E3AFF6EFA7D" box="[151,210,1390,1415]" pageId="23" pageNumber="24">Type</typeStatus>
material.
</emphasis>
<typeStatus id="DA258830890AFFDC20093E3BFE57FA7D" box="[340,491,1391,1415]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
, male, in
<collectionCode id="638FAE57890AFFDC23393E3BFD0FFA7D" box="[612,691,1391,1415]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="23" pageNumber="24">NZAC</collectionCode>
, board-mounted, undissected, labeled: /
<collectingCountry id="7D897602890AFFDC25D13E3BFA9AFA7D" box="[1164,1318,1390,1415]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
<collectorName id="A86B5344890AFFDC246F3E3BFEC6FA51" pageId="23" pageNumber="24">DN Glen Warren Res Oamaru</collectorName>
<collectingDate id="6164E9BA890AFFDC20DF3EC7FE76FA50" box="[386,458,1427,1451]" pageId="23" pageNumber="24" value="1919-01">
<date id="71201052890AFFDC20DF3EC7FE76FA50" box="[386,458,1427,1451]" pageId="23" pageNumber="24" value="1919-01">19 Jan</date>
</collectingDate>
0 7 / Washed soil sample.
</materialsCitation>
</paragraph>
<paragraph id="05213692890AFFDC219A3EE3FA21F95D" blockId="23.[151,1437,1390,2027]" pageId="23" pageNumber="24">
PARATYPES (55 specimens, 7 dissected), 8 males and 3 females labeled same as holotype;
<materialsCitation id="B5F63CCF890AFFDC245F3EE3FD48F9ED" ID-GBIF-Occurrence="1341350600" collectingDate="2008-01-12" collectorName="J. Nunn" country="New Zealand" elevation="150" location="Glen Warren Reserve" municipality="Oamaru" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="1398FD1B890AFFDC245F3EE3FAECFA35" box="[1282,1360,1462,1487]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
and
<specimenCount id="1398FD1B890AFFDC24D33EE3FF5FFA09" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
labeled: /
<collectingCountry id="7D897602890AFFDC20343E8FFDB5FA09" box="[361,521,1498,1523]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
: DN,
<location id="00416049890AFFDC23083E8EFCEFFA08" LSID="urn:lsid:plazi:treatment:8D378784890AFFD121CA3E50FCEFFE0E:00416049890AFFDC23083E8EFCEFFA08" box="[597,851,1498,1523]" country="New Zealand" municipality="Oamaru" name="Glen Warren Reserve" pageId="23" pageNumber="24">Glen Warren Reserve</location>
,
<collectingMunicipality id="E545ACE8890AFFDC223E3E8FFC7CFA09" box="[867,960,1499,1523]" pageId="23" pageNumber="24">Oamaru</collectingMunicipality>
,
<elevation id="8EB3D1A1890AFFDC228C3E8EFBA8FA09" box="[977,1044,1498,1523]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="m" value="150.0">
<quantity id="C2669B77890AFFDC228C3E8EFBA8FA09" box="[977,1044,1498,1523]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="m" value="150.0">150m</quantity>
</elevation>
, washed soil sample, dry coastal broadleaf forest, H45/864483
<collectorName id="A86B5344890AFFDC20A93EABFDF8F9EC" box="[500,580,1535,1558]" pageId="23" pageNumber="24">J.Nunn</collectorName>
<collectingDate id="6164E9BA890AFFDC23123EABFD62F9ED" box="[591,734,1535,1559]" pageId="23" pageNumber="24" value="2008-01-12">
<date id="71201052890AFFDC23123EABFD62F9ED" box="[591,734,1535,1559]" pageId="23" pageNumber="24" value="2008-01-12">12 Jan 2008</date>
</collectingDate>
/
</materialsCitation>
;
<materialsCitation id="B5F63CCF890AFFDC22593EABFC3FF9C1" ID-GBIF-Occurrence="1341350601" collectingDate="4 Jun" collectorName="DN Glen Warren Res Oamaru" country="New Zealand" location="New Zealand" pageId="23" pageNumber="24" specimenCount="13" specimenCount-female="4" specimenCount-male="9">
<specimenCount id="1398FD1B890AFFDC22593EABFCE2F9ED" box="[772,862,1534,1559]" pageId="23" pageNumber="24" type="male">9 males</specimenCount>
and
<specimenCount id="1398FD1B890AFFDC22C13EABFBB0F9ED" box="[924,1036,1534,1559]" pageId="23" pageNumber="24" type="female">4 females</specimenCount>
labeled: /
<collectingCountry id="7D897602890AFFDC25D03EABFA9BF9ED" box="[1165,1319,1534,1559]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
<collectorName id="A86B5344890AFFDC246E3EABFEC5F9C1" pageId="23" pageNumber="24">DN Glen Warren Res Oamaru</collectorName>
<collectingDate id="6164E9BA890AFFDC20DD3D77FE07F9C0" box="[384,443,1571,1595]" pageId="23" pageNumber="24">
<date id="71201052890AFFDC20DD3D77FE07F9C0" box="[384,443,1571,1595]" pageId="23" pageNumber="24">4 Jun</date>
</collectingDate>
0 6 / In soil sample from forest remnant /
</materialsCitation>
;
<materialsCitation id="B5F63CCF890AFFDC22D23D77FBBBF9A5" ID-GBIF-Occurrence="1341350615" collectingDate="2 Aug" collectorName="DN Roy Cooper Res Oamaru" country="New Zealand" location="New Zealand" pageId="23" pageNumber="24" specimenCount="26" specimenCount-female="8" specimenCount-male="18">
<specimenCount id="1398FD1B890AFFDC22D23D77FC4EF9C1" box="[911,1010,1570,1595]" pageId="23" pageNumber="24" type="male">18 males</specimenCount>
and
<specimenCount id="1398FD1B890AFFDC25753D77FB2FF9C1" box="[1064,1171,1570,1595]" pageId="23" pageNumber="24" type="female">8 females</specimenCount>
labeled:
<emphasis id="37EAEA80890AFFDC25A53D76FABCF9C1" bold="true" box="[1272,1280,1570,1595]" pageId="23" pageNumber="24">/</emphasis>
<collectingCountry id="7D897602890AFFDC245B3D77FA20F9C1" box="[1286,1436,1570,1595]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
<collectorName id="A86B5344890AFFDC21CA3D13FE63F9A5" box="[151,479,1607,1631]" pageId="23" pageNumber="24">DN Roy Cooper Res Oamaru</collectorName>
<collectingDate id="6164E9BA890AFFDC20BB3D13FD96F9A5" box="[486,554,1607,1631]" pageId="23" pageNumber="24">
<date id="71201052890AFFDC20BB3D13FD96F9A5" box="[486,554,1607,1631]" pageId="23" pageNumber="24">2 Aug</date>
</collectingDate>
0 7 / Washed soil sample broadleaf forest /
</materialsCitation>
;
<materialsCitation id="B5F63CCF890AFFDC254F3D13FC50F979" ID-GBIF-Occurrence="1341350633" collectingDate="1923-08" collectorName="DN Roy Cooper Res Oamaru" country="New Zealand" location="New Zealand" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="1398FD1B890AFFDC254F3D13FBE0F9A5" box="[1042,1116,1606,1631]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
and
<specimenCount id="1398FD1B890AFFDC25CF3D13FB4FF9A5" box="[1170,1267,1606,1631]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
labeled: /
<collectingCountry id="7D897602890AFFDC243A3D13FF4DF979" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
<collectorName id="A86B5344890AFFDC21A73D3FFDF0F979" box="[250,588,1643,1667]" pageId="23" pageNumber="24">DN Roy Cooper Res Oamaru</collectorName>
<collectingDate id="6164E9BA890AFFDC230B3D3FFD17F979" box="[598,683,1643,1667]" pageId="23" pageNumber="24" value="1923-08">
<date id="71201052890AFFDC230B3D3FFD17F979" box="[598,683,1643,1667]" pageId="23" pageNumber="24" value="1923-08">23 Aug</date>
</collectingDate>
0 2 / In earthy forest litter /
</materialsCitation>
;
<materialsCitation id="B5F63CCF890AFFDC22A73D3FFA21F95D" ID-GBIF-Occurrence="1341350628" collectingDate="2007-12-02" collectorName="J. Nunn" country="New Zealand" elevation="150" location="Allison Reserve" municipality="Akatore" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1">
<specimenCount id="1398FD1B890AFFDC22A73D3FFBFBF979" box="[1018,1095,1642,1667]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
labeled: /
<collectingCountry id="7D897602890AFFDC259E3D3FFADDF979" box="[1219,1377,1642,1667]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
: DN,
<location id="00416049890AFFDC21CA3DDAFEF1F95C" LSID="urn:lsid:plazi:treatment:8D378784890AFFD121CA3E50FCEFFE0E:00416049890AFFDC21CA3DDAFEF1F95C" box="[151,333,1678,1703]" country="New Zealand" municipality="Akatore" name="Allison Reserve" pageId="23" pageNumber="24">Allison Reserve</location>
,
<collectingMunicipality id="E545ACE8890AFFDC200A3DDAFE08F95D" box="[343,436,1678,1703]" pageId="23" pageNumber="24">Akatore</collectingMunicipality>
,
<elevation id="8EB3D1A1890AFFDC20E23DDAFDBEF95D" box="[447,514,1678,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="m" value="150.0">
<quantity id="C2669B77890AFFDC20E23DDAFDBEF95D" box="[447,514,1678,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="m" value="150.0">150m</quantity>
</elevation>
, washed soil sample, broadleaf /podocarp forest, M31/602701
<collectorName id="A86B5344890AFFDC25E33DDBFAB3F95C" box="[1214,1295,1679,1702]" pageId="23" pageNumber="24">J.Nunn</collectorName>
<collectingDate id="6164E9BA890AFFDC24483DDBFA25F95D" box="[1301,1433,1679,1703]" pageId="23" pageNumber="24" value="2007-12-02">
<date id="71201052890AFFDC24483DDBFA25F95D" box="[1301,1433,1679,1703]" pageId="23" pageNumber="24" value="2007-12-02">2 Dec 2007</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4D846519890AFFDC219A3DE6FB77F915" pageId="23" pageNumber="24" type="description">
<paragraph id="05213692890AFFDC219A3DE6FB77F915" blockId="23.[151,1437,1390,2027]" pageId="23" pageNumber="24">
<emphasis id="37EAEA80890AFFDC219A3DE6FE38F931" bold="true" box="[199,388,1714,1739]" pageId="23" pageNumber="24">Specific epithet.</emphasis>
The specific epithet honors the surname of Thomas Broun, famous
<collectingCountry id="7D897602890AFFDC252A3DE7FAB0F931" box="[1143,1292,1714,1739]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
coleopterist, the author of thousands of new species, including the first descriptions of
<collectingCountry id="7D897602890AFFDC228C3D83FBDBF915" box="[977,1127,1750,1775]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
Anillina
</paragraph>
</subSubSection>
<subSubSection id="4D846519890AFFDC219A3DAEFEDFF8A1" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="05213692890AFFDC219A3DAEFBE2F8E8" blockId="23.[151,1437,1390,2027]" box="[199,1118,1786,1811]" pageId="23" pageNumber="24">
<materialsCitation id="B5F63CCF890AFFDC219A3DAEFBE2F8E8" ID-GBIF-Occurrence="1341350631" box="[199,1118,1786,1811]" country="New Zealand" location="Oamaru" municipality="South Island" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
<emphasis id="37EAEA80890AFFDC219A3DAEFED9F8E9" bold="true" box="[199,357,1786,1811]" pageId="23" pageNumber="24">
<typeStatus id="DA258830890AFFDC219A3DAEFEBEF8E9" box="[199,258,1786,1811]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="7D897602890AFFDC20313DAFFDB9F8E9" box="[364,517,1786,1811]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
,
<collectingMunicipality id="E545ACE8890AFFDC234D3DAEFD23F8E9" box="[528,671,1786,1811]" pageId="23" pageNumber="24">South Island</collectingMunicipality>
,
<collectingRegion id="C75AF870890AFFDC23F63DAFFD4EF8E9" box="[683,754,1787,1811]" country="New Zealand" name="Otago" pageId="23" pageNumber="24">Otago</collectingRegion>
,
<location id="00416049890AFFDC23A03DAFFCE6F8E9" LSID="urn:lsid:plazi:treatment:8D378784890AFFD121CA3E50FCEFFE0E:00416049890AFFDC23A03DAFFCE6F8E9" box="[765,858,1787,1811]" country="New Zealand" municipality="South Island" name="Oamaru" pageId="23" pageNumber="24" stateProvince="Otago">Oamaru</location>
, Glen Warren Reserve.
</materialsCitation>
</paragraph>
<paragraph id="05213692890AFFDC219A3C4AFEDFF8A1" blockId="23.[151,1437,1390,2027]" pageId="23" pageNumber="24">
<emphasis id="37EAEA80890AFFDC219A3C4AFEE1F8CD" bold="true" box="[199,349,1822,1847]" pageId="23" pageNumber="24">Recognition.</emphasis>
Adults of this species (
<figureCitation id="9DA52A17890AFFDC233A3C4AFD0AF8CD" box="[615,694,1822,1847]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
F) are distinguished from other congeners by the characters listed in the species key.
</paragraph>
</subSubSection>
<subSubSection id="4D846519890AFFD1219A3C32FCEFFE0E" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24" type="description">
<paragraph id="05213692890AFFDC219A3C32FAEAF885" blockId="23.[151,1437,1390,2027]" box="[199,1366,1894,1919]" pageId="23" pageNumber="24">
<emphasis id="37EAEA80890AFFDC219A3C32FEE5F885" bold="true" box="[199,345,1894,1919]" pageId="23" pageNumber="24">Description.</emphasis>
Size. Medium to large for genus (SBL range
<quantity id="C2669B77890AFFDC220A3C32FC40F884" box="[855,1020,1894,1919]" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.95" metricValueMin="1.73" pageId="23" pageNumber="24" unit="mm" value="1.8399999999999999" valueMax="1.95" valueMin="1.73">1.731.95 mm</quantity>
, mean 1.81±
<quantity id="C2669B77890AFFDC25D43C32FABCF884" box="[1161,1280,1894,1919]" metricMagnitude="-5" metricUnit="m" metricValue="5.4" pageId="23" pageNumber="24" unit="mm" value="0.054">0.054 mm</quantity>
, n=20).
</paragraph>
<paragraph id="05213692890AFFDC219A3CDEFE17F811" blockId="23.[151,1437,1390,2027]" pageId="23" pageNumber="24">Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.36±0.011), head of moderate width for genus compared to pronotum (WH/WPm 0.72±0.015), pronotum rather narrow in comparison to elytra (WPm/WE 0.72±0.018).</paragraph>
<caption id="51E1661A8905FFD321CA3D4AFB2AF934" httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="24" pageNumber="25" targetBox="[159,1415,184,1535]" targetPageId="24">
<paragraph id="052136928905FFD321CA3D4AFB2AF934" blockId="24.[151,1436,1566,1742]" pageId="24" pageNumber="25">
<emphasis id="37EAEA808905FFD321CA3D4AFEA3F9CE" bold="true" box="[151,287,1566,1588]" pageId="24" pageNumber="25">FIGURE 15.</emphasis>
Line drawings of right ovipositor sclerite, lateral aspect, of
<taxonomicName id="C29E4D118905FFD322233D4AFC60F9CE" box="[894,988,1566,1588]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808905FFD322233D4AFC60F9CE" box="[894,988,1566,1588]" italics="true" pageId="24" pageNumber="25">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118905FFD325393D74FB55F9CE" box="[1124,1257,1566,1589]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808905FFD325393D74FB55F9CE" box="[1124,1257,1566,1589]" italics="true" pageId="24" pageNumber="25">Z. phyllobius</emphasis>
</taxonomicName>
(NZ, Canterbury, Ahuriri Scenic Reserve); B—
<taxonomicName id="C29E4D118905FFD320993D6AFDEAF9A9" box="[452,598,1598,1619]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808905FFD320993D6AFDEAF9A9" box="[452,598,1598,1619]" italics="true" pageId="24" pageNumber="25">Z. punctigerus</emphasis>
</taxonomicName>
(NZ, Canterbury, Sharplin Falls); C—
<taxonomicName id="C29E4D118905FFD322BD3D6AFB88F9A9" box="[992,1076,1598,1619]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA808905FFD322BD3D6AFB88F9A9" box="[992,1076,1598,1619]" italics="true" pageId="24" pageNumber="25">Z. nunni</emphasis>
</taxonomicName>
(NZ, Otago, Woodside Glen); D—
<taxonomicName id="C29E4D118905FFD321CA3D09FEBBF988" box="[151,263,1628,1650]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808905FFD321CA3D09FEBBF988" box="[151,263,1628,1650]" italics="true" pageId="24" pageNumber="25">Z. pallidus</emphasis>
</taxonomicName>
(NZ, Otago, Taieri Mouth); E—
<taxonomicName id="C29E4D118905FFD323393D09FD56F988" box="[612,746,1628,1650]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808905FFD323393D09FD56F988" box="[612,746,1628,1650]" italics="true" pageId="24" pageNumber="25">Z. pellucidus</emphasis>
</taxonomicName>
(NZ, Otago, Oamaru, Glen Warren Reserve); F—
<taxonomicName id="C29E4D118905FFD3245D3D09FADDF988" box="[1280,1377,1628,1650]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808905FFD3245D3D09FADDF988" box="[1280,1377,1628,1650]" italics="true" pageId="24" pageNumber="25">Z. brouni</emphasis>
</taxonomicName>
(NZ, Otago, Oamaru, Glen Warren Reserve); G—
<taxonomicName id="C29E4D118905FFD323363D28FD65F96B" box="[619,729,1659,1681]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="37EAEA808905FFD323363D28FD65F96B" box="[619,729,1659,1681]" italics="true" pageId="24" pageNumber="25">Z. carltoni</emphasis>
</taxonomicName>
(NZ, Otago, Mount Watkin); H—
<taxonomicName id="C29E4D118905FFD3251B3D28FB0BF96B" box="[1094,1207,1659,1681]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808905FFD3251B3D28FB0BF96B" box="[1094,1207,1659,1681]" italics="true" pageId="24" pageNumber="25">Z. lescheni</emphasis>
</taxonomicName>
(NZ, Otago, Waikaia Forest); I—
<taxonomicName id="C29E4D118905FFD320533DCEFE23F955" box="[270,415,1690,1711]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808905FFD320533DCEFE23F955" box="[270,415,1690,1711]" italics="true" pageId="24" pageNumber="25">Z. montivagus</emphasis>
</taxonomicName>
(NZ, Otago, North Rough Ridge); J—
<taxonomicName id="C29E4D118905FFD322763DCEFC38F955" box="[811,900,1690,1711]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="37EAEA808905FFD322763DCEFC38F955" box="[811,900,1690,1711]" italics="true" pageId="24" pageNumber="25">Z. nanus</emphasis>
</taxonomicName>
(NZ, Otago, Balclutha). Legend: es—ensiform seta; gc1—gonocoxite 1; gc2—gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05mm.
</paragraph>
</caption>
<paragraph id="052136928905FFD3219A3DA1FCF7F8F7" blockId="24.[151,1437,1781,2022]" box="[199,843,1781,1806]" pageId="24" pageNumber="25">Color. Body color rufotestaceous, appendages testaceous.</paragraph>
<paragraph id="052136928905FFD3219A3C4DFC60F880" blockId="24.[151,1437,1781,2022]" pageId="24" pageNumber="25">Prothorax. Pronotum slightly long (LP/LE 0.41±0.008) and moderately elongate (WPm/LP 1.25±0.026), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.56±0.068). Width between anterior angles moderately greater than between posterior angles (WPa/WPp 1.16±0.058).</paragraph>
<paragraph id="052136928905FFD3219A3CD1FE86F81C" blockId="24.[151,1437,1781,2022]" pageId="24" pageNumber="25">Elytra. Slightly depressed along suture, comparatively long (LE/SBL 0.52±0.013) and moderately wide (WE/ LE 0.70±0.025). Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.</paragraph>
<caption id="51E1661A8904FFD221CA3D14FAD7F92B" httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="25" pageNumber="26" targetBox="[171,1418,211,1571]" targetPageId="25">
<paragraph id="052136928904FFD221CA3D14FAD7F92B" blockId="25.[151,1436,1600,1745]" pageId="25" pageNumber="26">
<emphasis id="37EAEA808904FFD221CA3D14FEA4F9AF" bold="true" box="[151,280,1600,1622]" pageId="25" pageNumber="26">FIGURE 16</emphasis>
. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of
<taxonomicName id="C29E4D118904FFD2256E3D14FB2DF9AC" box="[1075,1169,1600,1622]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="37EAEA808904FFD2256E3D14FB2DF9AC" box="[1075,1169,1600,1622]" italics="true" pageId="25" pageNumber="26">Zeanillus</emphasis>
</taxonomicName>
species. A—
<taxonomicName id="C29E4D118904FFD224453D15FA20F9AC" box="[1304,1436,1600,1622]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="phyllobius">
<emphasis id="37EAEA808904FFD224453D15FA20F9AC" box="[1304,1436,1600,1622]" italics="true" pageId="25" pageNumber="26">Z. phyllobius</emphasis>
</taxonomicName>
(NZ, Canterbury, Ahuriri Scenic Reserve); B—
<taxonomicName id="C29E4D118904FFD223D73D34FC9CF98F" box="[650,800,1632,1653]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="37EAEA808904FFD223D73D34FC9CF98F" box="[650,800,1632,1653]" italics="true" pageId="25" pageNumber="26">Z. punctigerus</emphasis>
</taxonomicName>
(NZ, Canterbury, Sharplin Falls); C—
<taxonomicName id="C29E4D118904FFD225E63D34FAAEF98F" box="[1211,1298,1632,1653]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="37EAEA808904FFD225E63D34FAAEF98F" box="[1211,1298,1632,1653]" italics="true" pageId="25" pageNumber="26">Z. nunni</emphasis>
</taxonomicName>
(NZ, Otago, Woodside Glen); D—
<taxonomicName id="C29E4D118904FFD220253D2BFE41F969" box="[376,509,1661,1684]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="37EAEA808904FFD220253D2BFE41F969" box="[376,509,1661,1684]" italics="true" pageId="25" pageNumber="26">Z. pellucidus</emphasis>
</taxonomicName>
(NZ, Otago, Oamaru, Glen Warren Reserve); E—
<taxonomicName id="C29E4D118904FFD225573D2BFBC4F969" box="[1034,1144,1661,1684]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="37EAEA808904FFD225573D2BFBC4F969" box="[1034,1144,1661,1684]" italics="true" pageId="25" pageNumber="26">Z. pallidus</emphasis>
</taxonomicName>
(NZ, Otago, Taieri Mouth); F—
<taxonomicName id="C29E4D118904FFD221E33DC9FEA0F948" box="[190,284,1692,1714]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808904FFD221E33DC9FEA0F948" box="[190,284,1692,1714]" italics="true" pageId="25" pageNumber="26">Z. brouni</emphasis>
</taxonomicName>
(NZ, Otago, Oamaru, Glen Warren Reserve); G—
<taxonomicName id="C29E4D118904FFD2227D3DC9FC0CF948" box="[800,944,1693,1714]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808904FFD2227D3DC9FC0CF948" box="[800,944,1693,1714]" italics="true" pageId="25" pageNumber="26">Z. montivagus</emphasis>
</taxonomicName>
(NZ, Otago, North Rough Ridge); H—
<taxonomicName id="C29E4D118904FFD224193DC9FA20F948" box="[1348,1436,1693,1714]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="37EAEA808904FFD224193DC9FA20F948" box="[1348,1436,1693,1714]" italics="true" pageId="25" pageNumber="26">Z. nanus</emphasis>
</taxonomicName>
(NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05mm.
</paragraph>
</caption>
<paragraph id="052136928904FFD2219A3DAFFEA8F885" blockId="25.[151,1437,1787,2028]" pageId="25" pageNumber="26">
Male genitalia. Median lobe (
<figureCitation id="9DA52A178904FFD2234D3DAFFDE2F8E9" box="[528,606,1787,1812]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
Q) with convex ventral margin and rather long apex with tapered narrowly rounded tip. Sclerites of internal sac slightly sclerotized, separate, long, almost equal half of the length between apical and basal orifices. Left paramere with apical constriction (
<figureCitation id="9DA52A178904FFD222DB3C17FC67F8A1" box="[902,987,1859,1884]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
S). Ring sclerite without visible handle (
<figureCitation id="9DA52A178904FFD221C23C32FF4DF885" box="[159,241,1894,1919]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="25" pageNumber="26">Fig. 14</figureCitation>
H).
</paragraph>
<paragraph id="052136928904FFD2219A3CDFFC5FF816" blockId="25.[151,1437,1787,2028]" pageId="25" pageNumber="26">
Female internal genitalia. Goncoxite 2 with short ensiform setae (
<figureCitation id="9DA52A178904FFD222F33CDFFBBCF85E" box="[942,1024,1931,1956]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="25" pageNumber="26">Fig. 15</figureCitation>
F). Spermatheca sclerotized, slightly elongate, comparatively wide, with two compartments (
<figureCitation id="9DA52A178904FFD222783CFAFCC7F83D" box="[805,891,1966,1991]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="25" pageNumber="26">Fig. 16</figureCitation>
F). Spermathecal duct short and without coils. Attachments of spermathecal duct and gland to spermatheca close together.
</paragraph>
<paragraph id="052136928907FFD1219A3BC3FEB5FEE7" blockId="26.[151,1437,151,501]" pageId="26" pageNumber="27">
<emphasis id="37EAEA808907FFD1219A3BC3FDB5FF4A" bold="true" box="[199,521,151,176]" pageId="26" pageNumber="27">Geographical distribution.</emphasis>
This species is known from two distant coastal forest localities of North and South
<collectingRegion id="C75AF8708907FFD121BC3BE8FE96FF2E" box="[225,298,188,212]" country="New Zealand" name="Otago" pageId="26" pageNumber="27">Otago</collectingRegion>
: majority of specimens were collected in Oamaru (Glen Warren Reserve), and only one specimen is known from the Coastal
<collectingRegion id="C75AF8708907FFD120E23BB4FDBFFF02" box="[447,515,224,248]" country="New Zealand" name="Otago" pageId="26" pageNumber="27">Otago</collectingRegion>
Hills (Alison Reserve). The last locality needs confirmation (
<figureCitation id="9DA52A178907FFD125BE3B8BFA85FF02" box="[1251,1337,223,248]" captionStart="FIGURE 18" captionStartId="30.[151,250,1918,1940]" captionTargetBox="[198,1387,199,1885]" captionTargetId="figure@30.[189,1398,193,1897]" captionTargetPageId="30" captionText="FIGURE 18. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Zeanillus. A: white circles, Z. phyllobius; green circles, Z. punctigerus; red circles, Z. nunni; blue diamond, Z. pellucidus; green diamond, Z. pallidus. B: white triangles, Z. brouni; green triangles, Z. nanus; red triangles, Z. lescheni; yellow triangle, Z. carltoni; black triangle, Z. montivagus. Elevation scale bars are given in meters." httpUri="https://zenodo.org/record/167666/files/figure.png" pageId="26" pageNumber="27">Fig. 18</figureCitation>
B, white triangles).
</paragraph>
<paragraph id="052136928907FFD1219A3A73FA21FEBA" blockId="26.[151,1437,151,501]" box="[199,1437,295,320]" pageId="26" pageNumber="27">
<emphasis id="37EAEA808907FFD1219A3A73FEF3FEBA" bold="true" box="[199,335,295,320]" pageId="26" pageNumber="27">Way of life.</emphasis>
Specimens were collected in the coastal dry broadleaf forest and in the Podocarp/broadleaf forest.
</paragraph>
<paragraph id="052136928907FFD1219A3A18FCEFFE0E" blockId="26.[151,1437,151,501]" pageId="26" pageNumber="27">
<emphasis id="37EAEA808907FFD1219A3A18FECCFE9F" bold="true" box="[199,368,332,357]" pageId="26" pageNumber="27">Relationships.</emphasis>
In the structure of male median lobe, members of this species are distinct from the other members of subgenus. The shape of round sclerite is similar to the allopatric interior species
<taxonomicName id="C29E4D118907FFD125933A25FAF6FE72" box="[1230,1354,368,392]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808907FFD125933A25FAF6FE72" box="[1230,1354,368,392]" italics="true" pageId="26" pageNumber="27">Z. lescheni</emphasis>
</taxonomicName>
and
<taxonomicName id="C29E4D118907FFD124D83A25FEA4FE56" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808907FFD124D83A25FEA4FE56" italics="true" pageId="26" pageNumber="27">Z. montivagus</emphasis>
</taxonomicName>
. Also, females of
<taxonomicName id="C29E4D118907FFD120BE3AC1FDF5FE56" box="[483,585,404,428]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808907FFD120BE3AC1FDF5FE56" box="[483,585,404,428]" italics="true" pageId="26" pageNumber="27">Z. brouni</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="C29E4D118907FFD122463AC1FC04FE56" box="[795,952,405,428]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808907FFD122463AC1FC04FE56" box="[795,952,405,428]" italics="true" pageId="26" pageNumber="27">Z. montivagus</emphasis>
</taxonomicName>
in the shape of spermatheca (i.e., in having two compartments with the larger part short and wide). These morphological similarities suggest that
<taxonomicName id="C29E4D118907FFD1246F3AEDFA20FE2A" box="[1330,1436,440,464]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="37EAEA808907FFD1246F3AEDFA20FE2A" box="[1330,1436,440,464]" italics="true" pageId="26" pageNumber="27">Z. brouni</emphasis>
</taxonomicName>
shares a common ancestry with
<taxonomicName id="C29E4D118907FFD120A03A89FD20FE0E" box="[509,668,477,500]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="37EAEA808907FFD120A03A89FD20FE0E" box="[509,668,477,500]" italics="true" pageId="26" pageNumber="27">Z. montivagus</emphasis>
</taxonomicName>
and
<taxonomicName id="C29E4D118907FFD1238F3A89FCF0FE0E" box="[722,844,476,500]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="37EAEA808907FFD1238F3A89FCF0FE0E" box="[722,844,476,500]" italics="true" pageId="26" pageNumber="27">Z. lescheni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>