278 lines
35 KiB
XML
278 lines
35 KiB
XML
<document id="8E4B5CA99342CED096F0277C2612B61F" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D3787848906FFD721CA3DA5FD2CFA3A" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus lescheni Sokolov, 2016, sp.n." docType="treatment" docUuid="0C39ACB5-D6BD-4485-BF63-7BFDF5F76E36" docUuidSource="ZooBank" docVersion="9" lastPageNumber="29" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="28" updateTime="1698393955637" updateUser="plazi">
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<mods:mods id="2E8C1053D167E9DA84D105059DAF4A46" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D1AC8D6391295332296058B192CE8A37">
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<mods:title id="CEA23E15B2F7B01541905C990085C625">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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</mods:titleInfo>
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<mods:name id="8B0E6E18F0B77663887B0E994AD66785" type="personal">
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<mods:roleTerm id="F46BDA1B449CF4B5DD65EE89B887AAA7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A1A49656122F55E800A54D7A00B6AE21">Sokolov, Igor M.</mods:namePart>
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<mods:typeOfResource id="867C5DC94BB54ACDF4BEF694ABF797E5">text</mods:typeOfResource>
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<mods:title id="E99A83051E7A45A21E64A26ED4035A8D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B1564C45DCEA1F7C5B2FE8D0343CDD9B">
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<mods:date id="682FE4494F97A3C98D8A809540F84A1F">2016</mods:date>
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<mods:detail id="B406516FAE61EE7BC4E388C9734998FE" type="volume">
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<mods:number id="FFC97B2D78920224EC7CE69A6F6E3130">4196</mods:number>
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<mods:detail id="AEEA1684D90DDFBD89470D428BB468A0" type="issue">
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<mods:number id="2B123A96950E12AD1EDBD05A8464F679">1</mods:number>
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<mods:classification id="2FB84BC6D2EB03F81CE4DE7FB5F71278">journal article</mods:classification>
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<mods:identifier id="70D2ED45A0BE8A6BD34392336329796A" type="DOI">10.11646/zootaxa.4196.1.1</mods:identifier>
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<mods:identifier id="673B3EA40F7F1AFE424939304B390378" type="GBIF-Dataset">b7651a7b-df24-4347-9585-0633b88dba51</mods:identifier>
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<mods:identifier id="331B34007971D128511407CA6ACB24CE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D15B9D9FCF5514EE33CEDD2E86CB6A55" type="Zenodo-Dep">167648</mods:identifier>
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<mods:identifier id="29626B27E2E80C5DD66603D4F8A5941E" type="ZooBank">026D88BD-86C0-4811-814A-CB5CACB3104F</mods:identifier>
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<treatment id="8D3787848906FFD721CA3DA5FD2CFA3A" ID-DOI="http://doi.org/10.5281/zenodo.6089692" ID-GBIF-Taxon="124814322" ID-Zenodo-Dep="6089692" LSID="urn:lsid:zoobank.org:act:0C39ACB5-D6BD-4485-BF63-7BFDF5F76E36" httpUri="http://treatment.plazi.org/id/8D3787848906FFD721CA3DA5FD2CFA3A" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<subSubSection id="4D8465198906FFD021CA3DA5FBEEF8D7" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="052136928906FFD021CA3DA5FE01F8F1" blockId="27.[151,1106,1777,1837]" box="[151,445,1777,1803]" pageId="27" pageNumber="28">
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<heading id="5E6981FE8906FFD021CA3DA5FE01F8F1" bold="true" box="[151,445,1777,1803]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
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<emphasis id="37EAEA808906FFD021CA3DA5FE01F8F1" bold="true" box="[151,445,1777,1803]" pageId="27" pageNumber="28">
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<emphasis id="37EAEA808906FFD021CA3DA5FEC6F8F1" bold="true" box="[151,378,1777,1803]" italics="true" pageId="27" pageNumber="28">
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<taxonomicName id="C29E4D118906FFD021CA3DA5FEC8F8F1" ID-CoL="8M59T" box="[151,372,1777,1803]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">Zeanillus lescheni</taxonomicName>
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,
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||
</emphasis>
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<taxonomicNameLabel id="2CD957FB8906FFD020DF3DA5FE01F8F1" box="[386,445,1777,1803]" pageId="27" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="052136928906FFD021CA3C40FBEEF8D7" blockId="27.[151,1106,1777,1837]" box="[151,1106,1812,1837]" pageId="27" pageNumber="28">
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<figureCitation id="9DA52A178906FFD021CA3C40FF5CF8D7" box="[151,224,1812,1837]" captionStart="FIGURE 1" captionStartId="8.[151,250,1917,1939]" captionTargetBox="[172,1423,199,1885]" captionTargetId="figure@8.[164,1430,193,1891]" captionTargetPageId="8" captionText="FIGURE 1. SEM illustrations of head, dorsal aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. pallidus; D— Z. pellucidus; E— Z. lescheni; F— Z. nanus. Legend: cl—clypeus; cs—clypeal seta; fcc—fronto-clypeal carina; fs—frontal seta; lb—labrum; mp 3 — maxillary palpomere 3; mp 4 — maxillary palpomere 4; pos—postorbital seta; ssa—anterior supraorbital seta; ssp—posterior supraorbital seta. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/257540/files/figure.png" pageId="27" pageNumber="28">Figs. 1</figureCitation>
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E, 2D, 3G–H, 4E, 5E, 6E, 7G, 8D, 9D, 10JP, 11H, 13E–H,
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<date id="712010528906FFD022283C41FC21F8D7" box="[885,925,1813,1837]" pageId="27" pageNumber="28" value="2014-01">14I</date>
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||
, 15H, 17B, 18B
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4D8465198906FFD721CA3C08FDAEFEBA" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="052136928906FFD021CA3C08FB3AF862" blockId="27.[151,1437,1884,2017]" pageId="27" pageNumber="28">
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<materialsCitation id="B5F63CCF8906FFD021CA3C08FB3AF862" ID-GBIF-Occurrence="1341350606" collectingDate="1925-04" collectionCode="NZAC" collectorName="CO Whitecoomb Rd" country="New Zealand" location="Post Office Ck" pageId="27" pageNumber="28" specimenCount="2" typeStatus="holotype">
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||
<emphasis id="37EAEA808906FFD021CA3C08FEF3F88F" bold="true" box="[151,335,1884,1909]" pageId="27" pageNumber="28">
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||
<typeStatus id="DA2588308906FFD021CA3C08FF6EF88F" box="[151,210,1884,1909]" pageId="27" pageNumber="28">Type</typeStatus>
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material.
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||
</emphasis>
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||
<typeStatus id="DA2588308906FFD020023C09FE4AF88F" box="[351,502,1885,1909]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
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, male, in
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<collectionCode id="638FAE578906FFD023DD3C09FD73F88F" box="[640,719,1885,1909]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="27" pageNumber="28">NZAC</collectionCode>
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, board-mounted, undissected, labeled: /
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<collectingCountry id="7D8976028906FFD025993C09FAD8F88F" box="[1220,1380,1884,1909]" name="New Zealand" pageId="27" pageNumber="28">New Zealand</collectingCountry>
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||
<collectorName id="A86B53448906FFD024283C09FEF3F863" pageId="27" pageNumber="28">CO Whitecoomb Rd</collectorName>
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||
<quantity id="C2669B778906FFD0200B3CD4FE34F863" box="[342,392,1920,1945]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="km" value="2.0">2km</quantity>
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||
N of
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||
<location id="004160498906FFD020953CD5FDD3F863" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498906FFD020953CD5FDD3F863" box="[456,623,1920,1945]" country="New Zealand" name="Post Office Ck" pageId="27" pageNumber="28">Post Office Ck</location>
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/ Washed soil sample, beech forest
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||
<collectingDate id="6164E9BA8906FFD022A23CD4FBF2F863" box="[1023,1102,1920,1945]" pageId="27" pageNumber="28" value="1925-04">
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<date id="712010528906FFD022A23CD4FBF2F863" box="[1023,1102,1920,1945]" pageId="27" pageNumber="28" value="1925-04">25 Apr</date>
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</collectingDate>
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0 7 /.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="052136928906FFD7219A3CF1FDAEFEBA" blockId="27.[151,1437,1884,2017]" lastBlockId="28.[151,1437,151,1473]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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PARATYPES (13 specimens, 4 dissected), 1 male labeled same as holotype;
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||
<materialsCitation id="B5F63CCF8906FFD025603CF1FB29F81A" ID-GBIF-Occurrence="1341350598" collectingDate="1925-04" collectorName="CO Piano Flat Waikaia Forest" country="New Zealand" location="Piano Flat Waikaia Forest" municipality="Washed" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
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||
<specimenCount id="1398FD1B8906FFD025603CF1FB36F847" box="[1085,1162,1956,1981]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
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||
and
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||
<specimenCount id="1398FD1B8906FFD0259B3CF1FA95F847" box="[1222,1321,1956,1981]" pageId="27" pageNumber="28" type="female">1 female</specimenCount>
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||
labeled: /
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||
<collectingCountry id="7D8976028906FFD021CA3C9DFE91F81B" box="[151,301,1992,2017]" name="New Zealand" pageId="27" pageNumber="28">New Zealand</collectingCountry>
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<collectorName id="A86B53448906FFD020683C9CFD39F81A" box="[309,645,1992,2017]" pageId="27" pageNumber="28">
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CO
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<location id="004160498906FFD0203E3C9CFD39F81A" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498906FFD0203E3C9CFD39F81A" box="[355,645,1992,2017]" country="New Zealand" municipality="Washed" name="Piano Flat Waikaia Forest" pageId="27" pageNumber="28">Piano Flat Waikaia Forest</location>
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</collectorName>
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||
<collectingDate id="6164E9BA8906FFD023D03C9CFD60F81B" box="[653,732,1992,2017]" pageId="27" pageNumber="28" value="1925-04">
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||
<date id="712010528906FFD023D03C9CFD60F81B" box="[653,732,1992,2017]" pageId="27" pageNumber="28" value="1925-04">25 Apr</date>
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||
</collectingDate>
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||
0 7 /
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<collectingMunicipality id="E545ACE88906FFD022483C9CFCD1F81B" box="[789,877,1992,2017]" pageId="27" pageNumber="28">Washed</collectingMunicipality>
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||
soil sample beech forest /
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="B5F63CCF8906FFD725FF3C9CFF78FF2E" ID-GBIF-Occurrence="1341350593" country="New Zealand" lastPageId="28" lastPageNumber="29" location="Post Office Ck Waikaia Forest" pageId="27" pageNumber="28" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
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||
<specimenCount id="1398FD1B8906FFD025FF3C9CFB44F81B" box="[1186,1272,1992,2017]" pageId="27" pageNumber="28" type="male">3 males</specimenCount>
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||
and
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||
<specimenCount id="1398FD1B8906FFD0246D3C9CFA20F81B" box="[1328,1436,1992,2017]" pageId="27" pageNumber="28" type="female">3 females</specimenCount>
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labeled: /
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||
<collectingCountry id="7D8976028901FFD7205B3BCCFE20FF4A" box="[262,412,151,176]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
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||
CO
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<quantity id="C2669B778901FFD7208F3BC3FDB8FF4A" box="[466,516,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="km" value="2.0">2km</quantity>
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north of
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<location id="004160498901FFD723313BCCFC7EFF55" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498901FFD723313BCCFC7EFF55" box="[620,962,151,176]" country="New Zealand" name="Post Office Ck Waikaia Forest" pageId="28" pageNumber="29">Post Office Ck Waikaia Forest</location>
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/ Washed soil sample, beech forest 25/04/ 0 7 /
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||
</materialsCitation>
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||
;
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<materialsCitation id="B5F63CCF8901FFD7218D3BE8FF46FF02" ID-GBIF-Occurrence="1341350605" collectingDate="1925-04" collectorName="CO Waikaia Forest Whitecoomb Rd & Post Office Ck" country="New Zealand" location="Waikaia Forest" municipality="Washed" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1">
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||
<specimenCount id="1398FD1B8901FFD7218D3BE8FEA6FF2F" box="[208,282,188,213]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
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||
labeled: /
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||
<collectingCountry id="7D8976028901FFD720CD3BE9FD99FF2F" box="[400,549,188,213]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
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||
<collectorName id="A86B53448901FFD723713BE8FC7DFF2F" box="[556,961,188,213]" pageId="28" pageNumber="29">
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||
CO
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||
<location id="004160498901FFD723073BE8FCBEFF2E" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498901FFD723073BE8FCBEFF2E" box="[602,770,188,213]" country="New Zealand" municipality="Washed" name="Waikaia Forest" pageId="28" pageNumber="29">Waikaia Forest</location>
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||
Whitecoomb Rd
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||
</collectorName>
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||
at
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||
<collectorName id="A86B53448901FFD722BE3BE9FB34FF2F" box="[995,1160,188,213]" pageId="28" pageNumber="29">Post Office Ck</collectorName>
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||
<collectingDate id="6164E9BA8901FFD725D23BE8FB62FF2E" box="[1167,1246,188,213]" pageId="28" pageNumber="29" value="1925-04">
|
||
<date id="712010528901FFD725D23BE8FB62FF2E" box="[1167,1246,188,213]" pageId="28" pageNumber="29" value="1925-04">25 Apr</date>
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||
</collectingDate>
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||
0 7 /
|
||
<collectingMunicipality id="E545ACE88901FFD724483BE8FAD1FF2F" box="[1301,1389,188,213]" pageId="28" pageNumber="29">Washed</collectingMunicipality>
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||
soil sample /
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="B5F63CCF8901FFD720543BB4FDEDFEE6" ID-GBIF-Occurrence="1341350608" collectingDate="1925-04" collectorName="CO Waikaia Forest Whitecoomb Rd & Post Office Ck & Washed" country="New Zealand" location="Waikaia Forest" municipality="Nothophagus forest" pageId="28" pageNumber="29" specimenCount="2" specimenCount-male="2">
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||
<specimenCount id="1398FD1B8901FFD720543BB4FEDEFF02" box="[265,354,223,248]" pageId="28" pageNumber="29" type="male">2 males</specimenCount>
|
||
labeled: /
|
||
<collectingCountry id="7D8976028901FFD720BD3BB4FDC5FF02" box="[480,633,223,248]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
|
||
<collectorName id="A86B53448901FFD723DE3BB4FB98FF02" box="[643,1060,223,248]" pageId="28" pageNumber="29">
|
||
CO
|
||
<location id="004160498901FFD723E93B8BFCE3FF0D" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498901FFD723E93B8BFCE3FF0D" box="[692,863,223,248]" country="New Zealand" municipality="Nothophagus forest" name="Waikaia Forest" pageId="28" pageNumber="29">Waikaia Forest</location>
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||
Whitecoomb Rd
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||
</collectorName>
|
||
at
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||
<collectorName id="A86B53448901FFD725163BB4FB4BFF02" box="[1099,1271,223,248]" pageId="28" pageNumber="29">Post Office Ck</collectorName>
|
||
/
|
||
<collectorName id="A86B53448901FFD7244F3B8BFAD6FF02" box="[1298,1386,223,248]" pageId="28" pageNumber="29">Washed</collectorName>
|
||
soil sample
|
||
<collectingMunicipality id="E545ACE88901FFD721B03A50FE75FEE7" box="[237,457,260,285]" pageId="28" pageNumber="29">Nothophagus forest</collectingMunicipality>
|
||
<collectingDate id="6164E9BA8901FFD720923A50FDA2FEE6" box="[463,542,260,285]" pageId="28" pageNumber="29" value="1925-04">
|
||
<date id="712010528901FFD720923A50FDA2FEE6" box="[463,542,260,285]" pageId="28" pageNumber="29" value="1925-04">25 Apr</date>
|
||
</collectingDate>
|
||
0 7 /
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="B5F63CCF8901FFD723003A50FDAEFEBA" ID-GBIF-Occurrence="1341350602" collectingDate="1925-04" collectorName="CO Whitecoomb Rd & Washed" country="New Zealand" location="Post Office Ck" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="1398FD1B8901FFD723003A50FD1BFEE7" box="[605,679,260,285]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
|
||
labeled: /
|
||
<collectingCountry id="7D8976028901FFD722413A51FC0DFEE7" box="[796,945,260,285]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
|
||
<collectorName id="A86B53448901FFD722E53A50FB22FEE7" box="[952,1182,260,285]" pageId="28" pageNumber="29">CO Whitecoomb Rd</collectorName>
|
||
<quantity id="C2669B778901FFD725F93A50FB6AFEE7" box="[1188,1238,260,285]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="km" value="4.0">4km</quantity>
|
||
N
|
||
<location id="004160498901FFD725AA3A51FA21FEE7" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498901FFD725AA3A51FA21FEE7" box="[1271,1437,260,285]" country="New Zealand" name="Post Office Ck" pageId="28" pageNumber="29">Post Office Ck</location>
|
||
/
|
||
<collectorName id="A86B53448901FFD721F83A73FF41FEBA" box="[165,253,295,320]" pageId="28" pageNumber="29">Washed</collectorName>
|
||
soil sample
|
||
<collectingDate id="6164E9BA8901FFD720D73A73FE65FEBA" box="[394,473,295,320]" pageId="28" pageNumber="29" value="1925-04">
|
||
<date id="712010528901FFD720D73A73FE65FEBA" box="[394,473,295,320]" pageId="28" pageNumber="29" value="1925-04">25 Apr</date>
|
||
</collectingDate>
|
||
0 7 /.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D8465198901FFD7219A3A18FEDAFE57" pageId="28" pageNumber="29" type="description">
|
||
<paragraph id="052136928901FFD7219A3A18FEDAFE57" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3A18FE3AFE9F" bold="true" box="[199,390,332,357]" pageId="28" pageNumber="29">Specific epithet.</emphasis>
|
||
The specific epithet honors the surname of Richard A.B. Leschen, coleopterist of Landcare Research and Curator of the
|
||
<collectingCountry id="7D8976028901FFD720833A24FDC9FE72" box="[478,629,367,392]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
|
||
Arthropod Collection, who loaned the specimens of Anillina under his care for this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D8465198901FFD7219A3AE3FEDFFDE2" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="052136928901FFD7219A3AE3FB10FE35" blockId="28.[151,1437,151,1473]" box="[199,1196,439,464]" pageId="28" pageNumber="29">
|
||
<materialsCitation id="B5F63CCF8901FFD7219A3AE3FB14FE35" ID-GBIF-Occurrence="1341350607" box="[199,1192,439,464]" country="New Zealand" location="Waikaia Forest Conservation area" municipality="South Island" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Southland" typeStatus="holotype">
|
||
<emphasis id="37EAEA808901FFD7219A3AE3FED9FE2A" bold="true" box="[199,357,439,464]" pageId="28" pageNumber="29">
|
||
<typeStatus id="DA2588308901FFD7219A3AE3FEBEFE2A" box="[199,258,439,464]" pageId="28" pageNumber="29">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="7D8976028901FFD720313AECFDB9FE2A" box="[364,517,439,464]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="E545ACE88901FFD7234D3AE3FD23FE2A" box="[528,671,439,464]" pageId="28" pageNumber="29">South Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="C75AF8708901FFD723F63AE3FCA3FE2A" box="[683,799,439,464]" country="New Zealand" name="Southland" pageId="28" pageNumber="29">Southland</collectingRegion>
|
||
,
|
||
<location id="004160498901FFD722773AE3FB14FE35" LSID="urn:lsid:plazi:treatment:8D3787848906FFD721CA3DA5FD2CFA3A:004160498901FFD722773AE3FB14FE35" box="[810,1192,439,464]" country="New Zealand" municipality="South Island" name="Waikaia Forest Conservation area" pageId="28" pageNumber="29" stateProvince="Southland">Waikaia Forest Conservation area</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3A88FEDFFDE2" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3A88FEE1FE0F" bold="true" box="[199,349,476,501]" pageId="28" pageNumber="29">Recognition.</emphasis>
|
||
Adults of this species (
|
||
<figureCitation id="9DA52A178901FFD7233B3A88FD09FE0E" box="[614,693,476,501]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
|
||
H) are distinguished from other congeners by the characters listed in the species key.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D8465198901FFD7219A3970FD2CFA3A" pageId="28" pageNumber="29" type="description">
|
||
<paragraph id="052136928901FFD7219A3970FAEAFDC6" blockId="28.[151,1437,151,1473]" box="[199,1366,548,573]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3970FEE5FDC7" bold="true" box="[199,345,548,573]" pageId="28" pageNumber="29">Description.</emphasis>
|
||
Size. Medium to large for genus (SBL range
|
||
<quantity id="C2669B778901FFD7220A3970FC40FDC6" box="[855,1020,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.98" metricValueMin="1.72" pageId="28" pageNumber="29" unit="mm" value="1.85" valueMax="1.98" valueMin="1.72">1.72–1.98 mm</quantity>
|
||
, mean 1.84±
|
||
<quantity id="C2669B778901FFD725D73970FABCFDC6" box="[1162,1280,548,572]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="29" unit="mm" value="0.08">0.080 mm</quantity>
|
||
, n=14).
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3913FE17FD52" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.37±0.014), head of moderate width for genus compared to pronotum (WH/WPm 0.72±0.015), pronotum rather narrow in comparison to elytra (WPm/WE 0.71±0.017).</paragraph>
|
||
<paragraph id="052136928901FFD7219A39E0FCF7FD36" blockId="28.[151,1437,151,1473]" box="[199,843,692,717]" pageId="28" pageNumber="29">Color. Body color rufotestaceous, appendages testaceous.</paragraph>
|
||
<paragraph id="052136928901FFD7219A3983FC60FCC2" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">Prothorax. Pronotum slightly long (LP/LE 0.40±0.009) and moderately elongate (WPm/LP 1.28±0.031), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.52±0.070). Width between anterior angles moderately greater than between posterior angles (WPa/WPp 1.15±0.045).</paragraph>
|
||
<paragraph id="052136928901FFD7219A3810FE86FC5F" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">Elytra. Slightly depressed along suture, comparatively long (LE/SBL 0.53±0.012) and moderately wide (WE/ LE 0.69±0.019). Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.</paragraph>
|
||
<paragraph id="052136928901FFD7219A38FBFC44FBCE" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
Male genitalia. Median lobe (
|
||
<figureCitation id="9DA52A178901FFD7234138FBFDD2FC32" box="[540,622,943,968]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="28" pageNumber="29">Fig. 13</figureCitation>
|
||
E) with straight ventral margin and enlarged apex in dorsal projection. Sclerites of internal sac sclerotized, unequal in length, with long, thick ventral and short, narrow dorsal branches, united basally, almost equals to the half of the distance between apical and basal orifices. Left paramere with long apical constriction (
|
||
<figureCitation id="9DA52A178901FFD720293F48FE79FBCE" box="[372,453,1052,1077]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="28" pageNumber="29">Fig. 13</figureCitation>
|
||
G). Ring sclerite without visible handle (
|
||
<figureCitation id="9DA52A178901FFD722D33F48FC5CFBCE" box="[910,992,1052,1077]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="28" pageNumber="29">
|
||
Fig.
|
||
<date id="712010528901FFD7229F3F48FC5CFBCE" box="[962,992,1052,1076]" pageId="28" pageNumber="29" value="2014-01">14</date>
|
||
</figureCitation>
|
||
I).
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3F6BFBD8FB5A" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
Female internal genitalia. Goncoxite 2 with short ensiform setae (
|
||
<figureCitation id="9DA52A178901FFD722FF3F6BFC4EFBA2" box="[930,1010,1087,1112]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="28" pageNumber="29">Fig. 15</figureCitation>
|
||
H). Spermatheca sclerotized, gigantic, elongate and curved, with two compartments (
|
||
<figureCitation id="9DA52A178901FFD723FD3F30FD4FFB86" box="[672,755,1124,1149]" captionStart="FIGURE 17" captionStartId="27.[151,250,1593,1615]" captionTargetBox="[272,1296,204,1563]" captionTargetId="figure@27.[263,1309,193,1579]" captionTargetPageId="27" captionText="FIGURE 17. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. carltoni (NZ, Otago, Mount Watkin); B— Z. lescheni (NZ, Otago, Waikaia Forest); C— Z. nanus (NZ, Otago, Balclutha), the structure is the same as in Fig. 16 H and is repeated for purposes of comparison among species. Legend: sd—spermathecal duct. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167665/files/figure.png" pageId="28" pageNumber="29">Fig. 17</figureCitation>
|
||
B). Spermathecal duct short and with few coils. Attachments of spermathecal duct and gland to spermatheca at a moderate distance from each other.
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3FF8FC3EFB12" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3FF8FDBAFB3F" bold="true" box="[199,518,1196,1221]" pageId="28" pageNumber="29">Geographical distribution.</emphasis>
|
||
This species is known from several localities in the Waikaia Forest at the upper course of the Waikaia River of
|
||
<collectingRegion id="C75AF8708901FFD720A93F9BFDDAFB12" box="[500,614,1231,1256]" country="New Zealand" name="Southland" pageId="28" pageNumber="29">Southland</collectingRegion>
|
||
(
|
||
<figureCitation id="9DA52A178901FFD723283F9BFD7AFB12" box="[629,710,1231,1256]" captionStart="FIGURE 18" captionStartId="30.[151,250,1918,1940]" captionTargetBox="[198,1387,199,1885]" captionTargetId="figure@30.[189,1398,193,1897]" captionTargetPageId="30" captionText="FIGURE 18. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Zeanillus. A: white circles, Z. phyllobius; green circles, Z. punctigerus; red circles, Z. nunni; blue diamond, Z. pellucidus; green diamond, Z. pallidus. B: white triangles, Z. brouni; green triangles, Z. nanus; red triangles, Z. lescheni; yellow triangle, Z. carltoni; black triangle, Z. montivagus. Elevation scale bars are given in meters." httpUri="https://zenodo.org/record/167666/files/figure.png" pageId="28" pageNumber="29">Fig. 18</figureCitation>
|
||
B, red triangles).
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3FA0FBFBFAF7" blockId="28.[151,1437,151,1473]" box="[199,1095,1268,1293]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3FA0FEF3FAF7" bold="true" box="[199,335,1268,1293]" pageId="28" pageNumber="29">Way of life.</emphasis>
|
||
Specimens were collected from soil in a beech (
|
||
<taxonomicName id="C29E4D118901FFD722333FA0FC53FAF6" box="[878,1007,1268,1292]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="37EAEA808901FFD722333FA0FC53FAF6" box="[878,1007,1268,1292]" italics="true" pageId="28" pageNumber="29">Nothofagus</emphasis>
|
||
</taxonomicName>
|
||
) forest.
|
||
</paragraph>
|
||
<paragraph id="052136928901FFD7219A3E43FD2CFA3A" blockId="28.[151,1437,151,1473]" pageId="28" pageNumber="29">
|
||
<emphasis id="37EAEA808901FFD7219A3E43FECCFACA" bold="true" box="[199,368,1303,1328]" pageId="28" pageNumber="29">Relationships.</emphasis>
|
||
Males of this species are distinct from those of other members of subgenus in the structure of the median lobe. Hypertrophy of the apex of male median lobe and great increase in the size of spermatheca are shared with members of
|
||
<taxonomicName id="C29E4D118901FFD720F13E35FD9FFA82" box="[428,547,1376,1400]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carltoni">
|
||
<emphasis id="37EAEA808901FFD720F13E35FD9FFA82" box="[428,547,1376,1400]" italics="true" pageId="28" pageNumber="29">Z. carltoni</emphasis>
|
||
</taxonomicName>
|
||
. Males of
|
||
<taxonomicName id="C29E4D118901FFD723C73E35FCA8FA82" box="[666,788,1376,1400]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="37EAEA808901FFD723C73E35FCA8FA82" box="[666,788,1376,1400]" italics="true" pageId="28" pageNumber="29">Z. lescheni</emphasis>
|
||
</taxonomicName>
|
||
are also similar to those of the interior
|
||
<taxonomicName id="C29E4D118901FFD725903E35FAD0FA8D" box="[1229,1388,1376,1400]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="montivagus">
|
||
<emphasis id="37EAEA808901FFD725903E35FAD0FA8D" box="[1229,1388,1376,1400]" italics="true" pageId="28" pageNumber="29">Z. montivagus</emphasis>
|
||
</taxonomicName>
|
||
and coastal
|
||
<taxonomicName id="C29E4D118901FFD721B03ED1FEEAFA66" box="[237,342,1412,1436]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="brouni">
|
||
<emphasis id="37EAEA808901FFD721B03ED1FEEAFA66" box="[237,342,1412,1436]" italics="true" pageId="28" pageNumber="29">Z. brouni</emphasis>
|
||
</taxonomicName>
|
||
in the shape of round sclerite.. Based on the latter similarity, I suggest that
|
||
<taxonomicName id="C29E4D118901FFD725E33ED1FA85FA66" box="[1214,1337,1412,1436]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="37EAEA808901FFD725E33ED1FA85FA66" box="[1214,1337,1412,1436]" italics="true" pageId="28" pageNumber="29">Z. lescheni</emphasis>
|
||
</taxonomicName>
|
||
shares a common ancestry with the latter two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |